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<document id="6D3410EDBD3786675AE5C32C74832BE6" ID-DOI="10.11646/zootaxa.3635.1.9" ID-GBIF-Dataset="c2e1b77b-2a89-46f8-8b24-64ae71254792" ID-ISSN="1175-5326" ID-Zenodo-Dep="283539" ID-ZooBank="3B9C28A8-B64D-46AB-BE68-BF6CA85DBCAF" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460585776088" checkinUser="plazi" docAuthor="Shear, William A." docDate="2013" docId="03CE87AC9904FFD0F0C3114FFD3162B6" docLanguage="en" docName="zt03635p093.pdf" docOrigin="Zootaxa 3635 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cleidogona huebnerae Shear, 2013, n. sp." docType="treatment" docVersion="6" lastPageNumber="88" masterDocId="FFF7FFD49904FFD1F054164EFFBE657A" masterDocTitle="A new miniature species of the milliped genus Cleidogona from Costa Rica (Diplopoda, Chordeumatida, Cleidogonidae), with a review of Central American Cleidogonidae" masterLastPageNumber="93" masterPageNumber="87" pageNumber="87" updateTime="1698325116444" updateUser="plazi">
<mods:mods id="435DB05CDB87315CD17DA90D8368A5FA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D10E2DD1841E5CABA316F5CDF4417B80">
<mods:title id="5F542C76C0EFDF3F77748FE4CAEE53F2">A new miniature species of the milliped genus Cleidogona from Costa Rica (Diplopoda, Chordeumatida, Cleidogonidae), with a review of Central American Cleidogonidae</mods:title>
</mods:titleInfo>
<mods:name id="1288A6AAC09188FA48B5D042CF813315" type="personal">
<mods:role id="08EAA665222C833CBA4DEC1340D75CDA">
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<mods:namePart id="98ED73BC44DCF41C85A76400CB4A46A5">Shear, William A.</mods:namePart>
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<mods:title id="8F847C12D59CA2D811383C97AD7E1111">Zootaxa</mods:title>
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<mods:part id="B52772E31E3365D262E2C93269E27842">
<mods:date id="2066B374A8D9CF2FA3ABA7B6A5C338EB">2013</mods:date>
<mods:detail id="B04F25B4E1211FDA1877EE949C756FBA" type="volume">
<mods:number id="FBFE61B13E59E9A35BFBD7DA16DE625E">3635</mods:number>
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<mods:classification id="BEFC4443B8FC89BAFA01F89EBA9E8455">journal article</mods:classification>
<mods:identifier id="7690661B09D5859964E5C25080254CBC" type="DOI">10.11646/zootaxa.3635.1.9</mods:identifier>
<mods:identifier id="B0F348F978D252408187A91BEEF1D877" type="GBIF-Dataset">c2e1b77b-2a89-46f8-8b24-64ae71254792</mods:identifier>
<mods:identifier id="13035C98F243003DFB48FF521179B7BE" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03CE87AC9904FFD0F0C3114FFD3162B6" ID-DOI="http://doi.org/10.5281/zenodo.6164044" ID-GBIF-Taxon="127663650" ID-Zenodo-Dep="6164044" LSID="urn:lsid:plazi:treatment:03CE87AC9904FFD0F0C3114FFD3162B6" httpUri="http://treatment.plazi.org/id/03CE87AC9904FFD0F0C3114FFD3162B6" lastPageId="1" lastPageNumber="88" pageId="0" pageNumber="87">
<subSubSection id="C37D65319904FFD1F0C3114FFF4F62B6" pageId="0" pageNumber="87" type="nomenclature">
<paragraph id="8BD836BA9904FFD1F0C3114FFE446260" blockId="0.[151,506,1792,1852]" box="[151,506,1792,1819]" pageId="0" pageNumber="87">
<heading id="D09081D69904FFD1F0C3114FFE446260" bold="true" box="[151,506,1792,1819]" fontSize="11" level="1" pageId="0" pageNumber="87" reason="1">
<emphasis id="B913EAA89904FFD1F0C3114FFE446260" bold="true" box="[151,506,1792,1819]" pageId="0" pageNumber="87">
<emphasis id="B913EAA89904FFD1F0C3114FFE0E6261" bold="true" box="[151,432,1793,1819]" italics="true" pageId="0" pageNumber="87">
<taxonomicName id="4C674D399904FFD1F0C3114FFE146261" ID-CoL="VWXZ" box="[151,426,1793,1819]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="huebnerae" status="sp. nov.">Cleidogona huebnerae</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A22057D39904FFD1F1EC114EFE446260" box="[440,506,1792,1818]" pageId="0" pageNumber="87" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD836BA9904FFD1F0C3116AFF416246" blockId="0.[151,506,1792,1852]" box="[151,255,1828,1852]" pageId="0" pageNumber="87">
<figureCitation id="135C2A3F9904FFD1F0C3116AFF616246" box="[151,223,1828,1852]" captionStart="FIGURES 1 – 3" captionStartId="1.[151,264,758,781]" captionTargetBox="[167,1418,205,726]" captionTargetId="figure@1.[160,1426,195,734]" captionTargetPageId="1" captionText="FIGURES 1 – 3. Cleidogona huebnerae, n. sp., male. 1. Gonopods, posterior view. 2. Gonopods, anterior view. 3. Legpair 9, anterior view." httpUri="https://zenodo.org/record/283540/files/figure.png" pageId="0" pageNumber="87">
Figs.
<docIssue id="CF0449559904FFD1F081116AFF616246" box="[213,223,1828,1852]" pageId="0" pageNumber="87">1</docIssue>
</figureCitation>
–4
</paragraph>
<paragraph id="8BD836BA9904FFD1F0C31122FF4F62B6" blockId="0.[151,1436,1900,1996]" pageId="0" pageNumber="87">
<emphasis id="B913EAA89904FFD1F0C31122FF5D62FE" bold="true" box="[151,227,1900,1924]" pageId="0" pageNumber="87">
<typeStatus id="54DC88189904FFD1F0C31122FF6462FE" box="[151,218,1900,1924]" pageId="0" pageNumber="87">Types</typeStatus>
:
</emphasis>
Male
<typeStatus id="54DC88189904FFD1F1641122FE2C62FE" box="[304,402,1900,1924]" pageId="0" pageNumber="87" type="holotype">holotype</typeStatus>
and two male and female
<typeStatus id="54DC88189904FFD1F2951123FC9162FF" box="[705,815,1901,1925]" pageId="0" pageNumber="87" type="paratype">paratypes</typeStatus>
, with other, juvenile specimens from Tropenstation La Gamba,
<collectingCountry id="F370762A9904FFD1F0AC11C1FEC662DD" box="[248,376,1935,1959]" name="Costa Rica" pageId="0" pageNumber="87">Costa Rica</collectingCountry>
, collected in 2008 by Barbara Hübner.
<typeStatus id="54DC88189904FFD1F31411DEFC3C62D2" box="[832,898,1936,1960]" pageId="0" pageNumber="87">Types</typeStatus>
deposited in Naturhistorisches Museum Wien,
<collectingCountry id="F370762A9904FFD1F0C311FAFF5362B6" box="[151,237,1972,1996]" name="Austria" pageId="0" pageNumber="87">Austria</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF1866329905FFD0F0C314B8FE9D6651" httpUri="https://zenodo.org/record/283540/files/figure.png" pageId="1" pageNumber="88" targetBox="[167,1418,205,726]" targetPageId="1">
<paragraph id="8BD836BA9905FFD0F0C314B8FE9D6651" blockId="1.[151,1436,758,811]" pageId="1" pageNumber="88">
<emphasis id="B913EAA89905FFD0F0C314B8FE826676" bold="true" box="[151,316,758,781]" pageId="1" pageNumber="88">FIGURES 1–3.</emphasis>
<taxonomicName id="4C674D399905FFD0F11714B9FD926677" box="[323,556,759,781]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="huebnerae" status="sp. nov.">
<emphasis id="B913EAA89905FFD0F11714B9FD926677" box="[323,556,759,781]" italics="true" pageId="1" pageNumber="88">Cleidogona huebnerae</emphasis>
</taxonomicName>
,
<emphasis id="B913EAA89905FFD0F26E14B9FDC46676" bold="true" box="[570,634,759,780]" pageId="1" pageNumber="88">
<taxonomicNameLabel id="A22057D39905FFD0F26E14B9FDC86676" box="[570,630,759,780]" pageId="1" pageNumber="88" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
male. 1. Gonopods, posterior view. 2. Gonopods, anterior view. 3. Legpair 9, anterior view.
</paragraph>
</caption>
<subSubSection id="C37D65319905FFD0F0931519FEAC66C2" pageId="1" pageNumber="88" type="diagnosis">
<paragraph id="8BD836BA9905FFD0F0931519FEAC66C2" blockId="1.[151,1437,855,1996]" pageId="1" pageNumber="88">
<emphasis id="B913EAA89905FFD0F0931519FEFD660A" bold="true" box="[199,323,855,880]" pageId="1" pageNumber="88">Diagnosis:</emphasis>
Distinct from all other known Central American cleidogonid millipeds in the small size of mature specimens, being about 50% smaller than the next smallest congeneric species, and in the details of the gonopods (
<figureCitation id="135C2A3F9905FFD0F0CB15EEFEB766C2" box="[159,265,928,952]" captionStart="FIGURES 1 – 3" captionStartId="1.[151,264,758,781]" captionTargetBox="[167,1418,205,726]" captionTargetId="figure@1.[160,1426,195,734]" captionTargetPageId="1" captionText="FIGURES 1 – 3. Cleidogona huebnerae, n. sp., male. 1. Gonopods, posterior view. 2. Gonopods, anterior view. 3. Legpair 9, anterior view." httpUri="https://zenodo.org/record/283540/files/figure.png" pageId="1" pageNumber="88">Figs. 1, 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37D65319905FFD0F093158AFBCD6012" pageId="1" pageNumber="88" type="description">
<paragraph id="8BD836BA9905FFD0F093158AFBD7603E" blockId="1.[151,1437,855,1996]" pageId="1" pageNumber="88">
<emphasis id="B913EAA89905FFD0F093158AFEE566A7" bold="true" box="[199,347,964,989]" pageId="1" pageNumber="88">Description:</emphasis>
<emphasis id="B913EAA89905FFD0F132158AFE1666A6" box="[358,424,964,988]" italics="true" pageId="1" pageNumber="88">Male:</emphasis>
Length,
<quantity id="4C9F9B5F9905FFD0F242158BFDE066A6" box="[534,606,965,988]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="88" unit="mm" value="7.0">7 mm</quantity>
, greatest width
<quantity id="4C9F9B5F9905FFD0F34F158AFC3866A6" box="[795,902,964,988]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="1" pageNumber="88" unit="mm" value="0.62">0.62 mm</quantity>
. Ocelli 18, in five rows. Color light chestnut brown, dorsally mottled with darker purplish brown. Diplosegments with moderately developed lateral tubercles bearing lateralmost segmental setae; segmental setae about one-third width of body. Legpairs 1–7 only slightly more crassate than postgonopodal legs. Gonopods: in anterior view (
<figureCitation id="135C2A3F9905FFD0F3F6127EFC596132" box="[930,999,1072,1096]" captionStart="FIGURES 1 – 3" captionStartId="1.[151,264,758,781]" captionTargetBox="[167,1418,205,726]" captionTargetId="figure@1.[160,1426,195,734]" captionTargetPageId="1" captionText="FIGURES 1 – 3. Cleidogona huebnerae, n. sp., male. 1. Gonopods, posterior view. 2. Gonopods, anterior view. 3. Legpair 9, anterior view." httpUri="https://zenodo.org/record/283540/files/figure.png" pageId="1" pageNumber="88">Fig. 2</figureCitation>
) with broad sternum (
<emphasis id="B913EAA89905FFD0F4B1127EFB4E6132" bold="true" box="[1253,1264,1072,1096]" pageId="1" pageNumber="88">s</emphasis>
) not extending between angiocoxites, with paired, squarish median swellings; coxal knobs prominent. Coxae (
<emphasis id="B913EAA89905FFD0F486121BFB606117" bold="true" box="[1234,1246,1109,1133]" pageId="1" pageNumber="88">c</emphasis>
) with two setae. Angiocoxites (
<emphasis id="B913EAA89905FFD0F1691236FEE661EA" bold="true" box="[317,344,1144,1168]" pageId="1" pageNumber="88">ac</emphasis>
) short, broad, thin, with short, rounded apical process. In posterior view (
<figureCitation id="135C2A3F9905FFD0F4C21236FB6961EA" box="[1174,1239,1144,1168]" captionStart="FIGURES 1 – 3" captionStartId="1.[151,264,758,781]" captionTargetBox="[167,1418,205,726]" captionTargetId="figure@1.[160,1426,195,734]" captionTargetPageId="1" captionText="FIGURES 1 – 3. Cleidogona huebnerae, n. sp., male. 1. Gonopods, posterior view. 2. Gonopods, anterior view. 3. Legpair 9, anterior view." httpUri="https://zenodo.org/record/283540/files/figure.png" pageId="1" pageNumber="88">Fig. 1</figureCitation>
) angiocoxite tips extend prominently posteriorly, with fine cuticular spinules. Colpocoxites (
<emphasis id="B913EAA89905FFD0F38812D3FC4B61CF" bold="true" box="[988,1013,1181,1205]" pageId="1" pageNumber="88">cc</emphasis>
) very low, semirectangular, separated from angiocoxites by poorly sclerotized membrane. Ninth legpair (
<figureCitation id="135C2A3F9905FFD0F3D4128EFC7C61A2" box="[896,962,1216,1240]" captionStart="FIGURES 1 – 3" captionStartId="1.[151,264,758,781]" captionTargetBox="[167,1418,205,726]" captionTargetId="figure@1.[160,1426,195,734]" captionTargetPageId="1" captionText="FIGURES 1 – 3. Cleidogona huebnerae, n. sp., male. 1. Gonopods, posterior view. 2. Gonopods, anterior view. 3. Legpair 9, anterior view." httpUri="https://zenodo.org/record/283540/files/figure.png" pageId="1" pageNumber="88">Fig. 3</figureCitation>
) reduced as usual for genus, but each with coxa, trochanter, prefemur (
<emphasis id="B913EAA89905FFD0F18012AAFE476187" bold="true" box="[468,505,1252,1277]" pageId="1" pageNumber="88">cpf</emphasis>
) fused; coxal portion with small, perhaps vestigial gland openings surrounded by markedly thicker cuticle; femora (
<emphasis id="B913EAA89905FFD0F2721349FD91605A" bold="true" box="[550,559,1287,1312]" pageId="1" pageNumber="88">f</emphasis>
) distally thickened. Legpairs 10, 11 typical, coxae elongate, with extruded glands. Legpair 12 with forward-projecting sternal peg fitting
<emphasis id="B913EAA89905FFD0F31B1363FC2A603E" box="[847,916,1325,1348]" italics="true" pageId="1" pageNumber="88">in situ</emphasis>
between coxae 11.
</paragraph>
<paragraph id="8BD836BA9905FFD0F093131EFBCD6012" blockId="1.[151,1437,855,1996]" box="[199,1139,1360,1385]" pageId="1" pageNumber="88">
<emphasis id="B913EAA89905FFD0F093131EFE9A6012" box="[199,292,1360,1384]" italics="true" pageId="1" pageNumber="88">Female:</emphasis>
Length,
<quantity id="4C9F9B5F9905FFD0F1DC131EFE5E6012" box="[392,480,1360,1384]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="88" unit="mm" value="7.3">7.3 mm</quantity>
, greatest width
<quantity id="4C9F9B5F9905FFD0F2C4131EFD496012" box="[656,759,1360,1384]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="1" pageNumber="88" unit="mm" value="0.66">0.66 mm</quantity>
. Nonsexual characters as in male.
</paragraph>
</subSubSection>
<subSubSection id="C37D65319905FFD0F093133AFD4D60F6" box="[199,755,1396,1421]" pageId="1" pageNumber="88" type="distribution">
<paragraph id="8BD836BA9905FFD0F093133AFD4D60F6" blockId="1.[151,1437,855,1996]" box="[199,755,1396,1421]" pageId="1" pageNumber="88">
<emphasis id="B913EAA89905FFD0F093133AFEDC60F7" bold="true" box="[199,354,1396,1421]" pageId="1" pageNumber="88">Distribution:</emphasis>
Known only from the
<typeStatus id="54DC88189905FFD0F236133BFD2C60F7" box="[610,658,1397,1421]" pageId="1" pageNumber="88">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C37D65319905FFD0F09313D6FD3162B6" pageId="1" pageNumber="88" type="etymology">
<paragraph id="8BD836BA9905FFD0F09313D6FC9460CA" blockId="1.[151,1437,855,1996]" box="[199,810,1432,1456]" pageId="1" pageNumber="88">
<emphasis id="B913EAA89905FFD0F09313D6FEEF60CA" bold="true" box="[199,337,1432,1456]" pageId="1" pageNumber="88">Etymology:</emphasis>
Named for the collector, Barbara Hübner.
</paragraph>
<paragraph id="8BD836BA9905FFD0F09313F3FD3D631E" blockId="1.[151,1437,855,1996]" pageId="1" pageNumber="88">
<emphasis id="B913EAA89905FFD0F09313F3FEAD60AF" bold="true" box="[199,275,1469,1493]" pageId="1" pageNumber="88">Notes:</emphasis>
One of the juvenile specimens collected with the
<typeStatus id="54DC88189905FFD0F30013F3FC3160AF" box="[852,911,1469,1493]" pageId="1" pageNumber="88">types</typeStatus>
is a penultimate male, of interest because the ninth legpair, reduced in adult males, is of normal size in this instar. The gonopods appear as small primordia in place of the eighth legpair. While previously described Central American cleidogonids were primarily from higher elevations, this species comes from a lowland locality, suggesting that there may be additional species of
<taxonomicName id="4C674D399905FFD0F0C31002FEA4631E" box="[151,282,1612,1636]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAA89905FFD0F0C31002FEA4631E" box="[151,282,1612,1636]" italics="true" pageId="1" pageNumber="88">Cleidogona</emphasis>
</taxonomicName>
to be discovered in such places.
</paragraph>
<paragraph id="8BD836BA9905FFD0F093103EFAC263AA" blockId="1.[151,1437,855,1996]" pageId="1" pageNumber="88">
The gonopods recall species of
<taxonomicName id="4C674D399905FFD0F27E103EFD3963F2" box="[554,647,1648,1672]" class="Diplopoda" family="Cleidogonidae" genus="Dybasia" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAA89905FFD0F27E103EFD3963F2" box="[554,647,1648,1672]" italics="true" pageId="1" pageNumber="88">Dybasia</emphasis>
</taxonomicName>
(see below) in the short but obvious bifid sternal process and reduced colpocoxites, but are more complex at the angiocoxite tips. The ninth leg femora are distally enlarged, as in
<emphasis id="B913EAA89905FFD0F0C310F6FE9E63AA" box="[151,288,1720,1744]" italics="true" pageId="1" pageNumber="88">
<taxonomicName id="4C674D399905FFD0F0C310F6FEA363AA" box="[151,285,1720,1744]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Cleidogona</taxonomicName>
,
</emphasis>
and the coxoprefemora do not have processes. All in all, I think the species belongs in
<emphasis id="B913EAA89905FFD0F4A610F6FAC263AA" box="[1266,1404,1720,1744]" italics="true" pageId="1" pageNumber="88">
<taxonomicName id="4C674D399905FFD0F4A610F6FAC663AA" box="[1266,1400,1720,1744]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Cleidogona</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BD836BA9905FFD0F0931092FD3162B6" blockId="1.[151,1437,855,1996]" pageId="1" pageNumber="88">
Tropenstation La Gamba is a field station in southern
<collectingCountry id="F370762A9905FFD0F3601092FC0E638E" box="[820,944,1756,1780]" name="Costa Rica" pageId="1" pageNumber="88">Costa Rica</collectingCountry>
(
<geoCoordinate id="EE53507D9905FFD0F3951092FBDA638E" box="[961,1124,1756,1780]" direction="north" orientation="latitude" pageId="1" pageNumber="88" precision="1" value="8.701019">8°42’03.67”N</geoCoordinate>
, 83°12’
<geoCoordinate id="EE53507D9905FFD0F4E11092FAA8638E" box="[1205,1302,1756,1780]" direction="west" orientation="longitude" pageId="1" pageNumber="88" precision="555" value="-6.2">06.20W</geoCoordinate>
, elev.
<quantity id="4C9F9B5F9905FFD0F5351092FA2A638E" box="[1377,1428,1756,1780]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="88" unit="m" value="75.0">75m</quantity>
) operated by Universität Wien and founded by Regenwald der Österreicher (Rainforest of the Austrians). Specimens of
<taxonomicName id="4C674D399905FFD0F16C116BFE6F6246" box="[312,465,1828,1852]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="huebnerae">
<emphasis id="B913EAA89905FFD0F16C116BFE6F6246" box="[312,465,1828,1852]" italics="true" pageId="1" pageNumber="88">C. huebnerae</emphasis>
</taxonomicName>
were collected from leaf litter at two study sites, one a narrow ridge, and the other a steep slope down to a brook. Both were heavily vegetated with rainforest trees and understory plants but the canopy was less dense at the slope site. At both sites, the litter consisted mainly of fresh and undecomposed leaves, twigs, and fruits. The study focused on litter-trapping plants, but no specimens of
<taxonomicName id="4C674D399905FFD0F46011DFFB7262D2" box="[1076,1228,1936,1960]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="huebnerae">
<emphasis id="B913EAA89905FFD0F46011DFFB7262D2" box="[1076,1228,1936,1960]" italics="true" pageId="1" pageNumber="88">C. huebnerae</emphasis>
</taxonomicName>
were taken in this microhabitat (B. Hübner, pers. comm. 2013).
</paragraph>
</subSubSection>
</treatment>
</document>
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