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<mods:title id="DADD2FC913EC7485C812842A0AFC98CB">New records and four new species of Australian Thripidae (Thysanoptera) emphasise faunal relationships between northern Australia and Asia</mods:title>
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<paragraph id="8BD336A0FFEE513120FBA172FE12FC9B" blockId="5.[151,429,835,862]" box="[151,429,835,862]" pageId="5" pageNumber="40">
<heading id="D09B81CCFFEE513120FBA172FE12FC9B" bold="true" box="[151,429,835,862]" fontSize="11" level="1" pageId="5" pageNumber="40" reason="1">
<taxonomicName id="4C6C4D23FFEE513120FBA172FE12FC9B" ID-CoL="78DW" authority="Karny" authorityName="Karny" box="[151,429,835,862]" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB2FFEE513120FBA172FE12FC9B" bold="true" box="[151,429,835,862]" pageId="5" pageNumber="40">
<emphasis id="B918EAB2FFEE513120FBA172FEEBFC98" bold="true" box="[151,340,836,862]" italics="true" pageId="5" pageNumber="40">Rhamphothrips</emphasis>
Karny
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8BD336A0FFEE513120FBA1BCFC66FC66" blockId="5.[151,985,906,928]" box="[151,985,906,928]" pageId="5" pageNumber="40">
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<taxonomicName id="4C6C4D23FFEE513120FBA1BCFE52FC66" ID-CoL="78DW" authority="Karny, 1913: 123" authorityName="Karny" authorityPageNumber="123" authorityYear="1913" box="[151,493,906,928]" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB2FFEE513120FBA1BCFE8CFC66" box="[151,307,906,928]" italics="true" pageId="5" pageNumber="40">Rhamphothrips</emphasis>
<treatmentCitation id="0ACD10B1FFEE51312155A1BDFE52FC66" author="Karny" box="[313,493,907,928]" page="123" pageId="5" pageNumber="40" year="1913">
<bibRefCitation id="EFFD4B51FFEE51312155A1BDFE06FC66" author="Karny" box="[313,441,907,928]" pageId="5" pageNumber="47" refString="Karny, H. &amp; Docters van Leeuwen-Reijnvaan, W. &amp; J. (1913) Beitrage zur Kenntnis der Gallen von Java. 5. Uber die javanischen Thysanopteren-cecidien und deren Bewohner. Bulletin du Jardin Botanique de Buitenzorg 10, 1 - 126." type="journal article" year="1913">Karny, 1913</bibRefCitation>
: 123
</treatmentCitation>
</taxonomicName>
. Type-species
<taxonomicName id="4C6C4D23FFEE513122ECA1BCFC66FC66" authority="Karny." authorityName="Karny." box="[640,985,906,928]" class="Insecta" family="Thripidae" genus="Rhynchothrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="tenuirostris">
<emphasis id="B918EAB2FFEE513122ECA1BCFC31FC66" box="[640,910,906,928]" italics="true" pageId="5" pageNumber="40">Rhynchothrips tenuirostris</emphasis>
Karny.
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD336A0FFEE513120FBA1FBFC7FFB5F" blockId="5.[151,1437,973,2005]" pageId="5" pageNumber="40">
Adults of this genus of
<taxonomicName id="4C6C4D23FFEE513121CEA1FBFDB0FC23" ID-CoL="626PS" box="[418,527,973,997]" class="Insecta" family="Thripidae" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="family">Thripidae</taxonomicName>
have a particularly elongate mouth-cone (
<figureCitation id="13572A25FFEE51312381A1FBFB80FC23" box="[1005,1087,973,997]" captionStart="FIGURES 24 26" captionStartId="8.[151,264,1557,1580]" captionTargetBox="[152,1423,911,1535]" captionTargetId="figure@8.[151,1436,911,1535]" captionTargetPageId="8" captionText="FIGURES 24 26. Rhamphothrips cissus. (24) head &amp; pronotum; (25) meso &amp; metanotum, abdominal terga I II; (26) mouth cone and pterothoracic sterna." httpUri="https://zenodo.org/record/276843/files/figure.png" pageId="5" pageNumber="40">Fig. 26</figureCitation>
) and small head (
<figureCitation id="13572A25FFEE51312566A1FBFAE1FC23" box="[1290,1374,973,997]" captionStart="FIGURES 24 26" captionStartId="8.[151,264,1557,1580]" captionTargetBox="[152,1423,911,1535]" captionTargetId="figure@8.[151,1436,911,1535]" captionTargetPageId="8" captionText="FIGURES 24 26. Rhamphothrips cissus. (24) head &amp; pronotum; (25) meso &amp; metanotum, abdominal terga I II; (26) mouth cone and pterothoracic sterna." httpUri="https://zenodo.org/record/276843/files/figure.png" pageId="5" pageNumber="40">Fig. 24</figureCitation>
), and the males of some species exhibit remarkable differences in structure between large and small individuals (
<bibRefCitation id="EFFD4B51FFEE5131252CA1C7FF42FBEB" author="Tyagi" pageId="5" pageNumber="48" refString="Tyagi, K., Kumar, V. &amp; Mound, L. A. (2008) Sexual dimorphism among Thysanoptera Terebrantia, with a new species from Malaysia and remarkable species from India in Aeolothripidae and Thripidae. Insect Systematics and Evolution, 39, 155 - 170." type="journal article" year="2008">Tyagi et al., 2008</bibRefCitation>
). This
<typeStatus id="54D78802FFEE5131212BA620FEC8FBE8" box="[327,375,1046,1070]" pageId="5" pageNumber="40">type</typeStatus>
of male polymorphism is rare in
<taxonomicName id="4C6C4D23FFEE51312285A623FCE6FBEB" ID-CoL="626PS" box="[745,857,1045,1069]" class="Insecta" family="Thripidae" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="family">Thripidae</taxonomicName>
, although it is common among species of the family
<taxonomicName id="4C6C4D23FFEE513120FBA60FFEF9FB97" ID-CoL="625XG" box="[151,326,1081,1105]" class="Insecta" family="Phlaeothripidae" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="family">Phlaeothripidae</taxonomicName>
that exhibit male/male competitive behaviour (
<bibRefCitation id="EFFD4B51FFEE51312333A60FFC41FB97" author="Mound" box="[863,1022,1081,1105]" lastPageId="5" lastPageNumber="48" pageId="5" pageNumber="47" refString="Mound, L. A. (2005) Fighting, flight and fecundity: behavioural determinants of Thysanoptera structural diversity. Pp 81 - 105 in Ananthakrishnan TN &amp; Whitman D. [eds] Insects and phenotypic plasticity. Science Publishers Inc. Enfield, NH, USA." type="book chapter" year="2005">Mound, 2005</bibRefCitation>
). Unfortunately, for none of the species of
<taxonomicName id="4C6C4D23FFEE51312081A66BFE25FBB3" ID-CoL="78DW" box="[237,410,1117,1141]" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB2FFEE51312081A66BFE25FBB3" box="[237,410,1117,1141]" italics="true" pageId="5" pageNumber="40">Rhamphothrips</emphasis>
</taxonomicName>
is there any information on the behaviour of these large and small males, and moreover there has been almost no information concerning their host associations.
</paragraph>
</subSubSection>
<subSubSection id="C376652BFFEE513220A9A693FC87FE32" lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40" type="description">
<paragraph id="8BD336A0FFEE513120A9A693FDB2F9AB" blockId="5.[151,1437,973,2005]" pageId="5" pageNumber="40">
The genus
<taxonomicName id="4C6C4D23FFEE51312153A693FE53FB7B" box="[319,492,1189,1213]" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB2FFEE51312153A693FE53FB7B" box="[319,492,1189,1213]" italics="true" pageId="5" pageNumber="40">Rhamphothrips</emphasis>
</taxonomicName>
is closely related to one other Old World genus,
<taxonomicName id="4C6C4D23FFEE51312463A693FB5BFB7B" authority="Priesner." authorityName="Priesner." box="[1039,1252,1189,1213]" class="Insecta" family="Thripidae" genus="Exothrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB2FFEE51312463A693FBC3FB7B" box="[1039,1148,1189,1213]" italics="true" pageId="5" pageNumber="40">Exothrips</emphasis>
Priesner.
</taxonomicName>
Females of species in these two genera share one remarkable character state: on sternum VII the median two pairs of setae, S1 and S2, are close together medially, and far distant from S3, the lateral third pair (
<figureCitation id="13572A25FFEE51312468A6DBFBE4FAC3" box="[1028,1115,1261,1285]" captionStart="FIGURES 15 23" captionStartId="7.[151,264,1946,1969]" captionTargetBox="[151,1434,337,1924]" captionTargetId="figure@7.[151,1436,337,1925]" captionTargetPageId="7" captionText="FIGURES 15 23. Rhamphothrips species. R. amyae 15 19: (15) antenna; (16) head &amp; pronotum; (17) male terga V IX; (18) female sterna VI VII; (19) male fore tibia &amp; tarsus. R. cissus 20 21: (20) thoracic sterna; (21) male terga V IX. R. pandens (22) male terga IV IX. R. tenuirostris (23) male terga VI IX." httpUri="https://zenodo.org/record/276842/files/figure.png" pageId="5" pageNumber="40">Figs 18</figureCitation>
,
<figureCitation id="13572A25FFEE51312404A6DBFB37FAC3" box="[1128,1160,1261,1285]" captionStart="FIGURES 27 32" captionStartId="11.[151,264,1859,1882]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure@11.[151,1436,193,1838]" captionTargetPageId="11" captionText="FIGURES 27 32. Rhamphothrips and Tusothrips. R. tenuirostris 27 30: (27 28) large and small males, fore coxae and mesosternum; (29) meso &amp; metanotum and abdominal terga I II; (30) female sternum VII. T. setiprivus 31 32: (31) female sternum VII; (32) head, thorax and abdominal terga I II." httpUri="https://zenodo.org/record/276844/files/figure.png" pageId="5" pageNumber="40">30</figureCitation>
). Moreover, unlike most
<taxonomicName id="4C6C4D23FFEE513120FBA727FEB8FAEF" box="[151,263,1297,1321]" class="Insecta" family="Thripidae" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="subFamily" subFamily="Thripinae">Thripinae</taxonomicName>
, adults in these two genera have no elongate posteroangular setae on the pronotum.
<taxonomicName id="4C6C4D23FFEE513124C3A727FAA3FAEF" box="[1199,1308,1297,1321]" class="Insecta" family="Thripidae" genus="Exothrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB2FFEE513124C3A727FAA3FAEF" box="[1199,1308,1297,1321]" italics="true" pageId="5" pageNumber="40">Exothrips</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D23FFEE5131253DA727FEBCFA8B" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB2FFEE5131253DA727FEBCFA8B" italics="true" pageId="5" pageNumber="40">Rhamphothrips</emphasis>
</taxonomicName>
are not clearly distinguished from each other (
<bibRefCitation id="EFFD4B51FFEE5131234EA703FB87FA8B" author="Mound" box="[802,1080,1333,1357]" pageId="5" pageNumber="48" refString="Mound, L. A. &amp; Walker, A. K. (1987) Thysanoptera as tropical tramps: new records from New Zealand and The Pacific. New Zealand Entomologist, 9, 70 - 85." type="journal article" year="1987">Mound &amp; Walker, 1987</bibRefCitation>
), although species of the latter have the head unusually small and the mouth cone exceptionally long, whereas species of the former have the head and pronotum transverse as in most
<taxonomicName id="4C6C4D23FFEE5131225DA74BFD1DFA53" box="[561,674,1405,1429]" class="Insecta" family="Thripidae" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="subFamily" subFamily="Thripinae">Thripinae</taxonomicName>
. Unfortunately, the apparent length of the mouth cone depends to a large extent on the orientation of slide-mounted specimens, and the species described from Africa seem to have a shorter mouth cone than those from the Oriental and Australian regions. Among the remaining
<taxonomicName id="4C6C4D23FFEE51312480A7F3FAE3FA1B" box="[1260,1372,1477,1501]" class="Insecta" family="Thripidae" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="subFamily" subFamily="Thripinae">Thripinae</taxonomicName>
, only females of
<taxonomicName id="4C6C4D23FFEE5131217EA7DFFE36F9C7" box="[274,393,1513,1537]" class="Insecta" family="Thripidae" genus="Tusothrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB2FFEE5131217EA7DFFE36F9C7" box="[274,393,1513,1537]" italics="true" pageId="5" pageNumber="40">Tusothrips</emphasis>
</taxonomicName>
have the median two pairs of setae placed particularly close to each other (
<figureCitation id="13572A25FFEE513124A1A7DFFAA1F9C7" box="[1229,1310,1513,1537]" captionStart="FIGURES 27 32" captionStartId="11.[151,264,1859,1882]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure@11.[151,1436,193,1838]" captionTargetPageId="11" captionText="FIGURES 27 32. Rhamphothrips and Tusothrips. R. tenuirostris 27 30: (27 28) large and small males, fore coxae and mesosternum; (29) meso &amp; metanotum and abdominal terga I II; (30) female sternum VII. T. setiprivus 31 32: (31) female sternum VII; (32) head, thorax and abdominal terga I II." httpUri="https://zenodo.org/record/276844/files/figure.png" pageId="5" pageNumber="40">Fig. 31</figureCitation>
), but
<taxonomicName id="4C6C4D23FFEE51312531A7DCFF67F9E3" class="Insecta" family="Thripidae" genus="Tusothrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB2FFEE51312531A7DCFF67F9E3" italics="true" pageId="5" pageNumber="40">Tusothrips</emphasis>
</taxonomicName>
species have two pairs of prominent posteroangular setae on the pronotum (
<figureCitation id="13572A25FFEE51312454A43BFB32F9E3" box="[1080,1165,1549,1573]" captionStart="FIGURES 27 32" captionStartId="11.[151,264,1859,1882]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure@11.[151,1436,193,1838]" captionTargetPageId="11" captionText="FIGURES 27 32. Rhamphothrips and Tusothrips. R. tenuirostris 27 30: (27 28) large and small males, fore coxae and mesosternum; (29) meso &amp; metanotum and abdominal terga I II; (30) female sternum VII. T. setiprivus 31 32: (31) female sternum VII; (32) head, thorax and abdominal terga I II." httpUri="https://zenodo.org/record/276844/files/figure.png" pageId="5" pageNumber="40">Fig. 32</figureCitation>
). Despite this, there is a tendency among species of
<taxonomicName id="4C6C4D23FFEE513121A6A407FDDDF98F" box="[458,610,1585,1609]" class="Insecta" family="Thripidae" genus="Anaphothrips" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB2FFEE513121A6A407FDDDF98F" box="[458,610,1585,1609]" italics="true" pageId="5" pageNumber="40">Anaphothrips</emphasis>
</taxonomicName>
for the median two pairs of setae on sternite VII of females to be closer to each other than to the lateral pair.
</paragraph>
<paragraph id="8BD336A0FFEE513220A9A44FFA9FFEF9" blockId="5.[151,1437,973,2005]" lastBlockId="6.[151,1436,151,500]" lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40">
The
<typeStatus id="54D78802FFEE51312094A44CFE97F954" box="[248,296,1658,1682]" pageId="5" pageNumber="40">type</typeStatus>
species of
<taxonomicName id="4C6C4D23FFEE513121C5A44FFDE9F957" box="[425,598,1657,1681]" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB2FFEE513121C5A44FFDE9F957" box="[425,598,1657,1681]" italics="true" pageId="5" pageNumber="40">Rhamphothrips</emphasis>
</taxonomicName>
was based on a single female from Java, and
<bibRefCitation id="EFFD4B51FFEE5131240CA44FFABAF957" author="Bhatti" box="[1120,1285,1657,1681]" pageId="5" pageNumber="47" refString="Bhatti, J. S. (1978 b) Studies in the systematics of Rhamphothrips. Oriental Insects, 12, 281 - 303." type="journal article" year="1978" yearSuffix="b">Bhatti (1978b)</bibRefCitation>
has given an account of the confusing early references to this specimen. The genus currently includes 14 species, all from the Old World tropics.
<bibRefCitation id="EFFD4B51FFEE51312103A4F7FDBAF91F" author="Bhatti" box="[367,517,1729,1753]" pageId="5" pageNumber="47" refString="Bhatti, J. S. (1977) The genus Perissothrips in India. Oriental Insects, 11, 567 - 577." type="journal article" year="1977">Bhatti (1977)</bibRefCitation>
provided a key to distinguish four species from
<collectingCountry id="F37B7630FFEE51312449A4F7FBDEF91F" box="[1061,1121,1729,1753]" name="India" pageId="5" pageNumber="40">India</collectingCountry>
, under the synonymic name
<emphasis id="B918EAB2FFEE513120FBA4D3FE93F93B" box="[151,300,1765,1789]" italics="true" pageId="5" pageNumber="40">Perissothrips</emphasis>
, and
<bibRefCitation id="EFFD4B51FFEE51312107A4D3FDAEF93B" author="Bhatti" box="[363,529,1765,1789]" pageId="5" pageNumber="47" refString="Bhatti, J. S. (1978 b) Studies in the systematics of Rhamphothrips. Oriental Insects, 12, 281 - 303." type="journal article" year="1978" yearSuffix="b">Bhatti (1978b)</bibRefCitation>
, in establishing three generic synonyms of
<taxonomicName id="4C6C4D23FFEE51312396A4D3FB18F93B" box="[1018,1191,1765,1789]" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB2FFEE51312396A4D3FB18F93B" box="[1018,1191,1765,1789]" italics="true" pageId="5" pageNumber="40">Rhamphothrips</emphasis>
</taxonomicName>
, provided a full diagnosis of the genus together with a more extensive key to distinguish ten species. Subsequently,
<bibRefCitation id="EFFD4B51FFEE513124B1A53FFA23F8E7" author="Sakimura" box="[1245,1436,1801,1825]" pageId="5" pageNumber="48" refString="Sakimura, K. (1983) A new Rhamphothrips from Hawaii, Jamaica and Florida, and notes on R. pomeroyi (Thysanoptera: Thripidae). Proceedings of the entomological Society of Hawaii, 24, 299 - 303." type="journal article" year="1983">Sakimura (1983)</bibRefCitation>
described
<taxonomicName id="4C6C4D23FFEE51312166A518FE37F883" box="[266,392,1837,1861]" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="pandens">
<emphasis id="B918EAB2FFEE51312166A518FE37F883" box="[266,392,1837,1861]" italics="true" pageId="5" pageNumber="40">R. pandens</emphasis>
</taxonomicName>
, and this is known to be widespread from northern
<collectingCountry id="F37B7630FFEE513123BBA51BFB81F883" box="[983,1086,1837,1861]" name="Australia" pageId="5" pageNumber="40">Australia</collectingCountry>
across the Pacific and into the Caribbean. From northern
<collectingCountry id="F37B7630FFEE513121A6A567FDBBF8AF" box="[458,516,1873,1897]" name="India" pageId="5" pageNumber="40">India</collectingCountry>
Kulshrestha &amp; Vijay Veer (1984) described
<taxonomicName id="4C6C4D23FFEE51312466A564FB34F8AF" box="[1034,1163,1873,1897]" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="santokhi">
<emphasis id="B918EAB2FFEE51312466A564FB34F8AF" box="[1034,1163,1873,1897]" italics="true" pageId="5" pageNumber="40">R. santokhi</emphasis>
</taxonomicName>
as having the posterior abdominal segments dark brown, in contrast to the uniformly yellow colour of all other members of the genus. Specimens of
<taxonomicName id="4C6C4D23FFEE51312156A5ACFE05F877" box="[314,442,1945,1969]" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="santokhi">
<emphasis id="B918EAB2FFEE51312156A5ACFE05F877" box="[314,442,1945,1969]" italics="true" pageId="5" pageNumber="40">R. santokhi</emphasis>
</taxonomicName>
, identified by Richard zur Strassen and taken from flowers of
<taxonomicName id="4C6C4D23FFEE513124E3A5ACFAC2F877" box="[1167,1405,1946,1969]" class="Magnoliopsida" family="Euphorbiaceae" genus="Macaranga" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="gigantea">
<emphasis id="B918EAB2FFEE513124E3A5ACFAC2F877" box="[1167,1405,1946,1969]" italics="true" pageId="5" pageNumber="40">Macaranga gigantea</emphasis>
</taxonomicName>
in
<collectingCountry id="F37B7630FFEE513120FBA58BFF41F813" box="[151,254,1981,2005]" name="Malaysia" pageId="5" pageNumber="40">Malaysia</collectingCountry>
(Gombak near Kuala Lumpur), have the median pair of setae, S1, on the female seventh sternum much smaller than setae S2, a condition also found in another Indian species,
<taxonomicName id="4C6C4D23FFED513223D7A2AFFB1CFF69" authority="Hood" authorityName="Hood" box="[955,1187,151,176]" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="parviceps">
<emphasis id="B918EAB2FFED513223D7A2AFFBF8FF76" box="[955,1095,153,176]" italics="true" pageId="6" pageNumber="41">R. parviceps</emphasis>
(Hood)
</taxonomicName>
. Finally,
<bibRefCitation id="EFFD4B51FFED51322565A2AEFA23FF69" author="Wang" box="[1289,1436,151,176]" pageId="6" pageNumber="48" refString="Wang, C. - L. (1993) A new species, Rhamphothrips quintus (Thysanoptera: Thripidae) from Taiwan. Chinese Journal of Entomology, 13, 341 - 345." type="journal article" year="1993">Wang (1993)</bibRefCitation>
described
<taxonomicName id="4C6C4D23FFED51322160A28BFE3DFF12" box="[268,386,189,212]" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="quintus">
<emphasis id="B918EAB2FFED51322160A28BFE3DFF12" box="[268,386,189,212]" italics="true" pageId="6" pageNumber="41">R. quintus</emphasis>
</taxonomicName>
from
<collectingCountry id="F37B7630FFED513221A0A28AFD9DFF12" box="[460,546,188,212]" name="Taiwan" pageId="6" pageNumber="41">Taiwan</collectingCountry>
. Females of this species have the median two pairs of setae on sternum VII equally long (pers. comm. C-L Wang 2010), and thus similar to the condition in
<taxonomicName id="4C6C4D23FFED5132247AA2D7FB2EFF3E" box="[1046,1169,224,248]" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="pandens">
<emphasis id="B918EAB2FFED5132247AA2D7FB2EFF3E" box="[1046,1169,224,248]" italics="true" pageId="6" pageNumber="41">R. pandens</emphasis>
</taxonomicName>
and two further species described below from
<collectingCountry id="F37B7630FFED513221F8A332FE44FEDA" box="[404,507,260,284]" name="Australia" pageId="6" pageNumber="41">Australia</collectingCountry>
(
<figureCitation id="13572A25FFED51322266A332FDE1FEDA" box="[522,606,260,284]" captionStart="FIGURES 15 23" captionStartId="7.[151,264,1946,1969]" captionTargetBox="[151,1434,337,1924]" captionTargetId="figure@7.[151,1436,337,1925]" captionTargetPageId="7" captionText="FIGURES 15 23. Rhamphothrips species. R. amyae 15 19: (15) antenna; (16) head &amp; pronotum; (17) male terga V IX; (18) female sterna VI VII; (19) male fore tibia &amp; tarsus. R. cissus 20 21: (20) thoracic sterna; (21) male terga V IX. R. pandens (22) male terga IV IX. R. tenuirostris (23) male terga VI IX." httpUri="https://zenodo.org/record/276842/files/figure.png" pageId="6" pageNumber="41">Fig. 18</figureCitation>
). However,
<taxonomicName id="4C6C4D23FFED5132228BA333FCE6FEDA" box="[743,857,261,284]" class="Insecta" family="Thripidae" genus="Rhamphothrips" kingdom="Animalia" order="Thysanoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="quintus">
<emphasis id="B918EAB2FFED5132228BA333FCE6FEDA" box="[743,857,261,284]" italics="true" pageId="6" pageNumber="41">R. quintus</emphasis>
</taxonomicName>
differs in that the females have a small tooth at the inner apex of the fore tibia, and the males have extensive tooth-like craspeda laterally on terga IVVIII.
</paragraph>
<paragraph id="8BD336A0FFED513220A9A37BFC87FE32" blockId="6.[151,1436,151,500]" pageId="6" pageNumber="41">
Large and small males of the new species described below from
<collectingCountry id="F37B7630FFED513223FAA37AFC43FEA2" box="[918,1020,332,356]" name="Australia" pageId="6" pageNumber="41">Australia</collectingCountry>
differ considerably in structure. This intraspecific variation in males contrasts with the fact that females of these new species are scarcely distinguishable from each other. Males are chosen as
<typeStatus id="54D78802FFED5132222FA3A2FD0FFE6A" box="[579,688,404,428]" pageId="6" pageNumber="41" type="holotype">holotypes</typeStatus>
of the two new species described below, because of the structural similarities between females. Currently it is not possible to identify many female specimens of this genus from
<collectingCountry id="F37B7630FFED513220FBA3EAFF41FE32" box="[151,254,476,500]" name="Australia" pageId="6" pageNumber="41">Australia</collectingCountry>
that have been collected with no associated males.
</paragraph>
</subSubSection>
</treatment>
</document>