treatments-xml/data/03/C5/87/03C587F34921AE73FF7321DFFB751031.xml

<document id="FB86112F54AE802C39EB55D953831D46" ID-CLB-Dataset="31843" ID-DOI="10.11646/zootaxa.4282.2.6" ID-GBIF-Dataset="7a8f700f-f5a5-4055-8fd6-ed95699105ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010378" ID-ZooBank="0F3143CD-9D1A-43EB-8E01-0B08FEB46BAE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1498480921144" checkinUser="plazi" docAuthor="Shimada, Daisuke" docDate="2017" docId="03C587F34921AE73FF7321DFFB751031" docLanguage="en" docName="zootaxa.4282.2.6.pdf.imf" docOrigin="Zootaxa 4282 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tanaopsis japonica Shimada 2017, sp. nov." docType="treatment" docVersion="8" masterDocId="FFFCFF8B4920AE79FFE4271EFFD51325" masterDocTitle="A new species of Tanaopsis (Crustacea: Tanaidacea) from Japan, with remarks on the functions of serial ridges and grooves on the appendages" updateTime="1698457321784" updateUser="ExternalLinkService">
<mods:mods id="1F3FE32EEFC3400016E9257B280D3FDA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3D3268A9551EDB783D07D0800856E28B">A new species of Tanaopsis (Crustacea: Tanaidacea) from Japan, with remarks on the functions of serial ridges and grooves on the appendages</mods:title>
</mods:titleInfo>
<mods:name id="0AA9CBEDD344B145C76E390E643A63C0" type="personal">
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<mods:namePart id="91822C538DD70701D16A090E22660937">Shimada, Daisuke</mods:namePart>
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<mods:title id="979689668F0DE61116203180F9444D24">Zootaxa</mods:title>
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<mods:date id="B036F2853FE2C23EC7E9903AA75281A0">2017</mods:date>
<mods:detail id="80D6AA6D8B071020095F7F91B705CF71" type="pubDate">
<mods:number id="6E234DDF57C67E183FD83D52055304B3">2017-06-26</mods:number>
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<mods:number id="FAF84E4A8D325437837A11449F765C0D">4282</mods:number>
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<mods:identifier id="8D0F56E5DE3CB5ECD41A9F3A62E1C61C" type="CLB-Dataset">31843</mods:identifier>
<mods:identifier id="A022F1E9A6877159CBB15980CFA9FDCC" type="DOI">10.11646/zootaxa.4282.2.6</mods:identifier>
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<treatment id="03C587F34921AE73FF7321DFFB751031" ID-DOI="http://doi.org/10.5281/zenodo.5698066" ID-GBIF-Taxon="135435491" ID-Zenodo-Dep="5698066" LSID="urn:lsid:plazi:treatment:03C587F34921AE73FF7321DFFB751031" httpUri="http://treatment.plazi.org/id/03C587F34921AE73FF7321DFFB751031" lastPageId="10" pageId="1">
<subSubSection id="C376656E4921AE78FF7321DFFEC915D9" pageId="1" type="nomenclature">
<paragraph id="8BD336E54921AE78FF7321DFFE3115FE" blockId="1.[151,484,1729,1789]" box="[151,484,1729,1755]" pageId="1">
<heading id="D09B81894921AE78FF7321DFFE3115FE" bold="true" box="[151,484,1729,1755]" fontSize="11" level="1" pageId="1" reason="1">
<emphasis id="B918EAF74921AE78FF7321DFFE3115FE" bold="true" box="[151,484,1729,1755]" pageId="1">
<taxonomicName id="4C6C4D664921AE78FF7321DFFEAA15FE" ID-CoL="54NR3" authority="Shimada, 2017" authorityName="Shimada" authorityYear="2017" box="[151,383,1729,1755]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="1" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="B918EAF74921AE78FF7321DFFEAA15FE" bold="true" box="[151,383,1729,1755]" italics="true" pageId="1">Tanaopsis japonica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B578C4921AE78FE6321DFFE3115FE" box="[391,484,1729,1755]" pageId="1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD336E54921AE78FF7321FAFEC915D9" blockId="1.[151,484,1729,1789]" box="[151,284,1764,1789]" pageId="1">
<figureCitation id="13572A604921AE78FF7321FAFEC915D9" box="[151,284,1764,1789]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="2.[151,250,1853,1875]" captionStartId-1="3.[157,256,1810,1832]" captionStartId-2="4.[151,250,1898,1920]" captionStartId-3="5.[151,250,1827,1849]" captionStartId-4="6.[155,253,1102,1124]" captionStartId-5="7.[151,250,1751,1773]" captionStartId-6="8.[151,250,1548,1570]" captionStartId-7="9.[151,250,1983,2005]" captionTargetBox-0="[206,1380,373,1832]" captionTargetBox-1="[164,1422,625,1786]" captionTargetBox-2="[154,1432,339,1874]" captionTargetBox-3="[158,1428,481,1806]" captionTargetBox-4="[162,1425,193,1081]" captionTargetBox-5="[161,1425,697,1738]" captionTargetBox-6="[151,1436,193,1526]" captionTargetId-0="graphics@2.[259,1381,373,1832]" captionTargetId-1="graphics@3.[164,1422,625,1786]" captionTargetId-2="graphics@4.[154,1432,339,1874]" captionTargetId-3="graphics@5.[158,1428,481,1806]" captionTargetId-4="graphics@6.[162,1425,193,1080]" captionTargetId-5="figure@7.[161,1425,697,1738]" captionTargetId-6="figure@8.[151,1436,193,1526]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionText-0="FIGURE 1. Tanaopsis japonica sp. nov. A, holotype female; B, allotype male: A, B, body, dorsal view; a 1, b 1, body, left view; a 2, b 3, pleonite 5 and pleotelson, dorsal view; b 2, genital cones on pereonite 6, ventral view; a 3, b 4, pereonite 6 and pleon, left view. Scale bars: 0.5 mm." captionText-1="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." captionText-2="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." captionText-3="FIGURE 4. Tanaopsis japonica sp. nov., allotype male: A, right antennule; B, right antenna; C, anterior region of cephalothorax, left view; D, labium, ventral view; E, maxillipeds; F, right cheliped, outer view; f 1, same, distal region of fixed finger, outer view; G, left cheliped, inner view. Scale bars: 0.1 mm for A – D, F, G; 0.05 mm for E, f 1." captionText-4="FIGURE 5. Tanaopsis japonica sp. nov., allotype male: A – F, right pereopods 1 – 6, respectively, outer view; G, right pleopod 1, most setal ornamentation omitted; H, right uropod. Scale bars: 0.1 mm." captionText-5="FIGURE 6. Tanaopsis japonica sp. nov., paratype female, scanning electron microscopic images: A, terminal region of antennule, with double arrowhead indicating vestige of small cap-like article; B, cheliped-sclerite-cephalothorax connection, right view, with asterisk indicating acute dorsal process on pereopod- 1 coxa (other parts were detached); C – E, right chela, anterior (C), outer (D), and inner dorsal (E) views, with each white arrowhead indicating one ridge in outer series of denticulate ridges on dactylus, each black arrowhead indicating one ridge in series of smooth ridges on fixed finger, and each arrow indicating one fringed incision in inner hatching on dactylus; inserts in (D) and (E) are images magnified threefold from the corresponding boxes in the main panels. Scale bars: 0.01 mm for A; 0.05 mm for B – E." captionText-6="FIGURE 7. Tanaopsis japonica sp. nov. A, B, holotype female; C – F, paratype female; scanning electron microscopic images: A, part of right antenna, dorsal view, showing series of denticulate ridges on article 2; B, enlargement of ridges from (A); C, cephalothorax, anterior ventral view (left cheliped and antenna removed); D, E, proximal region of antennule and antenna in right view and right anterior ventral view, respecively; F, enlargement from (E) showing grooves and ridges. Each white arrowhead, dorsal denticulate ridge on antenna article 2; each black arrowhead, ventral groove on antennule article 1. Scale bars: 0.01 mm for A, B, F; 0.1 mm for C, E; 0.05 mm for D." captionText-7="FIGURE 8. Living female Tanaopsis japonica sp. nov. (paratype). Scale bar: 0.5 mm." httpUri-0="https://zenodo.org/record/1010380/files/figure.svg" httpUri-1="https://zenodo.org/record/1010382/files/figure.svg" httpUri-2="https://zenodo.org/record/1010384/files/figure.svg" httpUri-3="https://zenodo.org/record/1010386/files/figure.svg" httpUri-4="https://zenodo.org/record/1010388/files/figure.svg" httpUri-5="https://zenodo.org/record/1010390/files/figure.png" httpUri-6="https://zenodo.org/record/1010392/files/figure.png" pageId="1">Figures 1–8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C376656E4921AE7BFF732032FBD71265" lastPageId="2" pageId="1" type="materials_examined">
<paragraph id="8BD336E54921AE78FF732032FDB91495" blockId="1.[151,1436,1836,2005]" pageId="1">
<materialsCitation id="3B043CB84921AE78FF732032FDB91495" ID-GBIF-Occurrence="1671737252" collectingDate="2013-04-13" collectorName="D. Shimada." county="Material" latitude="42.286217" location="Asari-hama" longLatPrecision="1" longitude="142.46138" municipality="Female" pageId="1" specimenCount="1" stateProvince="Hokkaido" typeStatus="holotype">
<emphasis id="B918EAF74921AE78FF732032FE541460" bold="true" box="[151,385,1836,1861]" pageId="1">
<collectingCounty id="62B24E694921AE78FF732032FF2A1460" box="[151,255,1836,1861]" pageId="1">Material</collectingCounty>
examined.
</emphasis>
<typeStatus id="54D788474921AE78FE6C2032FE211460" box="[392,500,1836,1861]" pageId="1" type="holotype">Holotype</typeStatus>
. 
<collectingMunicipality id="6BB7AC9F4921AE78FE1B2032FD871460" box="[511,594,1836,1861]" pageId="1">Female</collectingMunicipality>
without developed or developing oostegites, ICHUM-5339 (BL 1.43, CW 0.32), 9 slides, 1 vial, and 1 SEM stub; 
<location id="8EB3603E4921AE78FDAA2051FD04144D" LSID="urn:lsid:plazi:treatment:03C587F34921AE73FF7321DFFB751031:8EB3603E4921AE78FDAA2051FD04144D" box="[590,721,1871,1896]" county="Material" latitude="42.286217" longLatPrecision="1" longitude="142.46138" municipality="Female" name="Asari-hama" pageId="1" stateProvince="Hokkaido">Asari-hama</location>
(
<location id="8EB3603E4921AE78FD3A2051FCC9144D" LSID="urn:lsid:plazi:treatment:03C587F34921AE73FF7321DFFB751031:8EB3603E4921AE78FD3A2051FCC9144D" box="[734,796,1871,1896]" county="Material" latitude="42.286217" longLatPrecision="1" longitude="142.46138" municipality="Female" name="Asari" pageId="1" stateProvince="Hokkaido">Asari</location>
beach), 
<location id="8EB3603E4921AE78FC9F2051FBE9144D" LSID="urn:lsid:plazi:treatment:03C587F34921AE73FF7321DFFB751031:8EB3603E4921AE78FC9F2051FBE9144D" box="[891,1084,1871,1896]" county="Material" latitude="42.286217" longLatPrecision="1" longitude="142.46138" municipality="Female" name="Higashi-shizunai" pageId="1" stateProvince="Hokkaido">Higashi-shizunai</location>
, 
<location id="8EB3603E4921AE78FBA32051FB1D144D" LSID="urn:lsid:plazi:treatment:03C587F34921AE73FF7321DFFB751031:8EB3603E4921AE78FBA32051FB1D144D" box="[1095,1224,1871,1896]" county="Material" latitude="42.286217" longLatPrecision="1" longitude="142.46138" municipality="Female" name="Shinhidaka" pageId="1" stateProvince="Hokkaido">Shinhidaka</location>
, 
<collectingRegion id="49A8F8074921AE78FB362051FA96144D" box="[1234,1347,1871,1896]" country="Japan" name="Hokkaido" pageId="1">Hokkaido</collectingRegion>
, 
<location id="8EB3603E4921AE78FAAA2051FF3714A9" LSID="urn:lsid:plazi:treatment:03C587F34921AE73FF7321DFFB751031:8EB3603E4921AE78FAAA2051FF3714A9" county="Material" latitude="42.286217" longLatPrecision="1" longitude="142.46138" municipality="Female" name="Pacific Ocean" pageId="1" stateProvince="Hokkaido">Pacific Ocean</location>
, 
<geoCoordinate id="EE5850224921AE78FF14206AFE5514A9" box="[240,384,1908,1932]" direction="north" orientation="latitude" pageId="1" precision="1" value="42.286217">42°17.173'N</geoCoordinate>
, 
<geoCoordinate id="EE5850224921AE78FE6B206AFDFF14A8" box="[399,554,1908,1933]" direction="east" orientation="longitude" pageId="1" precision="1" value="142.46138">142°27.683'E</geoCoordinate>
, muddy sand around roots of the seagrass 
<taxonomicName id="4C6C4D664921AE78FBC4206AFAF014A9" authorityName="Makino" authorityYear="1931" box="[1056,1317,1908,1933]" class="Liliopsida" family="Zosteraceae" genus="Phyllospadix" kingdom="Plantae" order="Alismatales" pageId="1" phylum="Tracheophyta" rank="species" species="iwatensis">
<emphasis id="B918EAF74921AE78FBC4206AFAF014A9" box="[1056,1317,1908,1933]" italics="true" pageId="1">Phyllospadix iwatensis</emphasis>
</taxonomicName>
, intertidal zone, 
<date id="FFD210254921AE78FF3E2086FEAF1495" box="[218,378,1943,1968]" pageId="1" value="2013-04-13">
<collectingDate id="EF96E9CD4921AE78FF3E2086FEAF1495" box="[218,378,1943,1968]" pageId="1" value="2013-04-13">13 April 2013</collectingDate>
</date>
, coll. by 
<collectorName id="269953334921AE78FE042086FDB91495" box="[480,620,1943,1968]" pageId="1">D. Shimada.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BD336E54921AE7BFF2320A2FCED13F0" blockId="1.[151,1436,1836,2005]" lastBlockId="2.[151,1436,151,320]" lastPageId="2" pageId="1">
<materialsCitation id="3B043CB84921AE7BFF2320A2FCED13F0" ID-GBIF-Occurrence="1671737251" collectingDate="2014-06-29" collectorName="A. Shibata." country="Japan" county="Esashi" lastPageId="2" latitude="41.86997" location="Male" longLatPrecision="1" longitude="140.11597" municipality="Kamome Island" pageId="1" specimenCount="1" stateProvince="Hokkaido" typeStatus="allotype">
<typeStatus id="54D788474921AE78FF2320A2FEF814F0" box="[199,301,1980,2005]" pageId="1" type="allotype">Allotype</typeStatus>
. 
<location id="8EB3603E4921AE78FEDF20A2FEA214F0" LSID="urn:lsid:plazi:treatment:03C587F34921AE73FF7321DFFB751031:8EB3603E4921AE78FEDF20A2FEA214F0" box="[315,375,1980,2005]" country="Japan" county="Esashi" latitude="41.86997" longLatPrecision="1" longitude="140.11597" municipality="Kamome Island" name="Male" pageId="1" stateProvince="Hokkaido">Male</location>
, ICHUM-5340 (BL 1.44, CW 0.32), 6 slides and 1 vial; 
<location id="8EB3603E4921AE78FBFB20A2FB1414F0" LSID="urn:lsid:plazi:treatment:03C587F34921AE73FF7321DFFB751031:8EB3603E4921AE78FBFB20A2FB1414F0" box="[1055,1217,1980,2005]" country="Japan" county="Esashi" latitude="41.86997" longLatPrecision="1" longitude="140.11597" municipality="Kamome Island" name="Kamome-jima" pageId="1" stateProvince="Hokkaido">Kamome-jima</location>
(
<collectingMunicipality id="6BB7AC9F4921AE78FB3020A2FA4514F0" box="[1236,1424,1980,2005]" pageId="1">Kamome Island</collectingMunicipality>
), 
<collectingCounty id="62B24E694922AE7BFF732789FF361395" box="[151,227,151,176]" pageId="2">Esashi</collectingCounty>
, 
<collectingRegion id="49A8F8074922AE7BFF0A2789FE8A1395" box="[238,351,151,176]" country="Japan" name="Hokkaido" pageId="2">Hokkaido</collectingRegion>
, Sea of 
<collectingCountry id="F37B76754922AE7BFE5C2786FE2E138A" box="[440,507,152,175]" name="Japan" pageId="2">Japan</collectingCountry>
, 
<geoCoordinate id="EE5850224922AE7BFDE32789FD421395" box="[519,663,151,176]" direction="north" orientation="latitude" pageId="2" precision="1" value="41.86997">41°52.198'N</geoCoordinate>
, 
<geoCoordinate id="EE5850224922AE7BFD462789FCFA1395" box="[674,815,151,176]" direction="east" orientation="longitude" pageId="2" precision="1" value="140.11597">140°6.958'E</geoCoordinate>
, muddy sand around roots of the seagrass 
<taxonomicName id="4C6C4D664922AE7BFAEE2789FF2A13F1" authorityName="Makino" authorityYear="1931" class="Liliopsida" family="Zosteraceae" genus="Phyllospadix" kingdom="Plantae" order="Alismatales" pageId="2" phylum="Tracheophyta" rank="species" species="iwatensis">
<emphasis id="B918EAF74922AE7BFAEE2789FF2A13F1" italics="true" pageId="2">Phyllospadix iwatensis</emphasis>
</taxonomicName>
, intertidal zone, 
<date id="FFD210254922AE7BFE5827A2FD8113F1" box="[444,596,188,212]" pageId="2" value="2014-06-29">
<collectingDate id="EF96E9CD4922AE7BFE5827A2FD8113F1" box="[444,596,188,212]" pageId="2" value="2014-06-29">29 June 2014</collectingDate>
</date>
, coll. by 
<collectorName id="269953334922AE7BFD5E27A2FCED13F0" box="[698,824,188,213]" pageId="2">A. Shibata.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BD336E54922AE7BFF2327FEFBD71265" blockId="2.[151,1436,151,320]" pageId="2">
<materialsCitation id="3B043CB84922AE7BFF2327FEFCF81238" ID-GBIF-Occurrence="1671737253" collectionCode="INSD" location="Female" pageId="2" specimenCount="1" typeStatus="paratype">
<typeStatus id="54D788474922AE7BFF2327FEFEF813DD" box="[199,301,224,248]" pageId="2" type="paratype">Paratype</typeStatus>
. 
<location id="8EB3603E4922AE7BFEDC27C1FE5F13DD" LSID="urn:lsid:plazi:treatment:03C587F34921AE73FF7321DFFB751031:8EB3603E4922AE7BFEDC27C1FE5F13DD" box="[312,394,223,248]" name="Female" pageId="2">Female</location>
without developed or developing oostegites, ICHUM-5341 (BL 1.75, CW 0.32), 5 slides, 1 vial, and 1 SEM stab; 
<collectionCode id="ED7DAE204922AE7BFE72261AFE021239" box="[406,471,260,284]" pageId="2">INSD</collectionCode>
accession number 
<accessionNumber id="943FAB064922AE7BFD56261AFCF81238" box="[690,813,260,285]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC210598" pageId="2">
LC 
<date id="FFD210254922AE7BFD32261AFCF81238" box="[726,813,260,285]" pageId="2" value="98-05-21">210598</date>
</accessionNumber>
</materialsCitation>
; 
<materialsCitation id="3B043CB84922AE7BFCDD261AFBD71265" ID-GBIF-Occurrence="1671737254" collectingDate="2016-08-27" collectionCode="SCUBA" collectorName="K. Kakui." country="Japan" elevation="3" latitude="43.214767" location="Oshoro" longLatPrecision="1" longitude="140.85507" municipality="Otaru" pageId="2" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<location id="8EB3603E4922AE7BFCDD261AFC5B1238" LSID="urn:lsid:plazi:treatment:03C587F34921AE73FF7321DFFB751031:8EB3603E4922AE7BFCDD261AFC5B1238" box="[825,910,260,285]" country="Japan" latitude="43.214767" longLatPrecision="1" longitude="140.85507" municipality="Otaru" name="Oshoro" pageId="2" stateProvince="Hokkaido">Oshoro</location>
, 
<collectingMunicipality id="6BB7AC9F4922AE7BFC7F261AFC0B1239" box="[923,990,260,284]" pageId="2">Otaru</collectingMunicipality>
, 
<collectingRegion id="49A8F8074922AE7BFC0E261AFB8E1238" box="[1002,1115,260,285]" country="Japan" name="Hokkaido" pageId="2">Hokkaido</collectingRegion>
, Sea of 
<collectingCountry id="F37B76754922AE7BFB5D261BFB291239" box="[1209,1276,261,284]" name="Japan" pageId="2">Japan</collectingCountry>
, 
<geoCoordinate id="EE5850224922AE7BFAEC261AFA4D1238" box="[1288,1432,260,285]" direction="north" orientation="latitude" pageId="2" precision="1" value="43.214767">43°12.886'N</geoCoordinate>
, 
<geoCoordinate id="EE5850224922AE7BFF732639FEE71265" box="[151,306,295,320]" direction="east" orientation="longitude" pageId="2" precision="1" value="140.85507">140°51.304'E</geoCoordinate>
, muddy sand, 
<quantity id="4C949B004922AE7BFE372636FDD5121A" box="[467,512,296,320]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="2" unit="m" value="3.0">
<elevation id="0041D1D64922AE7BFE372636FDD5121A" box="[467,512,296,320]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="2" unit="m" value="3.0">3 m</elevation>
</quantity>
, 
<collectionCode id="ED7DAE204922AE7BFDE82636FDB91265" box="[524,620,296,320]" pageId="2">SCUBA</collectionCode>
, 
<date id="FFD210254922AE7BFD932636FCF81265" box="[631,813,295,320]" pageId="2" value="2016-08-27">
<collectingDate id="EF96E9CD4922AE7BFD932636FCF81265" box="[631,813,295,320]" pageId="2" value="2016-08-27">27 August 2016</collectingDate>
</date>
, coll. by 
<collectorName id="269953334922AE7BFC772636FBD71265" box="[915,1026,295,320]" pageId="2">K. Kakui.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF13666D4922AE7BFF732023FE4914B4" httpUri="https://zenodo.org/record/1010380/files/figure.svg" pageId="2" targetBox="[206,1380,373,1832]" targetPageId="2">
<paragraph id="8BD336E54922AE7BFF732023FE4914B4" blockId="2.[151,1436,1853,1937]" pageId="2">
<emphasis id="B918EAF74922AE7BFF732023FEC71476" bold="true" box="[151,274,1853,1875]" pageId="2">FIGURE 1.</emphasis>
<taxonomicName id="4C6C4D664922AE7BFEFC2021FE091471" authority="Shimada, 2017" authorityName="Shimada" authorityYear="2017" box="[280,476,1855,1876]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="2" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="B918EAF74922AE7BFEFC2021FE091471" box="[280,476,1855,1876]" italics="true" pageId="2">Tanaopsis japonica</emphasis>
</taxonomicName>
<emphasis id="B918EAF74922AE7BFE062020FD9F1476" bold="true" box="[482,586,1854,1875]" pageId="2">
<taxonomicNameLabel id="A22B578C4922AE7BFE062020FDE71476" box="[482,562,1854,1875]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
, holotype female; 
<emphasis id="B918EAF74922AE7BFCE22020FCC21476" bold="true" box="[774,791,1854,1875]" pageId="2">B</emphasis>
, allotype male: 
<emphasis id="B918EAF74922AE7BFC5D2020FC1E1476" bold="true" box="[953,971,1854,1875]" pageId="2">A</emphasis>
, 
<emphasis id="B918EAF74922AE7BFC332020FC3D1476" bold="true" box="[983,1000,1854,1875]" pageId="2">B</emphasis>
, body, dorsal view; 
<emphasis id="B918EAF74922AE7BFB572020FB1E1476" bold="true" box="[1203,1227,1854,1875]" pageId="2">a1</emphasis>
, 
<emphasis id="B918EAF74922AE7BFB3C2020FB261476" bold="true" box="[1240,1267,1854,1875]" pageId="2">b1</emphasis>
, body, left view; 
<emphasis id="B918EAF74922AE7BFF732042FF651454" bold="true" box="[151,176,1884,1905]" pageId="2">a2</emphasis>
, 
<emphasis id="B918EAF74922AE7BFF582042FF021454" bold="true" box="[188,215,1884,1905]" pageId="2">b3</emphasis>
, pleonite 5 and pleotelson, dorsal view; 
<emphasis id="B918EAF74922AE7BFD8B2042FD5F1454" bold="true" box="[623,650,1884,1905]" pageId="2">b2</emphasis>
, genital cones on pereonite 6, ventral view; 
<emphasis id="B918EAF74922AE7BFBA32042FBB51454" bold="true" box="[1095,1120,1884,1905]" pageId="2">a3</emphasis>
, 
<emphasis id="B918EAF74922AE7BFB892042FB5D1454" bold="true" box="[1133,1160,1884,1905]" pageId="2">b4</emphasis>
, pereonite 6 and pleon, left view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C376656E4923AE7AFF232789FCFA13F0" pageId="3" type="diagnosis">
<paragraph id="8BD336E54923AE7AFF232789FCFA13F0" blockId="3.[151,1437,151,573]" pageId="3">
<emphasis id="B918EAF74923AE7AFF232789FE941395" bold="true" box="[199,321,151,176]" pageId="3">Diagnosis.</emphasis>
Pereonites 4 and 5 subequal in length (length ratio of pereonites 4/5 1.0); mandibular molar absent; uropod with biarticulate endopod and uniarticulate exopod.
</paragraph>
</subSubSection>
<subSubSection id="C376656E4923AE7AFF2327C1FBC513DD" box="[199,1040,223,248]" pageId="3" type="etymology">
<paragraph id="8BD336E54923AE7AFF2327C1FBC513DD" blockId="3.[151,1437,151,573]" box="[199,1040,223,248]" pageId="3">
<emphasis id="B918EAF74923AE7AFF2327C1FE9B13DD" bold="true" box="[199,334,223,248]" pageId="3">Etymology.</emphasis>
The specific name is an adjective referring to the 
<typeStatus id="54D788474923AE7AFC9B27FEFC7A13DD" box="[895,943,224,248]" pageId="3">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C376656E4923AE70FF23261AFC2517CD" lastPageId="9" pageId="3" type="description">
<paragraph id="8BD336E54923AE7AFF23261AFD791238" blockId="3.[151,1437,151,573]" box="[199,684,260,285]" pageId="3">
<emphasis id="B918EAF74923AE7AFF23261AFE181238" bold="true" box="[199,461,260,285]" pageId="3">Description of female.</emphasis>
Based on 
<typeStatus id="54D788474923AE7AFDA6261AFD7D1238" box="[578,680,260,285]" pageId="3" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BD336E54923AE7AFF232639FE501118" blockId="3.[151,1437,151,573]" pageId="3">
Body (
<figureCitation id="13572A604923AE7AFEF02639FE671265" box="[276,434,295,320]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1853,1875]" captionStartId-1="3.[157,256,1810,1832]" captionStartId-2="4.[151,250,1898,1920]" captionTargetBox-0="[206,1380,373,1832]" captionTargetBox-1="[164,1422,625,1786]" captionTargetBox-2="[154,1432,339,1874]" captionTargetId-0="graphics@2.[259,1381,373,1832]" captionTargetId-1="graphics@3.[164,1422,625,1786]" captionTargetId-2="graphics@4.[154,1432,339,1874]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Tanaopsis japonica sp. nov. A, holotype female; B, allotype male: A, B, body, dorsal view; a 1, b 1, body, left view; a 2, b 3, pleonite 5 and pleotelson, dorsal view; b 2, genital cones on pereonite 6, ventral view; a 3, b 4, pereonite 6 and pleon, left view. Scale bars: 0.5 mm." captionText-1="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." captionText-2="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri-0="https://zenodo.org/record/1010380/files/figure.svg" httpUri-1="https://zenodo.org/record/1010382/files/figure.svg" httpUri-2="https://zenodo.org/record/1010384/files/figure.svg" pageId="3">Fig. 1A, a1–3</figureCitation>
) dorsoventrally flattened, 4.4 times as long as CW, translucent, slightly yellowish when alive (
<figureCitation id="13572A604923AE7AFF3A2652FEF01241" box="[222,293,332,357]" captionStart="FIGURE 8" captionStartId="9.[151,250,1983,2005]" captionText="FIGURE 8. Living female Tanaopsis japonica sp. nov. (paratype). Scale bar: 0.5 mm." pageId="3">Fig. 8</figureCitation>
). Cephalothorax 0.2 times as long as BL, 0.9 times as long as wide, trapezoidal, with pair of lateral simple setae; eye lobes absent. Pereonites 1–6 with length ratio of 1.0:1.1:1.4:1.9:1.9:1.3; all wider than long, with one or two pairs of simple setae. Pleon 0.3 times as long as BL. Pleonites slightly wider than pereonite 6; all wider than long, similar in shape, with slight lamellar lateral projection (
<figureCitation id="13572A604923AE7AFC9D26A9FC1912F5" box="[889,972,439,464]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1875]" captionTargetBox="[206,1380,373,1832]" captionTargetId="graphics@2.[259,1381,373,1832]" captionTargetPageId="2" captionText="FIGURE 1. Tanaopsis japonica sp. nov. A, holotype female; B, allotype male: A, B, body, dorsal view; a 1, b 1, body, left view; a 2, b 3, pleonite 5 and pleotelson, dorsal view; b 2, genital cones on pereonite 6, ventral view; a 3, b 4, pereonite 6 and pleon, left view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1010380/files/figure.svg" pageId="3">Figs 1a</figureCitation>
<figureCitation id="13572A604923AE7AFC2826A6FC0E12F5" box="[972,987,440,464]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="3">3</figureCitation>
, 
<figureCitation id="13572A604923AE7AFC0326A6FC2312F5" box="[999,1014,440,464]" captionStart="FIGURE 8" captionStartId="9.[151,250,1983,2005]" captionText="FIGURE 8. Living female Tanaopsis japonica sp. nov. (paratype). Scale bar: 0.5 mm." pageId="3">8</figureCitation>
), and ventral keel (
<figureCitation id="13572A604923AE7AFB2A26A9FAC812F5" box="[1230,1309,439,464]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1875]" captionTargetBox="[206,1380,373,1832]" captionTargetId="graphics@2.[259,1381,373,1832]" captionTargetPageId="2" captionText="FIGURE 1. Tanaopsis japonica sp. nov. A, holotype female; B, allotype male: A, B, body, dorsal view; a 1, b 1, body, left view; a 2, b 3, pleonite 5 and pleotelson, dorsal view; b 2, genital cones on pereonite 6, ventral view; a 3, b 4, pereonite 6 and pleon, left view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1010380/files/figure.svg" pageId="3">Fig. 1a</figureCitation>
<figureCitation id="13572A604923AE7AFAF926A6FAF912F5" box="[1309,1324,440,464]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="3">3</figureCitation>
); pleonite 5 with two pairs of simple setae. Pleotelson 0.6 times as long as wide, narrower than pleonites, with pair of simple setae proximal to uropodal insertion, pair of simple setae and pair of PSS in subdistal region, and pair of simple setae in distal region.
</paragraph>
<caption id="DF13666D4923AE7AFF79200CFEB51440" httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="3" targetBox="[164,1422,625,1786]" targetPageId="3">
<paragraph id="8BD336E54923AE7AFF79200CFEB51440" blockId="3.[151,1437,1810,1893]" pageId="3">
<emphasis id="B918EAF74923AE7AFF79200CFECE1402" bold="true" box="[157,283,1810,1832]" pageId="3">FIGURE 2.</emphasis>
<taxonomicName id="4C6C4D664923AE7AFEC1200DFE38140D" authority="Shimada, 2017" authorityName="Shimada" authorityYear="2017" box="[293,493,1811,1832]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="3" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="B918EAF74923AE7AFEC1200DFE38140D" box="[293,493,1811,1832]" italics="true" pageId="3">Tanaopsis japonica</emphasis>
</taxonomicName>
<emphasis id="B918EAF74923AE7AFE12200CFD9C1402" bold="true" box="[502,585,1810,1831]" pageId="3">
<taxonomicNameLabel id="A22B578C4923AE7AFE12200CFD9C1402" box="[502,585,1810,1831]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female: 
<emphasis id="B918EAF74923AE7AFCF5200CFCF61402" bold="true" box="[785,803,1810,1831]" pageId="3">A</emphasis>
, left antennule; 
<emphasis id="B918EAF74923AE7AFC36200CFC361402" bold="true" box="[978,995,1810,1831]" pageId="3">B</emphasis>
, left antenna; 
<emphasis id="B918EAF74923AE7AFB98200CFB5B140D" bold="true" box="[1148,1166,1810,1832]" pageId="3">C</emphasis>
, proximal region of right antenna; 
<emphasis id="B918EAF74923AE7AFF17202FFED01463" bold="true" box="[243,261,1841,1862]" pageId="3">D</emphasis>
, labrum; 
<emphasis id="B918EAF74923AE7AFE8C202FFEAC1463" bold="true" box="[360,377,1841,1862]" pageId="3">E</emphasis>
, 
<emphasis id="B918EAF74923AE7AFE62202FFE461463" bold="true" box="[390,403,1841,1862]" pageId="3">F</emphasis>
, left and right mandibles; 
<emphasis id="B918EAF74923AE7AFD46202EFD661463" bold="true" box="[674,691,1840,1862]" pageId="3">G</emphasis>
, labium; 
<emphasis id="B918EAF74923AE7AFCF0202FFCFD1463" bold="true" box="[788,808,1841,1862]" pageId="3">H</emphasis>
, distal part of left maxillule; 
<emphasis id="B918EAF74923AE7AFBB7202FFB881463" bold="true" box="[1107,1117,1841,1862]" pageId="3">I</emphasis>
, maxillipeds; 
<emphasis id="B918EAF74923AE7AFB0A202FFB2E1463" bold="true" box="[1262,1275,1841,1862]" pageId="3">J</emphasis>
, right epignath. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BD336E54923AE7DFF23208CFE19121A" blockId="3.[151,1436,1938,2035]" lastBlockId="4.[151,1437,151,320]" lastPageId="4" pageId="3">
Antennule (
<figureCitation id="13572A604923AE7AFEB4208CFE78148F" box="[336,429,1938,1963]" captionStart="FIGURE 2" captionStartId="3.[157,256,1810,1832]" captionTargetBox="[164,1422,625,1786]" captionTargetId="graphics@3.[164,1422,625,1786]" captionTargetPageId="3" captionText="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="3">Fig. 2A</figureCitation>
) 0.9 times as long as cephalothorax; articles 1–4 with length ratio of 1.0:0.5:0.3:0.6. Article 1 with two distal simple setae, several PSS, and series of ventroproximal slight grooves (
<figureCitation id="13572A604923AE7AFB2020ABFAE914EB" box="[1220,1340,1973,1998]" captionStart="FIGURE 7" captionStartId="8.[151,250,1548,1570]" captionTargetBox="[151,1436,193,1526]" captionTargetId="figure@8.[151,1436,193,1526]" captionTargetPageId="8" captionText="FIGURE 7. Tanaopsis japonica sp. nov. A, B, holotype female; C – F, paratype female; scanning electron microscopic images: A, part of right antenna, dorsal view, showing series of denticulate ridges on article 2; B, enlargement of ridges from (A); C, cephalothorax, anterior ventral view (left cheliped and antenna removed); D, E, proximal region of antennule and antenna in right view and right anterior ventral view, respecively; F, enlargement from (E) showing grooves and ridges. Each white arrowhead, dorsal denticulate ridge on antenna article 2; each black arrowhead, ventral groove on antennule article 1. Scale bars: 0.01 mm for A, B, F; 0.1 mm for C, E; 0.05 mm for D." httpUri="https://zenodo.org/record/1010392/files/figure.png" pageId="3">Fig. 7D–F</figureCitation>
). Article 2 with two distal simple setae and several distal PSS. Article 3 with two distal simple setae. Article 4 with cap-like vestigial article (
<figureCitation id="13572A604924AE7DFEB52789FE7E1395" box="[337,427,151,176]" captionStart="FIGURE 6" captionStartId="7.[151,250,1751,1773]" captionTargetBox="[161,1425,697,1738]" captionTargetId="figure@7.[161,1425,697,1738]" captionTargetPageId="7" captionText="FIGURE 6. Tanaopsis japonica sp. nov., paratype female, scanning electron microscopic images: A, terminal region of antennule, with double arrowhead indicating vestige of small cap-like article; B, cheliped-sclerite-cephalothorax connection, right view, with asterisk indicating acute dorsal process on pereopod- 1 coxa (other parts were detached); C – E, right chela, anterior (C), outer (D), and inner dorsal (E) views, with each white arrowhead indicating one ridge in outer series of denticulate ridges on dactylus, each black arrowhead indicating one ridge in series of smooth ridges on fixed finger, and each arrow indicating one fringed incision in inner hatching on dactylus; inserts in (D) and (E) are images magnified threefold from the corresponding boxes in the main panels. Scale bars: 0.01 mm for A; 0.05 mm for B – E." httpUri="https://zenodo.org/record/1010390/files/figure.png" pageId="4">Fig. 6A</figureCitation>
), five simple setae and aesthetasc. Antenna (
<figureCitation id="13572A604924AE7DFC782789FBC61395" box="[924,1043,151,176]" captionStart="FIGURE 2" captionStartId="3.[157,256,1810,1832]" captionTargetBox="[164,1422,625,1786]" captionTargetId="graphics@3.[164,1422,625,1786]" captionTargetPageId="3" captionText="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="4">Fig. 2B, C</figureCitation>
) with six articles, 0.9 times as long as antennule; articles 1–6 with length ratio of 0.4:1.0:0.7:2.0:0.8:0.3. Article 1 naked. Article 2 with inner distal simple seta and series of dorsal denticulate ridges (
<figureCitation id="13572A604924AE7DFD2827C1FCF213DD" box="[716,807,223,248]" captionStart="FIGURE 2" captionStartId="3.[157,256,1810,1832]" captionTargetBox="[164,1422,625,1786]" captionTargetId="graphics@3.[164,1422,625,1786]" captionTargetPageId="3" captionText="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="4">Figs 2C</figureCitation>
, 
<figureCitation id="13572A604924AE7DFCD627FEFCA213DD" box="[818,887,224,248]" captionStart="FIGURE 7" captionStartId="8.[151,250,1548,1570]" captionTargetBox="[151,1436,193,1526]" captionTargetId="figure@8.[151,1436,193,1526]" captionTargetPageId="8" captionText="FIGURE 7. Tanaopsis japonica sp. nov. A, B, holotype female; C – F, paratype female; scanning electron microscopic images: A, part of right antenna, dorsal view, showing series of denticulate ridges on article 2; B, enlargement of ridges from (A); C, cephalothorax, anterior ventral view (left cheliped and antenna removed); D, E, proximal region of antennule and antenna in right view and right anterior ventral view, respecively; F, enlargement from (E) showing grooves and ridges. Each white arrowhead, dorsal denticulate ridge on antenna article 2; each black arrowhead, ventral groove on antennule article 1. Scale bars: 0.01 mm for A, B, F; 0.1 mm for C, E; 0.05 mm for D." httpUri="https://zenodo.org/record/1010392/files/figure.png" pageId="4">7A, B</figureCitation>
). Article 3 with inner distal simple seta. Article 4 with three distal simple setae and one mid-inner and several distal PSS. Article 5 with distal simple seta. Article 6 with six distal simple setae.
</paragraph>
<caption id="DF13666D4924AE7DFF732074FAB214F9" httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="4" targetBox="[154,1432,339,1874]" targetPageId="4">
<paragraph id="8BD336E54924AE7DFF732074FAB214F9" blockId="4.[151,1436,1898,2012]" pageId="4">
<emphasis id="B918EAF74924AE7DFF732074FEC6145A" bold="true" box="[151,275,1898,1920]" pageId="4">FIGURE 3.</emphasis>
<taxonomicName id="4C6C4D664924AE7DFEFF2075FE0914A5" authority="Shimada, 2017" authorityName="Shimada" authorityYear="2017" box="[283,476,1899,1920]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="4" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="B918EAF74924AE7DFEFF2075FE0914A5" box="[283,476,1899,1920]" italics="true" pageId="4">Tanaopsis japonica</emphasis>
</taxonomicName>
<emphasis id="B918EAF74924AE7DFE072075FDE614A5" bold="true" box="[483,563,1899,1920]" pageId="4">
<taxonomicNameLabel id="A22B578C4924AE7DFE072075FDE614A5" box="[483,563,1899,1920]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female: 
<emphasis id="B918EAF74924AE7DFD0B2074FCD4145A" bold="true" box="[751,769,1898,1919]" pageId="4">A</emphasis>
, right cheliped, outer view; 
<emphasis id="B918EAF74924AE7DFBF92074FBE3145A" bold="true" box="[1053,1078,1898,1919]" pageId="4">a1</emphasis>
, same, chela, inner view; 
<emphasis id="B918EAF74924AE7DFAD92074FA80145A" bold="true" box="[1341,1365,1898,1919]" pageId="4">a2</emphasis>
, same, distal region of fixed finger, outer view; 
<emphasis id="B918EAF74924AE7DFDCE2097FDEE14BB" bold="true" box="[554,571,1929,1950]" pageId="4">B</emphasis>
, left chelipedal dactylus; 
<emphasis id="B918EAF74924AE7DFCDE2097FC9914BA" bold="true" box="[826,844,1929,1951]" pageId="4">C</emphasis>
, 
<emphasis id="B918EAF74924AE7DFCBC2097FCBF14BB" bold="true" box="[856,874,1929,1950]" pageId="4">D</emphasis>
, right and left pereopods 1, respectively, outer views; 
<emphasis id="B918EAF74924AE7DFA612097FA4314BB" bold="true" box="[1413,1430,1929,1950]" pageId="4">E</emphasis>
, left pereopod 2, outer view; 
<emphasis id="B918EAF74924AE7DFE5620B6FE6A1498" bold="true" box="[434,447,1960,1981]" pageId="4">F</emphasis>
, left pereopod 3, inner view; 
<emphasis id="B918EAF74924AE7DFD0C20B6FCC4149B" bold="true" box="[744,785,1960,1982]" pageId="4">G–I</emphasis>
, right pereopods 4–6, respectively, outer view; 
<emphasis id="B918EAF74924AE7DFB0820B6FAD5149B" bold="true" box="[1260,1280,1960,1982]" pageId="4">i1</emphasis>
, serrate seta; 
<emphasis id="B918EAF74924AE7DFA6D20B6FA401498" bold="true" box="[1417,1429,1960,1981]" pageId="4">J</emphasis>
, right pleopod 1, most setal ornamentation omitted; 
<emphasis id="B918EAF74924AE7DFD7520D9FD7014F9" bold="true" box="[657,677,1991,2012]" pageId="4">K</emphasis>
, right uropod. Scale bars: 0.05 mm for a2, B, i1; 0.1 mm for the others.
</paragraph>
</caption>
<paragraph id="8BD336E54925AE7CFF232789FDBC1288" blockId="5.[151,1437,151,429]" pageId="5">
Labrum (
<figureCitation id="13572A604925AE7CFED42789FE5C1395" box="[304,393,151,176]" captionStart="FIGURE 2" captionStartId="3.[157,256,1810,1832]" captionTargetBox="[164,1422,625,1786]" captionTargetId="graphics@3.[164,1422,625,1786]" captionTargetPageId="3" captionText="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="5">Fig. 2D</figureCitation>
) naked. Mandibles (
<figureCitation id="13572A604925AE7CFD972789FD3D1395" box="[627,744,151,176]" captionStart="FIGURE 2" captionStartId="3.[157,256,1810,1832]" captionTargetBox="[164,1422,625,1786]" captionTargetId="graphics@3.[164,1422,625,1786]" captionTargetPageId="3" captionText="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="5">Fig. 2E, F</figureCitation>
) without molar. Incisor of left mandible (
<figureCitation id="13572A604925AE7CFB262789FACD1395" box="[1218,1304,151,176]" captionStart="FIGURE 2" captionStartId="3.[157,256,1810,1832]" captionTargetBox="[164,1422,625,1786]" captionTargetId="graphics@3.[164,1422,625,1786]" captionTargetPageId="3" captionText="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="5">Fig. 2E</figureCitation>
) with blunt tooth and several small teeth; lacinia mobilis widening distally, with several small teeth. Incisor of right mandible (
<figureCitation id="13572A604925AE7CFF7B27C1FF2613DD" box="[159,243,223,248]" captionStart="FIGURE 2" captionStartId="3.[157,256,1810,1832]" captionTargetBox="[164,1422,625,1786]" captionTargetId="graphics@3.[164,1422,625,1786]" captionTargetPageId="3" captionText="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="5">Fig. 2F</figureCitation>
) with blunt tooth and several smaller teeth. Labium (
<figureCitation id="13572A604925AE7CFCAA27C1FC7313DD" box="[846,934,223,248]" captionStart="FIGURE 2" captionStartId="3.[157,256,1810,1832]" captionTargetBox="[164,1422,625,1786]" captionTargetId="graphics@3.[164,1422,625,1786]" captionTargetPageId="3" captionText="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="5">Fig. 2G</figureCitation>
) naked. Maxillule (
<figureCitation id="13572A604925AE7CFB6127C1FB0B13DD" box="[1157,1246,223,248]" captionStart="FIGURE 2" captionStartId="3.[157,256,1810,1832]" captionTargetBox="[164,1422,625,1786]" captionTargetId="graphics@3.[164,1422,625,1786]" captionTargetPageId="3" captionText="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="5">Fig. 2H</figureCitation>
) endite with one thicker and three thinner spines; palp lost during dissection. Maxilla lost during dissection. Maxillipeds (
<figureCitation id="13572A604925AE7CFAA1261AFA411239" box="[1349,1428,260,285]" captionStart="FIGURE 2" captionStartId="3.[157,256,1810,1832]" captionTargetBox="[164,1422,625,1786]" captionTargetId="graphics@3.[164,1422,625,1786]" captionTargetPageId="3" captionText="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="5">
Fig. 
<date id="FFD210254925AE7CFA9F261AFA411239" box="[1403,1428,260,284]" pageId="5">2I</date>
</figureCitation>
) with bases, each bearing simple seta at insertion of palp; distal half of bases separated. Endites flared, separated, reaching beyond distal margin of palp article 1, each with simple seta. Palp article 1 naked; article 2 with two inner and one outer simple setae; article 3 with two inner simple setae; article 4 with three simple and three pinnate setae. Epignath (
<figureCitation id="13572A604925AE7CFEEF268AFE8F1289" box="[267,346,404,429]" captionStart="FIGURE 2" captionStartId="3.[157,256,1810,1832]" captionTargetBox="[164,1422,625,1786]" captionTargetId="graphics@3.[164,1422,625,1786]" captionTargetPageId="3" captionText="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="5">Fig. 2J</figureCitation>
) narrow, curved, naked.
</paragraph>
<caption id="DF13666D4925AE7CFF73203DFB411453" httpUri="https://zenodo.org/record/1010386/files/figure.svg" pageId="5" targetBox="[158,1428,481,1806]" targetPageId="5">
<paragraph id="8BD336E54925AE7CFF73203DFB411453" blockId="5.[151,1436,1827,1910]" pageId="5">
<emphasis id="B918EAF74925AE7CFF73203DFECF141D" bold="true" box="[151,282,1827,1849]" pageId="5">FIGURE 4.</emphasis>
<taxonomicName id="4C6C4D664925AE7CFECC203AFE21141C" authority="Shimada, 2017" authorityName="Shimada" authorityYear="2017" box="[296,500,1828,1849]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="5" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="B918EAF74925AE7CFECC203AFE21141C" box="[296,500,1828,1849]" italics="true" pageId="5">Tanaopsis japonica</emphasis>
</taxonomicName>
<emphasis id="B918EAF74925AE7CFDE6203AFD8C141C" bold="true" box="[514,601,1828,1849]" pageId="5">
<taxonomicNameLabel id="A22B578C4925AE7CFDE6203AFD8C141C" box="[514,601,1828,1849]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, allotype male: 
<emphasis id="B918EAF74925AE7CFCF0203DFCF3141D" bold="true" box="[788,806,1827,1848]" pageId="5">A</emphasis>
, right antennule; 
<emphasis id="B918EAF74925AE7CFC0B203AFBD5141C" bold="true" box="[1007,1024,1828,1849]" pageId="5">B</emphasis>
, right antenna; 
<emphasis id="B918EAF74925AE7CFB51203DFB12141C" bold="true" box="[1205,1223,1827,1849]" pageId="5">C</emphasis>
, anterior region of cephalothorax, left view; 
<emphasis id="B918EAF74925AE7CFE7E205CFE791472" bold="true" box="[410,428,1858,1879]" pageId="5">D</emphasis>
, labium, ventral view; 
<emphasis id="B918EAF74925AE7CFD73205CFD7D1472" bold="true" box="[663,680,1858,1879]" pageId="5">E</emphasis>
, maxillipeds; 
<emphasis id="B918EAF74925AE7CFCDD205CFC931472" bold="true" box="[825,838,1858,1879]" pageId="5">F</emphasis>
, right cheliped, outer view; 
<emphasis id="B918EAF74925AE7CFB81205CFBAC147D" bold="true" box="[1125,1145,1858,1880]" pageId="5">f1</emphasis>
, same, distal region of fixed finger, outer view; 
<emphasis id="B918EAF74925AE7CFEBC207EFEBC1453" bold="true" box="[344,361,1888,1910]" pageId="5">G</emphasis>
, left cheliped, inner view. Scale bars: 0.1 mm for A–D, F, G; 0.05 mm for E, f1.
</paragraph>
</caption>
<caption id="DF13666D4926AE7FFF7F2350FC5117A6" httpUri="https://zenodo.org/record/1010388/files/figure.svg" pageId="6" targetBox="[162,1425,193,1081]" targetPageId="6">
<paragraph id="8BD336E54926AE7FFF7F2350FC5117A6" blockId="6.[151,1436,1102,1155]" pageId="6">
<emphasis id="B918EAF74926AE7FFF7F2350FEC21741" bold="true" box="[155,279,1102,1124]" pageId="6">FIGURE 5.</emphasis>
<taxonomicName id="4C6C4D664926AE7FFEFA234EFE311740" authority="Shimada, 2017" authorityName="Shimada" authorityYear="2017" box="[286,484,1104,1125]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="6" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="B918EAF74926AE7FFEFA234EFE311740" box="[286,484,1104,1125]" italics="true" pageId="6">Tanaopsis japonica</emphasis>
</taxonomicName>
<emphasis id="B918EAF74926AE7FFE0F2351FDE91741" bold="true" box="[491,572,1103,1124]" pageId="6">
<taxonomicNameLabel id="A22B578C4926AE7FFE0F2351FDE91741" box="[491,572,1103,1124]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, allotype male: 
<emphasis id="B918EAF74926AE7FFD062350FCDB1746" bold="true" box="[738,782,1102,1123]" pageId="6">A–F</emphasis>
, right pereopods 1–6, respectively, outer view; 
<emphasis id="B918EAF74926AE7FFB112350FAD31741" bold="true" box="[1269,1286,1102,1124]" pageId="6">G</emphasis>
, right pleopod 1, most setal ornamentation omitted; 
<emphasis id="B918EAF74926AE7FFDF42370FDF117A6" bold="true" box="[528,548,1134,1155]" pageId="6">H</emphasis>
, right uropod. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BD336E54926AE7FFF2323B1FD1B16CD" blockId="6.[151,1437,1199,2017]" pageId="6">
Cheliped (
<figureCitation id="13572A604926AE7FFEDF23B1FE6017ED" box="[315,437,1199,1224]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="6">Figs 3A, a</figureCitation>
<figureCitation id="13572A604926AE7FFE5123AEFE1517ED" box="[437,448,1200,1224]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1875]" captionTargetBox="[206,1380,373,1832]" captionTargetId="graphics@2.[259,1381,373,1832]" captionTargetPageId="2" captionText="FIGURE 1. Tanaopsis japonica sp. nov. A, holotype female; B, allotype male: A, B, body, dorsal view; a 1, b 1, body, left view; a 2, b 3, pleonite 5 and pleotelson, dorsal view; b 2, genital cones on pereonite 6, ventral view; a 3, b 4, pereonite 6 and pleon, left view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1010380/files/figure.svg" pageId="6">1</figureCitation>
, a 
<figureCitation id="13572A604926AE7FFE3F23AEFE3F17ED" box="[475,490,1200,1224]" captionStart="FIGURE 2" captionStartId="3.[157,256,1810,1832]" captionTargetBox="[164,1422,625,1786]" captionTargetId="graphics@3.[164,1422,625,1786]" captionTargetPageId="3" captionText="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="6">2</figureCitation>
, B, 
<figureCitation id="13572A604926AE7FFDF323B1FD8D17ED" box="[535,600,1199,1224]" captionStart="FIGURE 6" captionStartId="7.[151,250,1751,1773]" captionTargetBox="[161,1425,697,1738]" captionTargetId="figure@7.[161,1425,697,1738]" captionTargetPageId="7" captionText="FIGURE 6. Tanaopsis japonica sp. nov., paratype female, scanning electron microscopic images: A, terminal region of antennule, with double arrowhead indicating vestige of small cap-like article; B, cheliped-sclerite-cephalothorax connection, right view, with asterisk indicating acute dorsal process on pereopod- 1 coxa (other parts were detached); C – E, right chela, anterior (C), outer (D), and inner dorsal (E) views, with each white arrowhead indicating one ridge in outer series of denticulate ridges on dactylus, each black arrowhead indicating one ridge in series of smooth ridges on fixed finger, and each arrow indicating one fringed incision in inner hatching on dactylus; inserts in (D) and (E) are images magnified threefold from the corresponding boxes in the main panels. Scale bars: 0.01 mm for A; 0.05 mm for B – E." httpUri="https://zenodo.org/record/1010390/files/figure.png" pageId="6">6B–E</figureCitation>
) with triangular articulation with cephalothorax via sclerite (
<figureCitation id="13572A604926AE7FFAE723B1FAB317ED" box="[1283,1382,1199,1224]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1875]" captionTargetBox="[206,1380,373,1832]" captionTargetId="graphics@2.[259,1381,373,1832]" captionTargetPageId="2" captionText="FIGURE 1. Tanaopsis japonica sp. nov. A, holotype female; B, allotype male: A, B, body, dorsal view; a 1, b 1, body, left view; a 2, b 3, pleonite 5 and pleotelson, dorsal view; b 2, genital cones on pereonite 6, ventral view; a 3, b 4, pereonite 6 and pleon, left view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1010380/files/figure.svg" pageId="6">Figs 1a1</figureCitation>
, 
<figureCitation id="13572A604926AE7FFA9723AEFA4217ED" box="[1395,1431,1200,1224]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="6">3A</figureCitation>
, 
<figureCitation id="13572A604926AE7FFF7323CAFF6F17C8" box="[151,186,1236,1261]" captionStart="FIGURE 6" captionStartId="7.[151,250,1751,1773]" captionTargetBox="[161,1425,697,1738]" captionTargetId="figure@7.[161,1425,697,1738]" captionTargetPageId="7" captionText="FIGURE 6. Tanaopsis japonica sp. nov., paratype female, scanning electron microscopic images: A, terminal region of antennule, with double arrowhead indicating vestige of small cap-like article; B, cheliped-sclerite-cephalothorax connection, right view, with asterisk indicating acute dorsal process on pereopod- 1 coxa (other parts were detached); C – E, right chela, anterior (C), outer (D), and inner dorsal (E) views, with each white arrowhead indicating one ridge in outer series of denticulate ridges on dactylus, each black arrowhead indicating one ridge in series of smooth ridges on fixed finger, and each arrow indicating one fringed incision in inner hatching on dactylus; inserts in (D) and (E) are images magnified threefold from the corresponding boxes in the main panels. Scale bars: 0.01 mm for A; 0.05 mm for B – E." httpUri="https://zenodo.org/record/1010390/files/figure.png" pageId="6">6B</figureCitation>
). Basis slightly longer than wide, with free posterior portion and dorsal simple seta. Merus with ventral simple seta. Carpus 1.5 times as long as wide, with one dorsomedial, one dorsodistal, and two ventral simple setae. Chela as long as carpus; propodal palm as long as fixed finger, with one outer, one longer inner, and three shorter inner simple setae at insertion of dactylus (
<figureCitation id="13572A604926AE7FFDDF2221FD5A167D" box="[571,655,1343,1368]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="6">Fig. 3a</figureCitation>
<figureCitation id="13572A604926AE7FFD6B225EFD4F167D" box="[655,666,1344,1368]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1875]" captionTargetBox="[206,1380,373,1832]" captionTargetId="graphics@2.[259,1381,373,1832]" captionTargetPageId="2" captionText="FIGURE 1. Tanaopsis japonica sp. nov. A, holotype female; B, allotype male: A, B, body, dorsal view; a 1, b 1, body, left view; a 2, b 3, pleonite 5 and pleotelson, dorsal view; b 2, genital cones on pereonite 6, ventral view; a 3, b 4, pereonite 6 and pleon, left view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1010380/files/figure.svg" pageId="6">1</figureCitation>
); fixed finger with two simple setae on ventral margin, three simple setae on cutting surface, series of outer smooth ridges (
<figureCitation id="13572A604926AE7FFCCA227AFC661659" box="[814,947,1380,1405]" captionStart="FIGURE 6" captionStartId="7.[151,250,1751,1773]" captionTargetBox="[161,1425,697,1738]" captionTargetId="figure@7.[161,1425,697,1738]" captionTargetPageId="7" captionText="FIGURE 6. Tanaopsis japonica sp. nov., paratype female, scanning electron microscopic images: A, terminal region of antennule, with double arrowhead indicating vestige of small cap-like article; B, cheliped-sclerite-cephalothorax connection, right view, with asterisk indicating acute dorsal process on pereopod- 1 coxa (other parts were detached); C – E, right chela, anterior (C), outer (D), and inner dorsal (E) views, with each white arrowhead indicating one ridge in outer series of denticulate ridges on dactylus, each black arrowhead indicating one ridge in series of smooth ridges on fixed finger, and each arrow indicating one fringed incision in inner hatching on dactylus; inserts in (D) and (E) are images magnified threefold from the corresponding boxes in the main panels. Scale bars: 0.01 mm for A; 0.05 mm for B – E." httpUri="https://zenodo.org/record/1010390/files/figure.png" pageId="6">Fig. 6C, D</figureCitation>
: black arrowheads), outer distal bifurcate process, and bifurcate claw; dactylus as long as fixed finger, with inner simple seta, three spiniform setae on cutting surface, series of denticulate outer dorsal ridges (
<figureCitation id="13572A604926AE7FFD0722B2FCB416E0" box="[739,865,1452,1477]" captionStart="FIGURE 6" captionStartId="7.[151,250,1751,1773]" captionTargetBox="[161,1425,697,1738]" captionTargetId="figure@7.[161,1425,697,1738]" captionTargetPageId="7" captionText="FIGURE 6. Tanaopsis japonica sp. nov., paratype female, scanning electron microscopic images: A, terminal region of antennule, with double arrowhead indicating vestige of small cap-like article; B, cheliped-sclerite-cephalothorax connection, right view, with asterisk indicating acute dorsal process on pereopod- 1 coxa (other parts were detached); C – E, right chela, anterior (C), outer (D), and inner dorsal (E) views, with each white arrowhead indicating one ridge in outer series of denticulate ridges on dactylus, each black arrowhead indicating one ridge in series of smooth ridges on fixed finger, and each arrow indicating one fringed incision in inner hatching on dactylus; inserts in (D) and (E) are images magnified threefold from the corresponding boxes in the main panels. Scale bars: 0.01 mm for A; 0.05 mm for B – E." httpUri="https://zenodo.org/record/1010390/files/figure.png" pageId="6">Fig. 6C–E</figureCitation>
: white arrowheads), inner hatching with fringed incisions (
<figureCitation id="13572A604926AE7FFEEE22D1FE5716CD" box="[266,386,1487,1512]" captionStart="FIGURE 6" captionStartId="7.[151,250,1751,1773]" captionTargetBox="[161,1425,697,1738]" captionTargetId="figure@7.[161,1425,697,1738]" captionTargetPageId="7" captionText="FIGURE 6. Tanaopsis japonica sp. nov., paratype female, scanning electron microscopic images: A, terminal region of antennule, with double arrowhead indicating vestige of small cap-like article; B, cheliped-sclerite-cephalothorax connection, right view, with asterisk indicating acute dorsal process on pereopod- 1 coxa (other parts were detached); C – E, right chela, anterior (C), outer (D), and inner dorsal (E) views, with each white arrowhead indicating one ridge in outer series of denticulate ridges on dactylus, each black arrowhead indicating one ridge in series of smooth ridges on fixed finger, and each arrow indicating one fringed incision in inner hatching on dactylus; inserts in (D) and (E) are images magnified threefold from the corresponding boxes in the main panels. Scale bars: 0.01 mm for A; 0.05 mm for B – E." httpUri="https://zenodo.org/record/1010390/files/figure.png" pageId="6">Fig. 6C–E</figureCitation>
: arrows), and triangular claw.
</paragraph>
<paragraph id="8BD336E54926AE7EFF2322EAFEA71241" blockId="6.[151,1437,1199,2017]" lastBlockId="7.[151,1437,151,645]" lastPageId="7" pageId="6">
Pereopods 1–6 cylindrical, with length ratio of 1.5:1.1:1.0:1.0:1.0:1.0. Pereopod 1 (
<figureCitation id="13572A604926AE7FFB6022EAFAD41529" box="[1156,1281,1524,1549]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="6">Fig. 3C, D</figureCitation>
) 0.4 times as long as BL, with length ratio of basis, ischium, merus, carpus, propodus, and dactylus-unguis 4.5:0.4:1.0:1.4:2.9:3.7. Coxa with dorsal simple seta and acute dorsal process. Basis cylindrical, narrow (4.5 times as long as wide), naked. Ischium with ventral simple seta. Merus naked. Carpus with three dorsodistal and one ventrodistal simple setae. Propodus serrated dorsally, with three dorso-subdistal and one ventro-subdistal setae, and dorsodistal serrations. Dactylus with dorsoproximal simple seta. Unguis twice as long as dactylus, naked. Pereopod 2 (
<figureCitation id="13572A604926AE7FFF5021D2FEDD15C1" box="[180,264,1740,1765]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="6">Fig. 3E</figureCitation>
) with length ratio of articles from basis to dactylus-unguis 5.1:0.4:1.0:1.5:3.1:3.6; similar to pereopod 1, except coxa without dorsal process, basis with one dorsal PSS, carpus with two dorsodistal simple setae, and propodus with two dorso-subdistal setae. Pereopod 3 (
<figureCitation id="13572A604926AE7FFD18200AFC851409" box="[764,848,1812,1837]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="6">Fig. 3F</figureCitation>
) with length ratio of articles from basis to dactylusunguis 5.3:0.5:1.0:1.4:3.2:3.1; similar to pereopod 2, except basis with one ventral PSS. Pereopod 4 (
<figureCitation id="13572A604926AE7FFADD2026FA461475" box="[1337,1427,1848,1873]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="6">Fig. 3G</figureCitation>
) without coxa. Length ratio of articles from basis to dactylus-unguis 3.5:0.3:1.0:1.3:1.7:1.7. Basis thick (2.5 times as long as wide), with one dorsal PSS. Ischium with ventral simple seta. Merus with one outer and one inner serratespiniform setae in ventrodistal region. Carpus with dorsodistal simple seta, and one outer and two inner serratespiniform setae in distal region. Propodus with one dorsal PSS, dorsal serrations on edge in distal half, and one dorsodistal, one outer ventrodistal, and one inner ventrodistal pinnate-spiniform setae. Dactylus naked. Unguis 0.6 times as long as dactylus. Pereopod 5 (
<figureCitation id="13572A604927AE7EFD9227A2FD0613F1" box="[630,723,188,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="7">Fig. 3H</figureCitation>
) with length ratio of articles from basis to dactylus-unguis 2.7:0.2:1.0:1.1:1.6:1.4; similar to pereopod 4 except basis with two dorsal and one ventral PSS, and ischium with two ventral simple setae. Pereopod 6 (
<figureCitation id="13572A604927AE7EFD86261AFD1A1238" box="[610,719,260,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="7">
Fig. 
<date id="FFD210254927AE7EFD7F261AFD601239" box="[667,693,260,284]" pageId="7">3I</date>
, 
<date id="FFD210254927AE7EFD22261AFD1A1238" box="[710,719,260,285]" pageId="7">i</date>
</figureCitation>
<figureCitation id="13572A604927AE7EFD2B261AFD0F1238" box="[719,730,260,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1875]" captionTargetBox="[206,1380,373,1832]" captionTargetId="graphics@2.[259,1381,373,1832]" captionTargetPageId="2" captionText="FIGURE 1. Tanaopsis japonica sp. nov. A, holotype female; B, allotype male: A, B, body, dorsal view; a 1, b 1, body, left view; a 2, b 3, pleonite 5 and pleotelson, dorsal view; b 2, genital cones on pereonite 6, ventral view; a 3, b 4, pereonite 6 and pleon, left view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1010380/files/figure.svg" pageId="7">1</figureCitation>
) with length ratio of articles from basis to dactylus-unguis 3.0:0.3:1.0:1.0:1.5:1.1; similar to pereopod 4 except propodus with two dorsodistal serrate setae (
<figureCitation id="13572A604927AE7EFAEC2639FA8D1265" box="[1288,1368,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="7">
Fig. 3 
<date id="FFD210254927AE7EFAAB2639FA8D1265" box="[1359,1368,295,320]" pageId="7">i</date>
</figureCitation>
<figureCitation id="13572A604927AE7EFABD2639FAB11265" box="[1369,1380,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1875]" captionTargetBox="[206,1380,373,1832]" captionTargetId="graphics@2.[259,1381,373,1832]" captionTargetPageId="2" captionText="FIGURE 1. Tanaopsis japonica sp. nov. A, holotype female; B, allotype male: A, B, body, dorsal view; a 1, b 1, body, left view; a 2, b 3, pleonite 5 and pleotelson, dorsal view; b 2, genital cones on pereonite 6, ventral view; a 3, b 4, pereonite 6 and pleon, left view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1010380/files/figure.svg" pageId="7">1</figureCitation>
) but lacking dorsal PSS.
</paragraph>
<paragraph id="8BD336E54927AE7EFF232671FDFF12D1" blockId="7.[151,1437,151,645]" pageId="7">
Pleopods (
<figureCitation id="13572A604927AE7EFEDA2671FE5812AD" box="[318,397,367,392]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="7">Fig. 3J</figureCitation>
) five pairs, all similar. Basal article naked. Endopod 2.2 times as long as basal article, with one inner subdistal and five outer plumose setae, and outer distal “step-tipped plumose seta” (see 
<bibRefCitation id="EFFD4B144927AE7EFB30268AFA4D1289" author="Kakui" box="[1236,1432,404,429]" pageId="7" refString="Kakui, K., Kajihara, H. &amp; Mawatari, S. F. (2010) A new species of Nesotanais Shiino, 1968 (Crustacea, Tanaidacea) from Japan, with a key to species and a note on male chelipeds. ZooKeys, 33, 1 - 17. https: // doi. org / 10.3897 / zookeys. 33.296" type="journal article" year="2010">
Kakui 
<emphasis id="B918EAF74927AE7EFAFA268BFA9B1288" box="[1310,1358,404,429]" italics="true" pageId="7">et al</emphasis>
. 2010
</bibRefCitation>
: fig. 5j2). Exopod 1.3 times as long as endopod, with 17–20 outer plumose setae; “vestigial proximal article” (see 
<bibRefCitation id="EFFD4B144927AE7EFF7326C2FEDF12D1" author="Bird" box="[151,266,476,501]" pageId="7" refString="Bird, G. J. (2012) A new leptochelioid family, Heterotanoididae (Crustacea: Peracarida: Tanaidacea), and a new species of Heterotanoides from New Zealand. Zootaxa, 3481, 1 - 26." type="journal article" year="2012">Bird 2012</bibRefCitation>
) with outer plumose seta.
</paragraph>
<paragraph id="8BD336E54927AE7EFF2326E1FE1911A1" blockId="7.[151,1437,151,645]" pageId="7">
Uropod (
<figureCitation id="13572A604927AE7EFECA26E1FE52113D" box="[302,391,511,536]" captionStart="FIGURE 3" captionStartId="4.[151,250,1898,1920]" captionTargetBox="[154,1432,339,1874]" captionTargetId="graphics@4.[154,1432,339,1874]" captionTargetPageId="4" captionText="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." httpUri="https://zenodo.org/record/1010384/files/figure.svg" pageId="7">Fig. 3K</figureCitation>
) with basal article naked. Endopod 2.5 times as long as basal article, biarticulate; article 1 with length/width ratio 1.7, with one distal PSS; article 2 as long as article 1, with one subdistal and four distal simple setae, and two distal PSS. Exopod uniarticulate, slightly shorter than endopodal article 1, with one subdistal and two distal simple setae.
</paragraph>
<caption id="DF13666D4927AE7EFF7321C9FBD01492" httpUri="https://zenodo.org/record/1010390/files/figure.png" pageId="7" targetBox="[161,1425,697,1738]" targetPageId="7">
<paragraph id="8BD336E54927AE7EFF7321C9FBD01492" blockId="7.[151,1436,1751,1975]" pageId="7">
<emphasis id="B918EAF74927AE7EFF7321C9FEC215C9" bold="true" box="[151,279,1751,1773]" pageId="7">FIGURE 6.</emphasis>
<taxonomicName id="4C6C4D664927AE7EFEC621C6FE3915C8" authority="Shimada, 2017" authorityName="Shimada" authorityYear="2017" box="[290,492,1752,1773]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="7" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="B918EAF74927AE7EFEC621C6FE3915C8" box="[290,492,1752,1773]" italics="true" pageId="7">Tanaopsis japonica</emphasis>
</taxonomicName>
<emphasis id="B918EAF74927AE7EFE1321C9FD9915C9" bold="true" box="[503,588,1751,1772]" pageId="7">
<taxonomicNameLabel id="A22B578C4927AE7EFE1321C9FD9915C9" box="[503,588,1751,1772]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype female, scanning electron microscopic images: 
<emphasis id="B918EAF74927AE7EFB5C21C9FB1F15C9" bold="true" box="[1208,1226,1751,1772]" pageId="7">A</emphasis>
, terminal region of antennule, with double arrowhead indicating vestige of small cap-like article; 
<emphasis id="B918EAF74927AE7EFC2421E7FC04142B" bold="true" box="[960,977,1785,1806]" pageId="7">B</emphasis>
, cheliped-sclerite-cephalothorax connection, right view, with asterisk indicating acute dorsal process on pereopod-1 coxa (other parts were detached); 
<emphasis id="B918EAF74927AE7EFB0C2004FACD1415" bold="true" box="[1256,1304,1818,1840]" pageId="7">C–E</emphasis>
, right chela, anterior (
<emphasis id="B918EAF74927AE7EFF102025FED31474" bold="true" box="[244,262,1851,1873]" pageId="7">C</emphasis>
), outer (
<emphasis id="B918EAF74927AE7EFEB92025FEBA1475" bold="true" box="[349,367,1851,1872]" pageId="7">D</emphasis>
), and inner dorsal (
<emphasis id="B918EAF74927AE7EFDD02025FD901475" bold="true" box="[564,581,1851,1872]" pageId="7">E</emphasis>
) views, with each white arrowhead indicating one ridge in outer series of denticulate ridges on dactylus, each black arrowhead indicating one ridge in series of smooth ridges on fixed finger, and each arrow indicating one fringed incision in inner hatching on dactylus; inserts in (D) and (E) are images magnified threefold from the corresponding boxes in the main panels. Scale bars: 0.01 mm for A; 0.05 mm for B–E.
</paragraph>
</caption>
<caption id="DF13666D4928AE71FF732112FD2C159E" httpUri="https://zenodo.org/record/1010392/files/figure.png" pageId="8" targetBox="[151,1436,193,1526]" targetPageId="8">
<paragraph id="8BD336E54928AE71FF732112FD2C159E" blockId="8.[151,1437,1548,1723]" pageId="8">
<emphasis id="B918EAF74928AE71FF732112FEC71504" bold="true" box="[151,274,1548,1570]" pageId="8">FIGURE 7.</emphasis>
<taxonomicName id="4C6C4D664928AE71FEFD2113FE0B1507" authority="Shimada, 2017" authorityName="Shimada" authorityYear="2017" box="[281,478,1549,1570]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="8" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="B918EAF74928AE71FEFD2113FE0B1507" box="[281,478,1549,1570]" italics="true" pageId="8">Tanaopsis japonica</emphasis>
</taxonomicName>
<emphasis id="B918EAF74928AE71FE002112FD9B1504" bold="true" box="[484,590,1548,1569]" pageId="8">
<taxonomicNameLabel id="A22B578C4928AE71FE002112FDE01504" box="[484,565,1548,1569]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
, 
<emphasis id="B918EAF74928AE71FDBF2112FDB91504" bold="true" box="[603,620,1548,1569]" pageId="8">B</emphasis>
, holotype female; 
<emphasis id="B918EAF74928AE71FCCE2112FC831507" bold="true" box="[810,854,1548,1570]" pageId="8">C–F</emphasis>
, paratype female; scanning electron microscopic images: 
<emphasis id="B918EAF74928AE71FF732134FF7C151A" bold="true" box="[151,169,1578,1599]" pageId="8">A</emphasis>
, part of right antenna, dorsal view, showing series of denticulate ridges on article 2; 
<emphasis id="B918EAF74928AE71FBF02135FBF01565" bold="true" box="[1044,1061,1579,1600]" pageId="8">B</emphasis>
, enlargement of ridges from (A); 
<emphasis id="B918EAF74928AE71FA672134FA401565" bold="true" box="[1411,1429,1578,1600]" pageId="8">C</emphasis>
, cephalothorax, anterior ventral view (left cheliped and antenna removed); 
<emphasis id="B918EAF74928AE71FC702157FC73157B" bold="true" box="[916,934,1609,1630]" pageId="8">D</emphasis>
, 
<emphasis id="B918EAF74928AE71FC502157FC10157B" bold="true" box="[948,965,1609,1630]" pageId="8">E</emphasis>
, proximal region of antennule and antenna in right view and right anterior ventral view, respecively; 
<emphasis id="B918EAF74928AE71FD002176FD241558" bold="true" box="[740,753,1640,1661]" pageId="8">F</emphasis>
, enlargement from (E) showing grooves and ridges. Each white arrowhead, dorsal denticulate ridge on antenna article 2; each black arrowhead, ventral groove on antennule article 1. Scale bars: 0.01 mm for A, B, F; 0.1 mm for C, E; 0.05 mm for D.
</paragraph>
</caption>
<paragraph id="8BD336E54928AE71FF2321F9FD5B1425" blockId="8.[151,1437,1767,2008]" box="[199,654,1767,1792]" pageId="8">
<emphasis id="B918EAF74928AE71FF2321F9FE621425" bold="true" box="[199,439,1767,1792]" pageId="8">Description of male.</emphasis>
Based on 
<typeStatus id="54D788474928AE71FDC921F9FD5E1425" box="[557,651,1767,1792]" pageId="8" type="allotype">allotype</typeStatus>
.
</paragraph>
<paragraph id="8BD336E54928AE71FF232015FECC144E" blockId="8.[151,1437,1767,2008]" pageId="8">
Body (
<figureCitation id="13572A604928AE71FEF12015FE631401" box="[277,438,1803,1828]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1853,1875]" captionStartId-1="3.[157,256,1810,1832]" captionStartId-2="4.[151,250,1898,1920]" captionStartId-3="5.[151,250,1827,1849]" captionTargetBox-0="[206,1380,373,1832]" captionTargetBox-1="[164,1422,625,1786]" captionTargetBox-2="[154,1432,339,1874]" captionTargetBox-3="[158,1428,481,1806]" captionTargetId-0="graphics@2.[259,1381,373,1832]" captionTargetId-1="graphics@3.[164,1422,625,1786]" captionTargetId-2="graphics@4.[154,1432,339,1874]" captionTargetId-3="graphics@5.[158,1428,481,1806]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Tanaopsis japonica sp. nov. A, holotype female; B, allotype male: A, B, body, dorsal view; a 1, b 1, body, left view; a 2, b 3, pleonite 5 and pleotelson, dorsal view; b 2, genital cones on pereonite 6, ventral view; a 3, b 4, pereonite 6 and pleon, left view. Scale bars: 0.5 mm." captionText-1="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." captionText-2="FIGURE 3. Tanaopsis japonica sp. nov., holotype female: A, right cheliped, outer view; a 1, same, chela, inner view; a 2, same, distal region of fixed finger, outer view; B, left chelipedal dactylus; C, D, right and left pereopods 1, respectively, outer views; E, left pereopod 2, outer view; F, left pereopod 3, inner view; G – I, right pereopods 4 – 6, respectively, outer view; i 1, serrate seta; J, right pleopod 1, most setal ornamentation omitted; K, right uropod. Scale bars: 0.05 mm for a 2, B, i 1; 0.1 mm for the others." captionText-3="FIGURE 4. Tanaopsis japonica sp. nov., allotype male: A, right antennule; B, right antenna; C, anterior region of cephalothorax, left view; D, labium, ventral view; E, maxillipeds; F, right cheliped, outer view; f 1, same, distal region of fixed finger, outer view; G, left cheliped, inner view. Scale bars: 0.1 mm for A – D, F, G; 0.05 mm for E, f 1." httpUri-0="https://zenodo.org/record/1010380/files/figure.svg" httpUri-1="https://zenodo.org/record/1010382/files/figure.svg" httpUri-2="https://zenodo.org/record/1010384/files/figure.svg" httpUri-3="https://zenodo.org/record/1010386/files/figure.svg" pageId="8">Fig. 1B, b1–4</figureCitation>
) similar to female but pleon proportionally longer. Pereonites 1–6 with length ratio of 1.0:1.2:1.7:2.4:2.3:1.5; all wider than long; pereonite 6 with pair of genital cones (
<figureCitation id="13572A604928AE71FBAA2031FB74146D" box="[1102,1185,1839,1864]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1875]" captionTargetBox="[206,1380,373,1832]" captionTargetId="graphics@2.[259,1381,373,1832]" captionTargetPageId="2" captionText="FIGURE 1. Tanaopsis japonica sp. nov. A, holotype female; B, allotype male: A, B, body, dorsal view; a 1, b 1, body, left view; a 2, b 3, pleonite 5 and pleotelson, dorsal view; b 2, genital cones on pereonite 6, ventral view; a 3, b 4, pereonite 6 and pleon, left view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1010380/files/figure.svg" pageId="8">Fig. 1b</figureCitation>
<figureCitation id="13572A604928AE71FB452031FB67146D" box="[1185,1202,1839,1864]" captionStart="FIGURE 2" captionStartId="3.[157,256,1810,1832]" captionTargetBox="[164,1422,625,1786]" captionTargetId="graphics@3.[164,1422,625,1786]" captionTargetPageId="3" captionText="FIGURE 2. Tanaopsis japonica sp. nov., holotype female: A, left antennule; B, left antenna; C, proximal region of right antenna; D, labrum; E, F, left and right mandibles; G, labium; H, distal part of left maxillule; I, maxillipeds; J, right epignath. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010382/files/figure.svg" pageId="8">2</figureCitation>
). Pleon 0.4 times as long as BL.
</paragraph>
<paragraph id="8BD336E54928AE70FF232069FBF913F0" blockId="8.[151,1437,1767,2008]" lastBlockId="9.[151,1437,151,1617]" lastPageId="9" pageId="8">
Antennule (
<figureCitation id="13572A604928AE71FEA82069FE7014B5" box="[332,421,1911,1936]" captionStart="FIGURE 4" captionStartId="5.[151,250,1827,1849]" captionTargetBox="[158,1428,481,1806]" captionTargetId="graphics@5.[158,1428,481,1806]" captionTargetPageId="5" captionText="FIGURE 4. Tanaopsis japonica sp. nov., allotype male: A, right antennule; B, right antenna; C, anterior region of cephalothorax, left view; D, labium, ventral view; E, maxillipeds; F, right cheliped, outer view; f 1, same, distal region of fixed finger, outer view; G, left cheliped, inner view. Scale bars: 0.1 mm for A – D, F, G; 0.05 mm for E, f 1." httpUri="https://zenodo.org/record/1010386/files/figure.svg" pageId="8">Fig. 4A</figureCitation>
) as long as cephalothorax, with six articles; thicker than that of female; articles 1–6 with length ratio of 1.0:0.4:0.1:0.1:0.2:0.2. Setation of articles 1–3 similar to those of female. Articles 4 and 5 with ventrodistal row of aesthetascs. Article 6 with six simple setae and aesthetasc; cap-like vestigial article not observed. Antenna (
<figureCitation id="13572A604929AE70FE9C2789FE1B1395" box="[376,462,151,176]" captionStart="FIGURE 4" captionStartId="5.[151,250,1827,1849]" captionTargetBox="[158,1428,481,1806]" captionTargetId="graphics@5.[158,1428,481,1806]" captionTargetPageId="5" captionText="FIGURE 4. Tanaopsis japonica sp. nov., allotype male: A, right antennule; B, right antenna; C, anterior region of cephalothorax, left view; D, labium, ventral view; E, maxillipeds; F, right cheliped, outer view; f 1, same, distal region of fixed finger, outer view; G, left cheliped, inner view. Scale bars: 0.1 mm for A – D, F, G; 0.05 mm for E, f 1." httpUri="https://zenodo.org/record/1010386/files/figure.svg" pageId="9">Fig. 4B</figureCitation>
) 0.8 times as long as antennule, similar to that of female except article 4 with two midouter PSS, and articles 5 and 6 with two and four distal simple setae, respectively.
</paragraph>
<paragraph id="8BD336E54929AE70FF2327C1FE7D1265" blockId="9.[151,1437,151,1617]" pageId="9">
Mouthparts (
<figureCitation id="13572A604929AE70FE8427C1FE3513DD" box="[352,480,223,248]" captionStart="FIGURE 4" captionStartId="5.[151,250,1827,1849]" captionTargetBox="[158,1428,481,1806]" captionTargetId="graphics@5.[158,1428,481,1806]" captionTargetPageId="5" captionText="FIGURE 4. Tanaopsis japonica sp. nov., allotype male: A, right antennule; B, right antenna; C, anterior region of cephalothorax, left view; D, labium, ventral view; E, maxillipeds; F, right cheliped, outer view; f 1, same, distal region of fixed finger, outer view; G, left cheliped, inner view. Scale bars: 0.1 mm for A – D, F, G; 0.05 mm for E, f 1." httpUri="https://zenodo.org/record/1010386/files/figure.svg" pageId="9">Fig. 4C–E</figureCitation>
) reduced. Vestigial labrum (
<figureCitation id="13572A604929AE70FCA527C1FC7113DD" box="[833,932,223,248]" captionStart="FIGURE 4" captionStartId="5.[151,250,1827,1849]" captionTargetBox="[158,1428,481,1806]" captionTargetId="graphics@5.[158,1428,481,1806]" captionTargetPageId="5" captionText="FIGURE 4. Tanaopsis japonica sp. nov., allotype male: A, right antennule; B, right antenna; C, anterior region of cephalothorax, left view; D, labium, ventral view; E, maxillipeds; F, right cheliped, outer view; f 1, same, distal region of fixed finger, outer view; G, left cheliped, inner view. Scale bars: 0.1 mm for A – D, F, G; 0.05 mm for E, f 1." httpUri="https://zenodo.org/record/1010386/files/figure.svg" pageId="9">Fig. 4C</figureCitation>
: arrowhead) observed. Labium (
<figureCitation id="13572A604929AE70FAD727C1FA4113DD" box="[1331,1428,223,248]" captionStart="FIGURE 4" captionStartId="5.[151,250,1827,1849]" captionTargetBox="[158,1428,481,1806]" captionTargetId="graphics@5.[158,1428,481,1806]" captionTargetPageId="5" captionText="FIGURE 4. Tanaopsis japonica sp. nov., allotype male: A, right antennule; B, right antenna; C, anterior region of cephalothorax, left view; D, labium, ventral view; E, maxillipeds; F, right cheliped, outer view; f 1, same, distal region of fixed finger, outer view; G, left cheliped, inner view. Scale bars: 0.1 mm for A – D, F, G; 0.05 mm for E, f 1." httpUri="https://zenodo.org/record/1010386/files/figure.svg" pageId="9">Fig. 4D</figureCitation>
) narrowing distally, naked. Maxillipeds (
<figureCitation id="13572A604929AE70FDBE261AFD651239" box="[602,688,260,285]" captionStart="FIGURE 4" captionStartId="5.[151,250,1827,1849]" captionTargetBox="[158,1428,481,1806]" captionTargetId="graphics@5.[158,1428,481,1806]" captionTargetPageId="5" captionText="FIGURE 4. Tanaopsis japonica sp. nov., allotype male: A, right antennule; B, right antenna; C, anterior region of cephalothorax, left view; D, labium, ventral view; E, maxillipeds; F, right cheliped, outer view; f 1, same, distal region of fixed finger, outer view; G, left cheliped, inner view. Scale bars: 0.1 mm for A – D, F, G; 0.05 mm for E, f 1." httpUri="https://zenodo.org/record/1010386/files/figure.svg" pageId="9">Fig. 4E</figureCitation>
) similar to those of female, except endites not flared and all setae on palp-article 4 simple.
</paragraph>
<paragraph id="8BD336E54929AE70FF232652FE911288" blockId="9.[151,1437,151,1617]" pageId="9">
Cheliped (
<figureCitation id="13572A604929AE70FED82652FE7C1240" box="[316,425,332,357]" captionStart="FIGURE 4" captionStartId="5.[151,250,1827,1849]" captionTargetBox="[158,1428,481,1806]" captionTargetId="graphics@5.[158,1428,481,1806]" captionTargetPageId="5" captionText="FIGURE 4. Tanaopsis japonica sp. nov., allotype male: A, right antennule; B, right antenna; C, anterior region of cephalothorax, left view; D, labium, ventral view; E, maxillipeds; F, right cheliped, outer view; f 1, same, distal region of fixed finger, outer view; G, left cheliped, inner view. Scale bars: 0.1 mm for A – D, F, G; 0.05 mm for E, f 1." httpUri="https://zenodo.org/record/1010386/files/figure.svg" pageId="9">Fig. 4F, f</figureCitation>
<figureCitation id="13572A604929AE70FE4D2652FE611240" box="[425,436,332,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1875]" captionTargetBox="[206,1380,373,1832]" captionTargetId="graphics@2.[259,1381,373,1832]" captionTargetPageId="2" captionText="FIGURE 1. Tanaopsis japonica sp. nov. A, holotype female; B, allotype male: A, B, body, dorsal view; a 1, b 1, body, left view; a 2, b 3, pleonite 5 and pleotelson, dorsal view; b 2, genital cones on pereonite 6, ventral view; a 3, b 4, pereonite 6 and pleon, left view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1010380/files/figure.svg" pageId="9">1</figureCitation>
, G) similar to that of female, except propodal palm with six shorter inner simple setae at insertion of dactylus, fixed finger with outer distal trifurcate process, and dactylus with two spiniform setae on cutting surface.
</paragraph>
<paragraph id="8BD336E54929AE70FF2326A9FB511119" blockId="9.[151,1437,151,1617]" pageId="9">
Pereopods 1–6 (
<figureCitation id="13572A604929AE70FE9B26A9FE2312F5" box="[383,502,439,464]" captionStart="FIGURE 5" captionStartId="6.[155,253,1102,1124]" captionTargetBox="[162,1425,193,1081]" captionTargetId="graphics@6.[162,1425,193,1080]" captionTargetPageId="6" captionText="FIGURE 5. Tanaopsis japonica sp. nov., allotype male: A – F, right pereopods 1 – 6, respectively, outer view; G, right pleopod 1, most setal ornamentation omitted; H, right uropod. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010388/files/figure.svg" pageId="9">Fig. 5A–F</figureCitation>
) with length ratio of 1.4:1.1:1.0:1.1:1.1:1.1 (note that pereopod-6 unguis broken); pereopod 1 length 0.38 times BL. Pereopods similar to those of female, but differ as follows: coxa of pereopod 1 without acute dorsal process; basis of both pereopods 4 and 5 with two ventral PSS; ischium of pereopods 4 and 6 with two ventral simple setae; carpus of pereopods 4–6 with two dorsodistal simple setae.
</paragraph>
<paragraph id="8BD336E54929AE70FF232559FABC118D" blockId="9.[151,1437,151,1617]" pageId="9">
Pleopods (
<figureCitation id="13572A604929AE70FED92559FE431145" box="[317,406,583,608]" captionStart="FIGURE 5" captionStartId="6.[155,253,1102,1124]" captionTargetBox="[162,1425,193,1081]" captionTargetId="graphics@6.[162,1425,193,1080]" captionTargetPageId="6" captionText="FIGURE 5. Tanaopsis japonica sp. nov., allotype male: A – F, right pereopods 1 – 6, respectively, outer view; G, right pleopod 1, most setal ornamentation omitted; H, right uropod. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010388/files/figure.svg" pageId="9">Fig. 5G</figureCitation>
) five pairs, all similar. Basal article naked. Endopod 1.8 times as long as basal article, with five longer and one shorter outer plumose setae, and one inner subdistal and one outer distal step-tipped plumose setae. Exopod 1.2 times as long as endopod, with 10–14 outer plumose setae; vestigial proximal article naked.
</paragraph>
<paragraph id="8BD336E54929AE70FF2325AAFC0511CA" blockId="9.[151,1437,151,1617]" pageId="9">
Uropod (
<figureCitation id="13572A604929AE70FEC825AAFE5111E8" box="[300,388,692,717]" captionStart="FIGURE 5" captionStartId="6.[155,253,1102,1124]" captionTargetBox="[162,1425,193,1081]" captionTargetId="graphics@6.[162,1425,193,1080]" captionTargetPageId="6" captionText="FIGURE 5. Tanaopsis japonica sp. nov., allotype male: A – F, right pereopods 1 – 6, respectively, outer view; G, right pleopod 1, most setal ornamentation omitted; H, right uropod. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1010388/files/figure.svg" pageId="9">Fig. 5H</figureCitation>
) similar to that of female, except endo- and exopod proportionally shorter (endopod twice as long as basal article) and endopodal article 2 with two distal simple setae.
</paragraph>
<paragraph id="8BD336E54929AE70FF2325E2FEBC1735" blockId="9.[151,1437,151,1617]" pageId="9">
<emphasis id="B918EAF74929AE70FF2325E2FE081030" bold="true" box="[199,477,764,789]" pageId="9">Variation and stability.</emphasis>
One additional female (
<typeStatus id="54D788474929AE70FD1025E2FC8D1031" box="[756,856,764,788]" pageId="9" type="paratype">paratype</typeStatus>
) was observed. The ratio of BL/CW was different among specimens, i.e., 4.4 (female 
<typeStatus id="54D788474929AE70FDC82401FD44101D" box="[556,657,799,824]" pageId="9" type="holotype">holotype</typeStatus>
; BL 1.43), 4.5 (
<typeStatus id="54D788474929AE70FCAC2401FC72101D" box="[840,935,799,824]" pageId="9" type="allotype">allotype</typeStatus>
; BL 1.44), and 5.5 (
<typeStatus id="54D788474929AE70FB6B243EFB20101D" box="[1167,1269,800,824]" pageId="9" type="paratype">paratype</typeStatus>
; BL 1.75), but pereonites 4 and 5 were subequal in length in all specimens, i.e., the length ratio of pereonites 4/5 was 1.0. In the two females, the numbers of simple, spiniform, serrate-spiniform, pinnate-spiniform, and serrate setae on the antenna, cheliped, and pereopods were constant. The number of setae on the pleopodal endopod was constant among the five pairs in each specimen and between the two females; that on the exopod varied, ranging from 
<quantity id="4C949B004929AE70FA8024B1FF0510C8" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="9" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">16 to 18 in</quantity>
the 
<typeStatus id="54D788474929AE70FEE424CBFEB110C8" box="[256,356,981,1005]" pageId="9" type="paratype">paratype</typeStatus>
. All specimens lacked setae on the basal article of all pleopods. All specimens had a biarticulate uropodal endopod.
</paragraph>
<paragraph id="8BD336E54929AE70FF232302FC2517CD" blockId="9.[151,1437,151,1617]" pageId="9">
<emphasis id="B918EAF74929AE70FF232302FE101710" bold="true" box="[199,453,1052,1077]" pageId="9">Genetic Information.</emphasis>
The nearly complete 18S-gene sequence (2275 nt) was determined from the paratype specimen; INSD accession number 
<accessionNumber id="943FAB064929AE70FDDF2321FD63177D" box="[571,694,1087,1112]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC210598" pageId="9">
LC 
<date id="FFD210254929AE70FDBB2321FD63177D" box="[607,694,1087,1112]" pageId="9" value="98-05-21">210598</date>
</accessionNumber>
. The three sequences in the International Nucleotide Sequence Database most similar to our sequence, as determined by BLAST searches (
<bibRefCitation id="EFFD4B144929AE70FC19237AFB331759" author="Altschul" box="[1021,1254,1124,1149]" pageId="9" refString="Altschul, S. F., Gish, W., Miller, W., Myers, E. W. &amp; Lipman, D. J. (1990) Basic local alignment search tool. Journal of Molecular Biology, 215, 403 - 410. https: // doi. org / 10.1016 / S 0022 - 2836 (05) 80360 - 2" type="journal article" year="1990">
Altschul 
<emphasis id="B918EAF74929AE70FB83237BFB4F1758" box="[1127,1178,1124,1149]" italics="true" pageId="9">et al</emphasis>
. 1990
</bibRefCitation>
), were from the tanaidaceans 
<taxonomicName id="4C6C4D664929AE70FED42399FE031785" authorityName="Dojiri &amp; Sieg" authorityYear="1997" box="[304,470,1159,1184]" class="Malacostraca" family="Akanthophoreidae" genus="Chauliopleona" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAF74929AE70FED42399FE031785" box="[304,470,1159,1184]" italics="true" pageId="9">Chauliopleona</emphasis>
</taxonomicName>
sp. KK-2011 (identity score [IS] 97%; query cover [QC] 65%; 
<taxonomicName id="4C6C4D664929AE70FB222399FA4C1785" authorityName="Sieg" authorityYear="1986" box="[1222,1433,1159,1184]" class="Malacostraca" family="Akanthophoreidae" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="family">Akanthophoreidae</taxonomicName>
; 
<bibRefCitation id="EFFD4B144929AE70FF7323B2FE8A17E1" author="Kakui" box="[151,351,1196,1221]" pageId="9" refString="Kakui, K. Katoh, T., Hiruta, S. F., Kobayashi, N. &amp; Kajihara, H. (2011) Molecular systematics of Tanaidacea (Crustacea: Peracarida) based on 18 S sequence data, with an amendment of suborder / superfamily-level classification. Zoological Science, 28, 749 - 757. https: // doi. org / 10.2108 / zsj. 28.749" type="journal article" year="2011">
Kakui 
<emphasis id="B918EAF74929AE70FF0023B3FEC017E0" box="[228,277,1196,1221]" italics="true" pageId="9">et al</emphasis>
. 2011
</bibRefCitation>
), 
<taxonomicName id="4C6C4D664929AE70FE9023B3FE2517E1" box="[372,496,1197,1220]" class="Malacostraca" family="Paratanaidae" genus="Paratanais" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAF74929AE70FE9023B3FE2517E1" box="[372,496,1197,1220]" italics="true" pageId="9">Paratanais</emphasis>
</taxonomicName>
sp. KK-2011 (IS 98%; QC 63%; 
<taxonomicName id="4C6C4D664929AE70FC9623B2FBD017E0" authorityName="Lang" authorityYear="1949" box="[882,1029,1196,1221]" class="Malacostraca" family="Paratanaidae" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="family">Paratanaidae</taxonomicName>
; 
<bibRefCitation id="EFFD4B144929AE70FBF523B2FB0F17E1" author="Kakui" box="[1041,1242,1196,1221]" pageId="9" refString="Kakui, K. Katoh, T., Hiruta, S. F., Kobayashi, N. &amp; Kajihara, H. (2011) Molecular systematics of Tanaidacea (Crustacea: Peracarida) based on 18 S sequence data, with an amendment of suborder / superfamily-level classification. Zoological Science, 28, 749 - 757. https: // doi. org / 10.2108 / zsj. 28.749" type="journal article" year="2011">
Kakui 
<emphasis id="B918EAF74929AE70FBBA23B3FB4517E0" box="[1118,1168,1196,1221]" italics="true" pageId="9">et al</emphasis>
. 2011
</bibRefCitation>
), and 
<taxonomicName id="4C6C4D664929AE70FAFB23B3FE4217CD" authority="Larsen, 2001" authorityName="Larsen" authorityYear="2001" class="Malacostraca" family="Paratanaidae" genus="Paratanais" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="species" species="malignus">
<emphasis id="B918EAF74929AE70FAFB23B3FF2B17CD" italics="true" pageId="9">Paratanais malignus</emphasis>
<bibRefCitation id="EFFD4B144929AE70FEE023CFFE4217CD" author="Larsen" box="[260,407,1232,1256]" pageId="9" refString="Larsen, K. (2001) Morphological and molecular investigation of polymorphism and cryptic species in tanaid crustaceans: implications for tanaid systematics and biodiversity estimates. Zoological Journal of the Linnean Society, 131, 353 - 379. https: // doi. org / 10.1111 / j. 1096 - 3642.2001. tb 02241. x" type="journal article" year="2001">Larsen, 2001</bibRefCitation>
</taxonomicName>
(IS 98%; QC 66%; 
<taxonomicName id="4C6C4D664929AE70FD9D23D1FCD917CD" authorityName="Lang" authorityYear="1949" box="[633,780,1231,1256]" class="Malacostraca" family="Paratanaidae" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="family">Paratanaidae</taxonomicName>
; 
<bibRefCitation id="EFFD4B144929AE70FCF323CEFC3117CD" author="Spears" box="[791,996,1231,1256]" pageId="9" refString="Spears, T., DeBry, R. W., Abele, L. G. &amp; Chodyla, K. (2005) Peracarid monophyly and interordinal phylogeny inferred from nuclear small-subunit ribosomal DNA sequences (Crustacea: Malacostraca: Peracarida). Proceedings of the Biological Society of Washington, 118, 117 - 157. https: // doi. org / 10.2988 / 0006 - 324 X (2005) 118 [117: PMAIPI] 2.0. CO; 2" type="journal article" year="2005">
Spears 
<emphasis id="B918EAF74929AE70FC8C23CFFC4F17CD" box="[872,922,1231,1256]" italics="true" pageId="9">et al</emphasis>
. 2005
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C376656E4929AE70FF2323EAFACE1629" box="[199,1307,1268,1293]" pageId="9" type="distribution">
<paragraph id="8BD336E54929AE70FF2323EAFACE1629" blockId="9.[151,1437,151,1617]" box="[199,1307,1268,1293]" pageId="9">
<emphasis id="B918EAF74929AE70FF2323EAFEB51628" bold="true" box="[199,352,1268,1293]" pageId="9">Distribution.</emphasis>
So far known only from three localities around 
<collectingRegion id="49A8F8074929AE70FC9F23EAFC3C1628" box="[891,1001,1268,1293]" country="Japan" name="Hokkaido" pageId="9">Hokkaido</collectingRegion>
Island, at depths of 
<quantity id="4C949B004929AE70FB2823EAFAC21629" box="[1228,1303,1268,1292]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="3.0" metricValueMin="0.0" pageId="9" unit="m" value="1.5" valueMax="3.0" valueMin="0.0">
0– 
<specimenCount id="9D6AFD6C4929AE70FB0E23EAFAC21629" box="[1258,1303,1268,1292]" pageId="9" type="generic">3 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C376656E4929AE73FF232209FB751031" lastPageId="10" pageId="9" type="discussion">
<paragraph id="8BD336E54929AE70FF232209FD7C1575" blockId="9.[151,1437,151,1617]" pageId="9">
<emphasis id="B918EAF74929AE70FF232209FEEE1615" bold="true" box="[199,315,1303,1328]" pageId="9">Remarks.</emphasis>
<taxonomicName id="4C6C4D664929AE70FEA52207FDCE1615" authority="Shimada, 2017" authorityName="Shimada" authorityYear="2017" box="[321,539,1305,1328]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="B918EAF74929AE70FEA52207FDCE1615" box="[321,539,1305,1328]" italics="true" pageId="9">Tanaopsis japonica</emphasis>
</taxonomicName>
<emphasis id="B918EAF74929AE70FDC62209FDAE1615" bold="true" box="[546,635,1303,1328]" pageId="9">
<taxonomicNameLabel id="A22B578C4929AE70FDC62209FDAE1615" box="[546,635,1303,1328]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the seventeenth species described in this genus and family. In having the uropod with the endopod biarticulate and the exopod uniarticulate, this species closely resembles 
<taxonomicName id="4C6C4D664929AE70FA632223FDC8165D" authority="Bird &amp; Bamber, 2000" authorityName="Bird &amp; Bamber" authorityYear="2000" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="species" species="chotkarakde">
<emphasis id="B918EAF74929AE70FA632223FEF7165D" italics="true" pageId="9">T. chotkarakde</emphasis>
<bibRefCitation id="EFFD4B144929AE70FECD2241FDC8165D" author="Bird" box="[297,541,1375,1400]" pageId="9" refString="Bird, G. J. &amp; Bamber, R. N. (2000) Additions to the tanaidomorph Tanaidacea (Crustacea: Peracarida) of Hong Kong. In: Morton, B. (Ed.), The Marine Flora and Fauna of Hong Kong and Southern China. Vol. V. Hong Kong University Press, Hong Kong, pp. 65 - 104." type="book chapter" year="2000">Bird &amp; Bamber, 2000</bibRefCitation>
</taxonomicName>
from around 
<collectingCountry id="F37B76754929AE70FD5D227FFCE9165C" box="[697,828,1377,1401]" name="Hong Kong" pageId="9">Hong Kong</collectingCountry>
and 
<taxonomicName id="4C6C4D664929AE70FC91227FFBBA165D" authority="Bird, 2011" authorityName="Bird" authorityYear="2011" box="[885,1135,1375,1400]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="species" species="rawhitia">
<emphasis id="B918EAF74929AE70FC91227FFC25165D" box="[885,1008,1375,1400]" italics="true" pageId="9">T. rawhitia</emphasis>
<bibRefCitation id="EFFD4B144929AE70FC132241FBBA165D" author="Bird" box="[1015,1135,1375,1400]" pageId="9" refString="Bird, G. J. (2011) Paratanaoidean tanaidaceans (Crustacea: Peracarida) from littoral and shallow sublittoral habitats in New Zealand, with descriptions of three new genera and seven new species. Zootaxa, 2891, 1 - 62." type="journal article" year="2011">Bird, 2011</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F37B76754929AE70FB50227FFA9E165D" box="[1204,1355,1375,1400]" name="New Zealand" pageId="9">New Zealand</collectingCountry>
waters (
<bibRefCitation id="EFFD4B144929AE70FF7A229AFE4F16B9" author="Bird" box="[158,410,1412,1437]" pageId="9" refString="Bird, G. J. &amp; Bamber, R. N. (2000) Additions to the tanaidomorph Tanaidacea (Crustacea: Peracarida) of Hong Kong. In: Morton, B. (Ed.), The Marine Flora and Fauna of Hong Kong and Southern China. Vol. V. Hong Kong University Press, Hong Kong, pp. 65 - 104." type="book chapter" year="2000">Bird &amp; Bamber 2000</bibRefCitation>
; 
<bibRefCitation id="EFFD4B144929AE70FE4E229AFDF416B9" author="Bird" box="[426,545,1412,1437]" pageId="9" refString="Bird, G. J. (2011) Paratanaoidean tanaidaceans (Crustacea: Peracarida) from littoral and shallow sublittoral habitats in New Zealand, with descriptions of three new genera and seven new species. Zootaxa, 2891, 1 - 62." type="journal article" year="2011">Bird 2011</bibRefCitation>
). Female 
<taxonomicName id="4C6C4D664929AE70FD70229BFCC016B9" authorityName="Shimada" authorityYear="2017" box="[660,789,1413,1436]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B918EAF74929AE70FD70229BFCC016B9" box="[660,789,1413,1436]" italics="true" pageId="9">T. japonica</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C6C4D664929AE70FC57229BFB8816B8" authorityName="Bird &amp; Bamber" authorityYear="2000" box="[947,1117,1412,1437]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="species" species="chotkarakde">
<emphasis id="B918EAF74929AE70FC57229BFB8816B8" box="[947,1117,1412,1437]" italics="true" pageId="9">T. chotkarakde</emphasis>
</taxonomicName>
in the following characters (character state of 
<taxonomicName id="4C6C4D664929AE70FE8E22B7FDC416E5" authorityName="Bird &amp; Bamber" authorityYear="2000" box="[362,529,1447,1472]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="species" species="chotkarakde">
<emphasis id="B918EAF74929AE70FE8E22B7FDC416E5" box="[362,529,1447,1472]" italics="true" pageId="9">T. chotkarakde</emphasis>
</taxonomicName>
in parentheses): the length ratio of pereonites 4/5 is 1.0 (1.2); antennal article 4 has three simple distal setae (two); the propodal palm of the cheliped has one longer and three shorter inner simple setae at the insertion of the dactylus (one longer and two shorter setae); the chelipedal dactylus has three spiniform setae on the cutting surface (one); and the pleopodal basis is naked (one outer seta) (
<bibRefCitation id="EFFD4B144929AE70FBAB210AFA901509" author="Bird" box="[1103,1349,1556,1581]" pageId="9" refString="Bird, G. J. &amp; Bamber, R. N. (2000) Additions to the tanaidomorph Tanaidacea (Crustacea: Peracarida) of Hong Kong. In: Morton, B. (Ed.), The Marine Flora and Fauna of Hong Kong and Southern China. Vol. V. Hong Kong University Press, Hong Kong, pp. 65 - 104." type="book chapter" year="2000">Bird &amp; Bamber 2000</bibRefCitation>
). Males have not yet been described for 
<taxonomicName id="4C6C4D664929AE70FE182127FD771575" authorityName="Bird &amp; Bamber" authorityYear="2000" box="[508,674,1591,1616]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="species" species="chotkarakde">
<emphasis id="B918EAF74929AE70FE182127FD771575" box="[508,674,1591,1616]" italics="true" pageId="9">T. chotkarakde</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF13666D4929AE70FF7320A1FBD114F0" box="[151,1028,1983,2006]" pageId="9">
<paragraph id="8BD336E54929AE70FF7320A1FBD114F0" blockId="9.[151,1028,1983,2006]" box="[151,1028,1983,2006]" pageId="9">
<emphasis id="B918EAF74929AE70FF7320A1FEC714F1" bold="true" box="[151,274,1983,2005]" pageId="9">FIGURE 8.</emphasis>
Living female 
<taxonomicName id="4C6C4D664929AE70FE4A20DFFDA614F3" authority="Shimada, 2017" authorityName="Shimada" authorityYear="2017" box="[430,627,1985,2006]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="9" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="B918EAF74929AE70FE4A20DFFDA614F3" box="[430,627,1985,2006]" italics="true" pageId="9">Tanaopsis japonica</emphasis>
</taxonomicName>
<emphasis id="B918EAF74929AE70FD9D20DEFD1C14F0" bold="true" box="[633,713,1984,2005]" pageId="9">
<taxonomicNameLabel id="A22B578C4929AE70FD9D20DEFD1C14F0" box="[633,713,1984,2005]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(paratype). Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BD336E5492AAE73FF232787FA5812AD" blockId="10.[151,1437,151,789]" pageId="10">
<taxonomicName id="4C6C4D66492AAE73FF232787FE761395" authorityName="Shimada" authorityYear="2017" box="[199,419,153,176]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B918EAF7492AAE73FF232787FE761395" box="[199,419,153,176]" italics="true" pageId="10">Tanaopsis japonica</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C6C4D66492AAE73FDDB2787FD681395" authorityName="Bird" authorityYear="2011" box="[575,701,151,176]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="species" species="rawhitia">
<emphasis id="B918EAF7492AAE73FDDB2787FD681395" box="[575,701,151,176]" italics="true" pageId="10">T. rawhitia</emphasis>
</taxonomicName>
as follows (character state of the latter in parentheses). In both sexes, 
<taxonomicName id="4C6C4D66492AAE73FF0027A3FEB013F1" authorityName="Shimada" authorityYear="2017" box="[228,357,189,212]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B918EAF7492AAE73FF0027A3FEB013F1" box="[228,357,189,212]" italics="true" pageId="10">T. japonica</emphasis>
</taxonomicName>
has the dactylus of pereopods 1–3 with a dorsoproximal simple seta (naked). In females, the mandible lacks a molar (point-tipped molar present); pereonites 4 and 5 are all wider than long (as wide as long), antennal article 2 lacks outer setae (one outer seta), and the pleopodal endopod has six outer setae in total (eight or nine). In males, 
<taxonomicName id="4C6C4D66492AAE73FEAA2637FE181265" authorityName="Shimada" authorityYear="2017" box="[334,461,297,320]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B918EAF7492AAE73FEAA2637FE181265" box="[334,461,297,320]" italics="true" pageId="10">T. japonica</emphasis>
</taxonomicName>
has the antennule with six articles bearing two rows of aesthetascs (seven articles and three rows), the propodal palm of the cheliped has one longer and six shorter inner simple setae at the insertion of the dactylus (two longer and 11 shorter setae), and the uropodal exopod is uniarticulate (biarticulate) (
<bibRefCitation id="EFFD4B14492AAE73FAEF2671FA5412AD" author="Bird" box="[1291,1409,367,392]" pageId="10" refString="Bird, G. J. (2011) Paratanaoidean tanaidaceans (Crustacea: Peracarida) from littoral and shallow sublittoral habitats in New Zealand, with descriptions of three new genera and seven new species. Zootaxa, 2891, 1 - 62." type="journal article" year="2011">Bird 2011</bibRefCitation>
).
</paragraph>
<paragraph id="8BD336E5492AAE73FF23268AFAF31145" blockId="10.[151,1437,151,789]" pageId="10">
Our male specimen of 
<taxonomicName id="4C6C4D66492AAE73FE20268BFD971289" authorityName="Shimada" authorityYear="2017" box="[452,578,405,428]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B918EAF7492AAE73FE20268BFD971289" box="[452,578,405,428]" italics="true" pageId="10">T. japonica</emphasis>
</taxonomicName>
is unique within 
<taxonomicName id="4C6C4D66492AAE73FCE7268AFC5A1288" authorityName="Blazewicz-Paszkowycz &amp; Bamber" authorityYear="2012" box="[771,911,404,429]" class="Malacostraca" family="Tanaopsidae" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="family">Tanaopsidae</taxonomicName>
in that its antennule consists of six articles and bears two rows of aesthetascs. Including male 
<taxonomicName id="4C6C4D66492AAE73FD4526A7FB4B12F5" authority="(Sars, 1882) sensu Sars (1886)" authorityName="(Sars, 1882) sensu Sars" authorityYear="1886" baseAuthorityName="Sars" baseAuthorityYear="1882" box="[673,1182,439,464]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="species" species="laticaudata">
<emphasis id="B918EAF7492AAE73FD4526A7FCE812F5" box="[673,829,439,464]" italics="true" pageId="10">T. laticaudata</emphasis>
(
<bibRefCitation id="EFFD4B14492AAE73FCA826A6FC1012F5" author="Sars" box="[844,965,440,464]" pageId="10" refString="Sars, G. O. (1882) Revision af gruppen: Isopoda chelifera med charakteristik af nye herhen horende arter og slaegter. Archiv for Mathematik og Naturvidenskab, 7, 1 - 54." type="journal article" year="1882">Sars, 1882</bibRefCitation>
) 
<emphasis id="B918EAF7492AAE73FC3026A7FBC712F5" box="[980,1042,441,464]" italics="true" pageId="10">sensu</emphasis>
<bibRefCitation id="EFFD4B14492AAE73FBFE26A6FB4B12F5" author="Sars" box="[1050,1182,439,464]" pageId="10" refString="Sars, G. O. (1886) Nye bidrag til kundskaben om Middelhavets invertebratfauna. III. Middelhavets Saxisopoder (Isopoda chelifera). Archiv for Mathematik og Naturvidenskab, 11, 263 - 368, 15 pls." type="journal article" year="1886">Sars (1886)</bibRefCitation>
</taxonomicName>
, which 
<bibRefCitation id="EFFD4B14492AAE73FB1226A6FF1F12D0" author="Lang" pageId="10" refString="Lang, K. (1967) Taxonomische und phylogenetische Untersuchungen uber die Tanaidaceen. 3. Der Umfang der Familien Tanaidae Sars, Lang und Paratanaidae Lang nebst Bemerkungen uber den taxonomischen Wert der Mandibeln und Maxillulae. Dazu eine taxonomisch-monographische Darstellung der Gattung Tanaopsis Sars. Arkiv for Zoologi, 19, 343 - 368, 1 pl." type="journal article" year="1967" yearSuffix="p">Lang (1967: p. 350)</bibRefCitation>
suggested might have been misidentified as to genus, males have previously been reported in nine congeners, all of which have seven-articulate antennule with three rows of aesthetascs (
<bibRefCitation id="EFFD4B14492AAE73FC19251EFBA6113D" author="Sars" box="[1021,1139,511,536]" pageId="10" refString="Sars, G. O. (1886) Nye bidrag til kundskaben om Middelhavets invertebratfauna. III. Middelhavets Saxisopoder (Isopoda chelifera). Archiv for Mathematik og Naturvidenskab, 11, 263 - 368, 15 pls." type="journal article" year="1886">Sars 1886</bibRefCitation>
; 
<bibRefCitation id="EFFD4B14492AAE73FB64251EFB28113D" author="Lang" box="[1152,1277,511,536]" pageId="10" refString="Lang, K. (1967) Taxonomische und phylogenetische Untersuchungen uber die Tanaidaceen. 3. Der Umfang der Familien Tanaidae Sars, Lang und Paratanaidae Lang nebst Bemerkungen uber den taxonomischen Wert der Mandibeln und Maxillulae. Dazu eine taxonomisch-monographische Darstellung der Gattung Tanaopsis Sars. Arkiv for Zoologi, 19, 343 - 368, 1 pl." type="journal article" year="1967">Lang 1967</bibRefCitation>
; 
<bibRefCitation id="EFFD4B14492AAE73FAEE26E1FA42113D" author="Shiino" box="[1290,1431,511,536]" pageId="10" refString="Shiino, S. M. (1970) Paratanaidae collected in Chile Bay, Greenwich Island by the XXII Chilean Antarctic Expedition, with an Apseudes from Porvenir Point, Tierra del Fuego Island. Instituto Antartico Chileno Serie Cientifica, 1, 77 - 122." type="journal article" year="1970">Shiino 1970</bibRefCitation>
; 
<bibRefCitation id="EFFD4B14492AAE73FF73253AFE6B1119" author="Kudinova-Pasternak" box="[151,446,548,573]" pageId="10" refString="Kudinova-Pasternak, R. K. (1984) The Tanaidacea (Crustacea: Malacostraca) of the Sea of Japan. Zoologicheskii Zhurnal, 63, 828 - 838. [in Russian with English abstract]" type="journal article" year="1984">Kudinova-Pasternak 1984</bibRefCitation>
; 
<bibRefCitation id="EFFD4B14492AAE73FE2D253AFD761119" author="Sieg" box="[457,675,548,573]" pageId="10" refString="Sieg, J. &amp; Dojiri, M. (1991) Two new species and a new genus of the suborder Tanaidomorpha (Crustacea: Tanaidacea) from Californian waters. Journal of Natural History, 25, 1493 - 1512. https: // doi. org / 10.1080 / 00222939100770951" type="journal article" year="1991">Sieg &amp; Dojiri 1991</bibRefCitation>
; 
<bibRefCitation id="EFFD4B14492AAE73FD4B253AFCF71119" author="Bird" box="[687,802,548,573]" pageId="10" refString="Bird, G. J. (2011) Paratanaoidean tanaidaceans (Crustacea: Peracarida) from littoral and shallow sublittoral habitats in New Zealand, with descriptions of three new genera and seven new species. Zootaxa, 2891, 1 - 62." type="journal article" year="2011">Bird 2011</bibRefCitation>
; 
<bibRefCitation id="EFFD4B14492AAE73FCCA253AFB401118" author="Segadilha" box="[814,1173,548,573]" pageId="10" refString="Segadilha, J. L. &amp; Araujo-Silva, C. L. (2015) Two new species of Tanaopsis (Tanaidacea: Tanaopsidae) from Admiralty Bay (Antarctica), with an identification key. Nauplius, 23, 31 - 45. https: // doi. org / 10.1590 / S 0104 - 64972015002315" type="journal article" year="2015">Segadilha &amp; Araújo-Silva 2015</bibRefCitation>
). To determine whether the six-articulate antennule is restricted to 
<taxonomicName id="4C6C4D66492AAE73FD972557FD251145" authorityName="Shimada" authorityYear="2017" box="[627,752,585,608]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B918EAF7492AAE73FD972557FD251145" box="[627,752,585,608]" italics="true" pageId="10">T. japonica</emphasis>
</taxonomicName>
, males must be examined from additional species.
</paragraph>
<paragraph id="8BD336E5492AAE73FF232572FB751031" blockId="10.[151,1437,151,789]" pageId="10">
Among congeners, 
<taxonomicName id="4C6C4D66492AAE73FE402573FD2B11A1" authority="Sieg &amp; Dojiri, 1991" authorityName="Sieg &amp; Dojiri" authorityYear="1991" box="[420,766,620,645]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="species" species="cadieni">
<emphasis id="B918EAF7492AAE73FE402573FDC411A0" box="[420,529,620,645]" italics="true" pageId="10">T. cadieni</emphasis>
<bibRefCitation id="EFFD4B14492AAE73FDF82572FD2B11A1" author="Sieg" box="[540,766,620,645]" pageId="10" refString="Sieg, J. &amp; Dojiri, M. (1991) Two new species and a new genus of the suborder Tanaidomorpha (Crustacea: Tanaidacea) from Californian waters. Journal of Natural History, 25, 1493 - 1512. https: // doi. org / 10.1080 / 00222939100770951" type="journal article" year="1991">Sieg &amp; Dojiri, 1991</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C6C4D66492AAE73FCD32573FBB211A1" authority="(Shiino, 1970)" authorityName="Shiino" authorityYear="1970" baseAuthorityName="Shiino" baseAuthorityYear="1970" box="[823,1127,620,645]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="species" species="gallardoi">
<emphasis id="B918EAF7492AAE73FCD32573FC6E11A0" box="[823,955,620,645]" italics="true" pageId="10">T. gallardoi</emphasis>
(
<bibRefCitation id="EFFD4B14492AAE73FC282572FB8A11A1" author="Shiino" box="[972,1119,620,645]" pageId="10" refString="Shiino, S. M. (1970) Paratanaidae collected in Chile Bay, Greenwich Island by the XXII Chilean Antarctic Expedition, with an Apseudes from Porvenir Point, Tierra del Fuego Island. Instituto Antartico Chileno Serie Cientifica, 1, 77 - 122." type="journal article" year="1970">Shiino, 1970</bibRefCitation>
)
</taxonomicName>
include descriptions of the male mouthparts. Although the maxillipedal endites are flared in these species, they are not flared in male 
<taxonomicName id="4C6C4D66492AAE73FA63258FFF2C11E9" authorityName="Shimada" authorityYear="2017" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B918EAF7492AAE73FA63258FFF2C11E9" italics="true" pageId="10">T. japonica</emphasis>
</taxonomicName>
. While 
<bibRefCitation id="EFFD4B14492AAE73FEB025AAFE2511E9" author="Shiino" box="[340,496,692,717]" pageId="10" refString="Shiino, S. M. (1970) Paratanaidae collected in Chile Bay, Greenwich Island by the XXII Chilean Antarctic Expedition, with an Apseudes from Porvenir Point, Tierra del Fuego Island. Instituto Antartico Chileno Serie Cientifica, 1, 77 - 122." type="journal article" year="1970">Shiino (1970)</bibRefCitation>
observed mandibles in male 
<taxonomicName id="4C6C4D66492AAE73FCD825ABFC6A11E8" baseAuthorityName="Shiino" baseAuthorityYear="1970" box="[828,959,692,717]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="species" species="gallardoi">
<emphasis id="B918EAF7492AAE73FCD825ABFC6A11E8" box="[828,959,692,717]" italics="true" pageId="10">T. gallardoi</emphasis>
</taxonomicName>
, we found only a simple opening in place of the mandibles (
<figureCitation id="13572A60492AAE73FE8F25C9FE1211D5" box="[363,455,727,752]" captionStart="FIGURE 4" captionStartId="5.[151,250,1827,1849]" captionTargetBox="[158,1428,481,1806]" captionTargetId="graphics@5.[158,1428,481,1806]" captionTargetPageId="5" captionText="FIGURE 4. Tanaopsis japonica sp. nov., allotype male: A, right antennule; B, right antenna; C, anterior region of cephalothorax, left view; D, labium, ventral view; E, maxillipeds; F, right cheliped, outer view; f 1, same, distal region of fixed finger, outer view; G, left cheliped, inner view. Scale bars: 0.1 mm for A – D, F, G; 0.05 mm for E, f 1." httpUri="https://zenodo.org/record/1010386/files/figure.svg" pageId="10">Fig. 4C</figureCitation>
). This suggests that, as observed in 
<taxonomicName id="4C6C4D66492AAE73FC8B25C9FBD111D5" authorityName="Sieg &amp; Heard" authorityYear="1985" box="[879,1028,727,752]" class="Malacostraca" family="Nototanaidae" genus="Nototanoides" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAF7492AAE73FC8B25C9FBD111D5" box="[879,1028,727,752]" italics="true" pageId="10">Nototanoides</emphasis>
</taxonomicName>
(
<taxonomicName id="4C6C4D66492AAE73FBF325C9FB6711D5" authorityName="Sieg" authorityYear="1976" box="[1047,1202,727,752]" class="Malacostraca" family="Nototanaidae" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="family">Nototanaidae</taxonomicName>
; 
<bibRefCitation id="EFFD4B14492AAE73FB2425C9FF061031" author="Kakui" pageId="10" refString="Kakui, K. &amp; Yamasaki, H. (2013) Nototanaids (Crustacea: Tanaidacea) from Japan, with the description of a new species of Nototanoides. Species Diversity, 18, 245 - 254. https: // doi. org / 10.12782 / sd. 18.2.245" type="journal article" year="2013">Kakui &amp; Yamasaki 2013</bibRefCitation>
), the degree of reduction of the male mouthparts can vary among species in 
<taxonomicName id="4C6C4D66492AAE73FBCC25E3FB4C1031" authorityName="Shimada" authorityYear="2017" box="[1064,1177,765,788]" class="Malacostraca" family="Tanaopsidae" genus="Tanaopsis" kingdom="Animalia" order="Tanaidacea" pageId="10" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAF7492AAE73FBCC25E3FB4C1031" box="[1064,1177,765,788]" italics="true" pageId="10">Tanaopsis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>