353 lines
49 KiB
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353 lines
49 KiB
XML
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<mods:title id="097B492EEC99C93EAF9499F14EF2F37D">A systematic revision of Draculoides (Schizomida: Hubbardiidae) of the Pilbara, Western Australia, Part I: the Western Pilbara</mods:title>
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<treatment id="03CE87D9FFA8FFE2CC8B0779FA61FB98" ID-DOI="http://doi.org/10.5281/zenodo.4428357" ID-GBIF-Taxon="176485727" ID-Zenodo-Dep="4428357" LSID="urn:lsid:zoobank.org:act:4E183B9C-D352-4805-82DA-60C5B9EFA783" httpUri="http://treatment.plazi.org/id/03CE87D9FFA8FFE2CC8B0779FA61FB98" lastPageId="49" lastPageNumber="50" pageId="46" pageNumber="47">
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<taxonomicName id="4C674D4CFFA8FFFDCC8B0779FD5BF893" authority="Abrams and Harvey" authorityName="Abrams and Harvey" authorityYear="2020" box="[151,699,1797,1824]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="immortalis" status="sp. nov.">
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<emphasis id="B913EADDFFA8FFFDCC8B0779FE53F8AC" bold="true" box="[151,435,1797,1824]" italics="true" pageId="46" pageNumber="47">Draculoides immortalis</emphasis>
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Abrams and Harvey
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,
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<taxonomicNameLabel id="A22057A6FFA8FFFDCED40779FCEAF893" box="[712,778,1797,1823]" pageId="46" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C37D6544FFA8FFFDCC8B0755FCBDF826" pageId="46" pageNumber="47" type="description">
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<paragraph id="8BD836CFFFA8FFFDCC8B0755FE86F8C8" blockId="46.[151,1312,1797,1896]" box="[151,358,1833,1860]" pageId="46" pageNumber="47">
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(
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<figureCitation id="135C2A4AFFA8FFFDCC830755FEEBF8CF" box="[159,267,1833,1859]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="16.[151,250,1868,1892]" captionStartId-1="17.[151,250,1910,1934]" captionStartId-2="18.[151,250,1547,1571]" captionStartId-3="19.[151,250,1882,1906]" captionStartId-4="20.[151,250,1216,1240]" captionStartId-5="21.[151,250,1709,1733]" captionStartId-6="22.[151,250,1755,1779]" captionStartId-7="23.[151,250,1919,1943]" captionTargetBox-0="[346,1243,188,1844]" captionTargetBox-1="[332,1254,181,1881]" captionTargetBox-2="[158,1420,181,1521]" captionTargetBox-3="[151,1425,182,1858]" captionTargetBox-4="[158,1424,183,1189]" captionTargetBox-5="[155,1423,181,1683]" captionTargetBox-6="[159,1425,181,1729]" captionTargetBox-7="[151,1430,181,1893]" captionTargetId-0="figure-20@16.[339,1249,181,1844]" captionTargetId-1="figure-17@17.[332,1254,181,1886]" captionTargetId-2="figure-20@18.[151,1436,181,1524]" captionTargetId-3="figure-17@19.[151,1436,181,1859]" captionTargetId-4="figure-20@20.[151,1436,181,1192]" captionTargetId-5="figure-17@21.[155,1432,181,1685]" captionTargetId-6="figure-20@22.[151,1436,181,1732]" captionTargetId-7="figure-17@23.[151,1436,181,1894]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="22" captionTargetPageId-7="23" captionText-0="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." captionText-1="FIGURE 2. Maximum Likelihood tree, based on the reduced dataset. See Methods for details. Each terminal represents a single species or OTU, with named species in bold. Clades are identified using dashed boxes and shaded boxes encompass the species discussed in this study. Nodes with bootstrap support <80 are not shown, except for the common ancestor of the Draculoides radiation." captionText-2="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-3="FIGURE 4. Maximum Likelihood tree based on COI mini-barcodes (Fig. 3) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-4="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-5="FIGURE 6. Maximum Likelihood tree based on COI mini-barcodes (Fig. 5) showing similarity of mini-barcode sequences relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-6="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-7="FIGURE 8. Maximum Likelihood tree based on ITS2 mini-barcodes (Fig. 7) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." figureDoi-0="http://doi.org/10.5281/zenodo.4416851" figureDoi-1="http://doi.org/10.5281/zenodo.4416855" figureDoi-2="http://doi.org/10.5281/zenodo.4416857" figureDoi-3="http://doi.org/10.5281/zenodo.4416859" figureDoi-4="http://doi.org/10.5281/zenodo.4416863" figureDoi-5="http://doi.org/10.5281/zenodo.4416865" figureDoi-6="http://doi.org/10.5281/zenodo.4416867" figureDoi-7="http://doi.org/10.5281/zenodo.4416869" httpUri-0="https://zenodo.org/record/4416851/files/figure.png" httpUri-1="https://zenodo.org/record/4416855/files/figure.png" httpUri-2="https://zenodo.org/record/4416857/files/figure.png" httpUri-3="https://zenodo.org/record/4416859/files/figure.png" httpUri-4="https://zenodo.org/record/4416863/files/figure.png" httpUri-5="https://zenodo.org/record/4416865/files/figure.png" httpUri-6="https://zenodo.org/record/4416867/files/figure.png" httpUri-7="https://zenodo.org/record/4416869/files/figure.png" pageId="46" pageNumber="47">Figs. 1–8</figureCitation>
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,
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<figureCitation id="135C2A4AFFA8FFFDCD0B0755FEBFF8C8" box="[279,351,1833,1860]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStartId-0="47.[151,250,1828,1852]" captionStartId-1="48.[151,250,1869,1893]" captionTargetBox-0="[156,1431,669,1803]" captionTargetBox-1="[149,1435,525,1843]" captionTargetId-0="figure-255@47.[156,1431,669,1804]" captionTargetId-1="figure-181@48.[149,1437,525,1845]" captionTargetPageId-0="47" captionTargetPageId-1="48" captionText-0="FIGURE 24. Draculoides immortalis sp. nov., holotype female (WAM T138476): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F." captionText-1="FIGURE 25. Draculoides immortalis sp. nov., holotype female (WAM T138476): A. Flagellum, dorsal; B. Flagellum, ventral; C. Flagellum, lateral; D. Spermathecae, ventral. Scale bars: A–C: 0.1 mm, D: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4416911" figureDoi-1="http://doi.org/10.5281/zenodo.4416913" httpUri-0="https://zenodo.org/record/4416911/files/figure.png" httpUri-1="https://zenodo.org/record/4416913/files/figure.png" pageId="46" pageNumber="47">24–25</figureCitation>
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)
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</paragraph>
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<paragraph id="8BD836CFFFA8FFFDCC8B0731FAC0F8E4" blockId="46.[151,1312,1797,1896]" box="[151,1312,1869,1896]" pageId="46" pageNumber="47">
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Zoobank Code: http://zoobank.org/NomenclaturalActs/
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<uuid id="FFC10C1AFFA8FFFDCF150731FAC0F8E4" box="[777,1312,1869,1896]" pageId="46" pageNumber="47">4E183B9C-D352-4805-82DA-60C5B9EFA783</uuid>
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</paragraph>
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<paragraph id="8BD836CFFFA8FFFDCC8B07EEFCBDF826" blockId="46.[151,861,1937,1962]" box="[151,861,1937,1962]" pageId="46" pageNumber="47">
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<taxonomicName id="4C674D4CFFA8FFFDCC8B07EEFE9CF825" authority="SCH" authorityName="SCH" box="[151,380,1937,1962]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis id="B913EADDFFA8FFFDCC8B07EEFEA1F826" box="[151,321,1938,1962]" italics="true" pageId="46" pageNumber="47">Paradraculoides</emphasis>
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SCH
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</taxonomicName>
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058:
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<bibRefCitation id="EFF64B3EFFA8FFFDCDB707EEFD8EF826" author="Abrams, K. M. & Huey, J. A. & Hillyer, M. J. & Humphreys, W. F. & Didham, R. K. & Harvey, M. S." box="[427,622,1938,1962]" pageId="46" pageNumber="47" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. & Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
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Abrams
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||
<emphasis id="B913EADDFFA8FFFDCE1E07EEFDCEF826" box="[514,558,1938,1962]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
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||
. 2019
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||
</bibRefCitation>
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||
MPE 106532: 8, fig. 2.
|
||
</paragraph>
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<emphasis id="B913EADDFFA8FFFDCC8B07A4FE60F87E" bold="true" box="[151,384,2008,2034]" pageId="46" pageNumber="47">Material examined.</emphasis>
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<materialsCitation id="3B0F3C92FFA8FFFDCD9B07A4FA7DF87E" ID-GBIF-Occurrence="3013449449" box="[391,1437,2008,2034]" country="Australia" location="Robe Valley" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
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<emphasis id="B913EADDFFA8FFFDCD9B07A4FDA6F87E" box="[391,582,2008,2034]" italics="true" pageId="46" pageNumber="47">
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<typeStatus id="54DC886DFFA8FFFDCD9B07A4FE0EF87E" box="[391,494,2008,2034]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
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female.
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</emphasis>
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<collectingCountry id="F370765FFFA8FFFDCE5607A4FD0DF87E" box="[586,749,2008,2034]" name="Australia" pageId="46" pageNumber="47">AUSTRALIA</collectingCountry>
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:
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<emphasis id="B913EADDFFA8FFFDCEEB07A5FC5FF87E" box="[759,959,2008,2034]" italics="true" pageId="46" pageNumber="47">
|
||
<collectingRegion id="49A3F82DFFA8FFFDCEEB07A5FC5FF87E" box="[759,959,2008,2034]" country="Australia" name="Western Australia" pageId="46" pageNumber="47">Western Australia</collectingRegion>
|
||
</emphasis>
|
||
: “Budgiebore”,
|
||
<location id="8EB86014FFA8FFFDC86A07A4FAE0F87E" LSID="urn:lsid:plazi:treatment:03CE87D9FFA8FFE2CC8B0779FA61FB98:8EB86014FFA8FFFDC86A07A4FAE0F87E" box="[1142,1280,2008,2034]" country="Australia" name="Robe Valley" pageId="46" pageNumber="47" stateProvince="Western Australia">Robe Valley</location>
|
||
, ca.
|
||
<quantity id="4C9F9B2AFFA8FFFDC92D07A4FA98F87E" box="[1329,1400,2008,2034]" metricMagnitude="4" metricUnit="m" metricValue="4.8" pageId="46" pageNumber="47" unit="km" value="48.0">48 km</quantity>
|
||
W.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFA9FFFCCC8B00EBFB6BFF59" blockId="47.[151,1436,150,645]" pageId="47" pageNumber="48">
|
||
<materialsCitation id="3B0F3C92FFA9FFFCCC8B00EBFB67FF59" ID-GBIF-Occurrence="3013449412" collectingDate="2015-06-06" collectionCode="WAM, COI" collectorName="J. Alexander & P. Brooshooft" latitude="-21.558722" longLatPrecision="1" longitude="115.86514" pageId="47" pageNumber="48" specimenCode="T138476, ITS2" specimenCount="1">
|
||
of Pannawonica,
|
||
<geoCoordinate id="EE535008FFA9FFFCCD4500EBFE15FF3D" box="[345,501,151,177]" degrees="21" direction="south" minutes="33" orientation="latitude" pageId="47" pageNumber="48" precision="1" seconds="31.4" value="-21.558722">21°33’31.4”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FFA9FFFCCE1D00EAFD5AFF3D" box="[513,698,150,177]" degrees="115" direction="east" minutes="51" orientation="longitude" pageId="47" pageNumber="48" precision="1" seconds="54.52" value="115.86514">115°51’54.52”E</geoCoordinate>
|
||
,
|
||
<date id="FFD9100FFFA9FFFCCED900EBFCB0FF3D" box="[709,848,150,177]" pageId="47" pageNumber="48" value="2015-06-06">
|
||
<collectingDate id="EF9DE9E7FFA9FFFCCED900EBFCB0FF3D" box="[709,848,150,177]" pageId="47" pageNumber="48" value="2015-06-06">6 June 2015</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="52264ED8FFA9FFFCCF4700EBFBF1FF3D" box="[859,1041,151,177]" pageId="47" pageNumber="48">troglofauna trap</collectingMethod>
|
||
,
|
||
<collectorName id="26925319FFA9FFFCC80100EBFB4BFF3D" box="[1053,1195,151,177]" pageId="47" pageNumber="48">J. Alexander</collectorName>
|
||
,
|
||
<collectorName id="26925319FFA9FFFCC8AA00EBFAAFFF3D" box="[1206,1359,151,177]" pageId="47" pageNumber="48">P. Brooshooft</collectorName>
|
||
(Biota Environmental Sciences, Budgiebore-20150604-06) (
|
||
<collectionCode id="ED76AE0AFFA9FFFCCEF100C7FCD4FF59" box="[749,820,187,213]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="47" pageNumber="48" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FFA9FFFCCF2600C7FC42FF5A" box="[826,930,187,214]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="47" pageNumber="48">T138476</specimenCode>
|
||
) (DNA:
|
||
<collectionCode id="ED76AE0AFFA9FFFCC81900C7FBD8FF59" box="[1029,1080,187,213]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="47" pageNumber="48" type="Herbarium">COI</collectionCode>
|
||
,
|
||
<specimenCode id="DBC19EB4FFA9FFFCC85F00C7FB60FF59" box="[1091,1152,187,213]" collectionCode="ITS" pageId="47" pageNumber="48">ITS2</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFA9FFFCCCDB00A2FC5CFDED" pageId="47" pageNumber="48" type="diagnosis">
|
||
<paragraph id="8BD836CFFFA9FFFCCCDB00A2FC5CFDED" blockId="47.[151,1436,150,645]" pageId="47" pageNumber="48">
|
||
<emphasis id="B913EADDFFA9FFFCCCDB00A2FEA2FF74" bold="true" box="[199,322,222,248]" pageId="47" pageNumber="48">Diagnosis.</emphasis>
|
||
Males are unknown. Females of this species can be distinguished from all other species of
|
||
<taxonomicName id="4C674D4CFFA9FFFCC95700A3FF3AFE91" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EADDFFA9FFFCC95700A3FF3AFE91" italics="true" pageId="47" pageNumber="48">Draculoides</emphasis>
|
||
</taxonomicName>
|
||
by the distinctive form of the spermathecae in which the inner lobe is noticeably smaller and narrower than the outer lobe. In all other
|
||
<taxonomicName id="4C674D4CFFA9FFFCCDD3015BFDB9FECD" box="[463,601,295,321]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EADDFFA9FFFCCDD3015BFDB9FECD" box="[463,601,295,321]" italics="true" pageId="47" pageNumber="48">Draculoides</emphasis>
|
||
</taxonomicName>
|
||
, the spermathecal lobes are roughly the same size or the inner lobes are larger than the outer lobes. In
|
||
<taxonomicName id="4C674D4CFFA9FFFCCDE40137FDB3FEE8" authorityName="Harvey" authorityYear="1992" box="[504,595,331,356]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="vinei">
|
||
<emphasis id="B913EADDFFA9FFFCCDE40137FDB3FEE8" box="[504,595,331,356]" italics="true" pageId="47" pageNumber="48">D. vinei</emphasis>
|
||
</taxonomicName>
|
||
the inner lobe is slightly smaller than the outer lobe but it can be easily distinguished from
|
||
<taxonomicName id="4C674D4CFFA9FFFCCD680113FDF1FE05" authorityName="Abrams and Harvey" authorityYear="2020" box="[372,529,367,393]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="immortalis">
|
||
<emphasis id="B913EADDFFA9FFFCCD680113FDF1FE05" box="[372,529,367,393]" italics="true" pageId="47" pageNumber="48">D. immortalis</emphasis>
|
||
</taxonomicName>
|
||
by the absence of a gonopod and the spermathecal lobes are connected basally before connection with the bursa in the former species while the gonopod is present and the spermathecal lobes are not connected in the latter.
|
||
<taxonomicName id="4C674D4CFFA9FFFCCDD601CBFD35FE5D" authorityName="Abrams and Harvey" authorityYear="2020" box="[458,725,439,465]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="immortalis">
|
||
<emphasis id="B913EADDFFA9FFFCCDD601CBFD35FE5D" box="[458,725,439,465]" italics="true" pageId="47" pageNumber="48">Draculoides immortalis</emphasis>
|
||
</taxonomicName>
|
||
can be diagnosed from all other
|
||
<taxonomicName id="4C674D4CFFA9FFFCC85601CBFB34FE5D" box="[1098,1236,439,465]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EADDFFA9FFFCC85601CBFB34FE5D" box="[1098,1236,439,465]" italics="true" pageId="47" pageNumber="48">Draculoides</emphasis>
|
||
</taxonomicName>
|
||
species that were sequenced at COI by the 50bp mini-barcode shown in
|
||
<figureCitation id="135C2A4AFFA9FFFCCEE701A7FCBCFE79" box="[763,860,475,501]" captionStart="FIGURE 3" captionStartId="18.[151,250,1547,1571]" captionTargetBox="[158,1420,181,1521]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416857" httpUri="https://zenodo.org/record/4416857/files/figure.png" pageId="47" pageNumber="48">Figure 3</figureCitation>
|
||
.
|
||
<taxonomicName id="4C674D4CFFA9FFFCCF7401A7FB93FE79" authorityName="Abrams and Harvey" authorityYear="2020" box="[872,1139,475,501]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="immortalis">
|
||
<emphasis id="B913EADDFFA9FFFCCF7401A7FB93FE79" box="[872,1139,475,501]" italics="true" pageId="47" pageNumber="48">Draculoides immortalis</emphasis>
|
||
</taxonomicName>
|
||
can be diagnosed from all other
|
||
<taxonomicName id="4C674D4CFFA9FFFCCCC40183FE82FD95" box="[216,354,511,537]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EADDFFA9FFFCCCC40183FE82FD95" box="[216,354,511,537]" italics="true" pageId="47" pageNumber="48">Draculoides</emphasis>
|
||
</taxonomicName>
|
||
species that were sequenced at ITS2 (
|
||
<figureCitation id="135C2A4AFFA9FFFCCF0E0183FCB5FD95" box="[786,853,511,537]" captionStart="FIGURE 7" captionStartId="22.[151,250,1755,1779]" captionTargetBox="[159,1425,181,1729]" captionTargetId="figure-20@22.[151,1436,181,1732]" captionTargetPageId="22" captionText="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416867" httpUri="https://zenodo.org/record/4416867/files/figure.png" pageId="47" pageNumber="48">Fig. 7</figureCitation>
|
||
) except for
|
||
<taxonomicName id="4C674D4CFFA9FFFCCFC20183FB6AFD95" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[990,1162,511,537]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="anachoretus">
|
||
<emphasis id="B913EADDFFA9FFFCCFC20183FB6AFD95" box="[990,1162,511,537]" italics="true" pageId="47" pageNumber="48">D. anachoretus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFA9FFFCC8850183FAECFD95" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1177,1292,511,537]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bythius">
|
||
<emphasis id="B913EADDFFA9FFFCC8850183FAECFD95" box="[1177,1292,511,537]" italics="true" pageId="47" pageNumber="48">D. bythius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFA9FFFCC9070183FA76FD94" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[1307,1430,511,536]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="eremius">
|
||
<emphasis id="B913EADDFFA9FFFCC9070183FA76FD94" box="[1307,1430,511,536]" italics="true" pageId="47" pageNumber="48">D. eremius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFA9FFFCCC8B025FFED6FDB1" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[151,310,547,573]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="gnophicola">
|
||
<emphasis id="B913EADDFFA9FFFCCC8B025FFED6FDB1" box="[151,310,547,573]" italics="true" pageId="47" pageNumber="48">D. gnophicola</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFA9FFFCCD5F025FFE56FDB1" authorityName="Harvey, Berry, Edward and Humphreys" authorityYear="2008" box="[323,438,547,573]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kryptus">
|
||
<emphasis id="B913EADDFFA9FFFCCD5F025FFE56FDB1" box="[323,438,547,573]" italics="true" pageId="47" pageNumber="48">D. kryptus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFA9FFFCCDDF025FFD76FDB1" authorityName="Abrams and Harvey" authorityYear="2020" box="[451,662,547,573]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="mckechnieorum">
|
||
<emphasis id="B913EADDFFA9FFFCCDDF025FFD76FDB1" box="[451,662,547,573]" italics="true" pageId="47" pageNumber="48">D. mckechnieorum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFA9FFFCCEBF025FFCA7FDB1" authorityName="Abrams and Harvey" authorityYear="2020" box="[675,839,547,573]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="warramboo">
|
||
<emphasis id="B913EADDFFA9FFFCCEBF025FFCA7FDB1" box="[675,839,547,573]" italics="true" pageId="47" pageNumber="48">D. warramboo</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFA9FFFCCF4F025FFC10FDB1" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[851,1008,547,573]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="belalugosii">
|
||
<emphasis id="B913EADDFFA9FFFCCF4F025FFC10FDB1" box="[851,1008,547,573]" italics="true" pageId="47" pageNumber="48">D. belalugosii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFA9FFFCCFE2025FFB29FDB1" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[1022,1225,547,573]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="christopherleei">
|
||
<emphasis id="B913EADDFFA9FFFCCFE2025FFB29FDB1" box="[1022,1225,547,573]" italics="true" pageId="47" pageNumber="48">D. christopherleei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFA9FFFCC8C9025FFA8EFDB1" authorityName="Abrams and Harvey" authorityYear="2020" box="[1237,1390,547,573]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="piscivultus">
|
||
<emphasis id="B913EADDFFA9FFFCC8C9025FFA8EFDB1" box="[1237,1390,547,573]" italics="true" pageId="47" pageNumber="48">D. piscivultus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C674D4CFFA9FFFCCC8B023BFEF5FDED" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[151,277,583,609]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="akashae">
|
||
<emphasis id="B913EADDFFA9FFFCCC8B023BFEF5FDED" box="[151,277,583,609]" italics="true" pageId="47" pageNumber="48">D. akashae</emphasis>
|
||
</taxonomicName>
|
||
, which are not distinguishable using the ITS2 mini-barcode.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFA9FFE2CCDB0216FE61FC05" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48" type="description">
|
||
<paragraph id="8BD836CFFFA9FFFCCCDB0216FAF9FD09" blockId="47.[151,1436,150,645]" box="[199,1305,618,645]" pageId="47" pageNumber="48">
|
||
<emphasis id="B913EADDFFA9FFFCCCDB0216FDE7FD08" bold="true" box="[199,519,618,644]" pageId="47" pageNumber="48">Description (adult female).</emphasis>
|
||
<emphasis id="B913EADDFFA9FFFCCE120217FDBDFD09" box="[526,605,619,645]" italics="true" pageId="47" pageNumber="48">Colour</emphasis>
|
||
. Yellow-brown; propeltidium and pedipalps somewhat darker.
|
||
</paragraph>
|
||
<caption id="DF186647FFA9FFFCCC8B0758FBD2F80F" ID-DOI="http://doi.org/10.5281/zenodo.4416911" ID-Zenodo-Dep="4416911" httpUri="https://zenodo.org/record/4416911/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,250,1828,1852]" targetBox="[156,1431,669,1803]" targetPageId="47">
|
||
<paragraph id="8BD836CFFFA9FFFCCC8B0758FBD2F80F" blockId="47.[151,1437,1828,1924]" pageId="47" pageNumber="48">
|
||
<emphasis id="B913EADDFFA9FFFCCC8B0758FEC0F8B0" bold="true" box="[151,288,1828,1852]" pageId="47" pageNumber="48">FIGURE 24.</emphasis>
|
||
<taxonomicName id="4C674D4CFFA9FFFCCD3A0758FDF8F8B0" authority="Abrams & Huey & Hillyer & Didham & Harvey, 2020" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[294,536,1828,1852]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="immortalis" status="sp. nov.">
|
||
<emphasis id="B913EADDFFA9FFFCCD3A0758FDF8F8B0" box="[294,536,1828,1852]" italics="true" pageId="47" pageNumber="48">Draculoides immortalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B913EADDFFA9FFFCCE020758FD8FF8B0" bold="true" box="[542,623,1828,1852]" pageId="47" pageNumber="48">
|
||
<taxonomicNameLabel id="A22057A6FFA9FFFCCE020758FD8FF8B0" box="[542,623,1828,1852]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype female (WAM T138476): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD836CFFFB6FFE3CCDB00EBFD54FF75" blockId="48.[151,1437,151,501]" pageId="48" pageNumber="49">
|
||
<emphasis id="B913EADDFFB6FFE3CCDB00EBFE8FFF3D" box="[199,367,151,177]" italics="true" pageId="48" pageNumber="49">Cephalothorax</emphasis>
|
||
. Propeltidium with 3 apical setae in a row on anterior process and 1 + 2 + 2 setae; eye spots absent. Mesopeltidia separated. Metapeltidium divided. Anterior sternum with 13 setae (including 2 sternapophysial setae); posterior sternum triangular with 6 setae.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFB6FFE3CCDB017FFEA2FE79" blockId="48.[151,1437,151,501]" pageId="48" pageNumber="49">
|
||
<emphasis id="B913EADDFFB6FFE3CCDB017FFED5FE91" box="[199,309,259,285]" italics="true" pageId="48" pageNumber="49">Chelicera</emphasis>
|
||
. Fixed finger with 2 large teeth plus 6 smaller teeth between these, proximal tooth with 1 small lateral tooth and distal tooth without small lateral teeth; membranous area between fixed and movable fingers with 3 large, lanceolate, terminally pilose setae (G1); G2 and G3 composed of 5 setae; internal face of chelicera with 5 short whip-like setae (G4); brush at base of fixed finger composed of 7 setae (G5A), each densely pilose in distal half and G5B composed of 10 setae; G6 with one seta; G7 composed of 4 setae. Movable finger serrula composed of 18 long lamellae, small blunt guard tooth present subdistally; 1 large accessory tooth present at two-thirds from base of serrula.
|
||
</paragraph>
|
||
<caption id="DF186647FFB6FFE3CC8B0731FBE1F805" ID-DOI="http://doi.org/10.5281/zenodo.4416913" ID-Zenodo-Dep="4416913" httpUri="https://zenodo.org/record/4416913/files/figure.png" pageId="48" pageNumber="49" startId="48.[151,250,1869,1893]" targetBox="[149,1435,525,1843]" targetPageId="48">
|
||
<paragraph id="8BD836CFFFB6FFE3CC8B0731FBE1F805" blockId="48.[151,1436,1869,1930]" pageId="48" pageNumber="49">
|
||
<emphasis id="B913EADDFFB6FFE3CC8B0731FEC0F8EA" bold="true" box="[151,288,1869,1894]" pageId="48" pageNumber="49">FIGURE 25.</emphasis>
|
||
<taxonomicName id="4C674D4CFFB6FFE3CD3A0732FDF7F8EA" authority="Abrams & Huey & Hillyer & Didham & Harvey, 2020" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[294,535,1870,1894]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="immortalis" status="sp. nov.">
|
||
<emphasis id="B913EADDFFB6FFE3CD3A0732FDF7F8EA" box="[294,535,1870,1894]" italics="true" pageId="48" pageNumber="49">Draculoides immortalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B913EADDFFB6FFE3CE010732FD8EF8EA" bold="true" box="[541,622,1870,1894]" pageId="48" pageNumber="49">
|
||
<taxonomicNameLabel id="A22057A6FFB6FFE3CE010732FD8EF8EA" box="[541,622,1870,1894]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype female (WAM T138476): A. Flagellum, dorsal; B. Flagellum, ventral; C. Flagellum, lateral; D. Spermathecae, ventral. Scale bars: A–C: 0.1 mm, D: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD836CFFFB7FFE2CCDB00EBFB69FED3" blockId="49.[151,1437,151,1044]" pageId="49" pageNumber="50">Fixed finger with 2 large teeth plus 6 smaller teeth between these; basal tooth with 1 small lateral tooth and distal tooth without small lateral teeth; brush at base of fixed finger composed of 7 setae (G5A), each densely pilose in distal half and G5B composed of 10 setae; membranous area between fixed and movable fingers with 3 large, lanceolate, terminally pilose setae (G1); internal face of chelicera with 5 short whip-like setae (G4); movable finger serrula composed of 18 long lamellae, small blunt guard tooth present subdistally; 1 large accessory tooth present at two-thirds from base of serrula. G2 and G3 composed of 5 setae; G7 composed of 4 setae.</paragraph>
|
||
<paragraph id="8BD836CFFFB7FFE2CCDB011BFA6FFE28" blockId="49.[151,1437,151,1044]" pageId="49" pageNumber="50">
|
||
<emphasis id="B913EADDFFB7FFE2CCDB011BFECCFE0D" box="[199,300,359,385]" italics="true" pageId="49" pageNumber="50">Pedipalp</emphasis>
|
||
. Without apophyses; trochanter with sharply produced ventro-distal extension, ventral margin with ca. 10 stout setae, without mesal spur; tarsus and tibia without spines; tarsal spur present; claw 0.5 × length of tarsus.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFB7FFE2CCDB01D1FB0EFE4A" blockId="49.[151,1437,151,1044]" box="[199,1262,428,455]" pageId="49" pageNumber="50">
|
||
<emphasis id="B913EADDFFB7FFE2CCDB01D1FEE4FE4A" box="[199,260,429,454]" italics="true" pageId="49" pageNumber="50">Legs.</emphasis>
|
||
Tarsus I with 6 segments; baso-dorsal margin of femur IV produced at about a 90° angle.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFB7FFE2CCDB01B3FC26FD87" blockId="49.[151,1437,151,1044]" pageId="49" pageNumber="50">
|
||
<emphasis id="B913EADDFFB7FFE2CCDB01B3FED1FE65" box="[199,305,463,489]" italics="true" pageId="49" pageNumber="50">Abdomen</emphasis>
|
||
. Chaetotaxy of tergite I: 2 macrosetae + 4 microsetae (microsetae diagonal), tergite II: 3 macrosetae + 6 microsetae (microsetae in column), tergites III–IX: 2: 2: 2: 2: 2: 2: 3.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFB7FFE2CCDB0268FE82FDFF" blockId="49.[151,1437,151,1044]" pageId="49" pageNumber="50">
|
||
<emphasis id="B913EADDFFB7FFE2CCDB0268FE66FDA2" box="[199,390,532,558]" italics="true" pageId="49" pageNumber="50">Female genitalia</emphasis>
|
||
. Two pairs of elongate spermathecae with outer lobe larger than inner lobe, each pair not connected basally (
|
||
<figureCitation id="135C2A4AFFB7FFE2CD55024BFE53FDDD" box="[329,435,566,593]" captionStart="FIGURE 25" captionStartId="48.[151,250,1869,1893]" captionTargetBox="[149,1435,525,1843]" captionTargetId="figure-181@48.[149,1437,525,1845]" captionTargetPageId="48" captionText="FIGURE 25. Draculoides immortalis sp. nov., holotype female (WAM T138476): A. Flagellum, dorsal; B. Flagellum, ventral; C. Flagellum, lateral; D. Spermathecae, ventral. Scale bars: A–C: 0.1 mm, D: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4416913" httpUri="https://zenodo.org/record/4416913/files/figure.png" pageId="49" pageNumber="50">Fig. 25D</figureCitation>
|
||
), distally round and smooth; sparsely covered with small pores; gonopod short, roughly triangular-shaped.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFB7FFE2CCDB0200FADBFCAD" blockId="49.[151,1437,151,1044]" pageId="49" pageNumber="50">
|
||
<emphasis id="B913EADDFFB7FFE2CCDB0200FEDAFD1A" box="[199,314,636,662]" italics="true" pageId="49" pageNumber="50">Flagellum</emphasis>
|
||
. 5 × longer than broad (
|
||
<figureCitation id="135C2A4AFFB7FFE2CE4C0200FD4BFD1A" box="[592,683,636,662]" captionStart="FIGURE 24" captionStartId="47.[151,250,1828,1852]" captionTargetBox="[156,1431,669,1803]" captionTargetId="figure-255@47.[156,1431,669,1804]" captionTargetPageId="47" captionText="FIGURE 24. Draculoides immortalis sp. nov., holotype female (WAM T138476): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F." figureDoi="http://doi.org/10.5281/zenodo.4416911" httpUri="https://zenodo.org/record/4416911/files/figure.png" pageId="49" pageNumber="50">Figs. 24</figureCitation>
|
||
D–F, 25A–C); seta dm1 situated close to posterior end of flagellomere II, posterior to vm2; setae dl1 situated anterior to dm4, dm4 situated midway between dl1 and dl3; dl3 situated almost at posterior margin, slightly more posterior than vl2, vm1 situated slightly posterior to vm2, vm3 situated closer to vm1 than to vm5, vm5 midway between vm3 and vl2, vl1 situated posterior to vm3 and anterior to dl1; one pair of microsetae at anterior end of flagellomere III, one pair of microsetae laterally between dl3 and vl2.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFB7FFE2CCDB0356FE61FC05" blockId="49.[151,1437,151,1044]" pageId="49" pageNumber="50">
|
||
<emphasis id="B913EADDFFB7FFE2CCDB0356FE71FCC8" box="[199,401,810,836]" italics="true" pageId="49" pageNumber="50">Dimensions (mm)</emphasis>
|
||
.
|
||
<materialsCitation id="3B0F3C92FFB7FFE2CDBF0355FE61FC05" ID-GBIF-Occurrence="3013449395" collectionCode="WAM" pageId="49" pageNumber="50" specimenCode="T138476" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54DC886DFFB7FFE2CDBF0355FDEBFCCF" box="[419,523,809,835]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
|
||
female (
|
||
<collectionCode id="ED76AE0AFFB7FFE2CE680355FD5BFCCF" box="[628,699,809,835]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="50" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FFB7FFE2CED80355FCCFFCC8" box="[708,815,809,836]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="49" pageNumber="50">T138476</specimenCode>
|
||
): Body length 3.99. Propeltidium 0.48/031. Chelicera 0.96. Flagellum 0.4/0.08. Pedipalp: trochanter 0.62, femur 0.71, patella 0.73, tibia 0.60, tarsus 0.31, claw 0.15, total excluding claw 2.96.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFB7FFE2CCDB03EDFCFAFC7D" pageId="49" pageNumber="50" type="discussion">
|
||
<paragraph id="8BD836CFFFB7FFE2CCDB03EDFD82FC43" blockId="49.[151,1437,151,1044]" pageId="49" pageNumber="50">
|
||
<emphasis id="B913EADDFFB7FFE2CCDB03EDFEDBFC27" bold="true" box="[199,315,913,939]" pageId="49" pageNumber="50">Remarks.</emphasis>
|
||
<taxonomicName id="4C674D4CFFB7FFE2CD5D03EDFDAAFC27" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[321,586,913,939]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="immortalis">
|
||
<emphasis id="B913EADDFFB7FFE2CD5D03EDFDAAFC27" box="[321,586,913,939]" italics="true" pageId="49" pageNumber="50">Draculoides immortalis</emphasis>
|
||
</taxonomicName>
|
||
is known from a single location ‘Budgiebore’ north of Mesa A situated in the Robe Valley,
|
||
<collectingRegion id="49A3F82DFFB7FFE2CD3003C9FE13FC42" box="[300,499,948,975]" country="Australia" name="Western Australia" pageId="49" pageNumber="50">Western Australia</collectingRegion>
|
||
(
|
||
<figureCitation id="135C2A4AFFB7FFE2CE1F03C8FDB5FC43" box="[515,597,948,975]" captionStart="FIGURE 1" captionStartId="16.[151,250,1868,1892]" captionTargetBox="[346,1243,188,1844]" captionTargetId="figure-20@16.[339,1249,181,1844]" captionTargetPageId="16" captionText="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." figureDoi="http://doi.org/10.5281/zenodo.4416851" httpUri="https://zenodo.org/record/4416851/files/figure.png" pageId="49" pageNumber="50">Fig.1A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFB7FFE2CCDB03AAFCFAFC7D" blockId="49.[151,1437,151,1044]" box="[199,794,982,1009]" pageId="49" pageNumber="50">
|
||
<emphasis id="B913EADDFFB7FFE2CCDB03AAFE81FC7D" bold="true" box="[199,353,982,1009]" pageId="49" pageNumber="50">Other names</emphasis>
|
||
. WAM SCH058 (
|
||
<bibRefCitation id="EFF64B3EFFB7FFE2CE3503ABFCEEFC7D" author="Abrams, K. M. & Huey, J. A. & Hillyer, M. J. & Humphreys, W. F. & Didham, R. K. & Harvey, M. S." box="[553,782,983,1009]" pageId="49" pageNumber="50" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. & Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
|
||
Abrams
|
||
<emphasis id="B913EADDFFB7FFE2CE9703ABFD5CFC7D" box="[651,700,983,1009]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
|
||
., 2019
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFB7FFE2CCDB0385FA61FB98" box="[199,1409,1017,1044]" pageId="49" pageNumber="50" type="etymology">
|
||
<paragraph id="8BD836CFFFB7FFE2CCDB0385FA61FB98" blockId="49.[151,1437,151,1044]" box="[199,1409,1017,1044]" pageId="49" pageNumber="50">
|
||
<emphasis id="B913EADDFFB7FFE2CCDB0385FEABFB9F" bold="true" box="[199,331,1017,1043]" pageId="49" pageNumber="50">Etymology.</emphasis>
|
||
The specific epithet is an adjective referring to the immortal nature of vampires (Latin, immortal).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |