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<mods:title id="0DBC94541C5527B92064106E494C0DC1">Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications</mods:title>
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<mods:namePart id="CD32AC33A20A43866E3AC0105B3E6D23">Ivanov, Martin</mods:namePart>
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<mods:namePart id="FB821CE43878B52AADE635669E15D6C8">Luján, Àngel Hernández</mods:namePart>
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<treatment id="03C587C7430DFFEDFEEBFD4A49A2F839" ID-DOI="http://doi.org/10.5281/zenodo.4488217" ID-GBIF-Taxon="177200716" ID-Zenodo-Dep="4488217" LSID="urn:lsid:plazi:treatment:03C587C7430DFFEDFEEBFD4A49A2F839" httpUri="http://treatment.plazi.org/id/03C587C7430DFFEDFEEBFD4A49A2F839" lastPageNumber="350" pageId="9" pageNumber="350">
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<paragraph id="8BD336D1430DFFEDFEEBFD4A4A78FD3D" blockId="9.[343,564,725,753]" box="[343,564,725,753]" pageId="9" pageNumber="350">
<heading id="D09B81BD430DFFEDFEEBFD4A4A78FD3D" box="[343,564,725,753]" centered="true" fontSize="9" level="2" pageId="9" pageNumber="350" reason="7">
?
<taxonomicName id="4C6C4D52430DFFEDFEDDFD4B49A6FD3C" authorityName="Oppel" authorityYear="1811" box="[353,490,727,753]" kingdom="Animalia" pageId="9" pageNumber="350" rank="superFamily" superFamily="Scincoidea">SCINCOIDEA</taxonomicName>
indet.
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<subSubSection id="C376655A430DFFEDFEC2FD694A41FCC2" box="[382,525,757,784]" pageId="9" pageNumber="350" type="description">
<paragraph id="8BD336D1430DFFEDFEC2FD694A41FCC2" blockId="9.[382,525,757,784]" box="[382,525,757,784]" pageId="9" pageNumber="350">
(
<figureCitation id="13572A54430DFFEDFE34FD694A48FCC2" box="[392,516,757,784]" captionStart="FIG" captionStartId="10.[132,143,1686,1704]" captionTargetBox="[132,1455,215,1644]" captionTargetId="figure-142@10.[132,1455,215,1645]" captionTargetPageId="10" captionText="FIG. 4. — Lacertoidea indet. (A, B) and? Scincoidea indet. (C, D) from the early Miocene (MN 4) of MWQ. Left dentary Pal. 1571 (2/2003 Reptile Joint) in (A) lateral and (B) medial aspects with detail of teeth. Left maxilla Pal. 1573 (2/2003 Reptile Joint) in (C) lateral and (D) medial aspects with detail of teeth. Abbreviations: dc, dental crest; cm, carina maxillaris; lf, labial foramina; Mg, Meckel`s groove; np, nasal process; sbs, subdental shelf; srs, supradental shelf; st, striations. Scale bars: A-D, 1 mm; B, D, 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4447573" httpUri="https://zenodo.org/record/4447573/files/figure.png" pageId="9" pageNumber="350">Fig. 4C, D</figureCitation>
)
</paragraph>
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<subSubSection id="C376655A430DFFEDFF38FCDB4A9EFCAB" pageId="9" pageNumber="350" type="materials_examined">
<paragraph id="8BD336D1430DFFEDFF38FCDB4A9EFCAB" blockId="9.[132,774,838,889]" pageId="9" pageNumber="350">
MATERIAL. —
<materialsCitation id="3B043C8C430DFFEDFEA1FCDB4A9EFCAB" ID-GBIF-Occurrence="3027945310" pageId="9" pageNumber="350">
MWQ, early Miocene, Burdigalian, Orleanian,
<collectionCode id="ED7DAE14430DFFEDFF38FCFD48E3FCAA" box="[132,175,865,888]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="9" pageNumber="350">MN</collectionCode>
4: 2/2003 Reptile Joint: one left maxilla (Pal. 1573).
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</paragraph>
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<subSubSection id="C376655A430DFFEDFF38FC294AC4FA9F" pageId="9" pageNumber="350" type="description">
<paragraph id="8BD336D1430DFFEDFF38FC294957FC1F" blockId="9.[131,777,949,1358]" box="[132,283,949,975]" pageId="9" pageNumber="350">DESCRIPTION</paragraph>
<paragraph id="8BD336D1430DFFEDFF38FC484896FC3C" blockId="9.[131,777,949,1358]" box="[132,218,980,1006]" pageId="9" pageNumber="350">
<emphasis id="B918EAC3430DFFEDFF38FC484896FC3C" box="[132,218,980,1006]" italics="true" pageId="9" pageNumber="350">Maxilla</emphasis>
</paragraph>
<paragraph id="8BD336D1430DFFEDFF38FC684AC4FA9F" blockId="9.[131,777,949,1358]" pageId="9" pageNumber="350">
Only a fragment of the left maxilla is preserved. It bears five tooth positions (only one and half teeth are attached). The supradental shelf is thin and straight in the preserved portion. The nasal process is partly preserved. On the medial side of the nasal process, there is a fine ridge (carina maxillaris
<emphasis id="B918EAC3430DFFEDFF40FB09497FFB7C" box="[252,307,1173,1198]" italics="true" pageId="9" pageNumber="350">sensu</emphasis>
<bibRefCitation id="EFFD4B20430DFFEDFE81FB084982FB7C" author="MULLER J." box="[317,462,1172,1198]" pageId="9" pageNumber="350" pagination="79 - 88" refId="ref25995" refString="MULLER J. 1996. - Eine neue Art der Echten Eidechsen (Reptilia: Lacertilia: Lacertidae) aus dem Unteren Miozan. Mainzer Geowissenschaftliche Mitteilungen 25: 79 - 88." type="journal article" year="1996">Müller 1996</bibRefCitation>
) that runs posterodorsally. The ridge originates from the subdental shelf. This gives an estimation that this fragment represents the anterior region of a maxilla. In the lower region of the external surface, three irregularly spaced labial foramina are located. Besides this, the rest of the surface is smooth.
</paragraph>
</subSubSection>
<subSubSection id="C376655A430DFFEDFF38FAE8497EF91E" pageId="9" pageNumber="350" type="description">
<paragraph id="8BD336D1430DFFEDFF38FAE848A2FA5F" blockId="9.[132,776,1396,1740]" box="[132,238,1396,1421]" pageId="9" pageNumber="350">
<emphasis id="B918EAC3430DFFEDFF38FAE848A2FA5F" box="[132,238,1396,1421]" italics="true" pageId="9" pageNumber="350">Dentition</emphasis>
</paragraph>
<paragraph id="8BD336D1430DFFEDFF38FA0F497EF91E" blockId="9.[132,776,1396,1740]" pageId="9" pageNumber="350">
The implantation is pleurodont. Only one tooth is more-orless completely preserved. The lingual aspect of the crown is bordered by the culmen lateralis anterior and culmen lateralis posterior (terms after
<bibRefCitation id="EFFD4B20430DFFEDFE53FA6F4AC7F9DF" author="RICHTER A." box="[495,651,1523,1549]" pageId="9" pageNumber="350" pagination="1 - 147" refId="ref27007" refString="RICHTER A. 1994. - Lacertilia aus der Unteren Kreide von Una und Galve (Spanien) und Anoual (Marokko). Berliner geowissenschaftliche Abhandlungen (E: Palaobiologie) 14: 1 - 147." type="journal article" year="1994">Richter 1994</bibRefCitation>
), between which the area is striated. This fine striation is formed by approximately six striae. The tooth crown termination is blunt. In medial aspect, it appears to be divided into rounded labial cusp, and a smaller, medially located lingual cusp. The tooth bases are pierced by small rounded resorption pits.
</paragraph>
</subSubSection>
<subSubSection id="C376655A430DFFEDFF38F96E49A2F839" pageId="9" pageNumber="350" type="discussion">
<paragraph id="8BD336D1430DFFEDFF38F96E48A0F8D8" blockId="9.[132,775,1778,2028]" box="[132,236,1778,1804]" pageId="9" pageNumber="350">REMARKS</paragraph>
<paragraph id="8BD336D1430DFFEDFF38F88E49A2F839" blockId="9.[132,775,1778,2028]" pageId="9" pageNumber="350">
The tooth morphology resembles that of Scincidae or Cordylidae (see
<bibRefCitation id="EFFD4B20430DFFEDFEF9F8AE4996F89E" author="KOSMA R." box="[325,474,1842,1868]" pageId="9" pageNumber="350" refId="ref25766" refString="KOSMA R. 2004. - The Dentition of Recent and Fossil Scincomorphan Lizards (Lacertilia, Squamata) - Systematics, Functional Morphology, Palecology. Unpublished PhD thesis, University of Hannover, Hannover. 187 p." type="book" year="2004">Kosma 2004</bibRefCitation>
). Both groups have been previously described from several lower Miocene sites of Central Europe (e.g.,
<bibRefCitation id="EFFD4B20430DFFEDFE19F8ED4A63F859" author="ROCEK Z." box="[421,559,1905,1931]" pageId="9" pageNumber="350" pagination="1 - 69" refId="ref27045" refString="ROCEK Z. 1984. - Lizards (Reptilia: Sauria) from the Lower Miocene locality Dolnice (Bohemia, Czechoslovakia). Rozpravy Ceskoslovenske Akademie Ved, Rada Matematickych a Prirodnich Ved 94: 1 - 69." type="journal article" year="1984">Roček 1984</bibRefCitation>
;
<bibRefCitation id="EFFD4B20430DFFEDFDFDF8EF4AB3F859" author="CERNANSKY A." box="[577,767,1905,1935]" pageId="9" pageNumber="350" pagination="837 - 847" refId="ref22955" refString="CERNANSKY A. 2012. - The oldest known European Neogene girdled lizard fauna (Squamata, Cordylidae), with comments on early Miocene immigration of African taxa. Geodiversitas 34 (4): 837 - 847. https: // doi. org / 10.5252 / g 2012 n 4 a 6" type="journal article" year="2012">Čerňanský 2012</bibRefCitation>
; Čerňanský 2016). Unfortunately, the character of preservation of this maxillary fragment from Mokrá does not allow a more specific allocation.
</paragraph>
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