257 lines
30 KiB
XML
257 lines
30 KiB
XML
<document id="07DBE534001811BEBD17715CD6E68B24" ID-CLB-Dataset="304560" ID-DOI="10.11646/zootaxa.5519.2.3" ID-GBIF-Dataset="a3266865-054f-4cff-bc0c-faf89ec8074c" ID-ISSN="1175-5326" ID-Zenodo-Dep="13915771" ID-ZooBank="BE0CA3E4-3695-403A-AD98-3C279238518B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1728585742752" checkinUser="plazi" docAuthor="Theischinger, Günther, Kalnins, Martins & Marinov, Milen" docDate="2024" docId="03C587C6CE7B987CFF24858F77A4F806" docLanguage="en" docName="zootaxa.5519.2.3.pdf" docOrigin="Zootaxa 5519 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5519.2.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anax tonga Theischinger, Kalnins & Marinov, 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="232" masterDocId="FFFCFFBECE74986DFFB3832C7740FF90" masterDocTitle="Two new species of Anax Leach, 1815 from the Solomon Islands and Tonga (Odonata: Aeshnidae)" masterLastPageNumber="242" masterPageNumber="215" pageNumber="230" updateTime="1729087811061" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="35C64DB3F42A91F15F13BD3803EB55F4">Two new species of Anax Leach, 1815 from the Solomon Islands and Tonga (Odonata: Aeshnidae)</mods:title>
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<mods:name id="64C31F63F4B0FCBC4B576DF7A6A2B7F1" type="personal">
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<mods:roleTerm id="6056BD85568DFCC4CB3E4E4C3BBD8D0C">Author</mods:roleTerm>
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<mods:namePart id="E5933B5AB2F2933CE873FF85D432C93F">Theischinger, Günther</mods:namePart>
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<mods:nameIdentifier id="28481E10B31FC0FAD49A92BA26782FD1" type="ORCID">0000-0002-5207-2626</mods:nameIdentifier>
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<mods:affiliation id="EA3F237E38EECE8EB5782EFC26B12CB7">Australian Museum, Entomology, 6 College Street, Sydney, NSW 2010</mods:affiliation>
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<mods:nameIdentifier id="E3FE4A6C8D5B2AF9904070B4237D454D" type="email">theischingergunther@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="51BD3D1AAFDEE659A20E2450E6E6AB3E" type="personal">
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<mods:roleTerm id="E0B18FBB310570BF2298194B2FBB7924">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2C0398CF0C5EFBF385CC006877EFE238">Kalnins, Martins</mods:namePart>
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<mods:nameIdentifier id="C55D3B817D939343016F83BD9E74F0C1" type="ORCID">0009-0008-8850-4444</mods:nameIdentifier>
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<mods:affiliation id="1DA3BD8B4AA19EBD2D20A970D76D49AF">Department of Biosystematics, Institute of Life Sciences and Technology, Daugavpils University, Parādes Street 1 A, Daugavpils, Latvia, LV- 5401</mods:affiliation>
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<mods:nameIdentifier id="D124E308A1D9CC78ED26954A6CA61F6A" type="email">martins.kalnins@biology.lv</mods:nameIdentifier>
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</mods:name>
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<mods:roleTerm id="8D95EEB39309FA580CA18F1AD62CEF00">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="45CA2E292CDECB3DF8B33C4CE36AC84B">Marinov, Milen</mods:namePart>
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<mods:nameIdentifier id="9D8A72B6F040C62A74488CE3D7095C8E" type="ORCID">0000-0003-3284-2555</mods:nameIdentifier>
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<mods:affiliation id="8DE58D5A535BF39410241E7982FA3D85">Biosecurity Surveillance & Incursion Investigation Plant Health Team, Ministry for Primary Industries, 14 Sir William Pickering Drive, Christchurch 8544, New Zealand</mods:affiliation>
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<mods:nameIdentifier id="A8B1C8D2F668A6C48133405DC5FB2BAC" type="email">milen.marinov@mpi.govt.nz</mods:nameIdentifier>
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<mods:title id="24644E554F75E7C2A9023F3ADC607AAF">Zootaxa</mods:title>
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<mods:part id="046B4F357E31E93427B1564DC11D693B">
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<mods:date id="F048B7F16EE31AAE46EBF8F0D497B5C6">2024</mods:date>
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<mods:number id="A5A780A6A905FA4B22C51CD9F033C1B7">2024-10-08</mods:number>
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<mods:number id="F1C5B942AABC8CD482AC5A514E6A8F7C">5519</mods:number>
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<mods:url id="06B91A283758956493A8E02CC3CFC6C3">http://dx.doi.org/10.11646/zootaxa.5519.2.3</mods:url>
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<mods:identifier id="230CF13E417A1ED5007E0C0D9DE9776B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C587C6CE7B987CFF24858F77A4F806" ID-DOI="http://doi.org/10.5281/zenodo.13940295" ID-GBIF-Taxon="243657026" ID-Zenodo-Dep="13940295" LSID="urn:lsid:plazi:treatment:03C587C6CE7B987CFF24858F77A4F806" httpUri="http://treatment.plazi.org/id/03C587C6CE7B987CFF24858F77A4F806" lastPageId="17" lastPageNumber="232" pageId="15" pageNumber="230">
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<subSubSection id="C376655BCE7B9862FF24858F76C9F92D" box="[151,393,1699,1725]" pageId="15" pageNumber="230" type="nomenclature">
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<paragraph id="8BD336D0CE7B9862FF24858F76C9F92D" blockId="15.[151,463,1699,1762]" box="[151,393,1699,1725]" pageId="15" pageNumber="230">
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<heading id="D09B81BCCE7B9862FF24858F76C9F92D" bold="true" box="[151,393,1699,1725]" fontSize="11" level="1" pageId="15" pageNumber="230" reason="1">
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<emphasis id="B918EAC2CE7B9862FF24858F76C9F92D" bold="true" box="[151,393,1699,1725]" pageId="15" pageNumber="230">
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<taxonomicName id="4C6C4D53CE7B9862FF24858F7664F92D" authorityName="Theischinger & Kalnins & Marinov" authorityYear="2024" box="[151,292,1699,1725]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="230" phylum="Arthropoda" rank="species" species="tonga" status="sp. nov.">
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<emphasis id="B918EAC2CE7B9862FF24858F7664F92D" bold="true" box="[151,292,1699,1725]" italics="true" pageId="15" pageNumber="230">Anax tonga</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22B57B9CE7B9862FE9F858F76C9F92D" box="[300,393,1699,1725]" pageId="15" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C376655BCE7B9862FF2485EB768FF971" box="[151,463,1735,1762]" pageId="15" pageNumber="230" type="description">
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<paragraph id="8BD336D0CE7B9862FF2485EB768FF971" blockId="15.[151,463,1699,1762]" box="[151,463,1735,1762]" pageId="15" pageNumber="230">
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(
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<figureCitation id="13572A55CE7B9862FF2C85EB77B6F972" box="[159,246,1735,1762]" captionStart="FIGURE 13–17" captionStartId="6.[151,250,476,500]" captionTargetBox="[409,1172,181,452]" captionTargetId="figure-22@6.[408,1180,181,452]" captionTargetPageId="6" captionText="FIGURE 13–17. Similarities: Anax spp., male terminalia, dorsal: (13) A. guttatus; (14) A. insulanus; (15) A. tonga; (16) A. maclachlani; (17) A. gibbosulus." figureDoi="http://doi.org/10.5281/zenodo.13915777" httpUri="https://zenodo.org/record/13915777/files/figure.png" pageId="15" pageNumber="230">Figs 15</figureCitation>
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,
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<figureCitation id="13572A55CE7B9862FEB185EB760BF971" box="[258,331,1735,1761]" captionStart="FIGURES 18–25" captionStartId="6.[151,264,1868,1892]" captionTargetBox="[151,1422,561,1845]" captionTargetId="figure-54@6.[151,1435,561,1845]" captionTargetPageId="6" captionText="FIGURES 18–25. Anax spp., male, dorsal: (18–21) S2–3; (22–25) terminalia: (18, 22) A. tonga; (19, 23) A. celebensis; (20, 24) A. insulanus; (21, 25) A. piraticus." figureDoi="http://doi.org/10.5281/zenodo.13940290" httpUri="https://zenodo.org/record/13940290/files/figure.png" pageId="15" pageNumber="230">18, 22</figureCitation>
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, 34, 35, 48)
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</paragraph>
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</subSubSection>
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<subSubSection id="C376655BCE7B9862FF24842374E3F8B9" box="[151,931,1807,1834]" pageId="15" pageNumber="230" type="materials_examined">
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<paragraph id="8BD336D0CE7B9862FF24842374E3F8B9" blockId="15.[151,1436,1807,1906]" box="[151,931,1807,1834]" pageId="15" pageNumber="230">
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<materialsCitation id="3B043C8DCE7B9862FF24842374E3F8B9" ID-GBIF-Occurrence="4953610304" box="[151,931,1807,1834]" collectingDate="1930-01" collectionCode="AM" collectorName="H. R. Rabone" country="Tonga" location="Tonga" pageId="15" pageNumber="230" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B918EAC2CE7B9862FF248423764DF8B9" bold="true" box="[151,269,1807,1833]" pageId="15" pageNumber="230">
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<typeStatus id="54D78872CE7B9862FF2484237648F8B9" box="[151,264,1807,1833]" pageId="15" pageNumber="230" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCount id="9D6AFD59CE7B9862FEA7843C7669F8BA" box="[276,297,1808,1834]" count="1" pageId="15" pageNumber="230" type="male">♂</specimenCount>
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(K.62615),
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<collectingCountry id="F37B7640CE7B9862FE01843C76BAF8BA" box="[434,506,1808,1834]" name="Tonga" pageId="15" pageNumber="230">Tonga</collectingCountry>
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,
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<date id="FFD21010CE7B9862FDB6843C752DF8B9" box="[517,621,1807,1834]" pageId="15" pageNumber="230" value="1930-01">
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<collectingDate id="EF96E9F8CE7B9862FDB6843C752DF8B9" box="[517,621,1807,1834]" pageId="15" pageNumber="230" value="1930-01">Jan 1930</collectingDate>
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</date>
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,
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<collectorName id="26995306CE7B9862FDCB843C744AF8B9" box="[632,778,1807,1834]" pageId="15" pageNumber="230">H.R. Rabone</collectorName>
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leg. (in
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<collectionCode id="ED7DAE15CE7B9862FCD6842374D7F8B9" box="[869,919,1807,1833]" country="Australia" name="Australian Museum" pageId="15" pageNumber="230" type="Museum">AM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C376655BCE7B9862FF74841F761EF8E2" pageId="15" pageNumber="230" type="etymology">
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<paragraph id="8BD336D0CE7B9862FF74841F761EF8E2" blockId="15.[151,1436,1807,1906]" pageId="15" pageNumber="230">
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<emphasis id="B918EAC2CE7B9862FF74841F760EF8DD" bold="true" box="[199,334,1843,1869]" pageId="15" pageNumber="230">Etymology.</emphasis>
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The name refers to the
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<typeStatus id="54D78872CE7B9862FDEF841875CCF8DE" box="[604,652,1844,1870]" pageId="15" pageNumber="230">type</typeStatus>
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locality, the island of
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<collectingCountry id="F37B7640CE7B9862FC3A84187492F8DE" box="[905,978,1844,1870]" name="Tonga" pageId="15" pageNumber="230">Tonga</collectingCountry>
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; tonga is used as a noun in apposition to the generic name.
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</paragraph>
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</subSubSection>
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<caption id="DF136658CE64987DFF2484BB74B6F844" ID-DOI="http://doi.org/10.5281/zenodo.13915800" ID-Zenodo-Dep="13915800" httpUri="https://zenodo.org/record/13915800/files/figure.png" pageId="16" pageNumber="231" startId="16.[151,264,1943,1967]" targetBox="[204,1384,181,1919]" targetPageId="16" targetType="figure">
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<paragraph id="8BD336D0CE64987DFF2484BB74B6F844" blockId="16.[151,1436,1943,2004]" pageId="16" pageNumber="231">
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<emphasis id="B918EAC2CE64987DFF2484BB763EF83F" bold="true" box="[151,382,1943,1967]" pageId="16" pageNumber="231">FIGURES 48–49. 48.</emphasis>
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<emphasis id="B918EAC2CE64987DFE3684B47542F83F" box="[389,514,1944,1967]" italics="true" pageId="16" pageNumber="231">
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<taxonomicName id="4C6C4D53CE64987DFE3684B476BDF83F" authorityName="Theischinger & Kalnins & Marinov" authorityYear="2024" box="[389,509,1944,1967]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="231" phylum="Arthropoda" rank="species" species="tonga">Anax tonga</taxonomicName>
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,
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</emphasis>
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male, holotype, dorsal: (a) habitus; (b) head; (c) S2–3; (d) terminalia.
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<emphasis id="B918EAC2CE64987DFB5384BB7240F83F" bold="true" box="[1248,1280,1943,1967]" pageId="16" pageNumber="231">49.</emphasis>
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<emphasis id="B918EAC2CE64987DFABB84B472DCF83F" box="[1288,1436,1944,1967]" italics="true" pageId="16" pageNumber="231">
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<taxonomicName id="4C6C4D53CE64987DFABB84B472D8F83F" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[1288,1432,1944,1967]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="231" phylum="Arthropoda" rank="species" species="guttatus">Anax guttatus</taxonomicName>
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,
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</emphasis>
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female (K.62616), from Tonga, dorsal: (a) habitus; (b) head; (c) S2–3; (d) terminalia.
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</paragraph>
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</caption>
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<subSubSection id="C376655BCE65987CFF2483BB76DEFB79" pageId="17" pageNumber="232" type="description">
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<paragraph id="8BD336D0CE65987CFF2483BB76FBFF21" blockId="17.[151,443,151,177]" box="[151,443,151,177]" pageId="17" pageNumber="232">
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<emphasis id="B918EAC2CE65987CFF2483BB76FBFF21" bold="true" box="[151,443,151,177]" pageId="17" pageNumber="232">
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Description of
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<typeStatus id="54D78872CE65987CFEFC83BB76FBFF21" box="[335,443,151,177]" pageId="17" pageNumber="232" type="holotype">holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="8BD336D0CE65987CFF2483F37209FE19" blockId="17.[151,1437,223,1942]" pageId="17" pageNumber="232">
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<emphasis id="B918EAC2CE65987CFF2483F3779BFF69" box="[151,219,223,249]" italics="true" pageId="17" pageNumber="232">Head.</emphasis>
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Labium and bases of mandibles pale to dark brown; remainder of mandibles and genae black; labrum basally brown, merging ventrally into brownish black; anteclypeus, postclypeus and widely angulate (c. 160–170°) antefrons pale orange brown; top of frons with a brownish black thick-stemmed T-mark contrasting with the brownish yellow ground area, black at base of ‘T’ extending laterally and ventrally, transverse anterior bar of ‘T’ laterally not well defined; antennae, vertex, occipital triangle and what can be detected of postgenae black. Eyes dark brown.
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</paragraph>
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<paragraph id="8BD336D0CE65987CFF7482BF779BFD89" blockId="17.[151,1437,223,1942]" pageId="17" pageNumber="232">
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<emphasis id="B918EAC2CE65987CFF7482BF7660FE3D" box="[199,288,403,429]" italics="true" pageId="17" pageNumber="232">Thorax.</emphasis>
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Prothorax largely brownish black and greyish brown. Pterothorax with pleura and antealar sinus pale orange brown without any distinct pattern recognizable. Terga largely yellowish brown to greyish brown. Sterna pale greyish brown. Legs: coxae, trochanters and base of femora brown, remainder of femora, tibiae, tarsi and claws black.
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</paragraph>
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<paragraph id="8BD336D0CE65987CFF74810874F9FD61" blockId="17.[151,1437,223,1942]" pageId="17" pageNumber="232">
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<emphasis id="B918EAC2CE65987CFF7481087653FDAD" box="[199,275,548,573]" italics="true" pageId="17" pageNumber="232">Wings.</emphasis>
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Humeral and axillary plates partly brownish black, partly yellowish grey. Venation greyish brown to black, costa with no median ray detectable. Membrane largely hyaline, Hw tinged very slightly pale brownish between anal loop and MA. Nodal index 12–22 / 22–
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<quantity id="4C949B35CE65987CFD468147746DFD15" box="[757,813,619,645]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="17" pageNumber="232" unit="in" value="12.0">12 in</quantity>
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Fw and 11–15 / 15–
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<quantity id="4C949B35CE65987CFBAB81477312FD15" box="[1048,1106,619,646]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="17" pageNumber="232" unit="in" value="12.0">12 in</quantity>
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Hw; Ax1 and Ax8 thickened in both wings; wing triangles with 6 cells in Fw, 5 cells in Hw; Arc between level of Ax2 and Ax3; Pt 4.5–5.0 mm, dark greyish brown, overlying 2–3 cells; 7–8 double cells between MP and CuA of Hw; anal loop with 11–13 cells; 4 rows of cells between dark greyish brown membranule and anal loop.
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</paragraph>
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<paragraph id="8BD336D0CE65987CFF7481D7749DFC7D" blockId="17.[151,1437,223,1942]" pageId="17" pageNumber="232">
|
||
<emphasis id="B918EAC2CE65987CFF7481D77679FC85" box="[199,313,763,789]" italics="true" pageId="17" pageNumber="232">Abdomen.</emphasis>
|
||
Dorsally largely black, ventrally in different shades of brown. S3 Index c. 5.0. S1 blackish brown; S2 with orange basal ring (presumed green in life) covering approximately 1/6 of segment length, its median 1/5 produced backward in a small distinct semicircle; remainder with small ill-defined yellow patches (presumed blue in life) each side of large black strongly pimpled median area; S3 black with short pale orange basal mid-dorsal patch and pair of barely detectable brownish latero-basal smudges and pair of subapical dorsolateral dull yellow spots; S4–9 black, laterally with only few barely detectable brownish smudges and with only pair of dull yellow subapical dorso-lateral spots each; S9 and S10 appearing unmarked, black.
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE65987CFF7480DB75EAFB31" blockId="17.[151,1437,223,1942]" pageId="17" pageNumber="232">
|
||
<emphasis id="B918EAC2CE65987CFF7480DB7609FB81" box="[199,329,1015,1041]" italics="true" pageId="17" pageNumber="232">Terminalia.</emphasis>
|
||
Cerci brown to black, with stalk moderately thick, short, slightly incurved; at least basal 2/3 of the remainder maximally almost twice as wide as in-curved tapered apical 1/3, the narrowing rather abruptly; apico-lateral spike also slightly in-curved and tiny, wide-based basal, inferior tooth. Epiproct largely brown, lateral edges black; basally markedly wider than long but barely 1/4 the length of cerci, apex very widely and shallowly emarginate; two pairs of small dorsal tubercles.
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE65987CFF7487807362FB56" blockId="17.[151,1437,223,1942]" box="[199,1058,1195,1222]" pageId="17" pageNumber="232">
|
||
<emphasis id="B918EAC2CE65987CFF7487807690FB55" box="[199,464,1195,1221]" italics="true" pageId="17" pageNumber="232">Measurements (in mm).</emphasis>
|
||
Abdomen (including cerci) 63.1, cerci 6.3, Hw 53.5.
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE65987CFF7487E376DEFB79" blockId="17.[151,1437,223,1942]" box="[199,414,1231,1257]" pageId="17" pageNumber="232">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376655BCE65987CFF7487DF77BEF8B9" pageId="17" pageNumber="232" type="diagnosis">
|
||
<paragraph id="8BD336D0CE65987CFF7487DF77BEF8B9" blockId="17.[151,1437,223,1942]" pageId="17" pageNumber="232">
|
||
<emphasis id="B918EAC2CE65987CFF7487DF7602FA9D" bold="true" box="[199,322,1267,1293]" pageId="17" pageNumber="232">Diagnosis.</emphasis>
|
||
A large dark rather short-bodied
|
||
<taxonomicName id="4C6C4D53CE65987CFD1E87DF75A7FA9D" authorityName="Leach" authorityYear="1815" box="[685,743,1267,1293]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="232" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EAC2CE65987CFD1E87DF75A7FA9D" box="[685,743,1267,1293]" italics="true" pageId="17" pageNumber="232">Anax</emphasis>
|
||
</taxonomicName>
|
||
(body length c.
|
||
<quantity id="4C949B35CE65987CFC2B87DF74A8FA9E" box="[920,1000,1267,1294]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="17" pageNumber="232" unit="mm" value="85.0">85 mm</quantity>
|
||
, wingspan c.
|
||
<quantity id="4C949B35CE65987CFBCF87DF7398FA9D" box="[1148,1240,1267,1293]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="232" unit="mm" value="110.0">110 mm</quantity>
|
||
) with dark brown to black T-mark (top bar ill-defined) on largely yellow top of frons and with high S3 Index (male c. 5.0). Mature male, preserved (
|
||
<figureCitation id="13572A55CE65987CFEEA861776ECFAC5" box="[345,428,1339,1365]" captionStart="FIGURES 48–49. 48" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[204,1384,181,1919]" captionTargetId="figure-22@16.[204,1384,181,1919]" captionTargetPageId="16" captionText="FIGURES 48–49. 48. Anax tonga, male, holotype, dorsal: (a) habitus; (b) head; (c) S2–3; (d) terminalia. 49. Anax guttatus, female (K.62616), from Tonga, dorsal: (a) habitus; (b) head; (c) S2–3; (d) terminalia." figureDoi="http://doi.org/10.5281/zenodo.13915800" httpUri="https://zenodo.org/record/13915800/files/figure.png" pageId="17" pageNumber="232">Fig. 48</figureCitation>
|
||
) with head and thorax mostly pale orange brown (expected to be green in life), abdomen base pale orange brown and black, remainder of abdomen largely black. Abdominal pattern generally made up by pairs of very small ill-defined dull pale yellow subapical dorso-lateral patches each on at least S3–8 or 9, none on S1, S2 and S10 and possibly S9. According to specifications for Papuan
|
||
<taxonomicName id="4C6C4D53CE65987CFC72868B7312FA51" box="[961,1106,1447,1473]" pageId="17" pageNumber="232">
|
||
<emphasis id="B918EAC2CE65987CFC72868B74BBFA51" box="[961,1019,1447,1473]" italics="true" pageId="17" pageNumber="232">Anax</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFFD4B21CE65987CFBCF868B723EFA52" author="Orr, A. G. & Kalkman, V. J." box="[1148,1406,1447,1474]" pageId="17" pageNumber="232" pagination="3 - 156" refId="ref12885" refString="Orr, A. G. & Kalkman, V. J. (2015) Field guide to the dragonflies of New Guinea. Brachytron, 17 (Supplement), 3 - 156." type="journal article" year="2015">Orr & Kalkman (2015)</bibRefCitation>
|
||
<taxonomicName id="4C6C4D53CE65987CFA37868B7797FA75" authorityName="Theischinger & Kalnins & Marinov" authorityYear="2024" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="232" phylum="Arthropoda" rank="species" species="tonga">
|
||
<emphasis id="B918EAC2CE65987CFA37868B7797FA75" italics="true" pageId="17" pageNumber="232">A. tonga</emphasis>
|
||
</taxonomicName>
|
||
is a shorter-bodied species (Abd
|
||
<quantity id="4C949B35CE65987CFDF986E775DAFA76" box="[586,666,1483,1510]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="17" pageNumber="232" unit="mm" value="63.0">63 mm</quantity>
|
||
, Hw
|
||
<quantity id="4C949B35CE65987CFD6786E77466FA76" box="[724,806,1483,1510]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="17" pageNumber="232" unit="mm" value="54.0">54 mm</quantity>
|
||
; ratio <1.3). With its long narrow waist the male appears most similar to
|
||
<taxonomicName id="4C6C4D53CE65987CFEFA86C3769CF999" authorityName="Lieftinck" authorityYear="1942" box="[329,476,1519,1545]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="232" phylum="Arthropoda" rank="species" species="celebensis">
|
||
<emphasis id="B918EAC2CE65987CFEFA86C3769CF999" box="[329,476,1519,1545]" italics="true" pageId="17" pageNumber="232">A. celebensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C4D53CE65987CFE5886C37536F999" authorityName="Theischinger & Kalnins & Marinov" authorityYear="2024" box="[491,630,1519,1545]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="232" phylum="Arthropoda" rank="species" species="insulanus">
|
||
<emphasis id="B918EAC2CE65987CFE5886C37536F999" box="[491,630,1519,1545]" italics="true" pageId="17" pageNumber="232">A. insulanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C4D53CE65987CFD1C86C37473F999" authorityName="Kennedy" authorityYear="1934" box="[687,819,1519,1545]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="232" phylum="Arthropoda" rank="species" species="piraticus">
|
||
<emphasis id="B918EAC2CE65987CFD1C86C37473F999" box="[687,819,1519,1545]" italics="true" pageId="17" pageNumber="232">A. piraticus</emphasis>
|
||
</taxonomicName>
|
||
. It can be distinguished from all of them by the apical 1/4 to 1/3 of the cerci noticeably arched inward and its tiny apico-lateral spike vs apical 1/3 of the cerci straight and with its apico-lateral spike larger and not in-curved. It can also be separated from
|
||
<taxonomicName id="4C6C4D53CE65987CFB9E851B7380F9C1" authorityName="Lieftinck" authorityYear="1942" box="[1069,1216,1591,1617]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="232" phylum="Arthropoda" rank="species" species="celebensis">
|
||
<emphasis id="B918EAC2CE65987CFB9E851B7380F9C1" box="[1069,1216,1591,1617]" italics="true" pageId="17" pageNumber="232">A. celebensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C4D53CE65987CFB45851B723AF9C1" authorityName="Kennedy" authorityYear="1934" box="[1270,1402,1591,1617]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="232" phylum="Arthropoda" rank="species" species="piraticus">
|
||
<emphasis id="B918EAC2CE65987CFB45851B723AF9C1" box="[1270,1402,1591,1617]" italics="true" pageId="17" pageNumber="232">A. piraticus</emphasis>
|
||
</taxonomicName>
|
||
by its very dark abdomen with only a few barely detectable pale smudges and a pair of very small whitish yellow spots on at least S3–8 or 9 vs abdomen with much richer and larger pale patching. It can also be distinguished from the male of
|
||
<taxonomicName id="4C6C4D53CE65987CFF40858F763DF92D" authorityName="Theischinger & Kalnins & Marinov" authorityYear="2024" box="[243,381,1699,1725]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="232" phylum="Arthropoda" rank="species" species="insulanus" status="sp. nov.">
|
||
<emphasis id="B918EAC2CE65987CFF40858F763DF92D" box="[243,381,1699,1725]" italics="true" pageId="17" pageNumber="232">A. insulanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B918EAC2CE65987CFE36858F769FF92D" bold="true" box="[389,479,1699,1725]" pageId="17" pageNumber="232">
|
||
<taxonomicNameLabel id="A22B57B9CE65987CFE36858F769FF92D" box="[389,479,1699,1725]" pageId="17" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the most similar species, by the presence in the male of a pair of dull whitish yellow subapical dorso-lateral spots in S3 and possibly S4, by a markedly wider abdomen (approximately
|
||
<quantity id="4C949B35CE65987CFB5985EB7200F972" box="[1258,1344,1735,1762]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="17" pageNumber="232" unit="mm" value="4.2">4.2 mm</quantity>
|
||
wide vs
|
||
<quantity id="4C949B35CE65987CFF2485C777BBF896" box="[151,251,1771,1798]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" pageId="17" pageNumber="232" unit="mm" value="3.85">3.85 mm</quantity>
|
||
wide at S9, and an S3 Index of 5 versus 6.25) and by an abruptly vs continuously narrowed apical third of the cerci.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376655BCE65987CFF74841F77A4F806" pageId="17" pageNumber="232" type="distribution">
|
||
<paragraph id="8BD336D0CE65987CFF74841F77A4F806" blockId="17.[151,1437,223,1942]" pageId="17" pageNumber="232">
|
||
<emphasis id="B918EAC2CE65987CFF74841F7621F8DD" bold="true" box="[199,353,1843,1869]" pageId="17" pageNumber="232">Distribution.</emphasis>
|
||
Known only from
|
||
<collectingCountry id="F37B7640CE65987CFDF6841875CDF8DE" box="[581,653,1844,1870]" name="Tonga" pageId="17" pageNumber="232">Tonga</collectingCountry>
|
||
, where it and a female of
|
||
<taxonomicName id="4C6C4D53CE65987CFC74841F7304F8DD" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[967,1092,1843,1869]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="232" phylum="Arthropoda" rank="species" species="guttatus">
|
||
<emphasis id="B918EAC2CE65987CFC74841F7304F8DD" box="[967,1092,1843,1869]" italics="true" pageId="17" pageNumber="232">A. guttatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13572A55CE65987CFBE4841F73ECF8DD" box="[1111,1196,1843,1869]" captionStart="FIGURES 48–49. 48" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[204,1384,181,1919]" captionTargetId="figure-22@16.[204,1384,181,1919]" captionTargetPageId="16" captionText="FIGURES 48–49. 48. Anax tonga, male, holotype, dorsal: (a) habitus; (b) head; (c) S2–3; (d) terminalia. 49. Anax guttatus, female (K.62616), from Tonga, dorsal: (a) habitus; (b) head; (c) S2–3; (d) terminalia." figureDoi="http://doi.org/10.5281/zenodo.13915800" httpUri="https://zenodo.org/record/13915800/files/figure.png" pageId="17" pageNumber="232">Fig. 49</figureCitation>
|
||
) from
|
||
<collectingCountry id="F37B7640CE65987CFB4C84187207F8DE" box="[1279,1351,1844,1870]" name="Tonga" pageId="17" pageNumber="232">Tonga</collectingCountry>
|
||
, Tubau College, Nafualu,
|
||
<collectingRegion id="49A8F832CE65987CFED4847476A0F8E2" box="[359,480,1880,1906]" country="Tonga" name="Tongatapu" pageId="17" pageNumber="232">Tongatapu</collectingRegion>
|
||
,
|
||
<date id="FFD21010CE65987CFE5F84747515F8E1" box="[492,597,1879,1906]" pageId="17" pageNumber="232" value="1930-01">Jan 1930</date>
|
||
, H.R. Rabone leg. (in AM: K.62616) were collected. Possibly endemic to
|
||
<collectingCountry id="F37B7640CE65987CFF24845077A0F806" box="[151,224,1916,1942]" name="Tonga" pageId="17" pageNumber="232">Tonga</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |