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<mods:title id="3BCAB1F3BEE0896C9A690C2ED60A5708">Taxonomic remarks, phylogeny and evolutionary notes on the leaf beetle species belonging to the Cryptocephalus sericeus complex (Coleoptera: Chrysomelidae: Cryptocephalinae)</mods:title>
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<subSubSection id="C3766520FFBD615FB89AFB70FD54FB67" box="[151,754,1193,1219]" pageId="1" pageNumber="334" type="nomenclature">
<paragraph id="8BD336ABFFBD615FB89AFB70FD54FB67" blockId="1.[151,754,1193,1219]" box="[151,754,1193,1219]" pageId="1" pageNumber="334">
<heading id="D09B81C7FFBD615FB89AFB70FD54FB67" bold="true" box="[151,754,1193,1219]" fontSize="11" level="1" pageId="1" pageNumber="334" reason="1">
<emphasis id="B918EAB9FFBD615FB89AFB70FD54FB67" bold="true" box="[151,754,1193,1219]" pageId="1" pageNumber="334">
<taxonomicName id="4C6C4D28FFBD615FB89AFB70FEF2FB67" ID-CoL="3WHF" box="[151,340,1193,1219]" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBD615FB89AFB70FEF2FB67" bold="true" box="[151,340,1193,1219]" italics="true" pageId="1" pageNumber="334">Cryptocephalus</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="4C6C4D28FFBD615FB9DBFB70FD54FB67" authority="Burlini, 1969" authorityName="Burlini" authorityYear="1969" box="[470,754,1193,1219]" class="Insecta" family="Chrysomelidae" genus="Cerodens" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBD615FB9DBFB70FDEEFB67" bold="true" box="[470,584,1193,1219]" italics="true" pageId="1" pageNumber="334">Cerodens</emphasis>
<bibRefCitation id="EFFD4B5AFFBD615FBA42FB70FD54FB67" author="Burlini" box="[591,754,1193,1219]" pageId="1" pageNumber="334" refString="Burlini, M. (1969) Cryptocephalus subgen. Cryptodontus n., Cryptoc. subgen. Cerodens n. nov. e tabella dei sottogeneri di Cryptocephalus. XXIV contributo alla conoscenza dei Cryptocephalini (Coleoptera Chrysomelidae). Memorie della Societa Entomologica Italiana, 48, 535 - 540." type="journal article" year="1969">Burlini, 1969</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<paragraph id="8BD336ABFFBD615FB89AFB29FDE6FA80" blockId="1.[151,1436,1264,1347]" pageId="1" pageNumber="334">
<treatmentCitationGroup id="AB7C1185FFBD615FB89AFB29FDE6FA80" pageId="1" pageNumber="334">
<taxonomicName id="4C6C4D28FFBD615FB89AFB29FE1DFAA2" authority="Burlini, 1969: 539" authorityName="Burlini" authorityPageNumber="539" authorityYear="1969" box="[151,443,1264,1286]" class="Insecta" family="Chrysomelidae" genus="Cerodens" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBD615FB89AFB29FF51FAA2" box="[151,247,1264,1286]" italics="true" pageId="1" pageNumber="334">Cerodens</emphasis>
<treatmentCitation id="0ACD10BAFFBD615FB8F3FB29FE1DFAA2" author="Burlini" box="[254,443,1264,1286]" page="539" pageId="1" pageNumber="334" year="1969">
<bibRefCitation id="EFFD4B5AFFBD615FB8F3FB29FE2CFAA2" author="Burlini" box="[254,394,1264,1286]" pageId="1" pageNumber="334" refString="Burlini, M. (1969) Cryptocephalus subgen. Cryptodontus n., Cryptoc. subgen. Cerodens n. nov. e tabella dei sottogeneri di Cryptocephalus. XXIV contributo alla conoscenza dei Cryptocephalini (Coleoptera Chrysomelidae). Memorie della Societa Entomologica Italiana, 48, 535 - 540." type="journal article" year="1969">Burlini, 1969</bibRefCitation>
: 539
</treatmentCitation>
</taxonomicName>
[subgenus].
<typeStatus id="54D78809FFBD615FBA4DFB28FDD4FAA2" box="[576,626,1265,1286]" pageId="1" pageNumber="334">Type</typeStatus>
species:
<taxonomicName id="4C6C4D28FFBD615FBADFFB29FBF0FAA2" authority="Burlini, 1953" authorityName="Burlini" authorityYear="1953" box="[722,1110,1264,1286]" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="species" species="kocheri">
<emphasis id="B918EAB9FFBD615FBADFFB29FC60FAA2" box="[722,966,1264,1286]" italics="true" pageId="1" pageNumber="334">Cryptocephalus kocheri</emphasis>
<bibRefCitation id="EFFD4B5AFFBD615FBBC3FB29FBF0FAA2" author="Burlini" box="[974,1110,1264,1286]" pageId="1" pageNumber="334" refString="Burlini, M. (1953) Une nouvelle espece de Cryptocephalus du Maroc: Cryptoc. Kocheri Burlini (Coleoptera Chrysomelidae). Comptes-Rendus des Seances Mensuelles de la Societe des Sciences Naturelles du Maroc, 5, 73 - 75." type="journal article" year="1953">Burlini, 1953</bibRefCitation>
</taxonomicName>
[homonym] (=
<taxonomicName id="4C6C4D28FFBD615FBCF1FB29FE04FA80" authority="Burlini, 1956" authorityName="Burlini" authorityYear="1956" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="species" species="emiliae">
<emphasis id="B918EAB9FFBD615FBCF1FB29FEB4FA80" italics="true" pageId="1" pageNumber="334">Cryptocephalus emiliae</emphasis>
<bibRefCitation id="EFFD4B5AFFBD615FB915FAD7FE04FA80" author="Burlini" box="[280,418,1294,1316]" pageId="1" pageNumber="334" refString="Burlini, M. (1956) Revisione dei Cryptocephalus Italiani e della maggior parte delle specie di Europa (Col. Chrysomelidae). Memorie della Societa Entomologica Italiana, 34 (1955 - 1956), 5 - 287." type="journal article" year="1956">Burlini, 1956</bibRefCitation>
</taxonomicName>
), by monotypy.
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</paragraph>
<paragraph id="8BD336ABFFBD615FB89AFAF4FAD3FAE7" blockId="1.[151,1436,1264,1347]" box="[151,1397,1325,1347]" pageId="1" pageNumber="334">
<treatmentCitationGroup id="AB7C1185FFBD615FB89AFAF4FAD3FAE7" box="[151,1397,1325,1347]" pageId="1" pageNumber="334">
<taxonomicName id="4C6C4D28FFBD615FB89AFAF4FE65FAE7" authority="Burlini, 1953: 75" authorityName="Burlini" authorityPageNumber="75" authorityYear="1953" box="[151,451,1325,1347]" class="Insecta" family="Chrysomelidae" genus="Ceropachys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBD615FB89AFAF4FEA9FAE7" box="[151,271,1325,1347]" italics="true" pageId="1" pageNumber="334">Ceropachys</emphasis>
<treatmentCitation id="0ACD10BAFFBD615FB918FAF4FE65FAE7" author="Burlini" box="[277,451,1325,1347]" page="75" pageId="1" pageNumber="334" year="1953">
<bibRefCitation id="EFFD4B5AFFBD615FB918FAF4FE06FAE7" author="Burlini" box="[277,416,1325,1347]" pageId="1" pageNumber="334" refString="Burlini, M. (1953) Une nouvelle espece de Cryptocephalus du Maroc: Cryptoc. Kocheri Burlini (Coleoptera Chrysomelidae). Comptes-Rendus des Seances Mensuelles de la Societe des Sciences Naturelles du Maroc, 5, 73 - 75." type="journal article" year="1953">Burlini, 1953</bibRefCitation>
: 75
</treatmentCitation>
</taxonomicName>
[subgenus]. Preoccupied genus name, not
<taxonomicName id="4C6C4D28FFBD615FBB79FAF4FBCDFAE7" authority="Costa, 1847" authorityName="Costa" authorityYear="1847" box="[884,1131,1325,1347]" class="Insecta" family="Chrysomelidae" genus="Ceropachys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBD615FBB79FAF4FC4AFAE7" box="[884,1004,1325,1347]" italics="true" pageId="1" pageNumber="334">Ceropachys</emphasis>
Costa, 1847
</taxonomicName>
[
<taxonomicName id="4C6C4D28FFBD615FBC7AFAF4FB4BFAE7" ID-CoL="C2L" box="[1143,1261,1325,1347]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C6C4D28FFBD615FBCFBFAF4FACDFAE7" ID-CoL="626T7" box="[1270,1387,1325,1347]" class="Insecta" family="Zopheridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="family">Colydiidae</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD336ABFFBD615FB89AFAB6FD85F969" blockId="1.[151,1437,1391,2029]" pageId="1" pageNumber="334">
<bibRefCitation id="EFFD4B5AFFBD615FB89AFAB6FE9EFA2C" author="Burlini" box="[151,312,1391,1416]" pageId="1" pageNumber="334" refString="Burlini, M. (1953) Une nouvelle espece de Cryptocephalus du Maroc: Cryptoc. Kocheri Burlini (Coleoptera Chrysomelidae). Comptes-Rendus des Seances Mensuelles de la Societe des Sciences Naturelles du Maroc, 5, 73 - 75." type="journal article" year="1953">Burlini (1953)</bibRefCitation>
established the monotypic subgenus
<taxonomicName id="4C6C4D28FFBD615FBAD1FAA9FCC7FA2C" box="[732,865,1392,1416]" class="Insecta" family="Chrysomelidae" genus="Ceropachys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBD615FBAD1FAA9FCC7FA2C" box="[732,865,1392,1416]" italics="true" pageId="1" pageNumber="334">Ceropachys</emphasis>
</taxonomicName>
for
<taxonomicName id="4C6C4D28FFBD615FBB9DFAA9FB07FA2C" authority="Burlini, 1956" authorityName="Burlini" authorityYear="1956" box="[912,1185,1391,1416]" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="species" species="emiliae">
<emphasis id="B918EAB9FFBD615FBB9DFAA9FBA2FA2C" box="[912,1028,1392,1416]" italics="true" pageId="1" pageNumber="334">C. emiliae</emphasis>
<bibRefCitation id="EFFD4B5AFFBD615FBC06FAB6FB07FA2C" author="Burlini" box="[1035,1185,1391,1416]" pageId="1" pageNumber="334" refString="Burlini, M. (1956) Revisione dei Cryptocephalus Italiani e della maggior parte delle specie di Europa (Col. Chrysomelidae). Memorie della Societa Entomologica Italiana, 34 (1955 - 1956), 5 - 287." type="journal article" year="1956">Burlini, 1956</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="4C6C4D28FFBD615FBCC4FAA9FF76FA09" authority="Burlini, 1953" authorityName="Burlini" authorityYear="1953" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="species" species="kocheri">
<emphasis id="B918EAB9FFBD615FBCC4FAA9FA98FA2C" box="[1225,1342,1392,1416]" italics="true" pageId="1" pageNumber="334">C. kocheri</emphasis>
<bibRefCitation id="EFFD4B5AFFBD615FBD48FAB6FF76FA09" author="Burlini" pageId="1" pageNumber="334" refString="Burlini, M. (1953) Une nouvelle espece de Cryptocephalus du Maroc: Cryptoc. Kocheri Burlini (Coleoptera Chrysomelidae). Comptes-Rendus des Seances Mensuelles de la Societe des Sciences Naturelles du Maroc, 5, 73 - 75." type="journal article" year="1953">Burlini, 1953</bibRefCitation>
</taxonomicName>
) on the basis of the peculiar shape of the antennomeres (fig. 5a), which in this species are significantly shortened, flattened and angled along the inner margin.
<bibRefCitation id="EFFD4B5AFFBD615FBB21FA6EFC77FA74" author="Burlini" box="[812,977,1463,1488]" pageId="1" pageNumber="334" refString="Burlini, M. (1969) Cryptocephalus subgen. Cryptodontus n., Cryptoc. subgen. Cerodens n. nov. e tabella dei sottogeneri di Cryptocephalus. XXIV contributo alla conoscenza dei Cryptocephalini (Coleoptera Chrysomelidae). Memorie della Societa Entomologica Italiana, 48, 535 - 540." type="journal article" year="1969">Burlini (1969)</bibRefCitation>
changed the name to
<taxonomicName id="4C6C4D28FFBD615FBCD1FA61FAE0FA74" box="[1244,1350,1464,1488]" class="Insecta" family="Chrysomelidae" genus="Cerodens" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBD615FBCD1FA61FAE0FA74" box="[1244,1350,1464,1488]" italics="true" pageId="1" pageNumber="334">Cerodens</emphasis>
</taxonomicName>
due to homonymy with
<taxonomicName id="4C6C4D28FFBD615FB96CFA05FD26FA50" authority="Costa, 1847" authorityName="Costa" authorityYear="1847" box="[353,640,1500,1524]" class="Insecta" family="Chrysomelidae" genus="Ceropachys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBD615FB96CFA05FE40FA50" box="[353,486,1500,1524]" italics="true" pageId="1" pageNumber="334">Ceropachys</emphasis>
Costa, 1847
</taxonomicName>
. In Palaearctic species of
<taxonomicName id="4C6C4D28FFBD615FBBB1FA05FBCBFA50" ID-CoL="3WHF" box="[956,1133,1500,1524]" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBD615FBBB1FA05FBCBFA50" box="[956,1133,1500,1524]" italics="true" pageId="1" pageNumber="334">Cryptocephalus</emphasis>
</taxonomicName>
antennomeres are mostly cylindrical, but their shape and length is rather variable in some unrelated groups (
<bibRefCitation id="EFFD4B5AFFBD615FBC5AFA26FAFDF9BC" author="Vela" box="[1111,1371,1535,1560]" pageId="1" pageNumber="334" refString="Vela, J. M. &amp; Bastazo, G. (2012) A new Cryptocephalus of the C. curvilinea- group (Coleoptera, Chrysomelidae) from the Atlantic coastal marshes of Southern Spain and Portugal. Boletin de la Asociacion Espanola de Entomologia, 36 (1 - 2), 31 - 41." type="journal article" year="2012">Vela &amp; Bastazo, 2012</bibRefCitation>
). The adaptive significance of this variability, if any, remains unknown. Even if in
<taxonomicName id="4C6C4D28FFBD615FBC33F9FDFB1EF998" box="[1086,1208,1572,1596]" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="species" species="emiliae">
<emphasis id="B918EAB9FFBD615FBC33F9FDFB1EF998" box="[1086,1208,1572,1596]" italics="true" pageId="1" pageNumber="334">C. emiliae</emphasis>
</taxonomicName>
this trait could be considered an utmost in a trend of shortening antennal articles, it is questionable that this condition is so distinct as to justify a separation at subgeneric level (fig. 5a vs. figs 5bh). This species is undoubtedly placed within the
<taxonomicName id="4C6C4D28FFBD615FBD8FF9B5FF55F903" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="species" species="sericeus">
<emphasis id="B918EAB9FFBD615FBD8FF9B5FF55F903" italics="true" pageId="1" pageNumber="334">C. sericeus</emphasis>
</taxonomicName>
species complex based on molecular data (
<bibRefCitation id="EFFD4B5AFFBD615FBADEF956FC5FF90C" author="Gomez-Zurita" box="[723,1017,1679,1704]" pageId="1" pageNumber="334" refString="Gomez-Zurita J., Sassi, D., Cardoso, A. &amp; Balke, M. (2011) Evolution of Cryptocephalus leaf beetles related to C. sericeus (Coleoptera: Chrysomelidae) and the role of hybridization in generating species mtDNA paraphyly. Zoologica Scripta, 41, 47 - 67. http: // dx. doi. org / 10.1111 / j. 1463 - 6409.2011.00500. x" type="journal article" year="2011">
Gómez-Zurita
<emphasis id="B918EAB9FFBD615FBB77F948FC17F90C" box="[890,945,1680,1704]" italics="true" pageId="1" pageNumber="334">et al.</emphasis>
, 2011
</bibRefCitation>
), thus the subgenus
<taxonomicName id="4C6C4D28FFBD615FBCD4F949FAE5F90C" box="[1241,1347,1680,1704]" class="Insecta" family="Chrysomelidae" genus="Cerodens" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBD615FBCD4F949FAE5F90C" box="[1241,1347,1680,1704]" italics="true" pageId="1" pageNumber="334">Cerodens</emphasis>
</taxonomicName>
renders
<taxonomicName id="4C6C4D28FFBD615FB89AF96DFEEFF968" ID-CoL="3WHF" box="[151,329,1716,1740]" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B918EAB9FFBD615FB89AF96DFEEFF968" box="[151,329,1716,1740]" italics="true" pageId="1" pageNumber="334">Cryptocephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57C2FFBD615FB942F96CFE2CF968" box="[335,394,1717,1740]" pageId="1" pageNumber="334" sensu="stricto">s. str.</taxonomicNameLabel>
paraphyletic.
</paragraph>
<paragraph id="8BD336ABFFBD615FB8CAF90EFA3AF849" blockId="1.[151,1437,1391,2029]" pageId="1" pageNumber="334">
Besides, on the basis of more traditional considerations, it seems inappropriate to keep the subgenus
<taxonomicName id="4C6C4D28FFBD615FBD3FF901FA3AF954" box="[1330,1436,1752,1776]" class="Insecta" family="Chrysomelidae" genus="Cerodens" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBD615FBD3FF901FA3AF954" box="[1330,1436,1752,1776]" italics="true" pageId="1" pageNumber="334">Cerodens</emphasis>
</taxonomicName>
as valid. The recommendations of
<bibRefCitation id="EFFD4B5AFFBD615FBA21F925FD7AF8B1" author="Platnick" box="[556,732,1788,1813]" pageId="1" pageNumber="334" refString="Platnick, N. I. (1976) Are monotypic genera possible? Systematic Zoology, 25 (2), 198 - 199. http: // dx. doi. org / 10.2307 / 2412749" type="journal article" year="1976">Platnick (1976)</bibRefCitation>
and
<bibRefCitation id="EFFD4B5AFFBD615FBB17F925FC14F8B0" author="Wiley" box="[794,946,1788,1813]" pageId="1" pageNumber="334" refString="Wiley, E. O. (1981) Phylogenetics. The Theory and Practice of Phylogenetic Systematics. John Wiley &amp; Sons. New York- Chichester- Brisbane-Toronto-Singapore, 439 pp." type="book" year="1981">Wiley (1981)</bibRefCitation>
sound particularly suitable for the matter. Supraspecific taxa must express exactly the evolutionary relationships (
<bibRefCitation id="EFFD4B5AFFBD615FBBBEF8C6FB99F89C" author="Wiley" box="[947,1087,1823,1848]" pageId="1" pageNumber="334" refString="Wiley, E. O. (1981) Phylogenetics. The Theory and Practice of Phylogenetic Systematics. John Wiley &amp; Sons. New York- Chichester- Brisbane-Toronto-Singapore, 439 pp." type="book" year="1981">Wiley, 1981</bibRefCitation>
). In the Darwinian paradigm of descent of species by divergence from a common ancestor, “any existing species must have at least one existing or extinct sister species … [therefore] … “monotypic [sub]genera seem impossible as they must always exclude at least one other species that is a descendent of the most recent ancestor (i. e. they must always be paraphyletic)” (
<bibRefCitation id="EFFD4B5AFFBD615FB893F869FEE1F86D" author="Platnick" box="[158,327,1968,1993]" pageId="1" pageNumber="334" refString="Platnick, N. I. (1976) Are monotypic genera possible? Systematic Zoology, 25 (2), 198 - 199. http: // dx. doi. org / 10.2307 / 2412749" type="journal article" year="1976">Platnick, 1976</bibRefCitation>
). Besides, according to
<bibRefCitation id="EFFD4B5AFFBD615FBA5BF869FD49F86C" author="Wiley" box="[598,751,1968,1993]" pageId="1" pageNumber="334" refString="Wiley, E. O. (1981) Phylogenetics. The Theory and Practice of Phylogenetic Systematics. John Wiley &amp; Sons. New York- Chichester- Brisbane-Toronto-Singapore, 439 pp." type="book" year="1981">Wiley (1981)</bibRefCitation>
, redundant taxa (i. e. taxa adding no additional evolutionary information) should not be produced. For all these reasons, I consider appropriate to propose
<taxonomicName id="4C6C4D28FFBD615FBCA5F80DFAFFF848" ID-CoL="3WHF" box="[1192,1369,2004,2028]" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBD615FBCA5F80DFAFFF848" box="[1192,1369,2004,2028]" italics="true" pageId="1" pageNumber="334">Cryptocephalus</emphasis>
</taxonomicName>
subg.
</paragraph>
<paragraph id="8BD336ABFFBE615CB89AFF41FE2CFF71" blockId="2.[151,1436,151,213]" pageId="2" pageNumber="335">
<taxonomicName id="4C6C4D28FFBE615CB89AFF41FE38FF14" authority="Burlini, 1969" authorityName="Burlini" authorityYear="1969" box="[151,414,151,176]" class="Insecta" family="Chrysomelidae" genus="Cerodens" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBE615CB89AFF41FEA7FF14" box="[151,257,152,176]" italics="true" pageId="2" pageNumber="335">Cerodens</emphasis>
<bibRefCitation id="EFFD4B5AFFBE615CB905FF4EFE38FF14" author="Burlini" box="[264,414,151,176]" pageId="2" pageNumber="335" refString="Burlini, M. (1969) Cryptocephalus subgen. Cryptodontus n., Cryptoc. subgen. Cerodens n. nov. e tabella dei sottogeneri di Cryptocephalus. XXIV contributo alla conoscenza dei Cryptocephalini (Coleoptera Chrysomelidae). Memorie della Societa Entomologica Italiana, 48, 535 - 540." type="journal article" year="1969">Burlini, 1969</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="4C6C4D28FFBE615CB9CBFF41FDDEFF14" ID-CoL="3WHF" box="[454,632,152,176]" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBE615CB9CBFF41FDDEFF14" box="[454,632,152,176]" italics="true" pageId="2" pageNumber="335">Cryptocephalus</emphasis>
</taxonomicName>
subg.
<taxonomicName id="4C6C4D28FFBE615CBACFFF41FC43FF14" authority="Burlini, 1953" authorityName="Burlini" authorityYear="1953" box="[706,997,151,176]" class="Insecta" family="Chrysomelidae" genus="Ceropachys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB9FFBE615CBACFFF41FCE1FF14" box="[706,839,152,176]" italics="true" pageId="2" pageNumber="335">Ceropachys</emphasis>
<bibRefCitation id="EFFD4B5AFFBE615CBB43FF4EFC43FF14" author="Burlini" box="[846,997,151,176]" pageId="2" pageNumber="335" refString="Burlini, M. (1953) Une nouvelle espece de Cryptocephalus du Maroc: Cryptoc. Kocheri Burlini (Coleoptera Chrysomelidae). Comptes-Rendus des Seances Mensuelles de la Societe des Sciences Naturelles du Maroc, 5, 73 - 75." type="journal article" year="1953">Burlini, 1953</bibRefCitation>
</taxonomicName>
) as
<emphasis id="B918EAB9FFBE615CBC1EFF4EFBC6FF14" bold="true" box="[1043,1120,151,176]" pageId="2" pageNumber="335">n. syn.</emphasis>
of
<emphasis id="B918EAB9FFBE615CBC8BFF41FB06FF0B" box="[1158,1184,152,175]" italics="true" pageId="2" pageNumber="335">C.</emphasis>
subg.
<taxonomicName id="4C6C4D28FFBE615CBCE6FF41FA3AFF14" ID-CoL="3WHF" box="[1259,1436,152,176]" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="335" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B918EAB9FFBE615CBCE6FF41FA3AFF14" box="[1259,1436,152,176]" italics="true" pageId="2" pageNumber="335">Cryptocephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57C2FFBE615CB89AFF64FF74FF70" box="[151,210,189,212]" pageId="2" pageNumber="335" sensu="stricto">s. str.</taxonomicNameLabel>
Geoffroy, 1762.
</paragraph>
</subSubSection>
</treatment>
</document>