418 lines
55 KiB
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418 lines
55 KiB
XML
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<mods:title id="48F86B5AEE8494FB5C15A04FC71F563F">A systematic revision of Draculoides (Schizomida: Hubbardiidae) of the Pilbara, Western Australia, Part I: the Western Pilbara</mods:title>
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<taxonomicName id="4C674D4CFFB8FFEDCC8B0483FD4CFA95" authority="Abrams and Harvey" authorityName="Abrams and Harvey" authorityYear="2020" box="[151,684,1279,1306]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="nosferatu" status="sp. nov.">
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<emphasis id="B913EADDFFB8FFEDCC8B0483FE45FA95" bold="true" box="[151,421,1279,1306]" italics="true" pageId="62" pageNumber="63">Draculoides nosferatu</emphasis>
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Abrams and Harvey
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,
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<paragraph id="8BD836CFFFB8FFEDCC8B055FFE86FAB2" blockId="62.[151,1302,1279,1378]" box="[151,358,1315,1342]" pageId="62" pageNumber="63">
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(
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<figureCitation id="135C2A4AFFB8FFEDCC83055FFEEBFAB1" box="[159,267,1315,1341]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="16.[151,250,1868,1892]" captionStartId-1="17.[151,250,1910,1934]" captionStartId-2="18.[151,250,1547,1571]" captionStartId-3="19.[151,250,1882,1906]" captionStartId-4="20.[151,250,1216,1240]" captionStartId-5="21.[151,250,1709,1733]" captionStartId-6="22.[151,250,1755,1779]" captionStartId-7="23.[151,250,1919,1943]" captionTargetBox-0="[346,1243,188,1844]" captionTargetBox-1="[332,1254,181,1881]" captionTargetBox-2="[158,1420,181,1521]" captionTargetBox-3="[151,1425,182,1858]" captionTargetBox-4="[158,1424,183,1189]" captionTargetBox-5="[155,1423,181,1683]" captionTargetBox-6="[159,1425,181,1729]" captionTargetBox-7="[151,1430,181,1893]" captionTargetId-0="figure-20@16.[339,1249,181,1844]" captionTargetId-1="figure-17@17.[332,1254,181,1886]" captionTargetId-2="figure-20@18.[151,1436,181,1524]" captionTargetId-3="figure-17@19.[151,1436,181,1859]" captionTargetId-4="figure-20@20.[151,1436,181,1192]" captionTargetId-5="figure-17@21.[155,1432,181,1685]" captionTargetId-6="figure-20@22.[151,1436,181,1732]" captionTargetId-7="figure-17@23.[151,1436,181,1894]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="22" captionTargetPageId-7="23" captionText-0="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." captionText-1="FIGURE 2. Maximum Likelihood tree, based on the reduced dataset. See Methods for details. Each terminal represents a single species or OTU, with named species in bold. Clades are identified using dashed boxes and shaded boxes encompass the species discussed in this study. Nodes with bootstrap support <80 are not shown, except for the common ancestor of the Draculoides radiation." captionText-2="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-3="FIGURE 4. Maximum Likelihood tree based on COI mini-barcodes (Fig. 3) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-4="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-5="FIGURE 6. Maximum Likelihood tree based on COI mini-barcodes (Fig. 5) showing similarity of mini-barcode sequences relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-6="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-7="FIGURE 8. Maximum Likelihood tree based on ITS2 mini-barcodes (Fig. 7) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." figureDoi-0="http://doi.org/10.5281/zenodo.4416851" figureDoi-1="http://doi.org/10.5281/zenodo.4416855" figureDoi-2="http://doi.org/10.5281/zenodo.4416857" figureDoi-3="http://doi.org/10.5281/zenodo.4416859" figureDoi-4="http://doi.org/10.5281/zenodo.4416863" figureDoi-5="http://doi.org/10.5281/zenodo.4416865" figureDoi-6="http://doi.org/10.5281/zenodo.4416867" figureDoi-7="http://doi.org/10.5281/zenodo.4416869" httpUri-0="https://zenodo.org/record/4416851/files/figure.png" httpUri-1="https://zenodo.org/record/4416855/files/figure.png" httpUri-2="https://zenodo.org/record/4416857/files/figure.png" httpUri-3="https://zenodo.org/record/4416859/files/figure.png" httpUri-4="https://zenodo.org/record/4416863/files/figure.png" httpUri-5="https://zenodo.org/record/4416865/files/figure.png" httpUri-6="https://zenodo.org/record/4416867/files/figure.png" httpUri-7="https://zenodo.org/record/4416869/files/figure.png" pageId="62" pageNumber="63">Figs. 1–8</figureCitation>
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,
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<figureCitation id="135C2A4AFFB8FFEDCD0B055FFEBEFAB2" box="[279,350,1315,1342]" captionStart-0="FIGURE 36" captionStart-1="FIGURE 37" captionStartId-0="63.[151,250,1135,1159]" captionStartId-1="63.[151,250,1875,1899]" captionTargetBox-0="[151,1436,525,1111]" captionTargetBox-1="[407,1176,1225,1845]" captionTargetId-0="figure-181@63.[151,1436,525,1111]" captionTargetId-1="figure-222@63.[407,1179,1219,1852]" captionTargetPageId-0="63" captionTargetPageId-1="63" captionText-0="FIGURE 36. Draculoides nosferatu sp. nov., holotype female (WAM T138516): A. Body, dorsal; B. Body, ventral; C. Body, lateral. The scale bar shown in image A is 1 mm and also applies to B and C." captionText-1="FIGURE 37. Draculoides nosferatu sp. nov., holotype female (WAM T138516): A. Spermathecae, ventral. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4416939" figureDoi-1="http://doi.org/10.5281/zenodo.4416941" httpUri-0="https://zenodo.org/record/4416939/files/figure.png" httpUri-1="https://zenodo.org/record/4416941/files/figure.png" pageId="62" pageNumber="63">36–37</figureCitation>
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)
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</paragraph>
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<paragraph id="8BD836CFFFB8FFEDCC8B053BFAF6FAEE" blockId="62.[151,1302,1279,1378]" box="[151,1302,1351,1378]" pageId="62" pageNumber="63">
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Zoobank Code: http://zoobank.org/NomenclaturalActs/
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<taxonomicName id="4C674D4CFFB8FFEDCC8B05F0FEB0FA28" authority="SCH" authorityName="SCH" box="[151,336,1420,1444]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
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<emphasis id="B913EADDFFB8FFEDCC8B05F0FEF4FA28" box="[151,276,1420,1444]" italics="true" pageId="62" pageNumber="63">Draculoides</emphasis>
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SCH
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061:
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<bibRefCitation id="EFF64B3EFFB8FFEDCD6205F0FDA1FA28" author="Abrams, K. M. & Huey, J. A. & Hillyer, M. J. & Humphreys, W. F. & Didham, R. K. & Harvey, M. S." box="[382,577,1420,1444]" pageId="62" pageNumber="63" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. & Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
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Abrams
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<emphasis id="B913EADDFFB8FFEDCDC905F0FDE1FA28" box="[469,513,1420,1444]" italics="true" pageId="62" pageNumber="63">et al</emphasis>
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. 2019
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</bibRefCitation>
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MPE 106532: 8, fig. 2.
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</paragraph>
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<emphasis id="B913EADDFFB8FFEDCC8B05AEFE60FA60" bold="true" box="[151,384,1490,1516]" pageId="62" pageNumber="63">Material examined.</emphasis>
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<typeStatus id="54DC886DFFB8FFEDCD9B05AEFE0EFA60" box="[391,494,1490,1516]" pageId="62" pageNumber="63" type="holotype">Holotype</typeStatus>
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female.
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:
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<collectingRegion id="49A3F82DFFB8FFEDCEEB05AFFC5EFA60" box="[759,958,1490,1516]" country="Australia" name="Western Australia" pageId="62" pageNumber="63">Western Australia</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D61FD46FFB8FFEDCFD105AEFC17FA60" box="[973,1015,1490,1516]" pageId="62" pageNumber="63" type="female">1 ♀</specimenCount>
|
||
,
|
||
<location id="8EB86014FFB8FFEDC81805AEFB6EFA60" LSID="urn:lsid:plazi:treatment:03CE87D9FFB8FF93CC8B0483FD02FCB5:8EB86014FFB8FFEDC81805AEFB6EFA60" box="[1028,1166,1490,1516]" country="Australia" latitude="-21.741034" longLatPrecision="1" longitude="116.55689" name="Robe Valley" pageId="62" pageNumber="63" stateProvince="Western Australia">Robe Valley</location>
|
||
, ca.
|
||
<quantity id="4C9F9B2AFFB8FFEDC8A305AEFAE5FA60" box="[1215,1285,1490,1516]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="62" pageNumber="63" unit="km" value="27.0">27 km</quantity>
|
||
ESE. of Pan- nawonica,
|
||
<geoCoordinate id="EE535008FFB8FFEDCD0C058AFE5AF99C" box="[272,442,1526,1552]" degrees="21" direction="south" minutes="44" orientation="latitude" pageId="62" pageNumber="63" precision="1" seconds="27.72" value="-21.741034">21°44’27.72”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FFB8FFEDCDD9058AFD9EF99C" box="[453,638,1526,1552]" degrees="116" direction="east" minutes="33" orientation="longitude" pageId="62" pageNumber="63" precision="1" seconds="24.81" value="116.55689">116°33’24.81”E</geoCoordinate>
|
||
,
|
||
<collectingDate id="EF9DE9E7FFB8FFEDCE95058AFC9CF99D" box="[649,892,1526,1553]" isEnumeration="true" pageId="62" pageNumber="63">
|
||
<date id="FFD9100FFFB8FFEDCE95058AFD1DF99D" box="[649,765,1526,1553]" pageId="62" pageNumber="63" value="2015-06-06">6/06/2015</date>
|
||
,
|
||
<date id="FFD9100FFFB8FFEDCF14058AFC9CF99D" box="[776,892,1526,1553]" pageId="62" pageNumber="63" value="2015-08-08">8/08/2015</date>
|
||
</collectingDate>
|
||
,
|
||
<collectingMethod id="52264ED8FFB8FFEDCF9B058AFBDDF99C" box="[903,1085,1526,1552]" pageId="62" pageNumber="63">troglofauna trap</collectingMethod>
|
||
,
|
||
<collectorName id="26925319FFB8FFEDC854058AFB34F99C" box="[1096,1236,1526,1552]" pageId="62" pageNumber="63">J. Alexander</collectorName>
|
||
(
|
||
<collectionCode id="ED76AE0AFFB8FFEDC8FD058AFAC8F99C" box="[1249,1320,1526,1552]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="62" pageNumber="63" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FFB8FFEDC932058AFA75F99D" box="[1326,1429,1526,1553]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="62" pageNumber="63">T138516</specimenCode>
|
||
) (DNA:
|
||
<collectionCode id="ED76AE0AFFB8FFEDCCF00666FEFFF9B8" box="[236,287,1562,1588]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="62" pageNumber="63" type="Herbarium">COI</collectionCode>
|
||
, 18S, 28S,
|
||
<specimenCode id="DBC19EB4FFB8FFEDCD810666FE3AF9B8" box="[413,474,1562,1588]" collectionCode="ITS" pageId="62" pageNumber="63">ITS2</specimenCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFB8FFEDCCDB0642FE14F880" blockId="62.[151,1437,1490,2020]" pageId="62" pageNumber="63">
|
||
<emphasis id="B913EADDFFB8FFEDCCDB0642FE98F9D4" box="[199,376,1598,1624]" italics="true" pageId="62" pageNumber="63">Other material.</emphasis>
|
||
<materialsCitation id="3B0F3C92FFB8FFEDCD630642FEB9F92C" ID-GBIF-Occurrence="3013449382" collectingDate="2015-06-06" collectingDateMax="2015-08-08" collectingDateMin="2015-06-06" collectionCode="WAM, COI" collectorName="J. Alexander" country="Australia" latitude="-21.741997" location="Pannawonica" longLatPrecision="1" longitude="116.56163" municipality="Robe Valley" pageId="62" pageNumber="63" specimenCode="T138518, ITS2" specimenCount="1" specimenCount-juvenile="1" stateProvince="Western Australia">
|
||
<collectingCountry id="F370765FFFB8FFEDCD630642FDC2F9D4" box="[383,546,1598,1624]" name="Australia" pageId="62" pageNumber="63">AUSTRALIA</collectingCountry>
|
||
:
|
||
<emphasis id="B913EADDFFB8FFEDCE320643FD18F9D4" box="[558,760,1598,1624]" italics="true" pageId="62" pageNumber="63">
|
||
<collectingRegion id="49A3F82DFFB8FFEDCE320643FD18F9D4" box="[558,760,1598,1624]" country="Australia" name="Western Australia" pageId="62" pageNumber="63">Western Australia</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D61FD46FFB8FFEDCF150642FC9DF9D4" box="[777,893,1598,1624]" pageId="62" pageNumber="63" type="juvenile">1 juvenile</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BBCACB5FFB8FFEDCF960642FBF7F9D4" box="[906,1047,1598,1624]" pageId="62" pageNumber="63">Robe Valley</collectingMunicipality>
|
||
, ca.
|
||
<quantity id="4C9F9B2AFFB8FFEDC8500642FB75F9D4" box="[1100,1173,1598,1624]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="62" pageNumber="63" unit="km" value="27.0">27 km</quantity>
|
||
ESE. of
|
||
<location id="8EB86014FFB8FFEDC91C0642FA78F9D4" LSID="urn:lsid:plazi:treatment:03CE87D9FFB8FF93CC8B0483FD02FCB5:8EB86014FFB8FFEDC91C0642FA78F9D4" box="[1280,1432,1598,1624]" country="Australia" latitude="-21.741997" longLatPrecision="1" longitude="116.56163" municipality="Robe Valley" name="Pannawonica" pageId="62" pageNumber="63" stateProvince="Western Australia">Pannawonica</location>
|
||
,
|
||
<geoCoordinate id="EE535008FFB8FFEDCC8B061EFEA1F9F0" box="[151,321,1634,1660]" degrees="21" direction="south" minutes="44" orientation="latitude" pageId="62" pageNumber="63" precision="1" seconds="31.19" value="-21.741997">21°44’31.19”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FFB8FFEDCD53061EFDE8F9F1" box="[335,520,1634,1661]" degrees="116" direction="east" minutes="33" orientation="longitude" pageId="62" pageNumber="63" precision="1" seconds="41.85" value="116.56163">116°33’41.85”E</geoCoordinate>
|
||
,
|
||
<date id="FFD9100FFFB8FFEDCE09061EFCF7F9F1" box="[533,791,1634,1661]" pageId="62" pageNumber="63" value="2015-06-06" valueMax="2015-08-08" valueMin="2015-06-06">
|
||
<collectingDate id="EF9DE9E7FFB8FFEDCE09061EFCF7F9F1" box="[533,791,1634,1661]" pageId="62" pageNumber="63" value="2015-06-06" valueMax="2015-08-08" valueMin="2015-06-06">6 June–8 August 2015</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="52264ED8FFB8FFEDCF38061EFC3CF9F0" box="[804,988,1634,1660]" pageId="62" pageNumber="63">troglofauna trap</collectingMethod>
|
||
,
|
||
<collectorName id="26925319FFB8FFEDCFF6061EFB98F9F0" box="[1002,1144,1634,1660]" pageId="62" pageNumber="63">J. Alexander</collectorName>
|
||
(
|
||
<collectionCode id="ED76AE0AFFB8FFEDC894061EFB2FF9F0" box="[1160,1231,1634,1660]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="62" pageNumber="63" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FFB8FFEDC8CB061EFADDF9F1" box="[1239,1341,1634,1661]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="62" pageNumber="63">T138518</specimenCode>
|
||
) (DNA:
|
||
<collectionCode id="ED76AE0AFFB8FFEDCC8B06FAFF2AF92C" box="[151,202,1670,1696]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="62" pageNumber="63" type="Herbarium">COI</collectionCode>
|
||
, 18S,
|
||
<specimenCode id="DBC19EB4FFB8FFEDCD0906FAFEB2F92C" box="[277,338,1670,1696]" collectionCode="ITS" pageId="62" pageNumber="63">ITS2</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0F3C92FFB8FFEDCD7406FAFB32F948" ID-GBIF-Occurrence="3013449407" collectingDate="2015-06-06" collectingDateMax="2015-08-08" collectingDateMin="2015-06-06" collectionCode="WAM, COI" collectorName="J. Alexander" country="Australia" latitude="-21.743767" location="Pannawonica" longLatPrecision="1" longitude="116.563644" municipality="Robe Valley" pageId="62" pageNumber="63" specimenCode="T138519, ITS2" specimenCount="1" specimenCount-juvenile="1" stateProvince="Western Australia">
|
||
<specimenCount id="9D61FD46FFB8FFEDCD7406FAFE3DF92C" box="[360,477,1670,1696]" pageId="62" pageNumber="63" type="juvenile">1 juvenile</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BBCACB5FFB8FFEDCDF006FAFD99F92C" box="[492,633,1670,1696]" pageId="62" pageNumber="63">Robe Valley</collectingMunicipality>
|
||
, ca.
|
||
<quantity id="4C9F9B2AFFB8FFEDCEAD06FAFD1BF92C" box="[689,763,1670,1696]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="62" pageNumber="63" unit="km" value="27.0">27 km</quantity>
|
||
ESE. of
|
||
<location id="8EB86014FFB8FFEDCF7506FAFBE1F92C" LSID="urn:lsid:plazi:treatment:03CE87D9FFB8FF93CC8B0483FD02FCB5:8EB86014FFB8FFEDCF7506FAFBE1F92C" box="[873,1025,1670,1696]" country="Australia" latitude="-21.743767" longLatPrecision="1" longitude="116.563644" municipality="Robe Valley" name="Pannawonica" pageId="62" pageNumber="63" stateProvince="Western Australia">Pannawonica</location>
|
||
,
|
||
<geoCoordinate id="EE535008FFB8FFEDC80C06FAFB5AF92D" box="[1040,1210,1670,1697]" degrees="21" direction="south" minutes="44" orientation="latitude" pageId="62" pageNumber="63" precision="1" seconds="37.56" value="-21.743767">21°44’37.56”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FFB8FFEDC8D406FAFA60F92C" box="[1224,1408,1670,1696]" degrees="116" direction="east" minutes="33" orientation="longitude" pageId="62" pageNumber="63" precision="1" seconds="49.11" value="116.563644">116°33’49.11”E</geoCoordinate>
|
||
,
|
||
<date id="FFD9100FFFB8FFEDC99206FAFE63F949" pageId="62" pageNumber="63" value="2015-06-06" valueMax="2015-08-08" valueMin="2015-06-06">
|
||
<collectingDate id="EF9DE9E7FFB8FFEDC99206FAFE63F949" pageId="62" pageNumber="63" value="2015-06-06" valueMax="2015-08-08" valueMin="2015-06-06">6 June–8 August 2015</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="52264ED8FFB8FFEDCD8C06D6FDA8F948" box="[400,584,1706,1732]" pageId="62" pageNumber="63">troglofauna trap</collectingMethod>
|
||
,
|
||
<collectorName id="26925319FFB8FFEDCE4A06D6FD04F948" box="[598,740,1706,1732]" pageId="62" pageNumber="63">J. Alexander</collectorName>
|
||
(
|
||
<collectionCode id="ED76AE0AFFB8FFEDCEE806D6FCDBF948" box="[756,827,1706,1732]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="62" pageNumber="63" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FFB8FFEDCF5F06D6FC4AF949" box="[835,938,1706,1733]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="62" pageNumber="63">T138519</specimenCode>
|
||
) (DNA:
|
||
<collectionCode id="ED76AE0AFFB8FFEDC80E06D6FBA5F948" box="[1042,1093,1706,1732]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="62" pageNumber="63" type="Herbarium">COI</collectionCode>
|
||
, 18S,
|
||
<specimenCode id="DBC19EB4FFB8FFEDC89206D6FB2AF948" box="[1166,1226,1706,1732]" collectionCode="ITS" pageId="62" pageNumber="63">ITS2</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0F3C92FFB8FFEDC8FC06D6FE10F880" ID-GBIF-Occurrence="3013449327" collectingDate="2015-06-06" collectingDateMax="2015-08-08" collectingDateMin="2015-06-06" collectionCode="WAM" collectorName="J. Alexander" country="Australia" latitude="-21.741997" location="Pannawonica" longLatPrecision="1" longitude="116.56163" municipality="Robe Valley" pageId="62" pageNumber="63" specimenCode="T139921" specimenCount="1" specimenCount-juvenile="1" stateProvince="Western Australia">
|
||
<specimenCount id="9D61FD46FFB8FFEDC8FC06D6FAB4F948" box="[1248,1364,1706,1732]" pageId="62" pageNumber="63" type="juvenile">1 juvenile</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BBCACB5FFB8FFEDC97D06D6FF3FF964" pageId="62" pageNumber="63">Robe Valley</collectingMunicipality>
|
||
ca.
|
||
<quantity id="4C9F9B2AFFB8FFEDCD1106B2FEB5F964" box="[269,341,1742,1768]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="62" pageNumber="63" unit="km" value="27.0">27 km</quantity>
|
||
ESE. of
|
||
<location id="8EB86014FFB8FFEDCDA606B2FDB2F964" LSID="urn:lsid:plazi:treatment:03CE87D9FFB8FF93CC8B0483FD02FCB5:8EB86014FFB8FFEDCDA606B2FDB2F964" box="[442,594,1742,1768]" country="Australia" latitude="-21.741997" longLatPrecision="1" longitude="116.56163" municipality="Robe Valley" name="Pannawonica" pageId="62" pageNumber="63" stateProvince="Western Australia">Pannawonica</location>
|
||
,
|
||
<geoCoordinate id="EE535008FFB8FFEDCE4206B2FCE8F964" box="[606,776,1742,1768]" degrees="21" direction="south" minutes="44" orientation="latitude" pageId="62" pageNumber="63" precision="1" seconds="31.19" value="-21.741997">21°44’31.19”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FFB8FFEDCF0806B2FC2DF965" box="[788,973,1742,1769]" degrees="116" direction="east" minutes="33" orientation="longitude" pageId="62" pageNumber="63" precision="1" seconds="41.85" value="116.56163">116°33’41.85”E</geoCoordinate>
|
||
,
|
||
<date id="FFD9100FFFB8FFEDCFC406B2FB36F965" box="[984,1238,1742,1769]" pageId="62" pageNumber="63" value="2015-06-06" valueMax="2015-08-08" valueMin="2015-06-06">
|
||
<collectingDate id="EF9DE9E7FFB8FFEDCFC406B2FB36F965" box="[984,1238,1742,1769]" pageId="62" pageNumber="63" value="2015-06-06" valueMax="2015-08-08" valueMin="2015-06-06">6 June–8 August 2015</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="52264ED8FFB8FFEDC8FD06B2FA78F964" box="[1249,1432,1742,1768]" pageId="62" pageNumber="63">troglofauna trap</collectingMethod>
|
||
,
|
||
<collectorName id="26925319FFB8FFEDCC8B068EFEC4F880" box="[151,292,1778,1804]" pageId="62" pageNumber="63">J. Alexander</collectorName>
|
||
(
|
||
<collectionCode id="ED76AE0AFFB8FFEDCD2D068EFE98F880" box="[305,376,1778,1804]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="62" pageNumber="63" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FFB8FFEDCD62068EFE09F880" box="[382,489,1778,1804]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="62" pageNumber="63">T139921</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFB8FFEDCCDB076AFDF6F868" pageId="62" pageNumber="63" type="diagnosis">
|
||
<paragraph id="8BD836CFFFB8FFEDCCDB076AFDF6F868" blockId="62.[151,1437,1490,2020]" pageId="62" pageNumber="63">
|
||
<emphasis id="B913EADDFFB8FFEDCCDB076AFEA2F8BC" bold="true" box="[199,322,1814,1840]" pageId="62" pageNumber="63">Diagnosis.</emphasis>
|
||
Males are unknown and the flagellum is missing from the only available female.
|
||
<taxonomicName id="4C674D4CFFB8FFEDC8C6076AFF3AF8D8" authorityName="Abrams and Harvey" authorityYear="2020" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="nosferatu">
|
||
<emphasis id="B913EADDFFB8FFEDC8C6076AFF3AF8D8" italics="true" pageId="62" pageNumber="63">Draculoides nosferatu</emphasis>
|
||
</taxonomicName>
|
||
is the only
|
||
<taxonomicName id="4C674D4CFFB8FFEDCD4B0746FE01F8D8" box="[343,481,1850,1876]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EADDFFB8FFEDCD4B0746FE01F8D8" box="[343,481,1850,1876]" italics="true" pageId="62" pageNumber="63">Draculoides</emphasis>
|
||
</taxonomicName>
|
||
species known that does not have a guard tooth on the movable finger of the chelicera.
|
||
<taxonomicName id="4C674D4CFFB8FFEDCC8B0722FE72F8F4" authorityName="Abrams and Harvey" authorityYear="2020" box="[151,402,1886,1912]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="nosferatu">
|
||
<emphasis id="B913EADDFFB8FFEDCC8B0722FE72F8F4" box="[151,402,1886,1912]" italics="true" pageId="62" pageNumber="63">Draculoides nosferatu</emphasis>
|
||
</taxonomicName>
|
||
can be diagnosed from all other
|
||
<taxonomicName id="4C674D4CFFB8FFEDCEE70722FC65F8F4" box="[763,901,1886,1912]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EADDFFB8FFEDCEE70722FC65F8F4" box="[763,901,1886,1912]" italics="true" pageId="62" pageNumber="63">Draculoides</emphasis>
|
||
</taxonomicName>
|
||
species that were sequenced at COI by the 50bp mini-barcode shown in
|
||
<figureCitation id="135C2A4AFFB8FFEDCDB807FEFDE5F810" box="[420,517,1922,1948]" captionStart="FIGURE 3" captionStartId="18.[151,250,1547,1571]" captionTargetBox="[158,1420,181,1521]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416857" httpUri="https://zenodo.org/record/4416857/files/figure.png" pageId="62" pageNumber="63">Figure 3</figureCitation>
|
||
.
|
||
<taxonomicName id="4C674D4CFFB8FFEDCE0E07FEFCEFF810" authorityName="Abrams and Harvey" authorityYear="2020" box="[530,783,1922,1948]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="nosferatu">
|
||
<emphasis id="B913EADDFFB8FFEDCE0E07FEFCEFF810" box="[530,783,1922,1948]" italics="true" pageId="62" pageNumber="63">Draculoides nosferatu</emphasis>
|
||
</taxonomicName>
|
||
can be diagnosed from all other
|
||
<taxonomicName id="4C674D4CFFB8FFEDC89B07FEFAF1F810" box="[1159,1297,1922,1948]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EADDFFB8FFEDC89B07FEFAF1F810" box="[1159,1297,1922,1948]" italics="true" pageId="62" pageNumber="63">Draculoides</emphasis>
|
||
</taxonomicName>
|
||
species that were sequenced at ITS2 (
|
||
<figureCitation id="135C2A4AFFB8FFEDCDAD07DAFE14F84C" box="[433,500,1958,1984]" captionStart="FIGURE 7" captionStartId="22.[151,250,1755,1779]" captionTargetBox="[159,1425,181,1729]" captionTargetId="figure-20@22.[151,1436,181,1732]" captionTargetPageId="22" captionText="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416867" httpUri="https://zenodo.org/record/4416867/files/figure.png" pageId="62" pageNumber="63">Fig. 7</figureCitation>
|
||
) except for
|
||
<taxonomicName id="4C674D4CFFB8FFEDCE6707DBFD1CF84C" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[635,764,1958,1984]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="claudiae">
|
||
<emphasis id="B913EADDFFB8FFEDCE6707DBFD1CF84C" box="[635,764,1958,1984]" italics="true" pageId="62" pageNumber="63">D. claudiae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFB8FFEDCF1707DBFC77F84C" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[779,919,1958,1984]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="carmillae">
|
||
<emphasis id="B913EADDFFB8FFEDCF1707DBFC77F84C" box="[779,919,1958,1984]" italics="true" pageId="62" pageNumber="63">D. carmillae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C674D4CFFB8FFEDCFD207DBFB8BF84C" authorityName="Harvey, Berry, Edward and Humphreys" authorityYear="2008" box="[974,1131,1959,1984]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="mesozeirus">
|
||
<emphasis id="B913EADDFFB8FFEDCFD207DBFB8BF84C" box="[974,1131,1959,1984]" italics="true" pageId="62" pageNumber="63">D. mesozeirus</emphasis>
|
||
</taxonomicName>
|
||
, which are not distinguishable using the ITS2 mini-barcode.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFB9FF93CCDB00EAFE61FDB1" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64" type="description">
|
||
<paragraph id="8BD836CFFFB9FFECCCDB00EAFB41FF3D" blockId="63.[151,1437,150,501]" box="[199,1185,150,177]" pageId="63" pageNumber="64">
|
||
<emphasis id="B913EADDFFB9FFECCCDB00EAFDE7FF3C" bold="true" box="[199,519,150,176]" pageId="63" pageNumber="64">Description (adult female).</emphasis>
|
||
<emphasis id="B913EADDFFB9FFECCE1200EBFDBDFF3D" box="[526,605,151,177]" italics="true" pageId="63" pageNumber="64">Colour</emphasis>
|
||
. Yellow-brown; propeltidium and pedipalps darker.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFB9FFECCCDB00C7FC60FE91" blockId="63.[151,1437,150,501]" pageId="63" pageNumber="64">
|
||
<emphasis id="B913EADDFFB9FFECCCDB00C7FE8FFF59" box="[199,367,187,213]" italics="true" pageId="63" pageNumber="64">Cephalothorax</emphasis>
|
||
. Propeltidium with 2+1 apical setae in a triangular formation on anterior process and 2 + 2 + 2 setae; eye spots absent. Mesopeltidia separated. Metapeltidium divided. Anterior sternum with 14 setae (including 2 sternapophysial setae); posterior sternum triangular with 6 setae.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFB9FFECCCDB015BFBDBFE79" blockId="63.[151,1437,150,501]" pageId="63" pageNumber="64">
|
||
<emphasis id="B913EADDFFB9FFECCCDB015BFED5FECD" box="[199,309,295,321]" italics="true" pageId="63" pageNumber="64">Chelicera</emphasis>
|
||
. Fixed finger with 2 large teeth plus 5 smaller teeth between these, proximal and distal teeth without small lateral teeth; membranous area between fixed and movable fingers with 3 large, lanceolate, terminally pilose setae (G1); G2 composed of 6 setae; G3 composed of 4 setae; internal face of chelicera with 5 short whip-like setae (G4); brush at base of fixed finger composed of 7 setae (G5A), each densely pilose in distal half and G5B composed of 10 setae; G6 with one seta; G7 composed of 2 setae. Movable finger serrula composed of ca. 18 long lamellae, guard tooth absent; 3 accessory teeth present, most distal large, most proximal tiny.
|
||
</paragraph>
|
||
<caption id="DF186647FFB9FFECCC8B0413FC41FB27" ID-DOI="http://doi.org/10.5281/zenodo.4416939" ID-Zenodo-Dep="4416939" httpUri="https://zenodo.org/record/4416939/files/figure.png" pageId="63" pageNumber="64" startId="63.[151,250,1135,1159]" targetBox="[151,1436,525,1111]" targetPageId="63">
|
||
<paragraph id="8BD836CFFFB9FFECCC8B0413FC41FB27" blockId="63.[151,1436,1135,1195]" pageId="63" pageNumber="64">
|
||
<emphasis id="B913EADDFFB9FFECCC8B0413FEC1FB0B" bold="true" box="[151,289,1135,1159]" pageId="63" pageNumber="64">FIGURE 36.</emphasis>
|
||
<taxonomicName id="4C674D4CFFB9FFECCD340413FDECFB0B" authority="Abrams & Huey & Hillyer & Didham & Harvey, 2020" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[296,524,1135,1159]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="nosferatu" status="sp. nov.">
|
||
<emphasis id="B913EADDFFB9FFECCD340413FDECFB0B" box="[296,524,1135,1159]" italics="true" pageId="63" pageNumber="64">Draculoides nosferatu</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B913EADDFFB9FFECCE0F0413FD85FB0B" bold="true" box="[531,613,1135,1159]" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="A22057A6FFB9FFECCE0F0413FD85FB0B" box="[531,613,1135,1159]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype female (WAM T138516): A. Body, dorsal; B. Body, ventral; C. Body, lateral. The scale bar shown in image A is 1 mm and also applies to B and C.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF186647FFB9FFECCC8B072FFF26F803" ID-DOI="http://doi.org/10.5281/zenodo.4416941" ID-Zenodo-Dep="4416941" httpUri="https://zenodo.org/record/4416941/files/figure.png" pageId="63" pageNumber="64" startId="63.[151,250,1875,1899]" targetBox="[407,1176,1225,1845]" targetPageId="63">
|
||
<paragraph id="8BD836CFFFB9FFECCC8B072FFF26F803" blockId="63.[151,1437,1875,1935]" pageId="63" pageNumber="64">
|
||
<emphasis id="B913EADDFFB9FFECCC8B072FFEC3F8E7" bold="true" box="[151,291,1875,1899]" pageId="63" pageNumber="64">FIGURE 37.</emphasis>
|
||
<taxonomicName id="4C674D4CFFB9FFECCD31072FFDF3F8E7" authority="Abrams & Huey & Hillyer & Didham & Harvey, 2020" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[301,531,1875,1899]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="nosferatu" status="sp. nov.">
|
||
<emphasis id="B913EADDFFB9FFECCD31072FFDF3F8E7" box="[301,531,1875,1899]" italics="true" pageId="63" pageNumber="64">Draculoides nosferatu</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B913EADDFFB9FFECCE010728FD91F8E0" bold="true" box="[541,625,1876,1900]" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="A22057A6FFB9FFECCE010728FD91F8E0" box="[541,625,1876,1900]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype female (WAM T138516): A. Spermathecae, ventral. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD836CFFFC6FF93CCDB00EBFA6FFF59" blockId="64.[151,1437,151,825]" pageId="64" pageNumber="65">
|
||
<emphasis id="B913EADDFFC6FF93CCDB00EBFECCFF3D" box="[199,300,151,177]" italics="true" pageId="64" pageNumber="65">Pedipalp</emphasis>
|
||
. Without apophyses; trochanter with sharply produced ventro-distal extension, ventral margin with ca. 9 stout setae, without mesal spur; tarsus and tibia without spines; tarsal spur present; claw 0.36 × length of tarsus.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFC6FF93CCDB00A3FB0EFF75" blockId="64.[151,1437,151,825]" box="[199,1262,223,249]" pageId="64" pageNumber="65">
|
||
<emphasis id="B913EADDFFC6FF93CCDB00A3FEE4FF74" box="[199,260,223,248]" italics="true" pageId="64" pageNumber="65">Legs.</emphasis>
|
||
Tarsus I with 6 segments; baso-dorsal margin of femur IV produced at about a 90° angle.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFC6FF93CCDB017FFBABFECD" blockId="64.[151,1437,151,825]" pageId="64" pageNumber="65">
|
||
<emphasis id="B913EADDFFC6FF93CCDB017FFED1FE91" box="[199,305,259,285]" italics="true" pageId="64" pageNumber="65">Abdomen</emphasis>
|
||
. Chaetotaxy of tergite I: 2 macrosetae + 4 microsetae (microsetae diagonal), tergite II:? (damaged) macrosetae + 6 microsetae (microsetae in column), tergites III–IX: 2: 2: 2: 2: 2: 2: 2.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFC6FF93CCDB0137FDA2FE21" blockId="64.[151,1437,151,825]" pageId="64" pageNumber="65">
|
||
<emphasis id="B913EADDFFC6FF93CCDB0137FE66FEE9" box="[199,390,331,357]" italics="true" pageId="64" pageNumber="65">Female genitalia</emphasis>
|
||
. Two pairs of spermathecae with short, equal-sized lobes, each pair connected basally before connection with bursa (
|
||
<figureCitation id="135C2A4AFFC6FF93CDB80113FDEDFE05" box="[420,525,367,393]" captionStart="FIGURE 37" captionStartId="63.[151,250,1875,1899]" captionTargetBox="[407,1176,1225,1845]" captionTargetId="figure-222@63.[407,1179,1219,1852]" captionTargetPageId="63" captionText="FIGURE 37. Draculoides nosferatu sp. nov., holotype female (WAM T138516): A. Spermathecae, ventral. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4416941" httpUri="https://zenodo.org/record/4416941/files/figure.png" pageId="64" pageNumber="65">Fig. 37A</figureCitation>
|
||
), distally round and smooth; sparsely covered with small pores; gonopod short, rectangular, distally slightly bifurcate.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFC6FF93CCDB01CBFD6AFE5D" blockId="64.[151,1437,151,825]" box="[199,650,439,465]" pageId="64" pageNumber="65">
|
||
<emphasis id="B913EADDFFC6FF93CCDB01CBFEDAFE5D" box="[199,314,439,465]" italics="true" pageId="64" pageNumber="65">Flagellum</emphasis>
|
||
. Collected without flagellum.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFC6FF93CCDB01A7FE61FDB1" blockId="64.[151,1437,151,825]" pageId="64" pageNumber="65">
|
||
<emphasis id="B913EADDFFC6FF93CCDB01A7FE70FE79" box="[199,400,475,501]" italics="true" pageId="64" pageNumber="65">Dimensions (mm)</emphasis>
|
||
.
|
||
<typeStatus id="54DC886DFFC6FF93CDBD01A7FDE9FE79" box="[417,521,475,501]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
|
||
female (WAM T138516): Body length 3.56. Propeltidium 1.21/0.62. Chelicera 0.69. Flagellum broken. Pedipalp: trochanter 0.46, femur 0.50, patella 0.54, tibia 0.44, tarsus 0.27, claw 0.10, total excluding claw 2.21.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFC6FF93CCDB023BFCFAFD7D" pageId="64" pageNumber="65" type="discussion">
|
||
<paragraph id="8BD836CFFFC6FF93CCDB023BFACAFD41" blockId="64.[151,1437,151,825]" pageId="64" pageNumber="65">
|
||
<emphasis id="B913EADDFFC6FF93CCDB023BFEDBFDED" bold="true" box="[199,315,583,609]" pageId="64" pageNumber="65">Remarks.</emphasis>
|
||
<taxonomicName id="4C674D4CFFC6FF93CD5F023BFDDFFDED" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[323,575,583,609]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="nosferatu">
|
||
<emphasis id="B913EADDFFC6FF93CD5F023BFDDFFDED" box="[323,575,583,609]" italics="true" pageId="64" pageNumber="65">Draculoides nosferatu</emphasis>
|
||
</taxonomicName>
|
||
is known from three locations within the most southern mesa of the Middle Robe mesas (
|
||
<figureCitation id="135C2A4AFFC6FF93CD330217FE69FD09" box="[303,393,619,645]" captionStart="FIGURE 1" captionStartId="16.[151,250,1868,1892]" captionTargetBox="[346,1243,188,1844]" captionTargetId="figure-20@16.[339,1249,181,1844]" captionTargetPageId="16" captionText="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." figureDoi="http://doi.org/10.5281/zenodo.4416851" httpUri="https://zenodo.org/record/4416851/files/figure.png" pageId="64" pageNumber="65">Fig. 1D</figureCitation>
|
||
). The phylogeny suggests that
|
||
<taxonomicName id="4C674D4CFFC6FF93CEF50217FC96FD09" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[745,886,619,645]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="nosferatu">
|
||
<emphasis id="B913EADDFFC6FF93CEF50217FC96FD09" box="[745,886,619,645]" italics="true" pageId="64" pageNumber="65">D. nosferatu</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B913EADDFFC6FF93CF630217FC6AFD08" box="[895,906,619,644]" italics="true" pageId="64" pageNumber="65">s</emphasis>
|
||
closest relatives are the other species from the Middle Robe area including
|
||
<taxonomicName id="4C674D4CFFC6FF93CDC402F3FDDFFD24" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[472,575,655,680]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="minae">
|
||
<emphasis id="B913EADDFFC6FF93CDC402F3FDDFFD24" box="[472,575,655,680]" italics="true" pageId="64" pageNumber="65">D. minae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFC6FF93CE5202F3FD30FD25" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[590,720,655,681]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="claudiae">
|
||
<emphasis id="B913EADDFFC6FF93CE5202F3FD30FD25" box="[590,720,655,681]" italics="true" pageId="64" pageNumber="65">D. claudiae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFC6FF93CEC302F3FC8CFD25" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[735,876,655,681]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="carmillae">
|
||
<emphasis id="B913EADDFFC6FF93CEC302F3FC8CFD25" box="[735,876,655,681]" italics="true" pageId="64" pageNumber="65">D. carmillae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C674D4CFFC6FF93CFBF02F3FBA2FD24" authorityName="Harvey, Berry, Edward and Humphreys" authorityYear="2008" box="[931,1090,655,680]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="mesozeirus">
|
||
<emphasis id="B913EADDFFC6FF93CFBF02F3FBA2FD24" box="[931,1090,655,680]" italics="true" pageId="64" pageNumber="65">D. mesozeirus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="135C2A4AFFC6FF93C84E02F3FB74FD25" box="[1106,1172,655,681]" captionStart="FIGURE 2" captionStartId="17.[151,250,1910,1934]" captionTargetBox="[332,1254,181,1881]" captionTargetId="figure-17@17.[332,1254,181,1886]" captionTargetPageId="17" captionText="FIGURE 2. Maximum Likelihood tree, based on the reduced dataset. See Methods for details. Each terminal represents a single species or OTU, with named species in bold. Clades are identified using dashed boxes and shaded boxes encompass the species discussed in this study. Nodes with bootstrap support <80 are not shown, except for the common ancestor of the Draculoides radiation." figureDoi="http://doi.org/10.5281/zenodo.4416855" httpUri="https://zenodo.org/record/4416855/files/figure.png" pageId="64" pageNumber="65">Fig. 2</figureCitation>
|
||
Clade H). The juvenile specimens listed above are associated with this species by locality and, in many cases, by sequence data.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFC6FF93CCDB02AAFCFAFD7D" blockId="64.[151,1437,151,825]" box="[199,794,726,753]" pageId="64" pageNumber="65">
|
||
<emphasis id="B913EADDFFC6FF93CCDB02AAFE81FD7D" bold="true" box="[199,353,726,753]" pageId="64" pageNumber="65">Other names</emphasis>
|
||
. WAM SCH061 (
|
||
<bibRefCitation id="EFF64B3EFFC6FF93CE3502ABFCEEFD7D" author="Abrams, K. M. & Huey, J. A. & Hillyer, M. J. & Humphreys, W. F. & Didham, R. K. & Harvey, M. S." box="[553,782,727,753]" pageId="64" pageNumber="65" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. & Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
|
||
Abrams
|
||
<emphasis id="B913EADDFFC6FF93CE9702ABFD5CFD7D" box="[651,700,727,753]" italics="true" pageId="64" pageNumber="65">et al</emphasis>
|
||
., 2019
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFC6FF93CCDB0287FD02FCB5" pageId="64" pageNumber="65" type="etymology">
|
||
<paragraph id="8BD836CFFFC6FF93CCDB0287FD02FCB5" blockId="64.[151,1437,151,825]" pageId="64" pageNumber="65">
|
||
<emphasis id="B913EADDFFC6FF93CCDB0287FEAEFC99" bold="true" box="[199,334,763,789]" pageId="64" pageNumber="65">Etymology.</emphasis>
|
||
The specific epithet refers to the seminal 1922 German horror film “Nosferatu: A Symphony of Horror,” and relates to the species living in the dark.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |