plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/C5/B2/03C5B216FFEBFFB4DC6BFE43697DFEB2.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

323 lines
42 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="EE2150FB245467A0A75B028577906B49" ID-DOI="10.5252/zoosystema2024v46a2" ID-ISSN="1638-9387" ID-ZooBank="4B49C1D9-1196-4942-969F-2E923B1FC12C" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1707561373589" checkinUser="plazi" docAuthor="Cupello, Mario" docDate="2024" docId="03C5B216FFEBFFB4DC6BFE43697DFEB2" docLanguage="en" docName="zoosystema.2024.46.2.pdf" docOrigin="Zoosystema 46 (2)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a2.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Gromphas lacordairii" docType="treatment" docVersion="6" lastPageNumber="34" masterDocId="FFFCCA6EFFE6FFB9DD69FFFA6C63FF8D" masterDocTitle="The genus Gromphas Dejean, 1836 (Coleoptera, Scarabaeinae): nomenclature, distribution, and conservation, including a contribution to the debate on electronic publications in zoology" masterLastPageNumber="59" masterPageNumber="23" pageNumber="34" updateTime="1708027481031" updateUser="jonas" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="83632E89220C95D271E1D72F638868BD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6E85E70976E94F293BD451832263C6D3">
<mods:title id="01FC5DADDF1B49B673223F675DC0666C">The genus Gromphas Dejean, 1836 (Coleoptera, Scarabaeinae): nomenclature, distribution, and conservation, including a contribution to the debate on electronic publications in zoology</mods:title>
</mods:titleInfo>
<mods:name id="A77DB18E621BA3F9D08D338DA4544858" type="personal">
<mods:role id="2C5B78B235260D2F0BF18EE8C960FC75">
<mods:roleTerm id="13A408A6F2C3AD7B245FC1D22ED291FC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2B3894A88202F0E2BC80E15A1CB621A0">Cupello, Mario</mods:namePart>
<mods:affiliation id="397A3C7AFA49FD5AE7E6F4A11F0CC345">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná (Brazil) &amp; Department of Entomology, Texas A &amp; M University, College Station, TX 77843 (United States)</mods:affiliation>
<mods:nameIdentifier id="F96FA259CADFA957EA574D31C141627D" type="email">mcupello@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="C35241B96F8F3DA9D626D4B1E4C26E16">text</mods:typeOfResource>
<mods:relatedItem id="2FF78BA706032A88AB2179705E7AD060" type="host">
<mods:titleInfo id="BD8FC11429C51D2193C1CA997D38F6F8">
<mods:title id="FC9A9E5796F3900D228264199935F23C">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="141E2EA336230675FDAC017DD34FDDBC">
<mods:date id="7C7589575DFB9ED73BEAEE59B8ABD447">2024</mods:date>
<mods:detail id="A839CE46497F8FB2D06A11F695C3AB69" type="pubDate">
<mods:number id="F33A7177FC196D7FC9770363E22B9378">2024-02-09</mods:number>
</mods:detail>
<mods:detail id="9769C9F10ED1CEC2D0C56A2254878287" type="volume">
<mods:number id="9837E2902C50002950CF6F01DC6D01DA">46</mods:number>
</mods:detail>
<mods:detail id="8CD0BEF85391FFF0DF1327F1F64C8EB8" type="issue">
<mods:number id="F222A228535FE5000F06B8D68B8040AE">2</mods:number>
</mods:detail>
<mods:extent id="35B4674253052B919DFE5AA5D728D0A6" unit="page">
<mods:start id="FAA4A881A6D51E3CFC86995996706D64">23</mods:start>
<mods:end id="3254437CA163B7B795C7E08221630348">59</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="82AEC5248C94E6C5D9F9DD81E779BE6B">
<mods:url id="1BF66B3377EA12FAFD1855B431679777">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a2.pdf</mods:url>
</mods:location>
<mods:classification id="9090B3DA13AC1E688F3BCE23F7F1ACCE">journal article</mods:classification>
<mods:identifier id="7944E7F462CBE5589056C96C15D002DA" type="DOI">10.5252/zoosystema2024v46a2</mods:identifier>
<mods:identifier id="B0D2A71BA7A56A9D654A700918CA07AA" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="B9E840499209742620752E2A7CABDF24" type="ZooBank">4B49C1D9-1196-4942-969F-2E923B1FC12C</mods:identifier>
</mods:mods>
<treatment id="03C5B216FFEBFFB4DC6BFE43697DFEB2" ID-DOI="http://doi.org/10.5281/zenodo.10643534" ID-Zenodo-Dep="10643534" LSID="urn:lsid:plazi:treatment:03C5B216FFEBFFB4DC6BFE43697DFEB2" httpUri="http://treatment.plazi.org/id/03C5B216FFEBFFB4DC6BFE43697DFEB2" lastPageNumber="34" pageId="13" pageNumber="34">
<subSubSection id="C376508BFFEBFFB4DC6BFE436EEAFE5E" box="[258,649,441,468]" pageId="13" pageNumber="34" type="nomenclature">
<paragraph id="8BD30300FFEBFFB4DC6BFE436EEAFE5E" blockId="13.[258,649,441,468]" box="[258,649,441,468]" pageId="13" pageNumber="34">
<heading id="D09BB46CFFEBFFB4DC6BFE436EEAFE5E" box="[258,649,441,468]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="34" reason="2">
<taxonomicName id="4C6C7883FFEBFFB4DC6BFE436EEAFE5E" ID-CoL="6L4H6" authority="(Oken, 1834)" baseAuthorityName="Oken" baseAuthorityYear="1834" box="[258,649,441,468]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="lacordairii">
<emphasis id="B918DF12FFEBFFB4DC6BFE436D8EFE59" bold="true" box="[258,493,441,468]" italics="true" pageId="13" pageNumber="34">Gromphas lacordairii</emphasis>
(
<bibRefCitation id="EFFD7EF1FFEBFFB4DC92FE436EE2FE5E" author="OKEN L." box="[507,641,441,467]" journalOrPublisher="Isis von Oken" pageId="13" pageNumber="34" pagination="1112 - 1127" part="11" publicationUrl="https://www.biodiversitylibrary.org/page/27510335" refId="ref34279" refString="[OKEN L.] 1834. - J. Th. Lacordaire, uber die Lebensweise der Kafer im mittaglichen America. Viele Kafer namentlich angefuhrt. Isis von Oken [volume for the year 1834] (11): 1112 - 1127. [The authorship of this review is not stated expressly anywhere, as far as I could find, but the context makes clear that the journal's editor, Lorenz Oken (1779 - 1851), the famous German naturalist of the Naturphilosophie movement, was the one who wrote it and the other literature reviews that appeared in the journal. For this reason, Oken should be credited with its authorship under Article 50.1 of the Code. In Cupello et al. (2022), we conservatively treated the review as published anonymously, but this seems unjustified. Kertesz (1986) tells the details of the history of Isis von Oken and confirms that Oken was the one who prepared the literature reviews] [publication date for nomenclatural purposes: 30 th November 1834. Neal Evenhuis (personal communication, 26 th January 2023) confirmed that Isis von Oken appeared in monthly issues. Oken's review of Lacordaire's paper appeared in the 11 th issue of 1834, therefore, in November of that year, with no more specific date known to me. Following Article 21.3. 1, I take the last day of that month as the publication date]. https: // www. biodiversitylibrary. org / page / 27510335" title="J. Th. Lacordaire, uber die Lebensweise der Kafer im mittaglichen America. Viele Kafer namentlich angefuhrt" type="journal article" year="1834">Oken, 1834</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C376508BFFEBFFB4DDEDFE086DDAFD36" pageId="13" pageNumber="34" type="reference_group">
<paragraph id="8BD30300FFEBFFB4DDEDFE086E9AFD86" blockId="13.[132,761,498,523]" box="[132,761,498,523]" pageId="13" pageNumber="34">
<treatmentCitationGroup id="AB7C242EFFEBFFB4DDEDFE086E9AFD86" box="[132,761,498,523]" pageId="13" pageNumber="34">
<treatmentCitation id="0ACD2511FFEBFFB4DDEDFE086E6BFD86" author="OKEN L." box="[132,520,498,523]" page="1117" pageId="13" pageNumber="34" year="1834">
<taxonomicName id="4C6C7883FFEBFFB4DDEDFE086E6BFD86" authority="Oken, 1834: 1117" authorityName="Oken" authorityPageNumber="1117" authorityYear="1834" box="[132,520,498,523]" class="Insecta" family="Scarabaeidae" genus="Coprobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="lacordairii">
<emphasis id="B918DF12FFEBFFB4DDEDFE086D29FD87" box="[132,330,498,522]" italics="true" pageId="13" pageNumber="34">Coprobius lacordairii</emphasis>
<bibRefCitation id="EFFD7EF1FFEBFFB4DC38FE086E6BFD86" author="OKEN L." box="[337,520,498,523]" journalOrPublisher="Isis von Oken" pageId="13" pageNumber="34" pagination="1112 - 1127" part="11" publicationUrl="https://www.biodiversitylibrary.org/page/27510335" refId="ref34279" refString="[OKEN L.] 1834. - J. Th. Lacordaire, uber die Lebensweise der Kafer im mittaglichen America. Viele Kafer namentlich angefuhrt. Isis von Oken [volume for the year 1834] (11): 1112 - 1127. [The authorship of this review is not stated expressly anywhere, as far as I could find, but the context makes clear that the journal's editor, Lorenz Oken (1779 - 1851), the famous German naturalist of the Naturphilosophie movement, was the one who wrote it and the other literature reviews that appeared in the journal. For this reason, Oken should be credited with its authorship under Article 50.1 of the Code. In Cupello et al. (2022), we conservatively treated the review as published anonymously, but this seems unjustified. Kertesz (1986) tells the details of the history of Isis von Oken and confirms that Oken was the one who prepared the literature reviews] [publication date for nomenclatural purposes: 30 th November 1834. Neal Evenhuis (personal communication, 26 th January 2023) confirmed that Isis von Oken appeared in monthly issues. Oken's review of Lacordaire's paper appeared in the 11 th issue of 1834, therefore, in November of that year, with no more specific date known to me. Following Article 21.3. 1, I take the last day of that month as the publication date]. https: // www. biodiversitylibrary. org / page / 27510335" title="J. Th. Lacordaire, uber die Lebensweise der Kafer im mittaglichen America. Viele Kafer namentlich angefuhrt" type="journal article" year="1834">Oken, 1834: 1117</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[
<date id="FFD225C0FFEBFFB4DF7DFE086E8CFD86" box="[532,751,498,523]" pageId="13" pageNumber="34" value="1834-11-30">30th November 1834</date>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD30300FFEBFFB4DDEDFDD86F65FDB6" blockId="13.[132,774,546,571]" box="[132,774,546,571]" pageId="13" pageNumber="34">
<treatmentCitationGroup id="AB7C242EFFEBFFB4DDEDFDD86F65FDB6" box="[132,774,546,571]" pageId="13" pageNumber="34">
<treatmentCitation id="0ACD2511FFEBFFB4DDEDFDD86D84FDB7" author="Harold" box="[132,487,546,570]" page="62" pageId="13" pageNumber="34" year="1869">
<taxonomicName id="4C6C7883FFEBFFB4DDEDFDD86D84FDB7" ID-CoL="6L4GZ" authority="Harold, 1869 b: 62" authorityName="Harold" authorityPageNumber="62" authorityYear="1869" box="[132,487,546,570]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B918DF12FFEBFFB4DDEDFDD86D4DFDB7" box="[132,302,546,570]" italics="true" pageId="13" pageNumber="34">Gromphas inermis</emphasis>
<bibRefCitation id="EFFD7EF1FFEBFFB4DC5AFDD86DD5FDB7" author="Harold" box="[307,438,546,570]" firstAuthor="Harold" pageId="13" pageNumber="34" refId="ref32645" refString="HAROLD E. 1869 a. - Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich: 979 - 1346 + [8]. [publication date for nomenclatural purposes: 27 th May 1969 (based on Bousquet 2016)] https: // www. biodiversitylibrary. org / page / 9641549" type="journal volume" year="1869">Harold, 1869</bibRefCitation>
b: 62
</taxonomicName>
</treatmentCitation>
[
<date id="FFD225C0FFEBFFB4DC9AFDD96EDBFDB7" box="[499,696,546,571]" pageId="13" pageNumber="34" value="1869-11-01">1st November 1869</date>
],
<emphasis id="B918DF12FFEBFFB4DFAFFDD96F65FDB6" bold="true" box="[710,774,547,571]" pageId="13" pageNumber="34">n. syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD30300FFEBFFB4DDEDFDA96DDAFD36" blockId="13.[131,775,594,699]" pageId="13" pageNumber="34">
<treatmentCitationGroup id="AB7C242EFFEBFFB4DDEDFDA96D3AFD0B" pageId="13" pageNumber="34">
<treatmentCitation id="0ACD2511FFEBFFB4DDEDFDA96EFCFDE7" author="d'Olsoufieff" box="[132,671,594,619]" page="59" pageId="13" pageNumber="34" year="1924">
<taxonomicName id="4C6C7883FFEBFFB4DDEDFDA96EFCFDE7" ID-CoL="7GVPV" authority="d'Olsoufieff, 1924: 59" authorityName="d'Olsoufieff" authorityPageNumber="59" authorityYear="1924" box="[132,671,594,619]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="lacordairii" subSpecies="bipunctata">
<emphasis id="B918DF12FFEBFFB4DDEDFDA96DD8FDE6" box="[132,443,595,619]" italics="true" pageId="13" pageNumber="34">Gromphas lacordairii bipunctata</emphasis>
d’Olsoufieff, 1924: 59
</taxonomicName>
</treatmentCitation>
[
<date id="FFD225C0FFEBFFB4DFC7FDA96D6BFD0B" pageId="13" pageNumber="34" value="1924-12-31">31st December 1924</date>
],
<emphasis id="B918DF12FFEBFFB4DC71FD946D3AFD0B" bold="true" box="[280,345,622,646]" pageId="13" pageNumber="34">n. syn.</emphasis>
</treatmentCitationGroup>
[subjective synonymy with
<taxonomicName id="4C6C7883FFEBFFB4DF0EFD946EA9FD08" ID-CoL="6L4GZ" authorityName="Harold" authorityYear="1869" box="[615,714,622,645]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B918DF12FFEBFFB4DF0EFD946EA9FD08" box="[615,714,622,645]" italics="true" pageId="13" pageNumber="34">G. inermis</emphasis>
</taxonomicName>
firstly established by
<bibRefCitation id="EFFD7EF1FFEBFFB4DC66FD726E1BFD2D" author="Cupello &amp; Vaz-de-Mello" box="[271,632,648,672]" firstAuthor="Cupello" page="464" pageId="13" pageNumber="34" pagination="439 - 482" refId="ref30610" refString="CUPELLO M. &amp; VAZ-DE-MELLO F. Z. 2013. - Taxonomic revision of the South American dung beetle genus Gromphas Brulle, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa 3722 (4): 439 - 482. https: // doi. org / 10.11646 / zootaxa. 3722.4.2" type="journal article" year="2013">Cupello &amp; Vaz-de-Mello (2013: 464)</bibRefCitation>
; here extended to
<taxonomicName id="4C6C7883FFEBFFB4DDF4FD5E6DCCFD36" ID-CoL="6L4H6" authority="(Oken, 1834)" baseAuthorityName="Oken" baseAuthorityYear="1834" box="[157,431,675,699]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="lacordairii">
<emphasis id="B918DF12FFEBFFB4DDF4FD5E6D7EFD36" box="[157,285,675,699]" italics="true" pageId="13" pageNumber="34">G. lacordairii</emphasis>
(
<bibRefCitation id="EFFD7EF1FFEBFFB4DC43FD596DCBFD36" author="OKEN L." box="[298,424,675,699]" journalOrPublisher="Isis von Oken" pageId="13" pageNumber="34" pagination="1112 - 1127" part="11" publicationUrl="https://www.biodiversitylibrary.org/page/27510335" refId="ref34279" refString="[OKEN L.] 1834. - J. Th. Lacordaire, uber die Lebensweise der Kafer im mittaglichen America. Viele Kafer namentlich angefuhrt. Isis von Oken [volume for the year 1834] (11): 1112 - 1127. [The authorship of this review is not stated expressly anywhere, as far as I could find, but the context makes clear that the journal's editor, Lorenz Oken (1779 - 1851), the famous German naturalist of the Naturphilosophie movement, was the one who wrote it and the other literature reviews that appeared in the journal. For this reason, Oken should be credited with its authorship under Article 50.1 of the Code. In Cupello et al. (2022), we conservatively treated the review as published anonymously, but this seems unjustified. Kertesz (1986) tells the details of the history of Isis von Oken and confirms that Oken was the one who prepared the literature reviews] [publication date for nomenclatural purposes: 30 th November 1834. Neal Evenhuis (personal communication, 26 th January 2023) confirmed that Isis von Oken appeared in monthly issues. Oken's review of Lacordaire's paper appeared in the 11 th issue of 1834, therefore, in November of that year, with no more specific date known to me. Following Article 21.3. 1, I take the last day of that month as the publication date]. https: // www. biodiversitylibrary. org / page / 27510335" title="J. Th. Lacordaire, uber die Lebensweise der Kafer im mittaglichen America. Viele Kafer namentlich angefuhrt" type="journal article" year="1834">Oken, 1834</bibRefCitation>
)
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="C376508BFFEBFFB4DDEDFD2E6EECFBCE" pageId="13" pageNumber="34" type="materials_examined">
<paragraph id="8BD30300FFEBFFB4DDEDFD2E6DB1FCAC" blockId="13.[132,775,723,962]" pageId="13" pageNumber="34">
<smallCapsWord id="8D3595DCFFEBFFB4DDEDFD2E6D79FD67" baselines="741,742,741,742" box="[132,282,724,747]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Name-bearing" pageId="13" pageNumber="34">NAME-BEARING</smallCapsWord>
<smallCapsWord id="8D3595DCFFEBFFB4DC49FD236D2EFD64" baselines="741" box="[288,333,729,745]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="type" pageId="13" pageNumber="34">TYPE</smallCapsWord>
. —
<materialsCitation id="3B04095DFFEBFFB4DC11FD296DADFCAC" pageId="13" pageNumber="34" specimenCount="1" typeStatus="syntype">
<taxonomicName id="4C6C7883FFEBFFB4DC11FD296E5EFD66" authorityName="Oken, 1834:" authorityYear="1117" box="[376,573,723,747]" class="Insecta" family="Scarabaeidae" genus="Coprobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="lacordairii">
<emphasis id="B918DF12FFEBFFB4DC11FD296E5EFD66" box="[376,573,723,747]" italics="true" pageId="13" pageNumber="34">
<originalDetermination id="A872BBFCFFEBFFB4DC11FD296E5EFD66" box="[376,573,723,747]" pageId="13" pageNumber="34">Coprobius lacordairii</originalDetermination>
</emphasis>
</taxonomicName>
:
<typeStatus id="54D7BDA2FFEBFFB4DF20FD2E6EFFFD61" box="[585,668,724,748]" pageId="13" pageNumber="34" type="syntype">syntypes</typeStatus>
(unknown sexes), unknown whereabouts, likely destroyed; allocation remains unambiguous (see the text above)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD30300FFEBFFB4DDEDFCDE6D02FCFF" blockId="13.[132,775,723,962]" pageId="13" pageNumber="34">
<materialsCitation id="3B04095DFFEBFFB4DDEDFCDE6D3EFCFF" collectionCode="MNHN" country="Argentina" determinerName="Cupello &amp; Vaz-de-Mello" location="Buenos Aires" municipality="Buenos Aires" pageId="13" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Buenos Aires">
<taxonomicName id="4C6C7883FFEBFFB4DDEDFCDE6D4DFCB1" ID-CoL="6L4GZ" authorityName="Harold" authorityYear="1869" box="[132,302,804,828]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B918DF12FFEBFFB4DDEDFCDE6D4DFCB1" box="[132,302,804,828]" italics="true" pageId="13" pageNumber="34">
<originalDetermination id="A872BBFCFFEBFFB4DDEDFCDE6D4DFCB1" box="[132,302,804,828]" pageId="13" pageNumber="34">Gromphas inermis</originalDetermination>
</emphasis>
</taxonomicName>
: lectotype (
<specimenCount id="9D6AC889FFEBFFB4DCCDFCDE6DB4FCB1" box="[420,471,804,828]" pageId="13" pageNumber="34" type="male">male</specimenCount>
), designated by
<bibRefCitation id="EFFD7EF1FFEBFFB4DF19FCDE6D3CFCDA" author="Cupello &amp; Vaz-de-Mello" firstAuthor="Cupello" page="464" pageId="13" pageNumber="34" pagination="439 - 482" refId="ref30610" refString="CUPELLO M. &amp; VAZ-DE-MELLO F. Z. 2013. - Taxonomic revision of the South American dung beetle genus Gromphas Brulle, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa 3722 (4): 439 - 482. https: // doi. org / 10.11646 / zootaxa. 3722.4.2" type="journal article" year="2013">
<determinerName id="7E0FEFC4FFEBFFB4DF19FCDE6EDDFCB1" box="[624,702,804,828]" pageId="13" pageNumber="34">Cupello</determinerName>
&amp;
<determinerName id="7E0FEFC4FFEBFFB4DFB5FCDF6CBEFCDA" pageId="13" pageNumber="34">Vaz-de-Mello</determinerName>
(2013: 464)
</bibRefCitation>
,
<collectionCode id="ED7D9BC5FFEBFFB4DC02FCC56DDDFCDB" box="[363,446,831,854]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="13" pageNumber="34">MNHN</collectionCode>
(ex Edgar von Harold and René Oberthür collections)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD30300FFEBFFB4DDEDFC8F6C90FC4F" blockId="13.[132,775,723,962]" pageId="13" pageNumber="34">
<materialsCitation id="3B04095DFFEBFFB4DDEDFC8F6C8CFC4F" collectionCode="MNHN" country="Brazil" determinerName="Cupello &amp; Vaz-de-Mello" location="Mato Grosso" pageId="13" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso">
<taxonomicName id="4C6C7883FFEBFFB4DDEDFC8F6DA3FC00" ID-CoL="7GVPV" authorityName="d'Olsoufieff" authorityYear="1924" box="[132,448,885,909]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="lacordairii" subSpecies="bipunctata">
<emphasis id="B918DF12FFEBFFB4DDEDFC8F6DA3FC00" box="[132,448,885,909]" italics="true" pageId="13" pageNumber="34">
<originalDetermination id="A872BBFCFFEBFFB4DDEDFC8F6DA3FC00" box="[132,448,885,909]" pageId="13" pageNumber="34">Gromphas lacordairii bipunctata</originalDetermination>
</emphasis>
</taxonomicName>
: lectotype (
<specimenCount id="9D6AC889FFEBFFB4DF5EFC8F6E08FC00" box="[567,619,885,909]" pageId="13" pageNumber="34" type="male">male</specimenCount>
), designated by
<bibRefCitation id="EFFD7EF1FFEBFFB4DDEDFC6A6D8CFC25" author="Cupello &amp; Vaz-de-Mello" box="[132,495,912,936]" firstAuthor="Cupello" page="464" pageId="13" pageNumber="34" pagination="439 - 482" refId="ref30610" refString="CUPELLO M. &amp; VAZ-DE-MELLO F. Z. 2013. - Taxonomic revision of the South American dung beetle genus Gromphas Brulle, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa 3722 (4): 439 - 482. https: // doi. org / 10.11646 / zootaxa. 3722.4.2" type="journal article" year="2013">
<determinerName id="7E0FEFC4FFEBFFB4DDEDFC6A6CB1FC25" box="[132,210,912,936]" pageId="13" pageNumber="34">Cupello</determinerName>
&amp;
<determinerName id="7E0FEFC4FFEBFFB4DD98FC6A6D17FC25" box="[241,372,912,936]" pageId="13" pageNumber="34">Vaz-de-Mello</determinerName>
(2013: 464)
</bibRefCitation>
,
<collectionCode id="ED7D9BC5FFEBFFB4DC91FC6A6E28FC2A" box="[504,587,912,935]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="13" pageNumber="34">MNHN</collectionCode>
(ex René Oberthür collection)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD30300FFEBFFB4DDEDFC216DAEFBA5" blockId="13.[132,775,987,1091]" pageId="13" pageNumber="34">
<smallCapsWord id="8D3595DCFFEBFFB4DDEDFC216CD5FC7D" baselines="1004,1004" box="[132,182,987,1010]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="13" pageNumber="34">
<typeStatus id="54D7BDA2FFEBFFB4DDEDFC216CD5FC7D" box="[132,182,987,1010]" pageId="13" pageNumber="34">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D3595DCFFEBFFB4DDD6FC1A6D7FFC7C" baselines="1005" box="[191,284,992,1009]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="13" pageNumber="34">LOCALITY</smallCapsWord>
. —
<materialsCitation id="3B04095DFFEBFFB4DC25FC216DABFBA5" country="Argentina" location="Buenos Aires" municipality="Buenos Aires" pageId="13" pageNumber="34" specimenCount="1" stateProvince="Buenos Aires">
<taxonomicName id="4C6C7883FFEBFFB4DC25FC216E7AFC7E" authorityName="Oken, 1834:" authorityYear="1117" box="[332,537,987,1011]" class="Insecta" family="Scarabaeidae" genus="Coprobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="lacordairii">
<emphasis id="B918DF12FFEBFFB4DC25FC216E7AFC7E" box="[332,537,987,1011]" italics="true" pageId="13" pageNumber="34">Coprobius lacordairii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C7883FFEBFFB4DF26FC216F63FC7F" authorityName="Harold" authorityYear="1869" box="[591,768,987,1011]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B918DF12FFEBFFB4DF26FC216F63FC7F" box="[591,768,987,1011]" italics="true" pageId="13" pageNumber="34">Gromphas inermis</emphasis>
</taxonomicName>
:
<collectingCountry id="F37B4390FFEBFFB4DDEDFC0F6C8AFB80" box="[132,233,1013,1037]" name="Argentina" pageId="13" pageNumber="34">Argentina</collectingCountry>
: either somewhere in the province of
<collectingRegion id="49A8CDE2FFEBFFB4DF01FC0C6E89FB80" box="[616,746,1013,1037]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="13" pageNumber="34">Buenos Aires</collectingRegion>
or the adjacent city of
<collectingMunicipality id="6BB7997AFFEBFFB4DC2FFBEB6DABFBA5" box="[326,456,1040,1064]" pageId="13" pageNumber="34">Buenos Aires</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD30300FFEBFFB4DDEDFBD16EECFBCE" blockId="13.[132,775,987,1091]" box="[132,655,1067,1091]" pageId="13" pageNumber="34">
<taxonomicName id="4C6C7883FFEBFFB4DDEDFBD16DD4FBCE" authorityName="d'Olsoufieff" authorityYear="1924" box="[132,439,1067,1091]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="lacordairii" subSpecies="bipunctata">
<emphasis id="B918DF12FFEBFFB4DDEDFBD16DD4FBCE" box="[132,439,1067,1091]" italics="true" pageId="13" pageNumber="34">Gromphas lacordairii bipunctata</emphasis>
</taxonomicName>
:
<materialsCitation id="3B04095DFFEBFFB4DCADFBD16EE8FBCE" box="[452,651,1067,1091]" country="Brazil" location="Mato Grosso" pageId="13" pageNumber="34" specimenCount="1" stateProvince="Mato Grosso">
<collectingCountry id="F37B4390FFEBFFB4DCADFBD16D9CFBCE" box="[452,511,1067,1091]" name="Brazil" pageId="13" pageNumber="34">Brazil</collectingCountry>
:
<collectingRegion id="49A8CDE2FFEBFFB4DF60FBD66EE8FBCE" box="[521,651,1067,1091]" country="Brazil" name="Mato Grosso" pageId="13" pageNumber="34">Mato Grosso</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C376508BFFEBFFB4DDEDFBA66EF5F98A" pageId="13" pageNumber="34" type="etymology">
<paragraph id="8BD30300FFEBFFB4DDEDFBA66EF5F98A" blockId="13.[130,777,1115,2027]" pageId="13" pageNumber="34">
<smallCapsWord id="8D3595DCFFEBFFB4DDEDFBA66C99FBFF" baselines="1133,1134" box="[132,250,1116,1139]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="13" pageNumber="34">ETYMOLOGY</smallCapsWord>
. —
<taxonomicName id="4C6C7883FFEBFFB4DC48FBA16DBCFBFE" authorityName="Oken, 1834:" authorityYear="1117" box="[289,479,1115,1139]" class="Insecta" family="Scarabaeidae" genus="Coprobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="lacordairii">
<emphasis id="B918DF12FFEBFFB4DC48FBA16DBCFBFE" box="[289,479,1115,1139]" italics="true" pageId="13" pageNumber="34">Coprobius lacordairii</emphasis>
</taxonomicName>
: the masculine singular genitive of the proper noun
<taxonomicName id="4C6C7883FFEBFFB4DC2CFB8C6DDBFB03" box="[325,440,1142,1166]" class="Insecta" family="Curculionidae" genus="Lacordairius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B918DF12FFEBFFB4DC2CFB8C6DDBFB03" box="[325,440,1142,1166]" italics="true" pageId="13" pageNumber="34">Lacordairius</emphasis>
</taxonomicName>
, the Latinised form of the French surname Lacordaire. It is an eponym after Belgian-French naturalist Jean Théodore Lacordaire (1801-1870), the collector of the
<typeStatus id="54D7BDA2FFEBFFB4DFB7FB566F64FB49" box="[734,775,1196,1220]" pageId="13" pageNumber="34">type</typeStatus>
material. The name is spelled as
<taxonomicName id="4C6C7883FFEBFFB4DCD0FB3D6E7CFB52" authorityName="Oken, 1834:" authorityYear="1117" box="[441,543,1223,1247]" class="Insecta" family="Scarabaeidae" genus="Coprobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="lacordairii">
‘
<emphasis id="B918DF12FFEBFFB4DCD6FB3D6E7CFB52" box="[447,543,1223,1247]" italics="true" pageId="13" pageNumber="34">lacordairii</emphasis>
</taxonomicName>
’, and not
<taxonomicName id="4C6C7883FFEBFFB4DFEBFB3D6E8AFB52" authorityName="Blanchard" authorityYear="1846" box="[642,745,1223,1247]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="lacordairei">
‘
<emphasis id="B918DF12FFEBFFB4DFE1FB3D6E8AFB52" box="[648,745,1223,1247]" italics="true" pageId="13" pageNumber="34">lacordairei</emphasis>
</taxonomicName>
’ as originally intended by its coiner
<bibRefCitation id="EFFD7EF1FFEBFFB4DCD8FB186E51FB77" author="Dejean" box="[433,562,1250,1274]" firstAuthor="Dejean" pageId="13" pageNumber="34" pagination="97 - 176" refId="ref31252" refString="DEJEAN P. F. M. A. 1833. - [livraison 2: 97 - 176], in Dejean P. F. M. A. 1833 - 1836. Catalogue des Coleopteres de la collection de M. le Comte Dejean. Mequignon-Marvis Pere et Fils, Paris, 443 p. [publication date for nomenclatural purposes: 27 th July 1833 (see Bousquet 2016)] https: // doi. org / 10.5962 / bhl. title. 8771" type="book chapter" year="1833">Dejean (1833</bibRefCitation>
,
<bibRefCitation id="EFFD7EF1FFEBFFB4DF54FB186E0CFB77" author="Dejean" box="[573,623,1250,1274]" firstAuthor="Dejean" pageId="13" pageNumber="34" pagination="1 - 384" refId="ref31341" refString="DEJEAN P. F. M. A. 1836. - [livraisons 1 - 4: 1 - 384], in DEJEAN P. F. M. A. 1836 - 1837. Catalogue des Coleopteres de la collection de M. le Comte Dejean. Troisieme edition, revue, corrigee et augmentee. Mequignon-Marvis Pere et Fils, Paris, v-xiv + 503 p. [publication date for nomenclatural purposes: 30 th July 1836 (see Bousquet 2016)]" type="book chapter" year="1836">1836</bibRefCitation>
) and so far cited in most of the literature, because Oken derived it not directly from Lacordaire’s name in modern French as Dejean did (i.e., the name Lacordaire + the Latin masculine genitive suffix -
<emphasis id="B918DF12FFEBFFB4DF24FAC96E37FAC7" box="[589,596,1331,1354]" italics="true" pageId="13" pageNumber="34">i</emphasis>
), but, as said, from its Latinized form,
<taxonomicName id="4C6C7883FFEBFFB4DC5AFAB76DC6FAE8" box="[307,421,1357,1381]" class="Insecta" family="Curculionidae" genus="Lacordairius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B918DF12FFEBFFB4DC5AFAB76DC6FAE8" box="[307,421,1357,1381]" italics="true" pageId="13" pageNumber="34">Lacordairius</emphasis>
</taxonomicName>
(i.e., by adding the masculine suffix -
<emphasis id="B918DF12FFEBFFB4DF97FAB46F66FAE8" box="[766,773,1358,1381]" italics="true" pageId="13" pageNumber="34">i</emphasis>
to the stem
<emphasis id="B918DF12FFEBFFB4DD84FA926D29FA0D" box="[237,330,1384,1408]" italics="true" pageId="13" pageNumber="34">Lacordairi</emphasis>
-; see Article 31.1.1 of the Code for more details). The
<taxonomicName id="4C6C7883FFEBFFB4DDC4FA796D77FA16" authorityName="Blanchard" authorityYear="1846" box="[173,276,1411,1435]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="lacordairei">
‘
<emphasis id="B918DF12FFEBFFB4DDDAFA796D77FA16" box="[179,276,1411,1435]" italics="true" pageId="13" pageNumber="34">lacordairei</emphasis>
</taxonomicName>
’ spelling is regarded here as an incorrect subsequent spelling in the terms of Article 33.3 of the Code; even though it is in prevailing usage, because this spelling has never been attributed to the publication of the original spelling (i.e., to
<bibRefCitation id="EFFD7EF1FFEBFFB4DF5DFA2E6EC2FA61" author="OKEN L." box="[564,673,1492,1516]" journalOrPublisher="Isis von Oken" pageId="13" pageNumber="34" pagination="1112 - 1127" part="11" publicationUrl="https://www.biodiversitylibrary.org/page/27510335" refId="ref34279" refString="[OKEN L.] 1834. - J. Th. Lacordaire, uber die Lebensweise der Kafer im mittaglichen America. Viele Kafer namentlich angefuhrt. Isis von Oken [volume for the year 1834] (11): 1112 - 1127. [The authorship of this review is not stated expressly anywhere, as far as I could find, but the context makes clear that the journal's editor, Lorenz Oken (1779 - 1851), the famous German naturalist of the Naturphilosophie movement, was the one who wrote it and the other literature reviews that appeared in the journal. For this reason, Oken should be credited with its authorship under Article 50.1 of the Code. In Cupello et al. (2022), we conservatively treated the review as published anonymously, but this seems unjustified. Kertesz (1986) tells the details of the history of Isis von Oken and confirms that Oken was the one who prepared the literature reviews] [publication date for nomenclatural purposes: 30 th November 1834. Neal Evenhuis (personal communication, 26 th January 2023) confirmed that Isis von Oken appeared in monthly issues. Oken's review of Lacordaire's paper appeared in the 11 th issue of 1834, therefore, in November of that year, with no more specific date known to me. Following Article 21.3. 1, I take the last day of that month as the publication date]. https: // www. biodiversitylibrary. org / page / 27510335" title="J. Th. Lacordaire, uber die Lebensweise der Kafer im mittaglichen America. Viele Kafer namentlich angefuhrt" type="journal article" year="1834">Oken 1834</bibRefCitation>
), it cannot be deemed a correct original spelling under Art. 33.3.1.
</paragraph>
</subSubSection>
<subSubSection id="C376508BFFEBFFB4DDEDF9F06DCCF867" pageId="13" pageNumber="34" type="discussion">
<paragraph id="8BD30300FFEBFFB4DDEDF9F06E89F9FF" blockId="13.[130,777,1115,2027]" pageId="13" pageNumber="34">
<emphasis id="B918DF12FFEBFFB4DDEDF9F06D55F9AC" box="[132,310,1546,1570]" italics="true" pageId="13" pageNumber="34">
<taxonomicName id="4C6C7883FFEBFFB4DDEDF9F06D52F9AC" ID-CoL="6L4GZ" authorityName="Harold" authorityYear="1869" box="[132,305,1546,1570]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="inermis">Gromphas inermis</taxonomicName>
:
</emphasis>
A Latin adjective in the nominative case meaning ‘unarmed’, ‘having no weapons’ (
<bibRefCitation id="EFFD7EF1FFEBFFB4DC86F9DE6EDAF9B0" author="Lewis &amp; Short" box="[495,697,1572,1597]" firstAuthor="Lewis" pageId="13" pageNumber="34" refId="ref33716" refString="LEWIS C. T. &amp; SHORT C. 1891. - Harpers' Latin dictionary. A new Latin dictionary founded on the translation of Freund's Latin-German lexicon edited by E. A. Andrews, LL. D. Revised, enlarged, and in great part rewritten. Harper &amp; Brothers, Franklin Square, New York, and Clarendon Press, Oxford, xiv + [2019] p." type="book" year="1891">Lewis &amp; Short 1891</bibRefCitation>
;
<bibRefCitation id="EFFD7EF1FFEBFFB4DFAFF9DF6CD4F9DA" author="Brown" firstAuthor="Brown" pageId="13" pageNumber="34" refId="ref29817" refString="BROWN R. W. 1954. - Composition of scientific words. A manual of methods and a lexicon of materials for the practice of logotechnics. Smithsonian Books, Washington, D. C., 882 p. https: // archive. org / details / compositionofsci 00 brow" type="book" year="1954">Brown 1954</bibRefCitation>
;
<bibRefCitation id="EFFD7EF1FFEBFFB4DDACF9C56D57F9DA" author="Glare" box="[197,308,1599,1623]" firstAuthor="Glare" pageId="13" pageNumber="34" refId="ref32292" refString="GLARE P. G. W. (ED.) 1968 - 1982. - Oxford Latin Dictionary. Clarendon Press, Oxford, xxiii + 2126 p." type="book" year="1968">Glare 1968</bibRefCitation>
-1982), a reference to the hornless condition of this species in contrast to the horned congeneric
<emphasis id="B918DF12FFEBFFB4DF08F9A16E89F9FF" box="[609,746,1627,1650]" italics="true" pageId="13" pageNumber="34">
<taxonomicName id="4C6C7883FFEBFFB4DF08F9A16E85F9FF" ID-CoL="6L4HD" baseAuthorityName="Perty" baseAuthorityYear="1830" box="[609,742,1627,1650]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="aeruginosa">G. aeruginosa</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BD30300FFEBFFB4DDEDF98F6DCCF867" blockId="13.[130,777,1115,2027]" pageId="13" pageNumber="34">
<taxonomicName id="4C6C7883FFEBFFB4DDEDF98F6DD9F900" ID-CoL="7GVPV" authorityName="d'Olsoufieff" authorityYear="1924" box="[132,442,1653,1677]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="lacordairii" subSpecies="bipunctata">
<emphasis id="B918DF12FFEBFFB4DDEDF98F6DD9F900" box="[132,442,1653,1677]" italics="true" pageId="13" pageNumber="34">Gromphas lacordairii bipunctata</emphasis>
</taxonomicName>
: A New Latin first-class adjective in the nominative case meaning ‘bi-punctate’, ‘bearing two punctures’, the combination of the Latin prefix
<emphasis id="B918DF12FFEBFFB4DF4FF9516E5AF94E" box="[550,569,1707,1731]" italics="true" pageId="13" pageNumber="34">bi</emphasis>
- for ‘two’, ‘occurring twice’, and the New Latin adjective
<emphasis id="B918DF12FFEBFFB4DCBAF93D6E4FF953" box="[467,556,1735,1758]" italics="true" pageId="13" pageNumber="34">punctatus</emphasis>
(-
<emphasis id="B918DF12FFEBFFB4DF29F93D6E28F953" box="[576,587,1735,1758]" italics="true" pageId="13" pageNumber="34">a</emphasis>
, -
<emphasis id="B918DF12FFEBFFB4DF37F93D6E1EF953" box="[606,637,1735,1758]" italics="true" pageId="13" pageNumber="34">um</emphasis>
) for ‘punctate’ (
<bibRefCitation id="EFFD7EF1FFEBFFB4DDE2F91B6D66F974" author="Brown" box="[139,261,1761,1785]" firstAuthor="Brown" pageId="13" pageNumber="34" refId="ref29817" refString="BROWN R. W. 1954. - Composition of scientific words. A manual of methods and a lexicon of materials for the practice of logotechnics. Smithsonian Books, Washington, D. C., 882 p. https: // archive. org / details / compositionofsci 00 brow" type="book" year="1954">Brown 1954</bibRefCitation>
;
<bibRefCitation id="EFFD7EF1FFEBFFB4DC78F91B6DA5F974" accessDate="2023-05-23" author="WIKTIONARY" box="[273,454,1761,1785]" pageId="13" pageNumber="34" publicationUrl="https://en.wiktionary.org/wiki/punctatus" refId="ref38003" refString="WIKTIONARY 2019 a. - punctatus. Available online at: https: // en. wiktionary. org / wiki / punctatus (last accessed on 23 rd May 2023)." title="punctatus" type="url" year="2019">Wiktionary 2019a</bibRefCitation>
). It alludes to the pair of tenuous posterior pronotal fossae that characterized the purported variety of
<taxonomicName id="4C6C7883FFEBFFB4DDEAF8ED6C9DF8A2" ID-CoL="6L4H6" baseAuthorityName="Oken" baseAuthorityYear="1834" box="[131,254,1815,1839]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="lacordairii">
<emphasis id="B918DF12FFEBFFB4DDEAF8ED6C9DF8A2" box="[131,254,1815,1839]" italics="true" pageId="13" pageNumber="34">G. lacordairii</emphasis>
</taxonomicName>
for which d’Olsoufieff (1924) established the name and which he knew exclusively from
<collectingRegion id="49A8CDE2FFEBFFB4DCA8F8C86E22F8C7" box="[449,577,1842,1866]" country="Brazil" name="Mato Grosso" pageId="13" pageNumber="34">Mato Grosso</collectingRegion>
(
<collectingCountry id="F37B4390FFEBFFB4DF27F8CB6EEFF8C4" box="[590,652,1841,1865]" name="Brazil" pageId="13" pageNumber="34">Brazil</collectingCountry>
).
<bibRefCitation id="EFFD7EF1FFEBFFB4DFF5F8CB6D32F8E9" author="Cupello &amp; Vaz-de-Mello" firstAuthor="Cupello" pageId="13" pageNumber="34" pagination="439 - 482" refId="ref30610" refString="CUPELLO M. &amp; VAZ-DE-MELLO F. Z. 2013. - Taxonomic revision of the South American dung beetle genus Gromphas Brulle, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa 3722 (4): 439 - 482. https: // doi. org / 10.11646 / zootaxa. 3722.4.2" type="journal article" year="2013">Cupello &amp; Vaz-de-Mello (2013)</bibRefCitation>
observed that specimens bearing this feature are always small and actually found throughout the range of the species, always in polymorphism with the much more abundant smooth individuals of all sizes. Due to this, and in keeping with the authors’ taxonomic criteria, they were prompted to invalidate d’Olsoufieff’s taxon and name.
</paragraph>
</subSubSection>
<subSubSection id="C376508BFFEBFFB4DE44FF22697DFEB2" pageId="13" pageNumber="34" type="distribution">
<paragraph id="8BD30300FFEBFFB4DE44FF22697DFEB2" blockId="13.[811,1456,215,319]" pageId="13" pageNumber="34">
<smallCapsWord id="8D3595DCFFEBFFB4DE44FF226FDBFF63" baselines="233,234" box="[813,952,216,239]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="13" pageNumber="34">DISTRIBUTION</smallCapsWord>
. — Open environments of middle South America in
<collectingCountry id="F37B4390FFEBFFB4DE42FF086F06FE87" box="[811,869,242,266]" name="Brazil" pageId="13" pageNumber="34">Brazil</collectingCountry>
,
<collectingCountry id="F37B4390FFEBFFB4DE04FF086FD0FE87" box="[877,947,242,266]" name="Bolivia" pageId="13" pageNumber="34">Bolivia</collectingCountry>
,
<collectingCountry id="F37B4390FFEBFFB4DED2FF086871FE87" box="[955,1042,242,266]" name="Paraguay" pageId="13" pageNumber="34">Paraguay</collectingCountry>
,
<collectingCountry id="F37B4390FFEBFFB4D973FF08681FFE87" box="[1050,1148,242,266]" name="Argentina" pageId="13" pageNumber="34">Argentina</collectingCountry>
, and
<collectingCountry id="F37B4390FFEBFFB4D9C5FF086962FE87" box="[1196,1281,242,266]" name="Uruguay" pageId="13" pageNumber="34">Uruguay</collectingCountry>
, inhabiting savannas, grasslands, floodplains, river sandbanks, pastures, and restingas as well as associated forest edges and gallery forests.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink