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<document id="2EB8D74F4D0BCE9D3CAA827A7B561A43" ID-DOI="10.11646/zootaxa.5361.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10084538" ID-ZooBank="D9AE18C9-A265-44F0-A0AF-E663A59B2EAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1698932362298" checkinUser="plazi" docAuthor="Zamani, Alireza, Darvishnia, Hamid &amp; Marusik, Yuri M." docDate="2023" docId="03C5CF42FF9B864F22B5F95BFD8C280C" docLanguage="en" docName="zootaxa.5361.3.3.pdf" docOrigin="Zootaxa 5361 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5361.3.3/52195" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tegenaria pallens Zamani &amp; Darvishnia &amp; Marusik 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="351" masterDocId="FFFCB73AFF9886492222FFEDFFCC2911" masterDocTitle="New data on cave spiders (Arachnida: Araneae) of Iran, with new species and records" masterLastPageNumber="366" masterPageNumber="345" pageNumber="348" updateTime="1699464889253" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="64E7516AE827E1737211A772DCE639EE">New data on cave spiders (Arachnida: Araneae) of Iran, with new species and records</mods:title>
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<mods:namePart id="482B6AD3A5C01D4B7D9C3A23585E5695">Zamani, Alireza</mods:namePart>
<mods:affiliation id="2A808743CBD9AA9A4140C522D77B65AD">Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Turku, Finland.</mods:affiliation>
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<mods:namePart id="515423A4E76BC2905C45E95F37A6C6F4">Darvishnia, Hamid</mods:namePart>
<mods:affiliation id="8341686FC5F83E7E8F02399AC29C5EB5">Department of Biology, Payame Noor University, Tehran, Iran.</mods:affiliation>
</mods:name>
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<mods:namePart id="DDB72CCBF3834E00AD136984BA1C6ED6">Marusik, Yuri M.</mods:namePart>
<mods:affiliation id="45EE574F3DF5FF7D08A64C5B12E18555">Institute for Biological Problems of the North, FEB RAS, Portovaya Str. 18. Magadan 685000, Russia. &amp; Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
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<mods:title id="A8C1AECC1A52F241E9DBFE530DBE5BBF">Zootaxa</mods:title>
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<mods:number id="AC9CEDF240DB22788D191336A21170F1">2023-11-02</mods:number>
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<mods:number id="6FE350C93640B706A9E466A3AF02C228">5361</mods:number>
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<treatment id="03C5CF42FF9B864F22B5F95BFD8C280C" LSID="urn:lsid:plazi:treatment:03C5CF42FF9B864F22B5F95BFD8C280C" httpUri="http://treatment.plazi.org/id/03C5CF42FF9B864F22B5F95BFD8C280C" lastPageId="6" lastPageNumber="351" pageId="3" pageNumber="348">
<subSubSection id="C3762DDFFF9B864A22B5F95BFD772FC1" box="[151,699,1718,1745]" pageId="3" pageNumber="348" type="nomenclature">
<paragraph id="8BD37E54FF9B864A22B5F95BFD772FC1" blockId="3.[151,699,1718,1780]" box="[151,699,1718,1745]" pageId="3" pageNumber="348">
<heading id="D09BC938FF9B864A22B5F95BFD772FC1" bold="true" box="[151,699,1718,1745]" fontSize="11" level="1" pageId="3" pageNumber="348" reason="1">
<emphasis id="B918A246FF9B864A22B5F95BFD772FC1" bold="true" box="[151,699,1718,1745]" pageId="3" pageNumber="348">
<taxonomicName id="4C6C05D7FF9B864A22B5F95BFDA72FC1" authority="Zamani &amp; Marusik" authorityName="Zamani &amp; Darvishnia &amp; Marusik" authorityYear="2023" box="[151,619,1718,1745]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="pallens" status="sp. nov.">
<emphasis id="B918A246FF9B864A22B5F95BFEA22FC0" bold="true" box="[151,366,1718,1745]" italics="true" pageId="3" pageNumber="348">Tegenaria pallens</emphasis>
Zamani &amp; Marusik
</taxonomicName>
,
<taxonomicNameLabel id="A22B1F3DFF9B864A205BF95BFD772FC1" box="[633,699,1718,1744]" pageId="3" pageNumber="348" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3762DDFFF9B864A22B5F937FE782FE5" box="[151,436,1754,1780]" pageId="3" pageNumber="348" type="description">
<paragraph id="8BD37E54FF9B864A22B5F937FE782FE5" blockId="3.[151,699,1718,1780]" box="[151,436,1754,1780]" pageId="3" pageNumber="348">
<figureCitation id="135762D1FF9B864A22B5F937FEDF2FE5" box="[151,275,1754,1780]" captionStart="FIGURE 1" captionStartId="1.[151,250,1254,1278]" captionTargetBox="[151,1436,345,1225]" captionTargetId="figure-87@1.[151,1436,345,1231]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Tegenaria pallens sp. n., dorsal view. A female; B male." figureDoi="http://doi.org/10.5281/zenodo.10084540" httpUri="https://zenodo.org/record/10084540/files/figure.png" pageId="3" pageNumber="348">Figs 1A–B</figureCitation>
,
<figureCitation id="135762D1FF9B864A233DF937FEA92FE5" box="[287,357,1754,1780]" captionStart="FIGURE 2" captionStartId="2.[151,250,1975,1999]" captionTargetBox="[161,1427,694,1948]" captionTargetId="figure-110@2.[151,1436,694,1951]" captionTargetPageId="2" captionText="FIGURE 2. Male palps of Tegenaria pallens sp. n. (A, B) and T. alamto (C).A, C ventral view; B retrolateral view. C reproduced from Zamani et al. (2018). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10084542" httpUri="https://zenodo.org/record/10084542/files/figure.png" pageId="3" pageNumber="348">2A–B</figureCitation>
,
<figureCitation id="135762D1FF9B864A2352F937FE782FE5" box="[368,436,1754,1780]" captionStart="FIGURE 3" captionStartId="3.[151,250,1082,1106]" captionTargetBox="[151,1436,181,1058]" captionTargetId="figure-18@3.[151,1436,181,1058]" captionTargetPageId="3" captionText="FIGURE 3. Epigyne of Tegenaria pallens sp. n. A macerated, ventral view; B vulva, dorsal view; C intact, ventral view. Scale bar: 0.2 mm. Abbreviation: Co—copulatory opening." figureDoi="http://doi.org/10.5281/zenodo.10084544" httpUri="https://zenodo.org/record/10084544/files/figure.png" pageId="3" pageNumber="348">3A‒C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3762DDFFF9B864A22B5F8CFFECD2E95" pageId="3" pageNumber="348" type="materials_examined">
<paragraph id="8BD37E54FF9B864A22B5F8CFFECD2E95" blockId="3.[151,1436,1826,1996]" pageId="3" pageNumber="348">
<emphasis id="B918A246FF9B864A22B5F8CFFE892E2D" bold="true" box="[151,325,1826,1852]" pageId="3" pageNumber="348">Type material.</emphasis>
<materialsCitation id="3B047409FF9B864A2369F8CFFCBD2E71" collectingDate="2017-11" collectionCode="ZMUT" collectorName="Kohgiluyeh &amp; M. J. Malek Hosseini" country="Iran" county="Gakal Cave" elevation="1100" latitude="30.312222" location="Boyer-Ahmad Province" longLatPrecision="1" longitude="51.15778" municipality="Gachsaran" pageId="3" pageNumber="348" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D7C0F6FF9B864A2369F8CFFE7F2E2D" box="[331,435,1826,1852]" pageId="3" pageNumber="348" type="holotype">Holotype</typeStatus>
♁ (
<collectionCode id="ED7DE691FF9B864A23FCF8CFFDFF2E2D" box="[478,563,1826,1852]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="3" pageNumber="348">ZMUT</collectionCode>
),
<collectingCountry id="F37B3EC4FF9B864A2067F8CFFD422E2D" box="[581,654,1826,1852]" name="Iran" pageId="3" pageNumber="348">IRAN</collectingCountry>
:
<emphasis id="B918A246FF9B864A20BAF8CFFB8E2E2D" box="[664,1090,1826,1852]" italics="true" pageId="3" pageNumber="348">
<collectorName id="26991B82FF9B864A20BAF8CFFCD52E2D" box="[664,793,1826,1852]" pageId="3" pageNumber="348">Kohgiluyeh</collectorName>
&amp;
<location id="8EB3288FFF9B864A211FF8CFFB8E2E2D" LSID="urn:lsid:plazi:treatment:03C5CF42FF9B864F22B5F95BFD8C280C:8EB3288FFF9B864A211FF8CFFB8E2E2D" box="[829,1090,1826,1852]" country="Iran" county="Gakal Cave" latitude="30.312222" longLatPrecision="1" longitude="51.15778" municipality="Gachsaran" name="Boyer-Ahmad Province" pageId="3" pageNumber="348">Boyer-Ahmad Province</location>
</emphasis>
:
<collectingMunicipality id="6BB7E42EFF9B864A2673F8CFFB062E2D" box="[1105,1226,1826,1852]" pageId="3" pageNumber="348">Gachsaran</collectingMunicipality>
,
<collectingCounty id="62B206D8FF9B864A26F6F8CFFA962E2D" box="[1236,1370,1826,1852]" pageId="3" pageNumber="348">Gakal Cave</collectingCounty>
,
<quantity id="4C94D3B1FF9B864A2747F8CFFF632E70" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="348" unit="m" value="1100.0">
<elevation id="00419967FF9B864A2747F8CFFF632E70" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="348" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<geoCoordinate id="EE581893FF9B864A229EF8ABFE9B2E71" box="[188,343,1862,1888]" degrees="30" direction="north" minutes="18" orientation="latitude" pageId="3" pageNumber="348" precision="1" seconds="44.0" value="30.312222">30°18'44.0&quot;N</geoCoordinate>
,
<geoCoordinate id="EE581893FF9B864A2346F8ABFE312E70" box="[356,509,1862,1889]" degrees="51" direction="east" minutes="09" orientation="longitude" pageId="3" pageNumber="348" precision="1" seconds="28.0" value="51.15778">51°09'28.0&quot;E</geoCoordinate>
,
<date id="FFD25894FF9B864A2028F8ABFDAA2E71" box="[522,614,1862,1888]" pageId="3" pageNumber="348" value="2017-11">
<collectingDate id="EF96A17CFF9B864A2028F8ABFDAA2E71" box="[522,614,1862,1888]" pageId="3" pageNumber="348" value="2017-11">XI.2017</collectingDate>
</date>
(
<collectorName id="26991B82FF9B864A2055F8ABFCA52E71" box="[631,873,1862,1888]" pageId="3" pageNumber="348">M.J. Malek Hosseini</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B047409FF9B864A215EF8AAFF312E95" collectingDate="2017-11" collectionCode="ZMUT" collectorName="Kohgiluyeh &amp; M. J. Malek Hosseini" country="Iran" county="Gakal Cave" elevation="1100" latitude="30.312222" location="Boyer-Ahmad Province" longLatPrecision="1" longitude="51.15778" municipality="Gachsaran" pageId="3" pageNumber="348" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54D7C0F6FF9B864A215EF8AAFC212E70" box="[892,1005,1863,1889]" pageId="3" pageNumber="348" type="paratype">Paratypes</typeStatus>
: 1♁
<specimenCount id="9D6AB5DDFF9B864A263EF8AAFBF32E73" box="[1052,1087,1863,1890]" count="1" pageId="3" pageNumber="348" type="female">1♀</specimenCount>
(
<collectionCode id="ED7DE691FF9B864A266DF8ABFB682E71" box="[1103,1188,1862,1888]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="3" pageNumber="348">ZMUT</collectionCode>
), same data as for the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3762DDFFF9B864A22E5F863FF3C2EDD" pageId="3" pageNumber="348" type="etymology">
<paragraph id="8BD37E54FF9B864A22E5F863FF3C2EDD" blockId="3.[151,1436,1826,1996]" pageId="3" pageNumber="348">
<emphasis id="B918A246FF9B864A22E5F863FE822EB9" bold="true" box="[199,334,1934,1960]" pageId="3" pageNumber="348">Etymology.</emphasis>
The specific epithet is a Latin adjective, referring to the pale coloration of both sexes of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3762DDFFF9C864D22E5FF7BFC372851" pageId="4" pageNumber="349" type="diagnosis">
<paragraph id="8BD37E54FF9C864D22E5FF7BFC372851" blockId="4.[151,1437,150,573]" pageId="4" pageNumber="349">
<emphasis id="B918A246FF9C864D22E5FF7BFE8E29A1" bold="true" box="[199,322,150,176]" pageId="4" pageNumber="349">Diagnosis.</emphasis>
This species is most similar to
<taxonomicName id="4C6C05D7FF9C864D2091FF7AFCED29A1" authorityName="Zamani &amp; Marusik" authorityYear="2019" box="[691,801,150,177]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="349" phylum="Arthropoda" rank="species" species="alamto">
<emphasis id="B918A246FF9C864D2091FF7AFD0F29A1" box="[691,707,151,176]" italics="true" pageId="4" pageNumber="349">T</emphasis>
.
<emphasis id="B918A246FF9C864D20F1FF7BFCED29A1" box="[723,801,150,176]" italics="true" pageId="4" pageNumber="349">alamto</emphasis>
</taxonomicName>
from western
<collectingCountry id="F37B3EC4FF9C864D21E8FF7AFC3629A0" box="[970,1018,151,177]" name="Iran" pageId="4" pageNumber="349">Iran</collectingCountry>
, from which it differs by the angled anterior edge of the conductor (
<emphasis id="B918A246FF9C864D23D9FF56FDDE29C5" box="[507,530,187,212]" italics="true" pageId="4" pageNumber="349">vs</emphasis>
. roundly bent), the bulb (from posterior edge to embolus) being longer than it is wide (
<emphasis id="B918A246FF9C864D22C2FF32FF3B29E9" box="[224,247,223,248]" italics="true" pageId="4" pageNumber="349">vs</emphasis>
. as long as wide), the embolus base positioned at 8:30 o’clock (
<emphasis id="B918A246FF9C864D21FAFF32FC2329E9" box="[984,1007,223,248]" italics="true" pageId="4" pageNumber="349">vs</emphasis>
. 9:30), the roundly bent embolus (
<emphasis id="B918A246FF9C864D275CFF32FA5929E9" box="[1406,1429,223,248]" italics="true" pageId="4" pageNumber="349">vs</emphasis>
. the embolus bent at right angle), the relatively long conductor (cf.
<figureCitation id="135762D1FF9C864D21A2FEEFFBE3280D" box="[896,1071,258,284]" captionStart="FIGURE 2" captionStartId="2.[151,250,1975,1999]" captionTargetBox="[161,1427,694,1948]" captionTargetId="figure-110@2.[151,1436,694,1951]" captionTargetPageId="2" captionText="FIGURE 2. Male palps of Tegenaria pallens sp. n. (A, B) and T. alamto (C).A, C ventral view; B retrolateral view. C reproduced from Zamani et al. (2018). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10084542" httpUri="https://zenodo.org/record/10084542/files/figure.png" pageId="4" pageNumber="349">Fig. 2A and 2C</figureCitation>
), and the converging receptacles forming a right angle (
<emphasis id="B918A246FF9C864D23B7FECAFE602851" box="[405,428,295,320]" italics="true" pageId="4" pageNumber="349">vs</emphasis>
. obtuse; cf.
<figureCitation id="135762D1FF9C864D2014FECBFD452851" box="[566,649,294,320]" captionStart="FIGURE 3" captionStartId="3.[151,250,1082,1106]" captionTargetBox="[151,1436,181,1058]" captionTargetId="figure-18@3.[151,1436,181,1058]" captionTargetPageId="3" captionText="FIGURE 3. Epigyne of Tegenaria pallens sp. n. A macerated, ventral view; B vulva, dorsal view; C intact, ventral view. Scale bar: 0.2 mm. Abbreviation: Co—copulatory opening." figureDoi="http://doi.org/10.5281/zenodo.10084544" httpUri="https://zenodo.org/record/10084544/files/figure.png" pageId="4" pageNumber="349">Fig. 3B</figureCitation>
and
<bibRefCitation id="EFFD03A5FF9C864D20E2FECBFC542851" author="Zamani, A. &amp; Marusik, Y. M. &amp; Malek-Hosseini, M. J." box="[704,920,294,321]" pageId="4" pageNumber="349" pagination="95 - 97" refId="ref9868" refString="Zamani, A., Marusik, Y. M. &amp; Malek-Hosseini, M. J. (2018) A new species of Tegenaria Latreille, 1804 (Araneae: Agelenidae) from western Iran. Zootaxa, 4444 (1), 95 - 97. https: // doi. org / 10.11646 / zootaxa. 4444.1.7" type="journal article" year="2018">
Zamani
<emphasis id="B918A246FF9C864D213FFECAFC822851" box="[797,846,294,320]" italics="true" pageId="4" pageNumber="349">et al</emphasis>
. 2018
</bibRefCitation>
: fig. 11).
</paragraph>
</subSubSection>
<subSubSection id="C3762DDFFF9C864C22E5FEA7FB8628BC" lastPageId="5" lastPageNumber="350" pageId="4" pageNumber="349" type="description">
<paragraph id="8BD37E54FF9C864D22E5FEA7FC022B2D" blockId="4.[151,1437,150,573]" pageId="4" pageNumber="349">
<emphasis id="B918A246FF9C864D22E5FEA7FE952875" bold="true" box="[199,345,330,356]" pageId="4" pageNumber="349">Description.</emphasis>
Male (
<typeStatus id="54D7C0F6FF9C864D238EFEA7FDD72875" box="[428,539,330,356]" pageId="4" pageNumber="349" type="holotype">Holotype</typeStatus>
). Habitus as in
<figureCitation id="135762D1FF9C864D20EFFEA7FCEA2875" box="[717,806,330,356]" captionStart="FIGURE 1" captionStartId="1.[151,250,1254,1278]" captionTargetBox="[151,1436,345,1225]" captionTargetId="figure-87@1.[151,1436,345,1231]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Tegenaria pallens sp. n., dorsal view. A female; B male." figureDoi="http://doi.org/10.5281/zenodo.10084540" httpUri="https://zenodo.org/record/10084540/files/figure.png" pageId="4" pageNumber="349">Fig. 1B</figureCitation>
. Total length 7.60. Carapace 3.40 long, 2.63 wide. Eye sizes: AME 0.14, ALE 0.17, PME 0.13, PLE 0.17. Cephalothorax and its appendages overall pale; pars cephalica, chelicerae, palps and femora I light brown. Chelicerae with 3 pro- and 3 retromarginal teeth.Abdomen and spinnerets light beige. Measurements of legs: I: 27.82 (7.50, 1.53, 7.50, 8.12, 3.17), II: 22.87 (6.20, 1.50, 5.76, 6.73, 2.68), III: 20.13 (5.50, 1.25, 4.78, 6.30, 2.30), IV: 25.37 (6.50, 1.45, 6.35, 8.37, 2.70). Spination: Palp: Fm: 3d. Legs: I: Fm: 2d, 4(5)pl, 2rl; Ti: 2pl, 2rl, 4v; Mt:
<quantity id="4C94D3B1FF9C864D2004FE13FDF32B09" box="[550,575,510,536]" metricMagnitude="3" metricUnit="kg" metricValue="8.0" pageId="4" pageNumber="349" unit="t" value="8.0">8t</quantity>
. II: Fm: 2d, 5pl, 2rl; Ti: 2pl, 2rl, 4v; Mt:
<quantity id="4C94D3B1FF9C864D262DFE13FBE42B09" box="[1039,1064,510,536]" metricMagnitude="3" metricUnit="kg" metricValue="9.0" pageId="4" pageNumber="349" unit="t" value="9.0">9t</quantity>
. III: Fm: 2d, 3pl, 2rl; Ti: 2d, 2pl, 2rl, 3v; Mt:
<quantity id="4C94D3B1FF9C864D233DFDCFFE892B2D" box="[287,325,546,572]" metricMagnitude="4" metricUnit="kg" metricValue="1.3" pageId="4" pageNumber="349" unit="t" value="13.0">13t</quantity>
. IV: Fm: 1(2)d, 1pl, 1(2)rl; Ti: 2d, 2pl, 2rl, 4(6)v; Mt:
<quantity id="4C94D3B1FF9C864D2186FDCFFC062B2D" box="[932,970,546,572]" metricMagnitude="4" metricUnit="kg" metricValue="1.3" pageId="4" pageNumber="349" unit="t" value="13.0">13t</quantity>
.
</paragraph>
<caption id="DF132EDCFF9C864D22B5F94CFB082FA8" ID-DOI="http://doi.org/10.5281/zenodo.10084546" ID-Zenodo-Dep="10084546" box="[151,1220,1697,1722]" httpUri="https://zenodo.org/record/10084546/files/figure.png" pageId="4" pageNumber="349" startId="4.[151,250,1697,1721]" targetBox="[161,1424,596,1665]" targetPageId="4" targetType="figure">
<paragraph id="8BD37E54FF9C864D22B5F94CFB082FA8" blockId="4.[151,1220,1697,1722]" box="[151,1220,1697,1722]" pageId="4" pageNumber="349">
<emphasis id="B918A246FF9C864D22B5F94CFEDF2FA8" bold="true" box="[151,275,1697,1721]" pageId="4" pageNumber="349">FIGURE 4.</emphasis>
Habitus of
<taxonomicName id="4C6C05D7FF9C864D23A8F94CFDF12FA8" authority="Zamani &amp; Darvishnia &amp; Marusik, 2023" authorityName="Zamani &amp; Darvishnia &amp; Marusik" authorityYear="2023" box="[394,573,1697,1721]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="349" phylum="Arthropoda" rank="species" species="elamana" status="sp. nov.">
<emphasis id="B918A246FF9C864D23A8F94CFDF12FA8" box="[394,573,1697,1721]" italics="true" pageId="4" pageNumber="349">Dysdera elamana</emphasis>
</taxonomicName>
<emphasis id="B918A246FF9C864D2066F94FFDB12FAB" bold="true" box="[580,637,1698,1722]" pageId="4" pageNumber="349">
<taxonomicNameLabel id="A22B1F3DFF9C864D2066F94FFDB12FAB" box="[580,637,1698,1722]" pageId="4" pageNumber="349" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(A) and
<taxonomicName id="4C6C05D7FF9C864D20F5F94FFC412FA8" authority="Zamani &amp; Darvishnia &amp; Marusik, 2023" authorityName="Zamani &amp; Darvishnia &amp; Marusik" authorityYear="2023" box="[727,909,1698,1721]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="349" phylum="Arthropoda" rank="species" species="zagros" status="sp. nov.">
<emphasis id="B918A246FF9C864D20F5F94FFC412FA8" box="[727,909,1698,1721]" italics="true" pageId="4" pageNumber="349">Harpactea zagros</emphasis>
</taxonomicName>
<emphasis id="B918A246FF9C864D21B1F94FFC002FAB" bold="true" box="[915,972,1698,1722]" pageId="4" pageNumber="349">
<taxonomicNameLabel id="A22B1F3DFF9C864D21B1F94FFC002FAB" box="[915,972,1698,1722]" pageId="4" pageNumber="349" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(B), males, dorsal view.
</paragraph>
</caption>
<paragraph id="8BD37E54FF9C864D22E5F905FA432EA7" blockId="4.[151,1437,1768,2010]" pageId="4" pageNumber="349">
Palp as in
<figureCitation id="135762D1FF9C864D2361F905FE732E13" box="[323,447,1768,1794]" captionStart="FIGURE 2" captionStartId="2.[151,250,1975,1999]" captionTargetBox="[161,1427,694,1948]" captionTargetId="figure-110@2.[151,1436,694,1951]" captionTargetPageId="2" captionText="FIGURE 2. Male palps of Tegenaria pallens sp. n. (A, B) and T. alamto (C).A, C ventral view; B retrolateral view. C reproduced from Zamani et al. (2018). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10084542" httpUri="https://zenodo.org/record/10084542/files/figure.png" pageId="4" pageNumber="349">Fig. 2A–B</figureCitation>
; tibia ca. 2 times longer than wide, 2 times shorter than cymbium, with 2 apophyses: membranous and roundly bent on tip retrolateral apophysis, and well sclerotized retro-dorsal digitiform apophysis with pointed tip; cymbium ca. 2.2 times longer than wide, with tip comprising ca. half of cymbium’s length; bulb longer than wide; sperm duct running along edge of tegulum; tegular apophysis elongate, ca. half of cymbium's width, originates at ca. 4 o’clock position; conductor as long as wide, with stem ca. 0.33 of conductor’s anterior width, anterior edge of conductor angled; embolus filiform, roundly bent, originating at ca. 8:30 o’clock position.
</paragraph>
<paragraph id="8BD37E54FF9C864C22E5F82DFDE72850" blockId="4.[151,1437,1768,2010]" lastBlockId="5.[151,1437,150,429]" lastPageId="5" lastPageNumber="350" pageId="4" pageNumber="349">
Female. Habitus as in
<figureCitation id="135762D1FF9C864D23E5F82DFDEC2ECB" box="[455,544,1984,2010]" captionStart="FIGURE 1" captionStartId="1.[151,250,1254,1278]" captionTargetBox="[151,1436,345,1225]" captionTargetId="figure-87@1.[151,1436,345,1231]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Tegenaria pallens sp. n., dorsal view. A female; B male." figureDoi="http://doi.org/10.5281/zenodo.10084540" httpUri="https://zenodo.org/record/10084540/files/figure.png" pageId="4" pageNumber="349">Fig. 1A</figureCitation>
. Total length 8.90. Carapace 3.87 long, 2.70 wide. Eye sizes: AME 0.11, ALE 0.15, PME 0.12, PLE 0.16. Colouration as in male. Chelicerae with 3 pro- and 3 retromarginal teeth. Measurements of legs: I: 21.92 (5.96, 1.53, 5.76, 6.14, 2.53), II: 18.51 (5.13, 1.40, 4.65, 5.20, 2.13), III: 16.73 (4.60, 1.33, 3.95, 5.00, 1.85), IV: 21.67 (5.80, 1.37, 5.50, 6.80, 2.20). Spination: Palp: Fm: 3d. Legs: I: Fm: 2d, 4pl, 2rl; Ti: 2pl, 3rl, 4v; Mt:
<quantity id="4C94D3B1FF9D864C22D3FEEEFEC6280C" box="[241,266,259,285]" metricMagnitude="3" metricUnit="kg" metricValue="7.0" pageId="5" pageNumber="350" unit="t" value="7.0">7t</quantity>
. II: Fm: 2d, 4pl, 2rl; Ti: 2pl, 2rl, 4v; Mt:
<quantity id="4C94D3B1FF9D864C20F6FEEEFD20280C" box="[724,748,259,285]" metricMagnitude="3" metricUnit="kg" metricValue="9.0" pageId="5" pageNumber="350" unit="t" value="9.0">9t</quantity>
. III: Fm: 2d, 2pl, 2rl; Ti: 2d, 2pl, 2rl, 5v; Mt:
<quantity id="4C94D3B1FF9D864C26C8FEEEFADC280C" box="[1258,1296,259,285]" metricMagnitude="4" metricUnit="kg" metricValue="1.1" pageId="5" pageNumber="350" unit="t" value="11.0">11t</quantity>
. IV: Fm: 2d, 1pl, 1rl; Ti: 2d, 2pl, 2rl, 5v; Mt:
<quantity id="4C94D3B1FF9D864C2022FECBFDEB2850" box="[512,551,294,321]" metricMagnitude="4" metricUnit="kg" metricValue="1.5" pageId="5" pageNumber="350" unit="t" value="15.0">15t</quantity>
.
</paragraph>
<paragraph id="8BD37E54FF9D864C22E5FEA6FB8628BC" blockId="5.[151,1437,150,429]" pageId="5" pageNumber="350">
Epigyne as in
<figureCitation id="135762D1FF9D864C234AFEA6FE2E2874" box="[360,482,331,357]" captionStart="FIGURE 3" captionStartId="3.[151,250,1082,1106]" captionTargetBox="[151,1436,181,1058]" captionTargetId="figure-18@3.[151,1436,181,1058]" captionTargetPageId="3" captionText="FIGURE 3. Epigyne of Tegenaria pallens sp. n. A macerated, ventral view; B vulva, dorsal view; C intact, ventral view. Scale bar: 0.2 mm. Abbreviation: Co—copulatory opening." figureDoi="http://doi.org/10.5281/zenodo.10084544" httpUri="https://zenodo.org/record/10084544/files/figure.png" pageId="5" pageNumber="350">Fig. 3A–C</figureCitation>
; epigynal plate over 3 times wider than long; fovea oval, transverse; teeth indistinct; copulatory openings (
<emphasis id="B918A246FF9D864C23B7FE82FE7A2898" box="[405,438,367,393]" italics="true" pageId="5" pageNumber="350">Co</emphasis>
) located on anterior edges of fovea; copulatory ducts seem coiled, forming 2 loops; receptacles oval, together with copulatory ducts converging and forming right angle.
</paragraph>
</subSubSection>
<caption id="DF132EDCFF9D864C22B5F84AFCAA2EF2" ID-DOI="http://doi.org/10.5281/zenodo.10084548" ID-Zenodo-Dep="10084548" httpUri="https://zenodo.org/record/10084548/files/figure.png" pageId="5" pageNumber="350" startId="5.[151,250,1959,1983]" targetBox="[194,1406,464,1936]" targetPageId="5" targetType="figure">
<paragraph id="8BD37E54FF9D864C22B5F84AFCAA2EF2" blockId="5.[151,1437,1959,2020]" pageId="5" pageNumber="350">
<emphasis id="B918A246FF9D864C22B5F84AFED92ED1" bold="true" box="[151,277,1959,1984]" pageId="5" pageNumber="350">FIGURE 5.</emphasis>
Male palp of
<taxonomicName id="4C6C05D7FF9D864C238CF845FDA82ED1" authority="Zamani &amp; Darvishnia &amp; Marusik, 2023" authorityName="Zamani &amp; Darvishnia &amp; Marusik" authorityYear="2023" box="[430,612,1960,1984]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="350" phylum="Arthropoda" rank="species" species="elamana" status="sp. nov.">
<emphasis id="B918A246FF9D864C238CF845FDA82ED1" box="[430,612,1960,1984]" italics="true" pageId="5" pageNumber="350">Dysdera elamana</emphasis>
</taxonomicName>
<emphasis id="B918A246FF9D864C204FF845FD642ED1" bold="true" box="[621,680,1960,1984]" pageId="5" pageNumber="350">
<taxonomicNameLabel id="A22B1F3DFF9D864C204FF845FD642ED1" box="[621,680,1960,1984]" pageId="5" pageNumber="350" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A retrolateral view; B dorsal view; C ventral view. Scale bars: 0.2 mm. Abbreviation:
<emphasis id="B918A246FF9D864C2308F821FE902EF2" box="[298,348,1996,2019]" italics="true" pageId="5" pageNumber="350">Ac—</emphasis>
anterior crest,
<emphasis id="B918A246FF9D864C23CFF821FDE82EF2" box="[493,548,1996,2019]" italics="true" pageId="5" pageNumber="350">Mc—</emphasis>
median crest,
<emphasis id="B918A246FF9D864C2093F821FD2E2EF2" box="[689,738,1996,2019]" italics="true" pageId="5" pageNumber="350">Vc—</emphasis>
ventral crest.
</paragraph>
</caption>
<subSubSection id="C3762DDFFF9E864F22E5FF7BFD4F29C4" pageId="6" pageNumber="351" type="discussion">
<paragraph id="8BD37E54FF9E864F22E5FF7BFD4F29C4" blockId="6.[151,1437,150,285]" pageId="6" pageNumber="351">
<emphasis id="B918A246FF9E864F22E5FF7BFE8829A1" bold="true" box="[199,324,150,176]" pageId="6" pageNumber="351">Comment.</emphasis>
This is the first known Iranian agelenid species displaying a certain level troglomorphism (i.e., elongated appendages and depigmentation).
</paragraph>
</subSubSection>
<subSubSection id="C3762DDFFF9E864F22E5FF33FD8C280C" pageId="6" pageNumber="351" type="distribution">
<paragraph id="8BD37E54FF9E864F22E5FF33FD8C280C" blockId="6.[151,1437,150,285]" pageId="6" pageNumber="351">
<emphasis id="B918A246FF9E864F22E5FF33FEAD29E9" bold="true" box="[199,353,222,248]" pageId="6" pageNumber="351">Distribution.</emphasis>
Known only from the
<typeStatus id="54D7C0F6FF9E864F2056FF32FD6829E8" box="[628,676,223,249]" pageId="6" pageNumber="351">type</typeStatus>
locality in Gakal Cave (
<figureCitation id="135762D1FF9E864F21E8FF32FBF829E8" box="[970,1076,222,249]" captionStart="FIGURE 15" captionStartId="16.[151,250,1977,2001]" captionTargetBox="[158,1430,525,1953]" captionTargetId="figure-141@16.[158,1430,525,1953]" captionTargetPageId="16" captionText="FIGURE 15. Some of the collection localities of the material reported in this study.A Barreh Zard; B Chehel Sotoon; C Gakal; D Kabootarlan; E Khoffash; F Malek. Photos by Mohammad Javad Malek-Hosseini." figureDoi="http://doi.org/10.5281/zenodo.10084568" httpUri="https://zenodo.org/record/10084568/files/figure.png" pageId="6" pageNumber="351">Fig. 15C</figureCitation>
), Kohgiluyeh &amp; Boyer-Ahmad Province, southwestern
<collectingCountry id="F37B3EC4FF9E864F2381FEEEFE1C280C" box="[419,464,259,285]" name="Iran" pageId="6" pageNumber="351">Iran</collectingCountry>
(
<figureCitation id="135762D1FF9E864F23C2FEEEFDFF280C" box="[480,563,259,285]" captionStart="FIGURE 17" captionStartId="18.[151,250,1232,1256]" captionTargetBox="[275,1311,345,1209]" captionTargetId="figure-69@18.[275,1311,345,1209]" captionTargetPageId="18" captionText="FIGURE 17. Partial map of Iran, showing the collection localities of the material reported in this study. 1 Ali Sheikh Cave; 2 Barreh Zard Cave; 3 Chehel Sotoon Cave; 4 Darreh-ye Ashkaft Cave; 5 Dengezlu Cave; 6 Gakal Cave; 7 Gelim Goosh Cave; 8 Hamzeh Zendan Cave; 9 Jabiglu Cave; 10 Kabootarlan Cave; 11 Kahman Cave; 12 Kerend Cave; 13 Khoffash Cave, Mar Kelga Cave; 14 Malek Cave, Markhareil Cave; 15 Mir Melas Cave; 16 Mooly Cave; 17 Pir Morad Cave; 18 Raad-e Gharbi Cave; 19 Tang-e Kaboutari; 20 Tang-e-Lor Cave; 21 Tang-e Zendan Cave; 22 Tashan Cave; 23 Torang Cave; 24 Var-e-Zamin Cave; 25 Yaran Cave." figureDoi="http://doi.org/10.5281/zenodo.10084572" httpUri="https://zenodo.org/record/10084572/files/figure.png" pageId="6" pageNumber="351">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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