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<document id="6FD1C14088A1F84467CAC207B6D3D088" ID-DOI="10.5281/zenodo.277061" ID-GBIF-Dataset="f298d254-83b1-43bb-9f8d-a4a7d984ebcc" ID-ISSN="1175-5326" ID-Zenodo-Dep="277061" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460268674169" checkinUser="plazi" docAuthor="Magalhães, Wagner F., Bailey, Julie H., Brock, - &amp; Davenport, Jennifer S." docDate="2011" docId="03C587D32247FFFEE4D32E7C766CFB68" docLanguage="en" docName="zt02804p014.pdf" docOrigin="Zootaxa 2804" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Raphidrilus harperi Magalhães, Bailey, Brock &amp; Davenport, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="8" masterDocId="FFFCFFAB2243FFF9E4442C10776EFFC0" masterDocTitle="On the genus Raphidrilus Monticelli, 1910 (Polychaeta: Ctenodrilidae) with description of two new species" masterLastPageNumber="14" masterPageNumber="1" pageNumber="5" updateTime="1698246225558" updateUser="plazi">
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<mods:title id="8EF5DF6010F0F0A297904EAB18F69BE4">On the genus Raphidrilus Monticelli, 1910 (Polychaeta: Ctenodrilidae) with description of two new species</mods:title>
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<mods:namePart id="C4687455F89D0B748747BE23BE1C6A1B">Magalhães, Wagner F.</mods:namePart>
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<mods:namePart id="2FB357048E259449B661F663D68504F2">Bailey, Julie H.</mods:namePart>
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<mods:namePart id="917CD3E76BDF7CF73B70B3A3E33B4062">Brock, -</mods:namePart>
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<mods:namePart id="0597E00FDECF3E34C4826F13D82ABEE7">Davenport, Jennifer S.</mods:namePart>
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<mods:date id="7E2021E304ED1917472670AAB6393A3B">2011</mods:date>
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<treatment id="03C587D32247FFFEE4D32E7C766CFB68" ID-DOI="http://doi.org/10.5281/zenodo.5631414" ID-GBIF-Taxon="119394606" ID-Zenodo-Dep="5631414" LSID="urn:lsid:plazi:treatment:03C587D32247FFFEE4D32E7C766CFB68" httpUri="http://treatment.plazi.org/id/03C587D32247FFFEE4D32E7C766CFB68" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5">
<subSubSection id="C376654E2247FFFDE4D32E7C76B0FD66" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BD336C52247FFFDE4D32E7C7682FD45" blockId="4.[151,492,619,678]" box="[151,492,619,646]" pageId="4" pageNumber="5">
<heading id="D09B81A92247FFFDE4D32E7C7682FD45" bold="true" box="[151,492,619,646]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="B918EAD72247FFFDE4D32E7C7682FD45" bold="true" box="[151,492,619,646]" pageId="4" pageNumber="5">
<taxonomicName id="4C6C4D462247FFFDE4D32E7C76E8FD46" box="[151,390,620,646]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72247FFFDE4D32E7C76E8FD46" bold="true" box="[151,390,620,646]" italics="true" pageId="4" pageNumber="5">Raphidrilus harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2247FFFDE5CA2E7B7682FD45" box="[398,492,619,645]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD336C52247FFFDE4D32E9E76B0FD66" blockId="4.[151,492,619,678]" box="[151,478,654,678]" pageId="4" pageNumber="5">
<figureCitation id="13572A402247FFFDE4D32E9E766EFD66" box="[151,256,654,678]" captionStart="FIGURE 2" captionStartId="5.[151,250,1584,1607]" captionTargetBox="[151,1436,443,1563]" captionTargetId="figure@5.[151,1436,443,1563]" captionTargetPageId="5" captionText="FIGURE 2. Raphidrilus harperi sp. nov. A, entire specimen in lateral view; B, mid-body segments with a segmented worm in the coelom; C, light microscopy of mid-body segments showing a segmented worm protruding from the coelom." httpUri="https://zenodo.org/record/277063/files/figure.png" pageId="4" pageNumber="5">Figures 2</figureCitation>
(AC) and 3 (AF)
</paragraph>
</subSubSection>
<subSubSection id="C376654E2247FFFDE4D32EC6724CFCBE" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BD336C52247FFFDE4D32EC6724CFCBE" blockId="4.[151,1437,726,1975]" pageId="4" pageNumber="5">
<emphasis id="B918EAD72247FFFDE4D32EC6756AFD2F" bold="true" box="[151,516,726,751]" pageId="4" pageNumber="5">
Material examined.
<typeStatus id="54D788672247FFFDE5CA2EC77691FD2F" box="[398,511,727,751]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED7DAE002247FFFDE64B2EC6750FFD2E" box="[527,609,726,750]" pageId="4" pageNumber="5">GIWW</collectionCode>
at Venice, Florida,
<collectingCountry id="F37B76552247FFFDE70E2EC674EBFD2E" box="[842,901,726,750]" name="United States of America" pageId="4" pageNumber="5">USA</collectingCountry>
,
<geoCoordinate id="EE5850022247FFFDE7D02EC67353FD2E" box="[916,1085,726,750]" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="27.10036">27º0601.3” N</geoCoordinate>
,
<geoCoordinate id="EE5850022247FFFDE0082EC67397FD2E" box="[1100,1273,726,750]" direction="west" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="-82.43569">82º2608.5” W</geoCoordinate>
, Station 5NB, coll. D. Seagle,
<date id="FFD210052247FFFDE50D2EEB756DFCD2" box="[329,515,762,787]" pageId="4" pageNumber="5" value="2009-08-28">August 28, 2009</date>
(
<collectionCode id="ED7DAE002247FFFDE6552EEA750AFCD2" LSID="urn:lsid:biocol.org:col:34871" box="[529,612,762,786]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5">USNM</collectionCode>
1150467).
<emphasis id="B918EAD72247FFFDE6A72EEB740AFCD3" bold="true" box="[739,868,763,787]" pageId="4" pageNumber="5">
<typeStatus id="54D788672247FFFDE6A72EEB7431FCD3" box="[739,863,763,787]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
:
</emphasis>
same location, date and collector as
<typeStatus id="54D788672247FFFDE0BB2EEA720AFCD2" box="[1279,1380,762,786]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
, Station 5NB (9,
<collectionCode id="ED7DAE002247FFFDE56B2F0E76EEFCF6" LSID="urn:lsid:biocol.org:col:34871" box="[303,384,798,822]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5">USNM</collectionCode>
1150469); Stations 5NB and 5NC (4,
<collectionCode id="ED7DAE002247FFFDE7732F0E7412FCF6" box="[823,892,798,822]" pageId="4" pageNumber="5">FSBC</collectionCode>
I
<date id="FFD210052247FFFDE7D22F0E7481FCF6" box="[918,1007,798,822]" pageId="4" pageNumber="5" value="50-07-22">072250</date>
); Station 5NB (4,
<collectionCode id="ED7DAE002247FFFDE0FA2F0E724BFCF6" LSID="urn:lsid:biocol.org:col:34986" box="[1214,1317,798,822]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34986" name="Museum and Art Gallery of the Northern Territory" pageId="4" pageNumber="5">MAGNT</collectionCode>
W23467 W23470); Station 5NB (1 mounted on stub,
<collectionCode id="ED7DAE002247FFFDE6D42F53758AFC9A" box="[656,740,835,858]" pageId="4" pageNumber="5">BMNH</collectionCode>
2011.7); Station 5NB (2 mounted on stub,
<collectionCode id="ED7DAE002247FFFDE0962F52724DFC9A" LSID="urn:lsid:biocol.org:col:34871" box="[1234,1315,834,858]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5">USNM</collectionCode>
1150468).
<emphasis id="B918EAD72247FFFDE4D32F777611FCBF" bold="true" box="[151,383,870,895]" pageId="4" pageNumber="5">
Non
<typeStatus id="54D788672247FFFDE4932F777664FCBF" box="[215,266,871,895]" pageId="4" pageNumber="5">type</typeStatus>
material:
</emphasis>
same location, date and collector as
<typeStatus id="54D788672247FFFDE75F2F767422FCBE" box="[795,844,870,894]" pageId="4" pageNumber="5">type</typeStatus>
series, Station 5NB (3); Station 5NC (1).
</paragraph>
</subSubSection>
<subSubSection id="C376654E2247FFFDE4832F9A73BCFA22" pageId="4" pageNumber="5" type="description">
<paragraph id="8BD336C52247FFFDE4832F9A7788FBCE" blockId="4.[151,1437,726,1975]" pageId="4" pageNumber="5">
<emphasis id="B918EAD72247FFFDE4832F9A7636FC63" bold="true" box="[199,344,906,931]" pageId="4" pageNumber="5">Description.</emphasis>
Complete specimens ranged from
<quantity id="4C949B202247FFFDE6AF2F9A741BFC62" box="[747,885,906,930]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="5.8" metricValueMin="3.1" pageId="4" pageNumber="5" unit="mm" value="4.45" valueMax="5.8" valueMin="3.1">3.15.8 mm</quantity>
in length,
<quantity id="4C949B202247FFFDE7B72F9A73E4FC62" box="[1011,1162,906,930]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="4" pageNumber="5" unit="mm" value="0.175" valueMax="0.25" valueMin="0.1">0.10.25 mm</quantity>
in width and possessed between 1727 chaetigers. Body thin, cylindrical, and elongated. First four chaetigers (thorax) and last few wider than long; abdominal chaetigers vary from 1.54 times longer than wide (
<figureCitation id="13572A402247FFFDE78B2FC2737CFC2A" box="[975,1042,978,1002]" captionStart="FIGURE 2" captionStartId="5.[151,250,1584,1607]" captionTargetBox="[151,1436,443,1563]" captionTargetId="figure@5.[151,1436,443,1563]" captionTargetPageId="5" captionText="FIGURE 2. Raphidrilus harperi sp. nov. A, entire specimen in lateral view; B, mid-body segments with a segmented worm in the coelom; C, light microscopy of mid-body segments showing a segmented worm protruding from the coelom." httpUri="https://zenodo.org/record/277063/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
A). Color in alcohol light yellow to brown.
</paragraph>
<paragraph id="8BD336C52247FFFDE481280A7649FAEE" blockId="4.[151,1437,726,1975]" pageId="4" pageNumber="5">
Prostomium short, broadly rounded with pair of posterolateral nuchal organs (
<figureCitation id="13572A402247FFFDE00B280A73FDFBF2" box="[1103,1171,1050,1074]" captionStart="FIGURE 3" captionStartId="6.[151,250,1749,1772]" captionTargetBox="[151,1436,337,1726]" captionTargetId="figure@6.[151,1436,337,1728]" captionTargetPageId="6" captionText="FIGURE 3. SEM of Raphidrilus harperi sp. nov. showing: A, complete specimen; B, anterior chaetigers in lateral view with two branchial filaments; C, anterior segments in lateral view; D, nuchal organ; E, serrate capillaries; F, pygidium." httpUri="https://zenodo.org/record/277064/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
C). Nuchal organs oval ciliary patches (~7 µm) with long cilia, situated in a shallow depression (
<figureCitation id="13572A402247FFFDE785282E736DFB96" box="[961,1027,1086,1110]" captionStart="FIGURE 3" captionStartId="6.[151,250,1749,1772]" captionTargetBox="[151,1436,337,1726]" captionTargetId="figure@6.[151,1436,337,1728]" captionTargetPageId="6" captionText="FIGURE 3. SEM of Raphidrilus harperi sp. nov. showing: A, complete specimen; B, anterior chaetigers in lateral view with two branchial filaments; C, anterior segments in lateral view; D, nuchal organ; E, serrate capillaries; F, pygidium." httpUri="https://zenodo.org/record/277064/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
D). Distinction between prostomium and peristomium inconspicuous laterally; peristomium single achaetous segment, appearing pearshaped together with the prostomium; a single biannulated achaetous segment following the peristomium (
<figureCitation id="13572A402247FFFDE0EE28967381FB5E" box="[1194,1263,1158,1182]" captionStart="FIGURE 3" captionStartId="6.[151,250,1749,1772]" captionTargetBox="[151,1436,337,1726]" captionTargetId="figure@6.[151,1436,337,1728]" captionTargetPageId="6" captionText="FIGURE 3. SEM of Raphidrilus harperi sp. nov. showing: A, complete specimen; B, anterior chaetigers in lateral view with two branchial filaments; C, anterior segments in lateral view; D, nuchal organ; E, serrate capillaries; F, pygidium." httpUri="https://zenodo.org/record/277064/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
B). Heart body usually begins in chaetiger 4, but sometimes projects anteriorly into the posterior region of chaetiger 3; usually extends to chaetiger 7, but occasionally continues to chaetiger
<quantity id="4C949B202247FFFDE71628DE7413FB26" box="[850,893,1230,1254]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="4" pageNumber="5" unit="in" value="9.0">9 in</quantity>
the material examined (
<figureCitation id="13572A402247FFFDE0CF28DE73A1FB26" box="[1163,1231,1230,1254]" captionStart="FIGURE 2" captionStartId="5.[151,250,1584,1607]" captionTargetBox="[151,1436,443,1563]" captionTargetId="figure@5.[151,1436,443,1563]" captionTargetPageId="5" captionText="FIGURE 2. Raphidrilus harperi sp. nov. A, entire specimen in lateral view; B, mid-body segments with a segmented worm in the coelom; C, light microscopy of mid-body segments showing a segmented worm protruding from the coelom." httpUri="https://zenodo.org/record/277063/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
A). Branchial filaments posterodorsal to notochaetae, easily broken and occurring in pairs or singly from chaetigers 3 through 11 (
<figureCitation id="13572A402247FFFDE4DB2906778FFAEE" box="[159,225,1302,1326]" captionStart="FIGURE 3" captionStartId="6.[151,250,1749,1772]" captionTargetBox="[151,1436,337,1726]" captionTargetId="figure@6.[151,1436,337,1728]" captionTargetPageId="6" captionText="FIGURE 3. SEM of Raphidrilus harperi sp. nov. showing: A, complete specimen; B, anterior chaetigers in lateral view with two branchial filaments; C, anterior segments in lateral view; D, nuchal organ; E, serrate capillaries; F, pygidium." httpUri="https://zenodo.org/record/277064/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
A, B).
</paragraph>
<paragraph id="8BD336C52247FFFDE481292A73BDFA7E" blockId="4.[151,1437,726,1975]" pageId="4" pageNumber="5">
Anterior chaetigers with 36 serrated capillaries in both noto and neurochaetal fascicles; posterior chaetigers with 14 capillaries per fascicle. Serrations of some capillaries evident using phase contrast microscopy with oil immersion at
<date id="FFD210052247FFFDE57729927617FA5A" box="[307,377,1410,1434]" pageId="4" pageNumber="5" value="1000-10">1000x</date>
magnification; SEM revealed fibrils along capillary edge with distance between the insertion point of two capillary fibrils approximately the same as half the width of a single fibril (
<figureCitation id="13572A402247FFFDE03429B673DCFA7E" box="[1136,1202,1446,1470]" captionStart="FIGURE 3" captionStartId="6.[151,250,1749,1772]" captionTargetBox="[151,1436,337,1726]" captionTargetId="figure@6.[151,1436,337,1728]" captionTargetPageId="6" captionText="FIGURE 3. SEM of Raphidrilus harperi sp. nov. showing: A, complete specimen; B, anterior chaetigers in lateral view with two branchial filaments; C, anterior segments in lateral view; D, nuchal organ; E, serrate capillaries; F, pygidium." httpUri="https://zenodo.org/record/277064/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
E).
</paragraph>
<paragraph id="8BD336C52247FFFDE48129DA73BCFA22" blockId="4.[151,1437,726,1975]" box="[197,1234,1482,1507]" pageId="4" pageNumber="5">
Pygidium elongated segment with dorsal anal aperture; fields of cilia not observed (
<figureCitation id="13572A402247FFFDE02B29DA73DCFA22" box="[1135,1202,1482,1506]" captionStart="FIGURE 3" captionStartId="6.[151,250,1749,1772]" captionTargetBox="[151,1436,337,1726]" captionTargetId="figure@6.[151,1436,337,1728]" captionTargetPageId="6" captionText="FIGURE 3. SEM of Raphidrilus harperi sp. nov. showing: A, complete specimen; B, anterior chaetigers in lateral view with two branchial filaments; C, anterior segments in lateral view; D, nuchal organ; E, serrate capillaries; F, pygidium." httpUri="https://zenodo.org/record/277064/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
F).
</paragraph>
</subSubSection>
<subSubSection id="C376654E2247FFFDE48129FF732EF98F" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BD336C52247FFFDE48129FF732EF98F" blockId="4.[151,1437,726,1975]" pageId="4" pageNumber="5">
<emphasis id="B918EAD72247FFFDE48129FF7625F9C7" bold="true" box="[197,331,1519,1543]" pageId="4" pageNumber="5">Etymology.</emphasis>
This species is named in honor of the third authors graduate advisor, Dr. Donald E. Harper, Jr., Professor Emeritus of Texas A&amp;M University at Galveston. Dr. Harper graciously introduced me to the world of polychaetes and has provided valuable guidance and encouragement over the years.
</paragraph>
</subSubSection>
<subSubSection id="C376654E2247FFFDE4812A4A732FF877" pageId="4" pageNumber="5" type="biology_ecology">
<paragraph id="8BD336C52247FFFDE4812A4A732FF877" blockId="4.[151,1437,726,1975]" pageId="4" pageNumber="5">
<emphasis id="B918EAD72247FFFDE4812A4A764BF9B3" bold="true" box="[197,293,1626,1651]" pageId="4" pageNumber="5">Biology.</emphasis>
<taxonomicName id="4C6C4D462247FFFDE56F2A4B7564F9B3" box="[299,522,1627,1651]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72247FFFDE56F2A4B7564F9B3" box="[299,522,1627,1651]" italics="true" pageId="4" pageNumber="5">Raphidrilus harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2247FFFDE6552A4B7504F9B3" box="[529,618,1627,1651]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B918EAD72247FFFDE6552A4B7504F9B3" bold="true" box="[529,618,1627,1651]" pageId="4" pageNumber="5">sp. nov.</emphasis>
</taxonomicNameLabel>
, was collected just north of a reverse osmosis plant outfall. Water quality data is typical for a shallow, estuarine waterbody during late summer in southwest Florida (
<tableCitation id="C6EE037E2247FFFDE0D52A6E738BF956" box="[1169,1253,1662,1686]" captionStart="TABLE 1" captionStartId="5.[151,235,152,173]" captionTargetBox="[151,1437,232,367]" captionTargetPageId="5" captionText="TABLE 1. Bottom water quality parameters for the sampling station where Raphidrilus harperi sp. nov., and R. nemasoma were collected." httpUri="http://table.plazi.org/id/DF13664D2246FFFCE4D32C88765FFF0C" pageId="4" pageNumber="5" tableUuid="DF13664D2246FFFCE4D32C88765FFF0C">Table 1</tableCitation>
). All specimens were sexually immature and regenerating specimens were not observed. Segmented worms were observed, however, in the coelom of several specimens and were oriented in both directions along the anteriorposterior axis of the host (
<figureCitation id="13572A402247FFFDE5402AFA7624F8C2" box="[260,330,1770,1794]" captionStart="FIGURE 2" captionStartId="5.[151,250,1584,1607]" captionTargetBox="[151,1436,443,1563]" captionTargetId="figure@5.[151,1436,443,1563]" captionTargetPageId="5" captionText="FIGURE 2. Raphidrilus harperi sp. nov. A, entire specimen in lateral view; B, mid-body segments with a segmented worm in the coelom; C, light microscopy of mid-body segments showing a segmented worm protruding from the coelom." httpUri="https://zenodo.org/record/277063/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
B, C). Segmented worms were dissected out of the host and neither chaetae nor branchiae were observed. These segmented worms may be an intracoelomic parasite or gestational larvae resulting from sexual reproduction.
<bibRefCitation id="EFFD4B342247FFFDE5772B237505F88A" author="Petersen" box="[307,619,1842,1866]" pageId="4" pageNumber="5" refString="Petersen, M. E. &amp; George, J. D. (1991) A new species of Raricirrus from northern Europe, with notes on its biology and a discussion of the affinities of the genus (Polychaeta: Ctenodrilidae), Pp. 185 - 208 in M. E. Petersen &amp; J. B. Kirkegaard, eds., Systematics, biology and morphology of world Polychaeta. Proceedings of the 2 nd International Polychaete Conference, Copenhagen 1986. Ophelia Supplement 5." type="journal volume" year="1991">Petersen and George (1991)</bibRefCitation>
indicated protandric hermaphroditism with internal gestation for
<taxonomicName id="4C6C4D462247FFFDE1012B24778EF8AF" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="species" species="nemasoma">
<emphasis id="B918EAD72247FFFDE1012B24778EF8AF" italics="true" pageId="4" pageNumber="5">R. nemasoma</emphasis>
</taxonomicName>
. Additionally,
<bibRefCitation id="EFFD4B342247FFFDE5CE2B4675C9F8AE" author="Sokolow" box="[394,679,1878,1902]" pageId="4" pageNumber="5" refString="Sokolow, I. (1911 a) Uber eine neue Ctenodrilus art und ihre Vermehrung. Zeitschrift fur wissenschaftliche Zoologie, Leipzig, 97 (3), 546 - 603." type="journal article" year="1911" yearSuffix="a">Sokolow (1911a, fig. 77)</bibRefCitation>
illustrated juveniles emerging from the coelom of a parent, which appear quite similar to the segmented worms observed in
<taxonomicName id="4C6C4D462247FFFDE75F2B6C74E0F853" box="[795,910,1915,1939]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72247FFFDE75F2B6C74E0F853" box="[795,910,1915,1939]" italics="true" pageId="4" pageNumber="5">R. harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2247FFFDE7D12B6B7480F853" box="[917,1006,1915,1939]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B918EAD72247FFFDE7D12B6B7480F853" bold="true" box="[917,1006,1915,1939]" pageId="4" pageNumber="5">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="13572A402247FFFDE7B92B6A732EF852" box="[1021,1088,1914,1938]" captionStart="FIGURE 2" captionStartId="5.[151,250,1584,1607]" captionTargetBox="[151,1436,443,1563]" captionTargetId="figure@5.[151,1436,443,1563]" captionTargetPageId="5" captionText="FIGURE 2. Raphidrilus harperi sp. nov. A, entire specimen in lateral view; B, mid-body segments with a segmented worm in the coelom; C, light microscopy of mid-body segments showing a segmented worm protruding from the coelom." httpUri="https://zenodo.org/record/277063/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
AC). Future
<emphasis id="B918EAD72247FFFDE0982B6C7249F853" box="[1244,1319,1916,1939]" italics="true" pageId="4" pageNumber="5">in vivo</emphasis>
investigations would help resolve the unknown reproductive processes of
<taxonomicName id="4C6C4D462247FFFDE72A2BB0748FF877" box="[878,993,1951,1975]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72247FFFDE72A2BB0748FF877" box="[878,993,1951,1975]" italics="true" pageId="4" pageNumber="5">R. harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2247FFFDE7AC2B8F732FF877" box="[1000,1089,1951,1975]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B918EAD72247FFFDE7AC2B8F732FF877" bold="true" box="[1000,1089,1951,1975]" pageId="4" pageNumber="5">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<caption id="DF13664D2246FFFCE4D32C88765FFF0C" ID-Table-UUID="DF13664D2246FFFCE4D32C88765FFF0C" httpUri="http://table.plazi.org/id/DF13664D2246FFFCE4D32C88765FFF0C" pageId="5" pageNumber="6" targetBox="[151,1437,232,367]" targetIsTable="true" targetPageId="5">
<paragraph id="8BD336C52246FFFCE4D32C88765FFF0C" blockId="5.[151,1437,152,204]" pageId="5" pageNumber="6">
<emphasis id="B918EAD72246FFFCE4D32C887668FF6D" bold="true" box="[151,262,152,173]" pageId="5" pageNumber="6">TABLE 1.</emphasis>
Bottom water quality parameters for the sampling station where
<taxonomicName id="4C6C4D462246FFFCE7EA2C887314FF6E" box="[942,1146,152,174]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72246FFFCE7EA2C887314FF6E" box="[942,1146,152,174]" italics="true" pageId="5" pageNumber="6">Raphidrilus harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2246FFFCE0C72C8873BBFF6D" box="[1155,1237,152,173]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B918EAD72246FFFCE0C72C8873BBFF6D" bold="true" box="[1155,1237,152,173]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
, and
<taxonomicName id="4C6C4D462246FFFCE1552C8972F3FF6E" box="[1297,1437,153,174]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="nemasoma">
<emphasis id="B918EAD72246FFFCE1552C8972F3FF6E" box="[1297,1437,153,174]" italics="true" pageId="5" pageNumber="6">R. nemasoma</emphasis>
</taxonomicName>
were collected.
</paragraph>
</caption>
<paragraph id="8BD336C52246FFFCE4DB2CF87277FEAF" pageId="5" pageNumber="6">
<table id="F96CC46522460006E4D32CF872F3FEAF" box="[151,1437,232,367]" gridcols="7" gridrows="3" pageId="5" pageNumber="6">
<tr id="355C348722460006E4D32CF872F3FEDD" box="[151,1437,232,285]" gridrow="0" pageId="5" pageNumber="6">
<th id="768D5DFB22460006E4D32CF87636FEDD" box="[151,344,232,285]" gridcol="0" gridrow="0" pageId="5" pageNumber="6">Species</th>
<th id="768D5DFB22460006E5D62CF875CCFEDD" box="[402,674,232,285]" gridcol="1" gridrow="0" pageId="5" pageNumber="6">Depth (m) Water Temper- ature (ºC)</th>
<th id="768D5DFB22460006E6F92CF87462FEDD" box="[701,780,232,285]" gridcol="2" gridrow="0" pageId="5" pageNumber="6">Salinity (ppt)</th>
<th id="768D5DFB22460006E7742CF874A1FEDD" box="[816,975,232,285]" gridcol="3" gridrow="0" pageId="5" pageNumber="6">Dissolved Oxy- gen (mg/L)</th>
<th id="768D5DFB22460006E7AE2CF873F5FEDD" box="[1002,1179,232,285]" gridcol="4" gridrow="0" pageId="5" pageNumber="6">Specific Conduc- tivity (μmho/cm)</th>
<th id="768D5DFB22460006E0852CF8738FFEDD" box="[1217,1249,232,285]" gridcol="5" gridrow="0" pageId="5" pageNumber="6">pH</th>
<th id="768D5DFB22460006E1482CF872F3FEDD" box="[1292,1437,232,285]" gridcol="6" gridrow="0" pageId="5" pageNumber="6">Chlorides (g/ Kg)</th>
</tr>
<tr id="355C348722460006E4D32D2072F3FE86" box="[151,1437,304,326]" gridrow="1" pageId="5" pageNumber="6">
<th id="768D5DFB22460006E4D32D207636FE86" box="[151,344,304,326]" gridcol="0" gridrow="1" pageId="5" pageNumber="6">
<taxonomicName id="4C6C4D462246FFFCE4DB2D21766FFE86" box="[159,257,305,326]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72246FFFCE4DB2D21766FFE86" box="[159,257,305,326]" italics="true" pageId="5" pageNumber="6">R. harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2246FFFCE5432D207636FE85" box="[263,344,304,325]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B918EAD72246FFFCE5432D207636FE85" bold="true" box="[263,344,304,325]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
</th>
<td id="768D5DFB22460006E5D62D2075CCFE86" box="[402,674,304,326]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">2.7 30.0</td>
<td id="768D5DFB22460006E6F92D207462FE86" box="[701,780,304,326]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">26.1</td>
<td id="768D5DFB22460006E7742D2074A1FE86" box="[816,975,304,326]" gridcol="3" gridrow="1" pageId="5" pageNumber="6">3.58</td>
<td id="768D5DFB22460006E7AE2D2073F5FE86" box="[1002,1179,304,326]" gridcol="4" gridrow="1" pageId="5" pageNumber="6">40,600</td>
<td id="768D5DFB22460006E0852D20738FFE86" box="[1217,1249,304,326]" gridcol="5" gridrow="1" pageId="5" pageNumber="6">7.3</td>
<td id="768D5DFB22460006E1482D2072F3FE86" box="[1292,1437,304,326]" gridcol="6" gridrow="1" pageId="5" pageNumber="6">14.4</td>
</tr>
<tr id="355C348722460006E4D32D4972F3FEAF" box="[151,1437,345,367]" gridrow="2" pageId="5" pageNumber="6">
<th id="768D5DFB22460006E4D32D497636FEAF" box="[151,344,345,367]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">
<taxonomicName id="4C6C4D462246FFFCE4DB2D4A7649FEAF" box="[159,295,346,367]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="nemasoma">
<emphasis id="B918EAD72246FFFCE4DB2D4A7649FEAF" box="[159,295,346,367]" italics="true" pageId="5" pageNumber="6">R. nemasoma</emphasis>
</taxonomicName>
</th>
<td id="768D5DFB22460006E5D62D4975CCFEAF" box="[402,674,345,367]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">6.4 14.0</td>
<td id="768D5DFB22460006E6F92D497462FEAF" box="[701,780,345,367]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">37.725</td>
<td id="768D5DFB22460006E7742D4974A1FEAF" box="[816,975,345,367]" gridcol="3" gridrow="2" pageId="5" pageNumber="6"></td>
<td id="768D5DFB22460006E7AE2D4973F5FEAF" box="[1002,1179,345,367]" gridcol="4" gridrow="2" pageId="5" pageNumber="6"></td>
<td id="768D5DFB22460006E0852D49738FFEAF" box="[1217,1249,345,367]" gridcol="5" gridrow="2" pageId="5" pageNumber="6"></td>
<td id="768D5DFB22460006E1482D4972F3FEAF" box="[1292,1437,345,367]" gridcol="6" gridrow="2" pageId="5" pageNumber="6"></td>
</tr>
</table>
</paragraph>
<caption id="DF13664D2246FFFCE4D32A207263F9A5" httpUri="https://zenodo.org/record/277063/files/figure.png" pageId="5" pageNumber="6" targetBox="[151,1436,443,1563]" targetPageId="5">
<paragraph id="8BD336C52246FFFCE4D32A207263F9A5" blockId="5.[151,1436,1584,1637]" pageId="5" pageNumber="6">
<emphasis id="B918EAD72246FFFCE4D32A20767DF986" bold="true" box="[151,275,1584,1607]" pageId="5" pageNumber="6">FIGURE 2.</emphasis>
<taxonomicName id="4C6C4D462246FFFCE55D2A20768DF986" box="[281,483,1584,1606]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72246FFFCE55D2A20768DF986" box="[281,483,1584,1606]" italics="true" pageId="5" pageNumber="6">Raphidrilus harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2246FFFCE5AD2A217554F986" box="[489,570,1585,1606]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B918EAD72246FFFCE5AD2A217554F986" bold="true" box="[489,570,1585,1606]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
A, entire specimen in lateral view; B, mid-body segments with a segmented worm in the coelom; C, light microscopy of mid-body segments showing a segmented worm protruding from the coelom.
</paragraph>
</caption>
<subSubSection id="C376654E2246FFFFE4812A817625FEDC" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8BD336C52246FFFFE4812A817625FEDC" blockId="5.[151,1436,1681,2030]" lastBlockId="6.[151,1436,151,284]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B918EAD72246FFFCE4812A817630F96A" bold="true" box="[197,350,1681,1706]" pageId="5" pageNumber="6">Distribution.</emphasis>
<taxonomicName id="4C6C4D462246FFFCE5222A827529F96A" box="[358,583,1682,1706]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72246FFFCE5222A827529F96A" box="[358,583,1682,1706]" italics="true" pageId="5" pageNumber="6">Raphidrilus harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2246FFFCE60B2A8275C4F96A" box="[591,682,1682,1706]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B918EAD72246FFFCE60B2A8275C4F96A" bold="true" box="[591,682,1682,1706]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
, is known only from the
<typeStatus id="54D788672246FFFCE7882A837492F96B" box="[972,1020,1683,1707]" pageId="5" pageNumber="6">type</typeStatus>
locality, the GIWW in Venice, Florida. The distribution of this species is suspected to extend further south into the Florida Keys (T. H. Perkins, pers. comm.) if it is the same species that
<bibRefCitation id="EFFD4B342246FFFCE6072ACB74EDF932" author="Petersen" box="[579,899,1754,1778]" pageId="5" pageNumber="6" refString="Petersen, M. E. &amp; George, J. D. (1991) A new species of Raricirrus from northern Europe, with notes on its biology and a discussion of the affinities of the genus (Polychaeta: Ctenodrilidae), Pp. 185 - 208 in M. E. Petersen &amp; J. B. Kirkegaard, eds., Systematics, biology and morphology of world Polychaeta. Proceedings of the 2 nd International Polychaete Conference, Copenhagen 1986. Ophelia Supplement 5." type="journal volume" year="1991">Petersen and George (1991)</bibRefCitation>
referred to in their study. Unfortunately, those specimens have been lost (T. H. Perkins, pers. comm.), and could not be observed for comparison. Based upon correspondence and associated drawings between T. H. Perkins and M. E. Petersen specimens from the Florida Keys superficially appear to be
<taxonomicName id="4C6C4D462246FFFCE58C2B577550F89E" box="[456,574,1862,1886]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72246FFFCE58C2B577550F89E" box="[456,574,1862,1886]" italics="true" pageId="5" pageNumber="6">R. harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2246FFFCE60D2B5675C8F89E" box="[585,678,1862,1886]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B918EAD72246FFFCE60D2B5675C8F89E" bold="true" box="[585,678,1862,1886]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
The general body shape, number of chaetae per fascicle and the description of chaetae all match
<taxonomicName id="4C6C4D462246FFFCE6432B7B7515F842" box="[519,635,1898,1922]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72246FFFCE6432B7B7515F842" box="[519,635,1898,1922]" italics="true" pageId="5" pageNumber="6">R. harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2246FFFCE6C62B7A75B3F842" box="[642,733,1898,1922]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B918EAD72246FFFCE6C62B7A75B3F842" bold="true" box="[642,733,1898,1922]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
The only difference is that no branchiae or scars of branchiae were observed in the specimens from the Florida Keys, whereas almost all specimens from Venice have at least a stub, a single branchial filament or multiple branchial filaments. Additional specimens from the FSBC I collections labeled as “
<taxonomicName id="4C6C4D462246FFFCE55E2BC676F0F82E" box="[282,414,2006,2030]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="genus">
<emphasis id="B918EAD72246FFFCE55E2BC676F0F82E" box="[282,414,2006,2030]" italics="true" pageId="5" pageNumber="6">Raphidrilus</emphasis>
</taxonomicName>
sp.” were examined for comparison (FSBC
<accessionNumber id="943FAB262246FFFCE7D02BC77485F82E" box="[916,1003,2006,2030]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/I45229" pageId="5" pageNumber="6" type="EnaNcbi">I 45229</accessionNumber>
and FSBC
<accessionNumber id="943FAB262246FFFCE02A2BC773A4F82E" box="[1134,1226,2006,2030]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/I45230" pageId="5" pageNumber="6" type="EnaNcbi">I 45230</accessionNumber>
). These specimens were collected from Broward County along the east coast of Florida and are not
<taxonomicName id="4C6C4D462245FFFFE06C2C8973F2FF70" box="[1064,1180,152,176]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72245FFFFE06C2C8973F2FF70" box="[1064,1180,152,176]" italics="true" pageId="6" pageNumber="7">R. harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2245FFFFE0E02C887390FF70" box="[1188,1278,152,176]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B918EAD72245FFFFE0E02C887390FF70" bold="true" box="[1188,1278,152,176]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
They possess pectinate falcigers in the middle and posterior chaetigers and are most likely an undescribed species of
<emphasis id="B918EAD72245FFFFE1642CAD72F2FF14" box="[1312,1436,189,212]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="4C6C4D462245FFFFE1642CAD72F6FF14" box="[1312,1432,189,212]" class="Polychaeta" family="Ctenodrilidae" genus="Raricirrus" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="genus">Raricirrus</taxonomicName>
.
</emphasis>
Future collection efforts along the Gulf coast of Florida would help determine the geographical distribution of
<taxonomicName id="4C6C4D462245FFFFE1C02CF17785FEDC" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72245FFFFE1C02CF17785FEDC" italics="true" pageId="6" pageNumber="7">R. harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2245FFFFE4B62D147625FEDC" box="[242,331,260,284]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B918EAD72245FFFFE4B62D147625FEDC" bold="true" box="[242,331,260,284]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<caption id="DF13664D2245FFFFE4D32AC5727BF8CA" httpUri="https://zenodo.org/record/277064/files/figure.png" pageId="6" pageNumber="7" targetBox="[151,1436,337,1726]" targetPageId="6">
<paragraph id="8BD336C52245FFFFE4D32AC5727BF8CA" blockId="6.[151,1436,1749,1802]" pageId="6" pageNumber="7">
<emphasis id="B918EAD72245FFFFE4D32AC5767DF92A" bold="true" box="[151,275,1749,1772]" pageId="6" pageNumber="7">FIGURE 3.</emphasis>
SEM of
<taxonomicName id="4C6C4D462245FFFFE5362AC57553F92B" box="[370,573,1749,1771]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72245FFFFE5362AC57553F92B" box="[370,573,1749,1771]" italics="true" pageId="6" pageNumber="7">Raphidrilus harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2245FFFFE6002AC675FBF92B" box="[580,661,1750,1771]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B918EAD72245FFFFE6002AC675FBF92B" bold="true" box="[580,661,1750,1771]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
showing: A, complete specimen; B, anterior chaetigers in lateral view with two branchial filaments; C, anterior segments in lateral view; D, nuchal organ; E, serrate capillaries; F, pygidium.
</paragraph>
</caption>
<subSubSection id="C376654E2245FFFEE4812B27766CFB68" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8BD336C52245FFFEE4812B2775A2FF37" blockId="6.[151,1436,1846,2016]" lastBlockId="7.[151,1436,151,535]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B918EAD72245FFFFE4812B277657F88F" bold="true" box="[197,313,1847,1871]" pageId="6" pageNumber="7">Remarks.</emphasis>
<tableCitation id="C6EE037E2245FFFFE5042B2676FAF88E" box="[320,404,1846,1870]" captionStart="TABLE 2" captionStartId="7.[151,235,584,605]" captionTargetBox="[151,1424,633,1107]" captionTargetPageId="7" captionText="TABLE 2. Taxonomic characters of four Raphidrilus species including one undescribed species from the western Pacific." httpUri="http://table.plazi.org/id/DF13664D2244FFFEE4D32E587207FD9E" pageId="6" pageNumber="7" tableUuid="DF13664D2244FFFEE4D32E587207FD9E">Table 2</tableCitation>
summarizes the morphological characters useful to separate species in the genus
<taxonomicName id="4C6C4D462245FFFFE16B2B2677D6F8B3" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="genus">
<emphasis id="B918EAD72245FFFFE16B2B2677D6F8B3" italics="true" pageId="6" pageNumber="7">Raphidrilus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C6C4D462245FFFFE4832B4B76C6F8B3" box="[199,424,1883,1907]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72245FFFFE4832B4B76C6F8B3" box="[199,424,1883,1907]" italics="true" pageId="6" pageNumber="7">Raphidrilus harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2245FFFFE5F52B4B7562F8B3" box="[433,524,1883,1907]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B918EAD72245FFFFE5F52B4B7562F8B3" bold="true" box="[433,524,1883,1907]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
, differs from
<taxonomicName id="4C6C4D462245FFFFE6E82B4C743FF8B3" box="[684,849,1883,1907]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="hawaiiensis" status="sp. nov.">
<emphasis id="B918EAD72245FFFFE6E82B4C743FF8B3" box="[684,849,1883,1907]" italics="true" pageId="6" pageNumber="7">R. hawaiiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2245FFFFE71E2B4B74DBF8B3" box="[858,949,1883,1907]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B918EAD72245FFFFE71E2B4B74DBF8B3" bold="true" box="[858,949,1883,1907]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
, and
<taxonomicName id="4C6C4D462245FFFFE7B22B4C73E1F8B3" box="[1014,1167,1884,1907]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="nemasoma">
<emphasis id="B918EAD72245FFFFE7B22B4C73E1F8B3" box="[1014,1167,1884,1907]" italics="true" pageId="6" pageNumber="7">R. nemasoma</emphasis>
</taxonomicName>
by the presence of 36 capillary chaetae per fascicle in anterior chaetigers, while in both
<taxonomicName id="4C6C4D462245FFFFE7C82B90735FF856" box="[908,1073,1918,1943]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="hawaiiensis" status="sp. nov.">
<emphasis id="B918EAD72245FFFFE7C82B90735FF856" box="[908,1073,1918,1943]" italics="true" pageId="6" pageNumber="7">R. hawaiiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2245FFFFE07F2B6F73F8F857" box="[1083,1174,1919,1943]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B918EAD72245FFFFE07F2B6F73F8F857" bold="true" box="[1083,1174,1919,1943]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
, and
<taxonomicName id="4C6C4D462245FFFFE09C2B90721EF857" box="[1240,1392,1920,1943]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="nemasoma">
<emphasis id="B918EAD72245FFFFE09C2B90721EF857" box="[1240,1392,1920,1943]" italics="true" pageId="6" pageNumber="7">R. nemasoma</emphasis>
</taxonomicName>
the number of capillaries per fascicle is never greater than 4. The elongated midbody and posterior segments in
<taxonomicName id="4C6C4D462245FFFFE1C02BB47785F81F" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72245FFFFE1C02BB47785F81F" italics="true" pageId="6" pageNumber="7">R. harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2245FFFFE4B02BD7763FF81F" box="[244,337,1991,2015]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B918EAD72245FFFFE4B02BD7763FF81F" bold="true" box="[244,337,1991,2015]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
, is also very distinct and lack subannulations present in
<taxonomicName id="4C6C4D462245FFFFE7B62BD873F6F81F" box="[1010,1176,1991,2015]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="hawaiiensis" status="sp. nov.">
<emphasis id="B918EAD72245FFFFE7B62BD873F6F81F" box="[1010,1176,1991,2015]" italics="true" pageId="6" pageNumber="7">R. hawaiiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2245FFFFE0E62BD77390F81F" box="[1186,1278,1991,2015]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B918EAD72245FFFFE0E62BD77390F81F" bold="true" box="[1186,1278,1991,2015]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
, and
<taxonomicName id="4C6C4D462245FFFEE1062BD8778EFF70" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" lastPageId="7" lastPageNumber="8" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="nemasoma">
<emphasis id="B918EAD72245FFFEE1062BD8778EFF70" italics="true" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">R. nemasoma</emphasis>
</taxonomicName>
. The heart body in
<taxonomicName id="4C6C4D462244FFFEE5822C897555FF70" box="[454,571,152,176]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72244FFFEE5822C897555FF70" box="[454,571,152,176]" italics="true" pageId="7" pageNumber="8">R. harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2244FFFEE6002C8875CEFF70" box="[580,672,152,176]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B918EAD72244FFFEE6002C8875CEFF70" bold="true" box="[580,672,152,176]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
, extends posteriorly to chaetigers 79, while in
<taxonomicName id="4C6C4D462244FFFEE0882C89721FFF70" box="[1228,1393,152,176]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="hawaiiensis" status="sp. nov.">
<emphasis id="B918EAD72244FFFEE0882C89721FFF70" box="[1228,1393,152,176]" italics="true" pageId="7" pageNumber="8">R. hawaiiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2244FFFEE13E2C8877A6FF14" pageId="7" pageNumber="8" rank="species">
<emphasis id="B918EAD72244FFFEE13E2C8877A6FF14" bold="true" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
, the heart body projects anteriorly to the middle of chaetiger 3 and in
<taxonomicName id="4C6C4D462244FFFEE7922CAD7303FF14" box="[982,1133,189,212]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="nemasoma">
<emphasis id="B918EAD72244FFFEE7922CAD7303FF14" box="[982,1133,189,212]" italics="true" pageId="7" pageNumber="8">R. nemasoma</emphasis>
</taxonomicName>
the heart body is restricted to the extension of chaetiger 4 (
<bibRefCitation id="EFFD4B342244FFFEE5B32CCF75AEFF37" author="Monticelli" box="[503,704,223,247]" pageId="7" pageNumber="8" refString="Monticelli, F. S. (1910 b) Raphidrilus nemasoma Montic. Nuovo Ctenodrilide del Golfo di Napoli (Revisione de'Ctenodrilidi). Archivio Zoologico, 4 (4), 401 - 436, plates 12 - 13." type="journal article" year="1910" yearSuffix="b">Monticelli 1910b</bibRefCitation>
).
</paragraph>
<paragraph id="8BD336C52244FFFEE4812D147483FDD7" blockId="7.[151,1436,151,535]" pageId="7" pageNumber="8">
The species of
<taxonomicName id="4C6C4D462244FFFEE5282D14769EFEDC" box="[364,496,260,284]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="genus">
<emphasis id="B918EAD72244FFFEE5282D14769EFEDC" box="[364,496,260,284]" italics="true" pageId="7" pageNumber="8">Raphidrilus</emphasis>
</taxonomicName>
referred to by
<bibRefCitation id="EFFD4B342244FFFEE6D02D1474E8FEDC" author="Qian" box="[660,902,260,284]" pageId="7" pageNumber="8" refString="Qian, P. - Y. &amp; Chia, F. - S. (1989) Sexual reproduction and larval development of Raphidrilus nemasoma Monticelli, 1910 (Polychaeta: Ctenodrilidae). Canadian Journal of Zoology, 67, 2345 - 2351." type="journal article" year="1989">Qian and Chia (1989)</bibRefCitation>
as
<emphasis id="B918EAD72244FFFEE7EF2D1473DBFEDC" box="[939,1205,260,284]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C6C4D462244FFFEE7EF2D1473DFFEDC" box="[939,1201,260,284]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="nemasoma">Raphidrilus nemasoma</taxonomicName>
,
</emphasis>
and later considered to be undescribed by
<bibRefCitation id="EFFD4B342244FFFEE5C22D3875ADFEFF" author="Petersen" box="[390,707,295,319]" pageId="7" pageNumber="8" refString="Petersen, M. E. &amp; George, J. D. (1991) A new species of Raricirrus from northern Europe, with notes on its biology and a discussion of the affinities of the genus (Polychaeta: Ctenodrilidae), Pp. 185 - 208 in M. E. Petersen &amp; J. B. Kirkegaard, eds., Systematics, biology and morphology of world Polychaeta. Proceedings of the 2 nd International Polychaete Conference, Copenhagen 1986. Ophelia Supplement 5." type="journal volume" year="1991">Petersen and George (1991)</bibRefCitation>
, is distinct from
<taxonomicName id="4C6C4D462244FFFEE73B2D39749CFE80" box="[895,1010,296,320]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72244FFFEE73B2D39749CFE80" box="[895,1010,296,320]" italics="true" pageId="7" pageNumber="8">R. harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2244FFFEE7BD2D38733DFE80" box="[1017,1107,296,320]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B918EAD72244FFFEE7BD2D38733DFE80" bold="true" box="[1017,1107,296,320]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C6C4D462244FFFEE0252D39726AFE80" box="[1121,1284,296,320]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="hawaiiensis" status="sp. nov.">
<emphasis id="B918EAD72244FFFEE0252D39726AFE80" box="[1121,1284,296,320]" italics="true" pageId="7" pageNumber="8">R. hawaiiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2244FFFEE1482D38720BFE80" box="[1292,1381,296,320]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B918EAD72244FFFEE1482D38720BFE80" bold="true" box="[1292,1381,296,320]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
, and
<taxonomicName id="4C6C4D462244FFFEE4D32D5D7641FEA4" box="[151,303,333,356]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="nemasoma">
<emphasis id="B918EAD72244FFFEE4D32D5D7641FEA4" box="[151,303,333,356]" italics="true" pageId="7" pageNumber="8">R. nemasoma</emphasis>
</taxonomicName>
. Even though adult worms of
<taxonomicName id="4C6C4D462244FFFEE6C82D5C7339FEA4" authority="Qian and Chia (1989)" authorityName="Qian and Chia" authorityYear="1989" box="[652,1111,332,356]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="genus">
<emphasis id="B918EAD72244FFFEE6C82D5C747EFEA4" box="[652,784,332,356]" italics="true" pageId="7" pageNumber="8">Raphidrilus</emphasis>
sensu
<bibRefCitation id="EFFD4B342244FFFEE71B2D5C7339FEA4" author="Qian" box="[863,1111,332,356]" pageId="7" pageNumber="8" refString="Qian, P. - Y. &amp; Chia, F. - S. (1989) Sexual reproduction and larval development of Raphidrilus nemasoma Monticelli, 1910 (Polychaeta: Ctenodrilidae). Canadian Journal of Zoology, 67, 2345 - 2351." type="journal article" year="1989">Qian and Chia (1989)</bibRefCitation>
</taxonomicName>
were not described by these authors, the many scattered short sensory cilia in addition to the nuchal organs present on the prostomium and peristomium, the sensory tufts posterodorsal to notochaetae, the short serrate neurochaetae (reported as being genital spines), and the terminal anus of juveniles worms (811 chaetigers) are unique characteristics not observed in the species from Florida, Hawaii, or the Mediterranean Sea. Adult specimens from the same locality sampled by
<bibRefCitation id="EFFD4B342244FFFEE1222DCC7622FDD7" author="Qian" pageId="7" pageNumber="8" refString="Qian, P. - Y. &amp; Chia, F. - S. (1989) Sexual reproduction and larval development of Raphidrilus nemasoma Monticelli, 1910 (Polychaeta: Ctenodrilidae). Canadian Journal of Zoology, 67, 2345 - 2351." type="journal article" year="1989">Qian and Chia (1989)</bibRefCitation>
need to be examined to confirm the status of a new species.
</paragraph>
<caption id="DF13664D2244FFFEE4D32E587207FD9E" ID-Table-UUID="DF13664D2244FFFEE4D32E587207FD9E" box="[151,1385,584,606]" httpUri="http://table.plazi.org/id/DF13664D2244FFFEE4D32E587207FD9E" pageId="7" pageNumber="8" targetBox="[151,1424,633,1107]" targetIsTable="true" targetPageId="7">
<paragraph id="8BD336C52244FFFEE4D32E587207FD9E" blockId="7.[151,1385,584,606]" box="[151,1385,584,606]" pageId="7" pageNumber="8">
<emphasis id="B918EAD72244FFFEE4D32E58766AFD9D" bold="true" box="[151,260,584,605]" pageId="7" pageNumber="8">TABLE 2.</emphasis>
Taxonomic characters of four
<taxonomicName id="4C6C4D462244FFFEE67F2E5875DDFD9E" box="[571,691,584,606]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="genus">
<emphasis id="B918EAD72244FFFEE67F2E5875DDFD9E" box="[571,691,584,606]" italics="true" pageId="7" pageNumber="8">Raphidrilus</emphasis>
</taxonomicName>
species including one undescribed species from the western Pacific.
</paragraph>
</caption>
<paragraph id="8BD336C52244FFFEE4DB2E6973F7FC3B" pageId="7" pageNumber="8">
<table id="F96CC46522440006E4D32E6972FEFB93" box="[151,1424,633,1107]" gridcols="5" gridrows="5" pageId="7" pageNumber="8">
<tr id="355C348722440006E4D32E6972FEFD6B" box="[151,1424,633,683]" gridrow="0" pageId="7" pageNumber="8">
<th id="768D5DFB22440006E4D32E69762FFD6B" box="[151,321,633,683]" gridcol="0" gridrow="0" pageId="7" pageNumber="8">Species</th>
<th id="768D5DFB22440006E5112E6975E2FD6B" box="[341,652,633,683]" gridcol="1" gridrow="0" pageId="7" pageNumber="8">Prostomium</th>
<th id="768D5DFB22440006E6D82E6974FBFD6B" box="[668,917,633,683]" gridcol="2" gridrow="0" pageId="7" pageNumber="8">Peristomium</th>
<th id="768D5DFB22440006E7F52E697315FD6B" box="[945,1147,633,683]" gridcol="3" gridrow="0" pageId="7" pageNumber="8">Achaetous segment</th>
<th id="768D5DFB22440006E0CA2E6972FEFD6B" box="[1166,1424,633,683]" gridcol="4" gridrow="0" pageId="7" pageNumber="8">Position and extent of heart body</th>
</tr>
<tr id="355C348722440006E4D32EAE72FEFCCE" box="[151,1424,702,782]" gridrow="1" pageId="7" pageNumber="8">
<th id="768D5DFB22440006E4D32EAE762FFCCE" box="[151,321,702,782]" gridcol="0" gridrow="1" pageId="7" pageNumber="8">
<taxonomicName id="4C6C4D462244FFFEE4DB2EAE766FFD13" box="[159,257,702,723]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="harperi" status="sp. nov.">
<emphasis id="B918EAD72244FFFEE4DB2EAE766FFD13" box="[159,257,702,723]" italics="true" pageId="7" pageNumber="8">R. harperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2244FFFEE4DB2ECB7785FD30" box="[159,235,731,752]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B918EAD72244FFFEE4DB2ECB7785FD30" bold="true" box="[159,235,731,752]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
</th>
<td id="768D5DFB22440006E5112EAE75E2FCCE" box="[341,652,702,782]" gridcol="1" gridrow="1" pageId="7" pageNumber="8">Rounded; pair of oval nuchal organs; pearshaped with peris- tomium</td>
<td id="768D5DFB22440006E6D82EAE74FBFCCE" box="[668,917,702,782]" gridcol="2" gridrow="1" pageId="7" pageNumber="8">Single annulus; not dor- sally delimited from pros- tomium</td>
<td id="768D5DFB22440006E7F52EAE7315FCCE" box="[945,1147,702,782]" gridcol="3" gridrow="1" pageId="7" pageNumber="8">One dorsally biannu- lated</td>
<td id="768D5DFB22440006E0CA2EAE72FEFCCE" box="[1166,1424,702,782]" gridcol="4" gridrow="1" pageId="7" pageNumber="8">Chaetiger 4; posteriorly directed to chaetiger 7</td>
</tr>
<tr id="355C348722440006E4D32F3172FEFCB1" box="[151,1424,801,881]" gridrow="2" pageId="7" pageNumber="8">
<th id="768D5DFB22440006E4D32F31762FFCB1" box="[151,321,801,881]" gridcol="0" gridrow="2" pageId="7" pageNumber="8">
<taxonomicName id="4C6C4D462244FFFEE4DB2F317644FCF6" box="[159,298,801,822]" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="hawaiiensis" status="sp. nov.">
<emphasis id="B918EAD72244FFFEE4DB2F317644FCF6" box="[159,298,801,822]" italics="true" pageId="7" pageNumber="8">R. hawaiiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57AC2244FFFEE4DB2F2E7785FC93" box="[159,235,830,851]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B918EAD72244FFFEE4DB2F2E7785FC93" bold="true" box="[159,235,830,851]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
</th>
<td id="768D5DFB22440006E5112F3175E2FCB1" box="[341,652,801,881]" gridcol="1" gridrow="2" pageId="7" pageNumber="8">Rounded; pair of oval nuchal organs; pearshaped with peris- tomium</td>
<td id="768D5DFB22440006E6D82F3174FBFCB1" box="[668,917,801,881]" gridcol="2" gridrow="2" pageId="7" pageNumber="8">Single annulus; not dor- sally delimited from pros- tomium</td>
<td id="768D5DFB22440006E7F52F317315FCB1" box="[945,1147,801,881]" gridcol="3" gridrow="2" pageId="7" pageNumber="8">12 dorsally biannu- lated</td>
<td id="768D5DFB22440006E0CA2F3172FEFCB1" box="[1166,1424,801,881]" gridcol="4" gridrow="2" pageId="7" pageNumber="8">Chaetiger 4; anteriorly directed to middle of chae- tiger 3</td>
</tr>
<tr id="355C348722440006E4D32F9472FEFC13" box="[151,1424,900,979]" gridrow="3" pageId="7" pageNumber="8">
<th id="768D5DFB22440006E4D32F94762FFC13" box="[151,321,900,979]" gridcol="0" gridrow="3" pageId="7" pageNumber="8">
<taxonomicName id="4C6C4D462244FFFEE4DB2F947651FC76" authority="Monticelli, 1910" authorityName="Monticelli" authorityYear="1910" class="Polychaeta" family="Ctenodrilidae" genus="Raphidrilus" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="nemasoma">
<emphasis id="B918EAD72244FFFEE4DB2F94764EFC59" box="[159,288,900,921]" italics="true" pageId="7" pageNumber="8">R. nemasoma</emphasis>
Monticelli, 1910
</taxonomicName>
</th>
<td id="768D5DFB22440006E5112F9475E2FC13" box="[341,652,900,979]" gridcol="1" gridrow="3" pageId="7" pageNumber="8">Broadly rounded; a pair of nuchal organs; thimbleshaped with peristomium</td>
<td id="768D5DFB22440006E6D82F9474FBFC13" box="[668,917,900,979]" gridcol="2" gridrow="3" pageId="7" pageNumber="8">Single annulus; not dor- sally delimited from pros- tomium</td>
<td id="768D5DFB22440006E7F52F947315FC13" box="[945,1147,900,979]" gridcol="3" gridrow="3" pageId="7" pageNumber="8">One dorsally biannu- lated</td>
<td id="768D5DFB22440006E0CA2F9472FEFC13" box="[1166,1424,900,979]" gridcol="4" gridrow="3" pageId="7" pageNumber="8">Chaetiger 4 only</td>
</tr>
<tr id="355C348722440006E4D32FF672FEFB93" box="[151,1424,998,1107]" gridrow="4" pageId="7" pageNumber="8">
<th id="768D5DFB22440006E4D32FF6762FFB93" box="[151,321,998,1107]" gridcol="0" gridrow="4" pageId="7" pageNumber="8">
<emphasis id="B918EAD72244FFFEE4DB2FF677BAFC3B" box="[159,212,998,1019]" italics="true" pageId="7" pageNumber="8">R. sp.</emphasis>
sensu Qian &amp; Chia, 1989*
</th>
<td id="768D5DFB22440006E5112FF675E2FB93" box="[341,652,998,1107]" gridcol="1" gridrow="4" pageId="7" pageNumber="8">Broadly rounded; a pair of nuchal organs and scattered sen- sorial tufts; thimbleshaped with peristomium</td>
<td id="768D5DFB22440006E6D82FF674FBFB93" box="[668,917,998,1107]" gridcol="2" gridrow="4" pageId="7" pageNumber="8">Single annulus?</td>
<td id="768D5DFB22440006E7F52FF67315FB93" box="[945,1147,998,1107]" gridcol="3" gridrow="4" pageId="7" pageNumber="8">One dorsally biannu- lated?</td>
<td id="768D5DFB22440006E0CA2FF672FEFB93" box="[1166,1424,998,1107]" gridcol="4" gridrow="4" pageId="7" pageNumber="8">?</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD336C52244FFFEE4D32864752BFB49" blockId="7.[151,581,1139,1192]" box="[151,581,1139,1162]" pageId="7" pageNumber="8">* based on juveniles with 811 chaetigers.</paragraph>
<paragraph id="8BD336C52244FFFEE4D32882766CFB68" blockId="7.[151,581,1139,1192]" box="[151,258,1170,1192]" pageId="7" pageNumber="8">continued.</paragraph>
</subSubSection>
</treatment>
</document>