223 lines
28 KiB
XML
223 lines
28 KiB
XML
<document id="7EBA952F704BB36056425B4A08EBD442" ID-CLB-Dataset="288370" ID-DOI="10.5252/zoosystema2024v46a2" ID-GBIF-Dataset="21b07f67-d540-4d6f-86fd-af0f84498c5f" ID-ISSN="1638-9387" ID-Zenodo-Dep="10667997" ID-ZooBank="4B49C1D9-1196-4942-969F-2E923B1FC12C" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1707561373589" checkinUser="plazi" docAuthor="Cupello, Mario" docDate="2024" docId="03C5B216FFE9FFB6DDEDF9696861FBCF" docLanguage="en" docName="zoosystema.2024.46.2.pdf" docOrigin="Zoosystema 46 (2)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a2.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Gromphas inermis Harold 1869" docType="treatment" docVersion="3" lastPageNumber="36" masterDocId="FFFCCA6EFFE6FFB9DD69FFFA6C63FF8D" masterDocTitle="The genus Gromphas Dejean, 1836 (Coleoptera, Scarabaeinae): nomenclature, distribution, and conservation, including a contribution to the debate on electronic publications in zoology" masterLastPageNumber="59" masterPageNumber="23" pageNumber="36" updateTime="1708028255164" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="59F4516B75963A281549021A1CE6463A">The genus Gromphas Dejean, 1836 (Coleoptera, Scarabaeinae): nomenclature, distribution, and conservation, including a contribution to the debate on electronic publications in zoology</mods:title>
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<mods:namePart id="EB37E998753464DCB7E85C5723662678">Cupello, Mario</mods:namePart>
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<mods:affiliation id="3DA0AD3B00F98B9114B1DEABFF592EBC">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná (Brazil) & Department of Entomology, Texas A & M University, College Station, TX 77843 (United States)</mods:affiliation>
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<mods:nameIdentifier id="A9FEF7593C5F6B85446A69A7FA805EAF" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:title id="7AFD7C66EB6800CCAEEA99D6094942B6">Zoosystema</mods:title>
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<mods:date id="A64DE53959256CD9BAB55C9534273546">2024</mods:date>
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<treatment id="03C5B216FFE9FFB6DDEDF9696861FBCF" ID-DOI="http://doi.org/10.5281/zenodo.10668037" ID-Zenodo-Dep="10668037" LSID="urn:lsid:plazi:treatment:03C5B216FFE9FFB6DDEDF9696861FBCF" httpUri="http://treatment.plazi.org/id/03C5B216FFE9FFB6DDEDF9696861FBCF" lastPageNumber="36" pageId="15" pageNumber="36">
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<subSubSection id="C376508BFFE9FFB6DDEDF9696D22F941" pageId="15" pageNumber="36" type="nomenclature">
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<paragraph id="8BD30300FFE9FFB6DDEDF9696D22F941" blockId="15.[132,683,1682,1740]" pageId="15" pageNumber="36">
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<collectionCode id="ED7D9BC5FFE9FFB6DDEDF9696DF4F921" box="[132,407,1682,1709]" pageId="15" pageNumber="36">THE TYPE SERIES OF</collectionCode>
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<taxonomicName id="4C6C7883FFE9FFB6DCF7F9686D22F941" ID-CoL="6L4GZ" authority="HAROLD, 1869" authorityName="Harold" authorityYear="1869" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="inermis">
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<emphasis id="B918DF12FFE9FFB6DCF7F9686EC8F921" box="[414,683,1682,1708]" italics="true" pageId="15" pageNumber="36">GROMPHAS INERMIS</emphasis>
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<bibRefCitation id="EFFD7EF1FFE9FFB6DDEDF9486D22F941" author="HAROLD" box="[132,321,1714,1740]" firstAuthor="HAROLD" pageId="15" pageNumber="36" refId="ref32645" refString="HAROLD E. 1869 a. - Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich: 979 - 1346 + [8]. [publication date for nomenclatural purposes: 27 th May 1969 (based on Bousquet 2016)] https: // www. biodiversitylibrary. org / page / 9641549" type="journal volume" year="1869">HAROLD, 1869</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C376508BFFE9FFB6DDEDF9086802FD5D" pageId="15" pageNumber="36" type="discussion">
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<paragraph id="8BD30300FFE9FFB6DDEDF9086802FD5D" blockId="15.[130,777,1778,2028]" lastBlockId="15.[811,1457,215,720]" pageId="15" pageNumber="36">
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<bibRefCitation id="EFFD7EF1FFE9FFB6DDEDF9086D8CF881" author="Cupello & Vaz-de-Mello" box="[132,495,1778,1804]" firstAuthor="Cupello" pageId="15" pageNumber="36" pagination="439 - 482" refId="ref30610" refString="CUPELLO M. & VAZ-DE-MELLO F. Z. 2013. - Taxonomic revision of the South American dung beetle genus Gromphas Brulle, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa 3722 (4): 439 - 482. https: // doi. org / 10.11646 / zootaxa. 3722.4.2" type="journal article" year="2013">Cupello & Vaz-de-Mello (2013)</bibRefCitation>
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designated the
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<typeStatus id="54D7BDA2FFE9FFB6DFCAF9086F64F881" box="[675,775,1778,1804]" pageId="15" pageNumber="36" type="lectotype">lectotype</typeStatus>
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of
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<taxonomicName id="4C6C7883FFE9FFB6DDCAF8E86D70F8A6" authorityName="Harold" authorityYear="1869" box="[163,275,1810,1836]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="inermis">
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<emphasis id="B918DF12FFE9FFB6DDCAF8E86D70F8A6" box="[163,275,1810,1836]" italics="true" pageId="15" pageNumber="36">G. inermis</emphasis>
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</taxonomicName>
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from the only
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<typeStatus id="54D7BDA2FFE9FFB6DCD4F8E86E6CF8A1" box="[445,527,1810,1836]" pageId="15" pageNumber="36" type="syntype">syntype</typeStatus>
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found by them in the
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<collectionCode id="ED7D9BC5FFE9FFB6DDEFF8C86C80F8C1" box="[134,227,1842,1868]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="36">MNHN</collectionCode>
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, but failed to illustrate it. I take the opportunity to do this here in
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<figureCitation id="13571F85FFE9FFB6DC4AF8AB6D1FF8E6" box="[291,380,1873,1899]" captionStart="FIG" captionStartId="16.[133,144,1962,1980]" captionTargetBox="[188,1394,219,1914]" captionTargetId="figure-15@16.[177,1413,196,1944]" captionTargetPageId="16" captionText="FIG. 6. — The lectotype of Gromphas inermis Harold, 1869, a junior subjective synonym of G. lacordairii (Oken, 1834): A, dorsal view; B, ventral view; C, lateral view; D, labels, note that the ‘Buen. Aires’ and ‘inermis / Harold’ labels are in Harold’s handwriting; E, posterior view; F, dorsal view of the head and protibiae. Photographs courtesy of Christophe Rivier (MNHN). Scale bars: A-C: 5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10668015" httpUri="https://zenodo.org/record/10668015/files/figure.png" pageId="15" pageNumber="36">Figure 6</figureCitation>
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. Since then, I have located one of the missing
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<typeStatus id="54D7BDA2FFE9FFB6DDB1F88B6D0CF806" box="[216,367,1905,1931]" pageId="15" pageNumber="36" type="paralectotype">paralectotypes</typeStatus>
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. It belongs to the
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<collectionCode id="ED7D9BC5FFE9FFB6DF48F8886E1EF801" box="[545,637,1906,1932]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="36">MNHN</collectionCode>
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, but has been on loan to my colleague Federico Ocampo, in
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<collectingCountry id="F37B4390FFE9FFB6DF34F86B6EABF826" box="[605,712,1937,1963]" name="Argentina" pageId="15" pageNumber="36">Argentina</collectingCountry>
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, since the 2000s. Many are the aspects that confirm that this specimen is indeed part of Harold’s type series.
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<bibRefCitation id="EFFD7EF1FFE9FFB6DF33F82B6E92F866" author="Harold" box="[602,753,2001,2027]" firstAuthor="Harold" pageId="15" pageNumber="36" refId="ref32645" refString="HAROLD E. 1869 a. - Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich: 979 - 1346 + [8]. [publication date for nomenclatural purposes: 27 th May 1969 (based on Bousquet 2016)] https: // www. biodiversitylibrary. org / page / 9641549" type="journal volume" year="1869">Harold (1869</bibRefCitation>
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b) described
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<taxonomicName id="4C6C7883FFE9FFB6DEF3FF22686AFF7C" authorityName="Harold" authorityYear="1869" box="[922,1033,216,242]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="inermis">
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<emphasis id="B918DF12FFE9FFB6DEF3FF22686AFF7C" box="[922,1033,216,242]" italics="true" pageId="15" pageNumber="36">G. inermis</emphasis>
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</taxonomicName>
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in a work dealing specifically with the material that he had seen during a visit to the
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<collectionCode id="ED7D9BC5FFE9FFB6D84EFF0269E2FE9F" box="[1319,1409,248,274]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="36">MNHN</collectionCode>
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(see
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<bibRefCitation id="EFFD7EF1FFE9FFB6DE47FEED6FA0FEBC" author="Cupello" box="[814,963,279,305]" firstAuthor="Cupello" pageId="15" pageNumber="36" pagination="15 - 24" refId="ref30581" refString="CUPELLO M. 2020. - The discovery of Edgar von Harold type material in the Museum of Zoology, Dresden. Scarabaeus 1: 15 - 24." type="journal article" year="2020">Cupello 2020</bibRefCitation>
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), and the specimen bears a label identifying it in Harold’s handwriting as
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<taxonomicName id="4C6C7883FFE9FFB6D927FECD68D8FEDC" authorityName="Harold" authorityYear="1869" box="[1102,1211,311,337]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="inermis">
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<emphasis id="B918DF12FFE9FFB6D927FECD68D8FEDC" box="[1102,1211,311,337]" italics="true" pageId="15" pageNumber="36">G. inermis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="13571F85FFE9FFB6D9A3FECD6969FEDC" box="[1226,1290,311,337]" captionStart="FIG" captionStartId="17.[134,143,899,917]" captionTargetBox="[133,777,216,856]" captionTargetId="figure-851@17.[104,805,184,889]" captionTargetPageId="17" captionText="FIG. 7. — The labels of the only known (and newly found) paralectotype of Gromphas inermis Harold, 1869. Notice that it is a d’Orbigny specimen from Corrientes (Argentina) and bears Blanchard’s nomen in litteris ‘G.aeneus’, which was only mentioned in the literature by Harold (1869b). The label at the top bearing the identification ‘Gromphas / inermis / Harold’ is in Harold’s handwriting. Courtesy of Federico Ocampo." figureDoi="http://doi.org/10.5281/zenodo.10668017" httpUri="https://zenodo.org/record/10668017/files/figure.png" pageId="15" pageNumber="36">Fig. 7</figureCitation>
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). Another label borne by the specimen, a rectangular, green one typical of South American specimens collected by Alcide d’Orbigny housed in the
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<collectionCode id="ED7D9BC5FFE9FFB6DEABFE6D687CFE3C" box="[962,1055,407,433]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="36">MNHN</collectionCode>
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, reads ‘
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<emphasis id="B918DF12FFE9FFB6D902FE6D68B7FE3D" box="[1131,1236,407,433]" italics="true" pageId="15" pageNumber="36">
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<taxonomicName id="4C6C7883FFE9FFB6D902FE6D68B3FE3D" box="[1131,1232,407,433]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="aeneus">G. aeneus</taxonomicName>
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,
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</emphasis>
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/
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<emphasis id="B918DF12FFE9FFB6D98FFE6C6956FE3D" box="[1254,1333,406,432]" italics="true" pageId="15" pageNumber="36">Blanch.</emphasis>
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/
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<emphasis id="B918DF12FFE9FFB6D821FE6D69CCFE3C" box="[1352,1455,407,433]" italics="true" pageId="15" pageNumber="36">
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<collectingRegion id="49A8CDE2FFE9FFB6D821FE6D69CCFE3C" box="[1352,1455,407,433]" country="Argentina" name="Corrientes" pageId="15" pageNumber="36">Corrientes</collectingRegion>
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</emphasis>
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/
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<emphasis id="B918DF12FFE9FFB6DE57FE4D6FB0FE5D" box="[830,979,438,464]" italics="true" pageId="15" pageNumber="36">M. D’Orbigny</emphasis>
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.’. This
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<emphasis id="B918DF12FFE9FFB6D94EFE4D68B7FE5D" box="[1063,1236,438,464]" italics="true" pageId="15" pageNumber="36">nomen in litteris</emphasis>
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was curiously never mentioned by
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<bibRefCitation id="EFFD7EF1FFE9FFB6DEAFFE2C681CFE7D" author="Blanchard" box="[966,1151,470,496]" firstAuthor="Blanchard" pageId="15" pageNumber="36" pagination="169 - 184" refId="ref29449" refString="BLANCHARD E. 1846. - [livraison 80: 169 - 184, pl. 31], in BLANCHARD E. & BRULLE G. A. 1835 - 1846. Voyage dans L'Amerique Meridionale (le Bresil, la Republique orientale de L'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833, par Alcide d'Orbigny. Tome Sixieme. 2 e partie: Insectes. P. Bertrand, Paris, and Ve. Lerrault, Strasbourg, 222 p. [publication date for nomenclatural purposes: 18 th December 1846 (Evenhuis, personal communication, 3 rd September 2019)] https: // www. biodiversitylibrary. org / page / 2531073" type="book chapter" year="1846">Blanchard (1846)</bibRefCitation>
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in his study of the d’Orbigny South American material or anyone else in the literature except for
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<bibRefCitation id="EFFD7EF1FFE9FFB6DE38FDEC6F80FDBD" author="Harold" box="[849,995,534,560]" firstAuthor="Harold" pageId="15" pageNumber="36" refId="ref32645" refString="HAROLD E. 1869 a. - Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich: 979 - 1346 + [8]. [publication date for nomenclatural purposes: 27 th May 1969 (based on Bousquet 2016)] https: // www. biodiversitylibrary. org / page / 9641549" type="journal volume" year="1869">Harold (1869</bibRefCitation>
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b), who cited it as one of the (to use modern jargon) rejected unavailable synonyms of
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<taxonomicName id="4C6C7883FFE9FFB6D9BBFDCC6958FDDD" authorityName="Harold" authorityYear="1869" box="[1234,1339,566,592]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="inermis">
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<emphasis id="B918DF12FFE9FFB6D9BBFDCC6958FDDD" box="[1234,1339,566,592]" italics="true" pageId="15" pageNumber="36">G. inermis</emphasis>
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</taxonomicName>
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along with ‘
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<taxonomicName id="4C6C7883FFE9FFB6DE58FDAC6FA6FDFD" box="[817,965,598,624]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="larcordairei">
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<emphasis id="B918DF12FFE9FFB6DE58FDAC6FA6FDFD" box="[817,965,598,624]" italics="true" pageId="15" pageNumber="36">G. larcordairei</emphasis>
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</taxonomicName>
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Dejean’. The only way that Harold could have learnt of the existence of this name was from this label, thus making clear that he did examine this d’Orbigny specimen for the description of
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<taxonomicName id="4C6C7883FFE9FFB6DE87FD4C6839FD42" authorityName="Harold" authorityYear="1869" box="[1006,1114,694,720]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="inermis">
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<emphasis id="B918DF12FFE9FFB6DE87FD4C6839FD42" box="[1006,1114,694,720]" italics="true" pageId="15" pageNumber="36">G. inermis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="C376508BFFE9FFB6DE44FD1D6861FBCF" pageId="15" pageNumber="36" type="materials_examined">
|
||
<paragraph id="8BD30300FFE9FFB6DE44FD1D6901FC13" blockId="15.[811,1456,743,1090]" pageId="15" pageNumber="36">
|
||
<materialsCitation id="3B04095DFFE9FFB6DE44FD1D6901FC13" collectionCode="MNHN" collectorName="F. Z. Vaz-de-Mello" country="Argentina" determinerName="Cupello & Vaz-de-Mello" municipality="Buenos Aires" pageId="15" pageNumber="36" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
|
||
<smallCapsWord id="8D3595DCFFE9FFB6DE44FD1D6F3EFD70" baselines="760,761" box="[813,861,743,766]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="15" pageNumber="36">TYPE</smallCapsWord>
|
||
<smallCapsWord id="8D3595DCFFE9FFB6DE0CFD176FA0FD73" baselines="762" box="[869,963,749,766]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="15" pageNumber="36">MATERIAL</smallCapsWord>
|
||
<smallCapsWord id="8D3595DCFFE9FFB6DEA3FD17684DFD73" baselines="762" box="[970,1070,749,766]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="examined" pageId="15" pageNumber="36">EXAMINED</smallCapsWord>
|
||
. —
|
||
<emphasis id="B918DF12FFE9FFB6D932FD1D68DEFD72" bold="true" box="[1115,1213,743,767]" pageId="15" pageNumber="36">
|
||
<typeStatus id="54D7BDA2FFE9FFB6D932FD1D68DEFD72" box="[1115,1213,743,767]" pageId="15" pageNumber="36" type="lectotype">Lectotype</typeStatus>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D6AC889FFE9FFB6D9BFFD116889FC8F" box="[1238,1258,747,770]" count="1" pageId="15" pageNumber="36" type="male">♂</specimenCount>
|
||
;
|
||
<collectionCode id="ED7D9BC5FFE9FFB6D99EFD1D6929FD73" box="[1271,1354,743,766]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="36">MNHN</collectionCode>
|
||
(ex Edgar von Harold and René Oberthür collections); designated by
|
||
<bibRefCitation id="EFFD7EF1FFE9FFB6D80BFCFB6821FCB9" author="Cupello & Vaz-de-Mello" firstAuthor="Cupello" page="464" pageId="15" pageNumber="36" pagination="439 - 482" refId="ref30610" refString="CUPELLO M. & VAZ-DE-MELLO F. Z. 2013. - Taxonomic revision of the South American dung beetle genus Gromphas Brulle, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa 3722 (4): 439 - 482. https: // doi. org / 10.11646 / zootaxa. 3722.4.2" type="journal article" year="2013">
|
||
<determinerName id="7E0FEFC4FFE9FFB6D80BFCFB69CCFC94" box="[1378,1455,769,793]" pageId="15" pageNumber="36">Cupello</determinerName>
|
||
&
|
||
<determinerName id="7E0FEFC4FFE9FFB6DE2FFCE66FABFCB9" box="[838,968,796,820]" pageId="15" pageNumber="36">Vaz-de-Mello</determinerName>
|
||
(2013: 464)
|
||
</bibRefCitation>
|
||
: “
|
||
<emphasis id="B918DF12FFE9FFB6D93CFCE768DEFCBE" box="[1109,1213,796,820]" italics="true" pageId="15" pageNumber="36">Buen. Aires</emphasis>
|
||
” [Harold’s handwriting],
|
||
<taxonomicName id="4C6C7883FFE9FFB6DE42FCCC6F18FCC3" authorityName="Harold" authorityYear="1869" box="[811,891,822,846]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="inermis">
|
||
“
|
||
<emphasis id="B918DF12FFE9FFB6DE5EFCCD6F18FCC3" box="[823,891,823,846]" italics="true" pageId="15" pageNumber="36">inermis</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<emphasis id="B918DF12FFE9FFB6DEE2FCCC6FB7FCC3" box="[907,980,822,846]" italics="true" pageId="15" pageNumber="36">Harold.</emphasis>
|
||
” [Harold’s handwriting], “Ex-Musaeo / E.Harold”, “LECTOTYPE”, “MNHN, Paris / EC12059 / [QR code]”, “HOLO
|
||
<emphasis id="B918DF12FFE9FFB6DE27FC966FFFFC0E" box="[846,924,876,899]" italics="true" pageId="15" pageNumber="36">LECTO</emphasis>
|
||
TYPE /
|
||
<taxonomicName id="4C6C7883FFE9FFB6DE87FC91682DFC0E" authorityName=", Dejean" authorityYear="1836" box="[1006,1102,875,899]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918DF12FFE9FFB6DE87FC91682DFC0E" box="[1006,1102,875,899]" italics="true" pageId="15" pageNumber="36">Gromphas</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C6C7883FFE9FFB6D90AFC9668CBFC0E" authorityName="Harold" authorityYear="1869" box="[1123,1192,876,899]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="inermis">
|
||
<emphasis id="B918DF12FFE9FFB6D90AFC9668CBFC0E" box="[1123,1192,876,899]" italics="true" pageId="15" pageNumber="36">inermis</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<emphasis id="B918DF12FFE9FFB6D9D7FC966888FC0E" box="[1214,1259,876,899]" italics="true" pageId="15" pageNumber="36">Har.</emphasis>
|
||
/
|
||
<collectorName id="269966D6FFE9FFB6D996FC9669CCFC0E" box="[1279,1455,875,899]" pageId="15" pageNumber="36">F.Z. Vaz-de-Mello</collectorName>
|
||
<emphasis id="B918DF12FFE9FFB6DE44FC7C6F03FC13" box="[813,864,902,926]" italics="true" pageId="15" pageNumber="36">2013</emphasis>
|
||
” [Fernando Z. Vaz-de-Mello’s handwriting] (
|
||
<figureCitation id="13571F85FFE9FFB6D873FC7C6936FC13" box="[1306,1365,902,926]" captionStart="FIG" captionStartId="16.[133,144,1962,1980]" captionTargetBox="[188,1394,219,1914]" captionTargetId="figure-15@16.[177,1413,196,1944]" captionTargetPageId="16" captionText="FIG. 6. — The lectotype of Gromphas inermis Harold, 1869, a junior subjective synonym of G. lacordairii (Oken, 1834): A, dorsal view; B, ventral view; C, lateral view; D, labels, note that the ‘Buen. Aires’ and ‘inermis / Harold’ labels are in Harold’s handwriting; E, posterior view; F, dorsal view of the head and protibiae. Photographs courtesy of Christophe Rivier (MNHN). Scale bars: A-C: 5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10668015" httpUri="https://zenodo.org/record/10668015/files/figure.png" pageId="15" pageNumber="36">Fig. 6</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD30300FFE9FFB6DE44FC5A6861FBCF" blockId="15.[811,1456,743,1090]" pageId="15" pageNumber="36">
|
||
<materialsCitation id="3B04095DFFE9FFB6DE44FC5A6F9DFBCF" collectionCode="MNHN" collectorName="M. D'Orbigny" country="Argentina" location="Corrientes" pageId="15" pageNumber="36" specimenCount="1" stateProvince="Corrientes" typeStatus="paralectotype">
|
||
<emphasis id="B918DF12FFE9FFB6DE44FC5A6FD0FC35" bold="true" box="[813,947,928,952]" pageId="15" pageNumber="36">
|
||
<typeStatus id="54D7BDA2FFE9FFB6DE44FC5A6FD0FC35" box="[813,947,928,952]" pageId="15" pageNumber="36" type="paralectotype">Paralectotype</typeStatus>
|
||
</emphasis>
|
||
• unsexed;
|
||
<collectionCode id="ED7D9BC5FFE9FFB6D94AFC5B681AFC35" box="[1059,1145,929,952]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="36">MNHN</collectionCode>
|
||
: green disk [obverse] / not seen, but probably aged white, stating an accession number, likely from 1834 given that it is a d’Orbigny specimen [reverse], “
|
||
<emphasis id="B918DF12FFE9FFB6D82AFC2F693FFC61" box="[1347,1372,981,1004]" italics="true" pageId="15" pageNumber="36">G.</emphasis>
|
||
ae
|
||
<emphasis id="B918DF12FFE9FFB6D818FC2C69C3FC60" box="[1393,1440,982,1005]" italics="true" pageId="15" pageNumber="36">neus,</emphasis>
|
||
/
|
||
<emphasis id="B918DF12FFE9FFB6DE44FC156F0EFB8A" box="[813,877,1007,1031]" italics="true" pageId="15" pageNumber="36">Blanch</emphasis>
|
||
/
|
||
<emphasis id="B918DF12FFE9FFB6DE17FC0A6F81FB8A" box="[894,994,1008,1031]" italics="true" pageId="15" pageNumber="36">
|
||
<collectingRegion id="49A8CDE2FFE9FFB6DE17FC0A6FBDFB8A" box="[894,990,1008,1031]" country="Argentina" name="Corrientes" pageId="15" pageNumber="36">Corrientes</collectingRegion>
|
||
.
|
||
</emphasis>
|
||
/
|
||
<emphasis id="B918DF12FFE9FFB6DE9BFC0A6819FB8A" box="[1010,1146,1007,1031]" italics="true" pageId="15" pageNumber="36">
|
||
<collectorName id="269966D6FFE9FFB6DE9BFC0A6816FB8A" box="[1010,1141,1007,1031]" pageId="15" pageNumber="36">M. D’Orbigny</collectorName>
|
||
.
|
||
</emphasis>
|
||
” [unknown handwriting], “
|
||
<emphasis id="B918DF12FFE9FFB6D8E8FC0A69C5FB85" box="[1409,1446,1008,1032]" italics="true" pageId="15" pageNumber="36">558</emphasis>
|
||
” [unknown handwriting], “
|
||
<taxonomicName id="4C6C7883FFE9FFB6D958FBF068F2FBAF" authorityName=", Dejean" authorityYear="1836" box="[1073,1169,1034,1058]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918DF12FFE9FFB6D958FBF068F2FBAF" box="[1073,1169,1034,1058]" italics="true" pageId="15" pageNumber="36">Gromphas</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C6C7883FFE9FFB6D9CFFBF16888FBAF" authorityName="Harold" authorityYear="1869" box="[1190,1259,1035,1058]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="inermis">
|
||
<emphasis id="B918DF12FFE9FFB6D9CFFBF16888FBAF" box="[1190,1259,1035,1058]" italics="true" pageId="15" pageNumber="36">inermis</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<emphasis id="B918DF12FFE9FFB6D868FBF06927FBAF" box="[1281,1348,1034,1058]" italics="true" pageId="15" pageNumber="36">Harold</emphasis>
|
||
” [Harold’s handwriting] (
|
||
<figureCitation id="13571F85FFE9FFB6DED3FBD06F96FBCF" box="[954,1013,1066,1090]" captionStart="FIG" captionStartId="17.[134,143,899,917]" captionTargetBox="[133,777,216,856]" captionTargetId="figure-851@17.[104,805,184,889]" captionTargetPageId="17" captionText="FIG. 7. — The labels of the only known (and newly found) paralectotype of Gromphas inermis Harold, 1869. Notice that it is a d’Orbigny specimen from Corrientes (Argentina) and bears Blanchard’s nomen in litteris ‘G.aeneus’, which was only mentioned in the literature by Harold (1869b). The label at the top bearing the identification ‘Gromphas / inermis / Harold’ is in Harold’s handwriting. Courtesy of Federico Ocampo." figureDoi="http://doi.org/10.5281/zenodo.10668017" httpUri="https://zenodo.org/record/10668017/files/figure.png" pageId="15" pageNumber="36">Fig. 7</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |