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<document id="E10EF3DC8CE61D41C9BF0385DA1A5582" ID-DOI="10.5281/zenodo.277308" ID-GBIF-Dataset="61dd45a6-918a-4b12-8157-6090f7514613" ID-ISSN="1175-5326" ID-Zenodo-Dep="277308" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460286829391" checkinUser="plazi" docAuthor="Rosa, Simone Policena" docDate="2011" docId="03C58797BE02292AFF57FE43FBCBF480" docLanguage="en" docName="zt02831p022.pdf" docOrigin="Zootaxa 2831" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Triplonychus cruspinosus Rosa, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="6" masterDocId="FFFCFFEFBE00292FFFC0FF89FFB7F724" masterDocTitle="New species of Triplonychus Candèze and Globothorax Fleutiaux from Brazil (Coleoptera, Elateridae, Cardiophorinae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="3" updateTime="1698250676213" updateUser="plazi">
<mods:mods id="4D5A99BD567B54CEEB32B387ADAAB02A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C998508E3CA4EB58FB2A57D08C67C5C8">
<mods:title id="A2AF261334D8D80BC74E110FE948C975">New species of Triplonychus Candèze and Globothorax Fleutiaux from Brazil (Coleoptera, Elateridae, Cardiophorinae)</mods:title>
</mods:titleInfo>
<mods:name id="15CC713B18FD7ADF917C6640F4947B2F" type="personal">
<mods:role id="3A4FEEEDB069369FA1F0C1DF5B27B8E2">
<mods:roleTerm id="197DC7C7A230D6C6A9421FB4E233D373">Author</mods:roleTerm>
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<mods:namePart id="158129591CE76B1B537D6D0F0448CDD6">Rosa, Simone Policena</mods:namePart>
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<mods:title id="2562434927ED49FD8865E11DEA838746">Zootaxa</mods:title>
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<mods:part id="F1E94DD42712406D555BA979BDD567D4">
<mods:date id="39F3A2FAA1370B8832253EEC338F622B">2011</mods:date>
<mods:detail id="6A550A5737676B66F7B2B65537487D1F" type="volume">
<mods:number id="24E60F22D9C8B86791E2E33B9760528D">2831</mods:number>
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<mods:classification id="2889B6D5FE2CFFA0B77381CB2DEB4430">journal article</mods:classification>
<mods:identifier id="BB04523FA0D11F260765253F7D950F54" type="DOI">10.5281/zenodo.277308</mods:identifier>
<mods:identifier id="C9DF53D26D5A92EB97B259608DC6D641" type="GBIF-Dataset">61dd45a6-918a-4b12-8157-6090f7514613</mods:identifier>
<mods:identifier id="D017E4C6B99AF08205CD4952476CF07A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7C91FCD98302A9AA15AFF880CFE5B3B0" type="Zenodo-Dep">277308</mods:identifier>
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<treatment id="03C58797BE02292AFF57FE43FBCBF480" ID-DOI="http://doi.org/10.5281/zenodo.6188944" ID-GBIF-Taxon="119397854" ID-Zenodo-Dep="6188944" LSID="urn:lsid:plazi:treatment:03C58797BE02292AFF57FE43FBCBF480" httpUri="http://treatment.plazi.org/id/03C58797BE02292AFF57FE43FBCBF480" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
<subSubSection id="C376650ABE02292DFF57FE43FD65F521" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BD33681BE02292DFF57FE43FD83F6C0" blockId="2.[151,722,458,517]" box="[151,564,458,484]" pageId="2" pageNumber="3">
<heading id="D09B81EDBE02292DFF57FE43FD83F6C0" bold="true" box="[151,564,458,484]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B918EA93BE02292DFF57FE43FD83F6C0" bold="true" box="[151,564,458,484]" pageId="2" pageNumber="3">
<taxonomicName id="4C6C4D02BE02292DFF57FE43FE78F6C0" box="[151,463,458,484]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cruspinosus" status="sp. nov.">
<emphasis id="B918EA93BE02292DFF57FE43FE78F6C0" bold="true" box="[151,463,458,484]" italics="true" pageId="2" pageNumber="3">Triplonychus cruspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE02292DFE16FE43FD83F6C0" box="[470,564,458,484]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD33681BE02292DFF57FE64FD65F521" blockId="2.[151,722,458,517]" box="[151,722,493,517]" pageId="2" pageNumber="3">
(
<figureCitation id="13572A04BE02292DFF5FFE64FEAFF521" box="[159,280,493,517]" captionStart-0="FIGURES 1 – 8" captionStart-1="FIGURES 9 – 14" captionStartId-0="3.[151,264,1819,1842]" captionStartId-1="4.[151,264,1977,2000]" captionTargetBox-0="[154,1426,218,1746]" captionTargetBox-1="[191,1374,202,1951]" captionTargetId-0="figure@3.[152,1437,211,1765]" captionTargetId-1="figure@4.[181,1386,190,1952]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." captionText-1="FIGURES 9 – 14. Triplonychus cruspinosus sp. nov. 9, tergite 8; 10, sternite 8; 11, sternite 9; 12, tergites 9 and 10; 13, 14, aedeagus (ventral, dorsal). Abbreviation: vs, ventral sclerite of penis." httpUri-0="https://zenodo.org/record/277309/files/figure.png" httpUri-1="https://zenodo.org/record/277310/files/figure.png" pageId="2" pageNumber="3">Figs. 1–14</figureCitation>
,
<figureCitation id="13572A04BE02292DFEE5FE64FE21F521" box="[293,406,493,517]" captionStart="FIGURES 68 – 76" captionStartId="15.[151,264,1443,1466]" captionTargetBox="[151,1436,193,1421]" captionTargetId="figure@15.[151,1436,193,1422]" captionTargetPageId="15" captionText="FIGURES 68 – 76. Triplonychus cruspinosus sp. nov. 68, maxilla; 69, detail of maxilla; 70, labium. T. tibialatus sp. nov. 71, maxilla; 72, detail of maxilla; 73, labium. Head: 74, T. cruspinosus sp. nov.; 75, T. tibialatus sp. nov.; 76, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277317/files/figure.png" pageId="2" pageNumber="3">68–70, 74</figureCitation>
,
<figureCitation id="13572A04BE02292DFE62FE64FDA3F521" box="[418,532,493,517]" captionStart="FIGURES 77 – 90" captionStartId="16.[151,264,1935,1958]" captionTargetBox="[184,1388,193,1898]" captionTargetId="figure@16.[184,1389,193,1898]" captionTargetPageId="16" captionText="FIGURES 77 – 90. Anterior, median and posterior legs (outer surface): 77 – 79, Triplonychus cruspinosus sp. nov.; 80 – 82, T. crassifemoris sp. nov.; 83 – 85, T. tibialatus sp. nov.; 86 – 88, Globothorax latidens sp. nov. Claws: 89, T. cruspinosus sp. nov.; 90, G. latidens sp. nov." httpUri="https://zenodo.org/record/277318/files/figure.png" pageId="2" pageNumber="3">77–79, 89</figureCitation>
,
<figureCitation id="13572A04BE02292DFDE0FE64FDDFF521" box="[544,616,493,517]" captionStart="FIGURES 91 – 98" captionStartId="17.[151,264,1951,1974]" captionTargetBox="[403,1181,190,1931]" captionTargetId="figure@17.[396,1192,185,1935]" captionTargetPageId="17" captionText="FIGURES 91 – 98. Right elytron: 91, Triplonychus cruspinosus sp. nov.; 92, T. crassifemoris sp. nov.; 93, T. tibialatus sp. nov.; 94, Globothorax latidens sp. nov. Mesoscutum and scutellar shield: 95, Triplonychus cruspinosus sp. nov.; 96, T. crassifemoris sp. nov.; 97, T. tibialatus sp. nov.; 98, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277319/files/figure.png" pageId="2" pageNumber="3">91, 95</figureCitation>
,
<figureCitation id="13572A04BE02292DFDB4FE64FD24F521" box="[628,659,493,517]" captionStart="FIGURES 99 – 100" captionStartId="18.[151,264,865,888]" captionTargetBox="[312,1336,193,841]" captionTargetId="figure@18.[237,1349,193,844]" captionTargetPageId="18" captionText="FIGURES 99 – 100. Lateral view of mesoscutum and scutellar shield: 99, Triplonychus cruspinosus sp. nov.; 100, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277320/files/figure.png" pageId="2" pageNumber="3">99</figureCitation>
,
<figureCitation id="13572A04BE02292DFD5EFE64FD7EF521" box="[670,713,493,517]" captionStart="FIGURES 101 – 104" captionStartId="19.[151,264,1959,1982]" captionTargetBox="[222,1358,193,1919]" captionTargetId="figure@19.[222,1370,193,1925]" captionTargetPageId="19" captionText="FIGURES 101 – 104. Dorsal habitus of holotypes: 101, Triplonychus cruspinosus sp. nov. (4,5 mm length); 102, T. crassifemoris sp. nov. (7 mm length); 103, T. tibialatus sp. nov. (8 mm length); 104, Globothorax latidens sp. nov. (6 mm length)." httpUri="https://zenodo.org/record/277321/files/figure.png" pageId="2" pageNumber="3">101</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C376650ABE02292DFF57FDBCFB61F569" box="[151,1238,565,589]" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8BD33681BE02292DFF57FDBCFB61F569" blockId="2.[151,1437,565,2033]" box="[151,1238,565,589]" pageId="2" pageNumber="3">
<emphasis id="B918EA93BE02292DFF57FDBCFEA9F569" bold="true" box="[151,286,565,589]" pageId="2" pageNumber="3">Etymology.</emphasis>
From Latin,
<emphasis id="B918EA93BE02292DFE71FDBFFE56F569" box="[433,481,566,589]" italics="true" pageId="2" pageNumber="3">crus</emphasis>
= leg;
<emphasis id="B918EA93BE02292DFDF0FDBFFD26F569" box="[560,657,566,589]" italics="true" pageId="2" pageNumber="3">spinosus</emphasis>
= spiny, alluding to the spinose legs of this species.
</paragraph>
</subSubSection>
<subSubSection id="C376650ABE02292DFF05FDD0FDB8F5F9" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BD33681BE02292DFF05FDD0FDB8F5F9" blockId="2.[151,1437,565,2033]" pageId="2" pageNumber="3">
<emphasis id="B918EA93BE02292DFF05FDD0FE88F556" bold="true" box="[197,319,601,626]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
Dorsal tegument evenly colored; antennae surpassing the hind angles of pronotum by two or three antennomeres (
<figureCitation id="13572A04BE02292DFE81FDF4FE15F5B1" box="[321,418,637,661]" captionStart="FIGURES 101 – 104" captionStartId="19.[151,264,1959,1982]" captionTargetBox="[222,1358,193,1919]" captionTargetId="figure@19.[222,1370,193,1925]" captionTargetPageId="19" captionText="FIGURES 101 – 104. Dorsal habitus of holotypes: 101, Triplonychus cruspinosus sp. nov. (4,5 mm length); 102, T. crassifemoris sp. nov. (7 mm length); 103, T. tibialatus sp. nov. (8 mm length); 104, Globothorax latidens sp. nov. (6 mm length)." httpUri="https://zenodo.org/record/277321/files/figure.png" pageId="2" pageNumber="3">Fig. 101</figureCitation>
); pronotum wider than long with single well impressed punctation (
<figureCitation id="13572A04BE02292DFB53FDF4FB61F5B1" box="[1171,1238,637,661]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[154,1426,218,1746]" captionTargetId="figure@3.[152,1437,211,1765]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
); metacoxal plate triangular with posterior margin rounded (
<figureCitation id="13572A04BE02292DFDB5FD28FD0DF59D" box="[629,698,673,697]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[154,1426,218,1746]" captionTargetId="figure@3.[152,1437,211,1765]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
); legs slender, tibiae with rows of long spiniform setae on dorsal and ventral margin (
<figureCitation id="13572A04BE02292DFEB9FD4CFDB4F5F9" box="[377,515,709,733]" captionStart="FIGURES 77 – 90" captionStartId="16.[151,264,1935,1958]" captionTargetBox="[184,1388,193,1898]" captionTargetId="figure@16.[184,1389,193,1898]" captionTargetPageId="16" captionText="FIGURES 77 – 90. Anterior, median and posterior legs (outer surface): 77 – 79, Triplonychus cruspinosus sp. nov.; 80 – 82, T. crassifemoris sp. nov.; 83 – 85, T. tibialatus sp. nov.; 86 – 88, Globothorax latidens sp. nov. Claws: 89, T. cruspinosus sp. nov.; 90, G. latidens sp. nov." httpUri="https://zenodo.org/record/277318/files/figure.png" pageId="2" pageNumber="3">Figs. 77–79</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C376650ABE02292AFF05FD60FECAF533" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3" type="description">
<paragraph id="8BD33681BE02292DFF05FD60FED0F491" blockId="2.[151,1437,565,2033]" pageId="2" pageNumber="3">
<emphasis id="B918EA93BE02292DFF05FD60FEE0F426" bold="true" box="[197,343,745,770]" pageId="2" pageNumber="3">Description.</emphasis>
Male (
<figureCitation id="13572A04BE02292DFE68FD60FDBCF425" box="[424,523,745,769]" captionStart="FIGURES 101 – 104" captionStartId="19.[151,264,1959,1982]" captionTargetBox="[222,1358,193,1919]" captionTargetId="figure@19.[222,1370,193,1925]" captionTargetPageId="19" captionText="FIGURES 101 – 104. Dorsal habitus of holotypes: 101, Triplonychus cruspinosus sp. nov. (4,5 mm length); 102, T. crassifemoris sp. nov. (7 mm length); 103, T. tibialatus sp. nov. (8 mm length); 104, Globothorax latidens sp. nov. (6 mm length)." httpUri="https://zenodo.org/record/277321/files/figure.png" pageId="2" pageNumber="3">Fig. 101</figureCitation>
). Body convex, densely covered with fine and short setae; setae light brown and semidecumbent to erect on dorsal surface, yellow and decumbent on ventral surface. Integument bright; head, pronotum and elytra yellow or light to dark brown, with antennae and legs yellow; ventral surface lighter than pronotum and elytra. Pronotal and elytral punctures darker than the surrounding area. A few entirely yellow specimens are probably teneral adults. Total length: 4.0–6.0 mm; elytra 2.5–2.7 times longer than prothorax; elytral base as wide as prothorax.
</paragraph>
<paragraph id="8BD33681BE02292DFF05FC48FAA8F265" blockId="2.[151,1437,565,2033]" pageId="2" pageNumber="3">
Head (
<figureCitation id="13572A04BE02292DFED0FC48FED1F4FD" box="[272,358,961,985]" captionStart="FIGURES 68 – 76" captionStartId="15.[151,264,1443,1466]" captionTargetBox="[151,1436,193,1421]" captionTargetId="figure@15.[151,1436,193,1422]" captionTargetPageId="15" captionText="FIGURES 68 – 76. Triplonychus cruspinosus sp. nov. 68, maxilla; 69, detail of maxilla; 70, labium. T. tibialatus sp. nov. 71, maxilla; 72, detail of maxilla; 73, labium. Head: 74, T. cruspinosus sp. nov.; 75, T. tibialatus sp. nov.; 76, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277317/files/figure.png" pageId="2" pageNumber="3">Fig. 74</figureCitation>
). Frons concave; anterior margin carinate, rounded and produced. Frontoclypeal region steeply declivous to base of labrum and 2 times wider than long. Punctures smaller than those of pronotal disk, umbilicate, 0.5–1.0 diameter apart. Antenna (
<figureCitation id="13572A04BE02292DFDCFFB80FDF8F305" box="[527,591,1033,1057]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[154,1426,218,1746]" captionTargetId="figure@3.[152,1437,211,1765]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
) surpassing the hind angles of pronotum at antennomere 8 or 9; antennomere 3 1.6–2.0 times longer than 2; 4 1.1–1.3 times longer than 3. Dorsal index of eye prominence 0.4. Labrum semielliptical, strongly punctate with short and long setae. Mandible (
<figureCitation id="13572A04BE02292DFC66FBD8FC5DF34D" box="[934,1002,1105,1129]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[154,1426,218,1746]" captionTargetId="figure@3.[152,1437,211,1765]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
) short (only the tips superimpose each other) unidentate, evenly arcuate, with rows of small setae on mesobasal margin, a longitudinal short carina dorsally and a stouter and longer seta than the fine setae of the lateroventral margin. Maxilla (
<figureCitation id="13572A04BE02292DFB5DFB10FB47F395" box="[1181,1264,1177,1201]" captionStart="FIGURES 68 – 76" captionStartId="15.[151,264,1443,1466]" captionTargetBox="[151,1436,193,1421]" captionTargetId="figure@15.[151,1436,193,1422]" captionTargetPageId="15" captionText="FIGURES 68 – 76. Triplonychus cruspinosus sp. nov. 68, maxilla; 69, detail of maxilla; 70, labium. T. tibialatus sp. nov. 71, maxilla; 72, detail of maxilla; 73, labium. Head: 74, T. cruspinosus sp. nov.; 75, T. tibialatus sp. nov.; 76, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277317/files/figure.png" pageId="2" pageNumber="3">Fig. 68</figureCitation>
) with basistipe fused to medistipe (
<figureCitation id="13572A04BE02292DFEB9FB34FE7BF3F1" box="[377,460,1213,1237]" captionStart="FIGURES 68 – 76" captionStartId="15.[151,264,1443,1466]" captionTargetBox="[151,1436,193,1421]" captionTargetId="figure@15.[151,1436,193,1422]" captionTargetPageId="15" captionText="FIGURES 68 – 76. Triplonychus cruspinosus sp. nov. 68, maxilla; 69, detail of maxilla; 70, labium. T. tibialatus sp. nov. 71, maxilla; 72, detail of maxilla; 73, labium. Head: 74, T. cruspinosus sp. nov.; 75, T. tibialatus sp. nov.; 76, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277317/files/figure.png" pageId="2" pageNumber="3">Fig. 69</figureCitation>
) with a long stout seta and a few fine short setae; galea triangular with long setae on outer angle and shorter, denser setae on inner angle. Labium (
<figureCitation id="13572A04BE02292DFC86FB68FC20F3DD" box="[838,919,1249,1273]" captionStart="FIGURES 68 – 76" captionStartId="15.[151,264,1443,1466]" captionTargetBox="[151,1436,193,1421]" captionTargetId="figure@15.[151,1436,193,1422]" captionTargetPageId="15" captionText="FIGURES 68 – 76. Triplonychus cruspinosus sp. nov. 68, maxilla; 69, detail of maxilla; 70, labium. T. tibialatus sp. nov. 71, maxilla; 72, detail of maxilla; 73, labium. Head: 74, T. cruspinosus sp. nov.; 75, T. tibialatus sp. nov.; 76, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277317/files/figure.png" pageId="2" pageNumber="3">Fig. 70</figureCitation>
) with mentum trapezoidal with a few setae on posterior angles; prementum notched at anteromedian margin, lateroanterior angles rounded and produced; second labial palpomere with long spiniform setae apically; apical maxillary and labial palpomeres securiform.
</paragraph>
<paragraph id="8BD33681BE02292DFF05FAC4FA2AF0D4" blockId="2.[151,1437,565,2033]" pageId="2" pageNumber="3">
Thorax. Prothorax (
<figureCitation id="13572A04BE02292DFE61FAC4FDBCF241" box="[417,523,1357,1381]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[154,1426,218,1746]" captionTargetId="figure@3.[152,1437,211,1765]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="2" pageNumber="3">Figs. 3–4</figureCitation>
) about 1.2 times wider than long, lateral margins slightly rounded, carinate on the posterior 3/4. Pronotum convex, punctures on disk large 0.5–1.5 diameters apart, on anterior margin smaller 0.2– 1.0 diameter apart, on posterior margin very small 1–2 diameters apart. Pronotal hind angles flat, short, obliquely truncate; posterior margin between hind angle and median line with one or two short notches. Hypomera (
<figureCitation id="13572A04BE02292DFA8FFA30FA24F2F5" box="[1359,1427,1465,1489]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[154,1426,218,1746]" captionTargetId="figure@3.[152,1437,211,1765]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
) with punctation double; notosternal suture curved. Prosternum (
<figureCitation id="13572A04BE02292DFCA6FA54FC1EF2D1" box="[870,937,1501,1525]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[154,1426,218,1746]" captionTargetId="figure@3.[152,1437,211,1765]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
) 1.3 times longer than wide, with punctures as large as those of pronotal disk, umbilicate, 1–3 diameters apart on the median region and 0.5–1.0 diameter apart on lateral margin. Prosternal process (
<figureCitation id="13572A04BE02292DFD85F9ACFD3EF119" box="[581,649,1573,1597]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[154,1426,218,1746]" captionTargetId="figure@3.[152,1437,211,1765]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
) 2.1–2.5 times longer than coxal diameter with ventral surface of the posterior half nearly as wide as the dorsal surface and steeply sloping dorsad at apex (
<figureCitation id="13572A04BE02292DFB49F9C0FB65F145" box="[1161,1234,1609,1633]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[154,1426,218,1746]" captionTargetId="figure@3.[152,1437,211,1765]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
). Procoxal cavity closed. Scutellar shield (
<figureCitation id="13572A04BE02292DFE6DF9E4FDB7F1A1" box="[429,512,1645,1669]" captionStart="FIGURES 91 – 98" captionStartId="17.[151,264,1951,1974]" captionTargetBox="[403,1181,190,1931]" captionTargetId="figure@17.[396,1192,185,1935]" captionTargetPageId="17" captionText="FIGURES 91 – 98. Right elytron: 91, Triplonychus cruspinosus sp. nov.; 92, T. crassifemoris sp. nov.; 93, T. tibialatus sp. nov.; 94, Globothorax latidens sp. nov. Mesoscutum and scutellar shield: 95, Triplonychus cruspinosus sp. nov.; 96, T. crassifemoris sp. nov.; 97, T. tibialatus sp. nov.; 98, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277319/files/figure.png" pageId="2" pageNumber="3">Fig. 95</figureCitation>
) pentagonal to cordiform with a short median notch on anterior margin, abruptly elevated above the level of mesoscutum (
<figureCitation id="13572A04BE02292DFDAEF91DFD73F188" box="[622,708,1684,1708]" captionStart="FIGURES 99 – 100" captionStartId="18.[151,264,865,888]" captionTargetBox="[312,1336,193,841]" captionTargetId="figure@18.[237,1349,193,844]" captionTargetPageId="18" captionText="FIGURES 99 – 100. Lateral view of mesoscutum and scutellar shield: 99, Triplonychus cruspinosus sp. nov.; 100, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277320/files/figure.png" pageId="2" pageNumber="3">Fig. 99</figureCitation>
). Mesoventrite (
<figureCitation id="13572A04BE02292DFCBEF91DFC75F188" box="[894,962,1684,1708]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[154,1426,218,1746]" captionTargetId="figure@3.[152,1437,211,1765]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
) with posterior region inclined 45o above the level of anterior region; lateral lobes gradually and moderately narrowed apicad; mesoventral cavity widened up to anterior border of mesocoxae then abruptly convergent. Mesocoxal cavity oval, closed. Mesepisternum with a transverse carina near the anterior inner angle; anterior inner angle acute. Meso-metaventral suture weakly impressed. Metaventrite (
<figureCitation id="13572A04BE02292DFE7EF8ADFD86F018" box="[446,561,1828,1852]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[154,1426,218,1746]" captionTargetId="figure@3.[152,1437,211,1765]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="2" pageNumber="3">Figs. 7, 8</figureCitation>
) with punctation double; metacoxal plate triangular with posterior margin rounded. Elytra (
<figureCitation id="13572A04BE02292DFE97F8C1FE1EF044" box="[343,425,1864,1888]" captionStart="FIGURES 91 – 98" captionStartId="17.[151,264,1951,1974]" captionTargetBox="[403,1181,190,1931]" captionTargetId="figure@17.[396,1192,185,1935]" captionTargetPageId="17" captionText="FIGURES 91 – 98. Right elytron: 91, Triplonychus cruspinosus sp. nov.; 92, T. crassifemoris sp. nov.; 93, T. tibialatus sp. nov.; 94, Globothorax latidens sp. nov. Mesoscutum and scutellar shield: 95, Triplonychus cruspinosus sp. nov.; 96, T. crassifemoris sp. nov.; 97, T. tibialatus sp. nov.; 98, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277319/files/figure.png" pageId="2" pageNumber="3">Fig. 91</figureCitation>
) slightly rounded or parallel-sided up to posterior 1/3 then tapering to apex; elytral striae each with 2 rows of punctures predominantly paired side by side; interstices 3, 5 and 7 more convex than 2, 4, 6 and 8 on apical region, slightly convex on anterior region; interstice 9 carinate on posterior 2/3. Legs (
<figureCitation id="13572A04BE02292DFAC7F819FA26F08C" box="[1287,1425,1936,1960]" captionStart="FIGURES 77 – 90" captionStartId="16.[151,264,1935,1958]" captionTargetBox="[184,1388,193,1898]" captionTargetId="figure@16.[184,1389,193,1898]" captionTargetPageId="16" captionText="FIGURES 77 – 90. Anterior, median and posterior legs (outer surface): 77 – 79, Triplonychus cruspinosus sp. nov.; 80 – 82, T. crassifemoris sp. nov.; 83 – 85, T. tibialatus sp. nov.; 86 – 88, Globothorax latidens sp. nov. Claws: 89, T. cruspinosus sp. nov.; 90, G. latidens sp. nov." httpUri="https://zenodo.org/record/277318/files/figure.png" pageId="2" pageNumber="3">Figs. 77–79</figureCitation>
). Femur narrow, tibia slightly widened apicad, pilose, with rows of spiniform setae on dorsal and apical margin longer than the fine setae, tibial spurs stout about 0.5 as long as tarsomere 1; tarsomeres densely pilose with fine setae.
</paragraph>
<caption id="DF136609BE03292CFF57F892FD2DF074" httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="3" pageNumber="4" targetBox="[154,1426,218,1746]" targetPageId="3">
<paragraph id="8BD33681BE03292CFF57F892FD2DF074" blockId="3.[151,1436,1819,1872]" pageId="3" pageNumber="4">
<emphasis id="B918EA93BE03292CFF57F892FE8DF015" bold="true" box="[151,314,1819,1842]" pageId="3" pageNumber="4">FIGURES 1–8.</emphasis>
<taxonomicName id="4C6C4D02BE03292CFE80F895FDF3F016" box="[320,580,1820,1842]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cruspinosus" status="sp. nov.">
<emphasis id="B918EA93BE03292CFE80F895FDF3F016" box="[320,580,1820,1842]" italics="true" pageId="3" pageNumber="4">Triplonychus cruspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE03292CFD8BF895FD2CF016" box="[587,667,1820,1842]" pageId="3" pageNumber="4" rank="species">
<emphasis id="B918EA93BE03292CFD8BF895FD22F015" bold="true" box="[587,661,1820,1841]" pageId="3" pageNumber="4">sp. nov</emphasis>
.
</taxonomicNameLabel>
1, antenna; 2, mandible (dorsal); 3–5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral).
</paragraph>
</caption>
<caption id="DF136609BE04292BFF57F830FC93F0CA" httpUri="https://zenodo.org/record/277310/files/figure.png" pageId="4" pageNumber="5" targetBox="[191,1374,202,1951]" targetPageId="4">
<paragraph id="8BD33681BE04292BFF57F830FC93F0CA" blockId="4.[151,1436,1977,2030]" pageId="4" pageNumber="5">
<emphasis id="B918EA93BE04292BFF57F830FEF1F0EB" bold="true" box="[151,326,1977,2000]" pageId="4" pageNumber="5">FIGURES 9–14.</emphasis>
<taxonomicName id="4C6C4D02BE04292BFE8CF830FDF8F0EB" box="[332,591,1977,1999]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cruspinosus" status="sp. nov.">
<emphasis id="B918EA93BE04292BFE8CF830FDF8F0EB" box="[332,591,1977,1999]" italics="true" pageId="4" pageNumber="5">Triplonychus cruspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE04292BFD95F833FD12F0EB" box="[597,677,1978,1999]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B918EA93BE04292BFD95F833FD28F0EB" bold="true" box="[597,671,1978,1999]" pageId="4" pageNumber="5">sp. nov</emphasis>
.
</taxonomicNameLabel>
9, tergite 8; 10, sternite 8; 11, sternite 9; 12, tergites 9 and 10; 13, 14, aedeagus (ventral, dorsal). Abbreviation: vs, ventral sclerite of penis.
</paragraph>
</caption>
<paragraph id="8BD33681BE05292AFF05FF1EFB72F6A3" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="6">
Abdomen (
<figureCitation id="13572A04BE05292AFE84FF1EFE3DF78B" box="[324,394,151,175]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[154,1426,218,1746]" captionTargetId="figure@3.[152,1437,211,1765]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation>
) rounded laterally; punctures fine, umbilicate, 0.5 diameter apart, denser on lateral margin. First ventrite 5.3–6.5 times longer on lateral border than at middle; last ventrite pentagonal. Tergite 8 (
<figureCitation id="13572A04BE05292AFA8DFF35FA23F7F0" box="[1357,1428,188,212]" captionStart="FIGURES 9 – 14" captionStartId="4.[151,264,1977,2000]" captionTargetBox="[191,1374,202,1951]" captionTargetId="figure@4.[181,1386,190,1952]" captionTargetPageId="4" captionText="FIGURES 9 – 14. Triplonychus cruspinosus sp. nov. 9, tergite 8; 10, sternite 8; 11, sternite 9; 12, tergites 9 and 10; 13, 14, aedeagus (ventral, dorsal). Abbreviation: vs, ventral sclerite of penis." httpUri="https://zenodo.org/record/277310/files/figure.png" pageId="5" pageNumber="6">Fig. 9</figureCitation>
) evenly sclerotized, with a few long and stout setae on lateroposterior and posterior margins; anterior sclerotized border U-shaped emarginated. Sternite 8 (
<figureCitation id="13572A04BE05292AFDACFE8DFD08F638" box="[620,703,260,284]" captionStart="FIGURES 9 – 14" captionStartId="4.[151,264,1977,2000]" captionTargetBox="[191,1374,202,1951]" captionTargetId="figure@4.[181,1386,190,1952]" captionTargetPageId="4" captionText="FIGURES 9 – 14. Triplonychus cruspinosus sp. nov. 9, tergite 8; 10, sternite 8; 11, sternite 9; 12, tergites 9 and 10; 13, 14, aedeagus (ventral, dorsal). Abbreviation: vs, ventral sclerite of penis." httpUri="https://zenodo.org/record/277310/files/figure.png" pageId="5" pageNumber="6">Fig. 10</figureCitation>
), pentagonal, evenly sclerotized, with short setae laterally and on apical region, apex narrow emarginated at middle. Sternite 9 (
<figureCitation id="13572A04BE05292AFC92FEAEFC15F61B" box="[850,930,295,319]" captionStart="FIGURES 9 – 14" captionStartId="4.[151,264,1977,2000]" captionTargetBox="[191,1374,202,1951]" captionTargetId="figure@4.[181,1386,190,1952]" captionTargetPageId="4" captionText="FIGURES 9 – 14. Triplonychus cruspinosus sp. nov. 9, tergite 8; 10, sternite 8; 11, sternite 9; 12, tergites 9 and 10; 13, 14, aedeagus (ventral, dorsal). Abbreviation: vs, ventral sclerite of penis." httpUri="https://zenodo.org/record/277310/files/figure.png" pageId="5" pageNumber="6">Fig. 11</figureCitation>
) with sides rounded tapering to apex; lateral and posterior borders sclerotized, anterior and median region translucent, apex with short and elongate setae. Tergite 9 (
<figureCitation id="13572A04BE05292AFF17FEE6FE9EF6A3" box="[215,297,367,391]" captionStart="FIGURES 9 – 14" captionStartId="4.[151,264,1977,2000]" captionTargetBox="[191,1374,202,1951]" captionTargetId="figure@4.[181,1386,190,1952]" captionTargetPageId="4" captionText="FIGURES 9 – 14. Triplonychus cruspinosus sp. nov. 9, tergite 8; 10, sternite 8; 11, sternite 9; 12, tergites 9 and 10; 13, 14, aedeagus (ventral, dorsal). Abbreviation: vs, ventral sclerite of penis." httpUri="https://zenodo.org/record/277310/files/figure.png" pageId="5" pageNumber="6">Fig. 12</figureCitation>
) with anterior margin sinuous, posterior lobes acute, tergite 10 (
<figureCitation id="13572A04BE05292AFC37FEE6FBFFF6A3" box="[1015,1096,367,391]" captionStart="FIGURES 9 – 14" captionStartId="4.[151,264,1977,2000]" captionTargetBox="[191,1374,202,1951]" captionTargetId="figure@4.[181,1386,190,1952]" captionTargetPageId="4" captionText="FIGURES 9 – 14. Triplonychus cruspinosus sp. nov. 9, tergite 8; 10, sternite 8; 11, sternite 9; 12, tergites 9 and 10; 13, 14, aedeagus (ventral, dorsal). Abbreviation: vs, ventral sclerite of penis." httpUri="https://zenodo.org/record/277310/files/figure.png" pageId="5" pageNumber="6">Fig. 12</figureCitation>
) semioval.
</paragraph>
<paragraph id="8BD33681BE05292AFF05FE1DFECAF533" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="6">
Aedeagus (
<figureCitation id="13572A04BE05292AFE84FE1DFE79F688" box="[324,462,404,428]" captionStart="FIGURES 9 – 14" captionStartId="4.[151,264,1977,2000]" captionTargetBox="[191,1374,202,1951]" captionTargetId="figure@4.[181,1386,190,1952]" captionTargetPageId="4" captionText="FIGURES 9 – 14. Triplonychus cruspinosus sp. nov. 9, tergite 8; 10, sternite 8; 11, sternite 9; 12, tergites 9 and 10; 13, 14, aedeagus (ventral, dorsal). Abbreviation: vs, ventral sclerite of penis." httpUri="https://zenodo.org/record/277310/files/figure.png" pageId="5" pageNumber="6">Figs. 13, 14</figureCitation>
). Parameres with apex acute directed lateroventrad, without setae, articulated to penis by a transparent membrane. Penis with a ventral sclerite oval about 3.4 times longer than wide, basal struts 0.4 times as long as its total length, median region between basal struts with a triangular process; lateral margins tapering to rounded apex.
</paragraph>
</subSubSection>
<subSubSection id="C376650ABE05292AFF05FDADFD53F478" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BD33681BE05292AFF05FDADFDCDF57B" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="6">
<emphasis id="B918EA93BE05292AFF05FDADFEC3F518" bold="true" box="[197,372,547,572]" pageId="5" pageNumber="6">
<typeStatus id="54D78823BE05292AFF05FDADFEB7F518" box="[197,256,548,572]" pageId="5" pageNumber="6">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54D78823BE05292AFEBCFDADFE5FF518" box="[380,488,548,572]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
. [
<collectingCountry id="F37B7611BE05292AFE3AFDADFDF5F518" box="[506,578,548,572]" name="Brazil" pageId="5" pageNumber="6">Brasil</collectingCountry>
, Reserva Ducke, 26 Km NE Manaus, Barbosa, M.G.V.], [Plot A, malaise 1, DEZ/1995], [62], [0031799], male (
<collectionCode id="ED7DAE44BE05292AFDE9FDCEFDD8F57B" box="[553,623,583,607]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="6">INPA</collectionCode>
).
</paragraph>
<paragraph id="8BD33681BE05292AFF05FDE4FB76F413" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="6">
<typeStatus id="54D78823BE05292AFF05FDE4FE82F5A1" box="[197,309,621,645]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
. [
<collectingCountry id="F37B7611BE05292AFE8BFDE5FE24F5A0" box="[331,403,620,644]" name="Brazil" pageId="5" pageNumber="6">Brasil</collectingCountry>
, Reserva Ducke, 26 Km NE Manaus, Barbosa, M.G.V.], malaise trap,
<date id="FFD21041BE05292AFB1CFDE5FAF9F5A0" box="[1244,1358,620,644]" pageId="5" pageNumber="6" value="1995-02-05">
II.1995,
<specimenCount id="9D6AFD08BE05292AFA80FDE5FAF9F5A0" box="[1344,1358,620,644]" pageId="5" pageNumber="6" type="male">5</specimenCount>
</date>
males (
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),
<specimenCount id="9D6AFD08BE05292AFF39FD06FEE6F583" box="[249,337,655,679]" pageId="5" pageNumber="6" type="male">2 males</specimenCount>
(
<collectionCode id="ED7DAE44BE05292AFEA3FD06FE70F583" box="[355,455,655,679]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="6">MZUSP</collectionCode>
);
<date id="FFD21041BE05292AFE1DFD06FDD5F583" box="[477,610,655,679]" pageId="5" pageNumber="6" value="1995-04-10">
IV.1995,
<specimenCount id="9D6AFD08BE05292AFD86FD06FDD5F583" box="[582,610,655,679]" pageId="5" pageNumber="6" type="male">10</specimenCount>
</date>
males (
<collectionCode id="ED7DAE44BE05292AFD7EFD06FCB4F583" box="[702,771,655,679]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="6">INPA</collectionCode>
),
<specimenCount id="9D6AFD08BE05292AFCD9FD19FCC6F583" box="[793,881,655,679]" pageId="5" pageNumber="6" type="male">7 males</specimenCount>
(
<collectionCode id="ED7DAE44BE05292AFC43FD06FC50F583" box="[899,999,655,679]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="6">MZUSP</collectionCode>
);
<date id="FFD21041BE05292AFC3CFD19FBC3F583" box="[1020,1140,655,679]" pageId="5" pageNumber="6" value="1995-05-01">
V. 1995,
<specimenCount id="9D6AFD08BE05292AFBA6FD06FBC3F583" box="[1126,1140,655,679]" pageId="5" pageNumber="6" type="male">1</specimenCount>
</date>
male (
<collectionCode id="ED7DAE44BE05292AFB06FD06FABCF583" box="[1222,1291,655,679]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="6">INPA</collectionCode>
);
<date id="FFD21041BE05292AFAE1FD06FA2BF583" box="[1313,1436,655,679]" pageId="5" pageNumber="6" value="1995-11-04">
XI.1995,
<specimenCount id="9D6AFD08BE05292AFA4EFD06FA2BF583" box="[1422,1436,655,679]" pageId="5" pageNumber="6" type="male">4</specimenCount>
</date>
males (
<collectionCode id="ED7DAE44BE05292AFF27FD3DFE9AF5E8" box="[231,301,692,716]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="6">INPA</collectionCode>
);
<date id="FFD21041BE05292AFE81FD3DFE66F5E8" box="[321,465,692,716]" pageId="5" pageNumber="6" value="1995-12-30">
XII.1995,
<specimenCount id="9D6AFD08BE05292AFE75FD3DFE66F5E8" box="[437,465,692,716]" pageId="5" pageNumber="6" type="male">30</specimenCount>
</date>
males (
<collectionCode id="ED7DAE44BE05292AFDE9FD3DFDD8F5E8" box="[553,623,692,716]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="6">INPA</collectionCode>
),
<specimenCount id="9D6AFD08BE05292AFDBCFD3DFD68F5E8" box="[636,735,692,716]" pageId="5" pageNumber="6" type="male">12 males</specimenCount>
(
<collectionCode id="ED7DAE44BE05292AFD2FFD3DFCE4F5E8" box="[751,851,692,716]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="6">MZUSP</collectionCode>
);
<date id="FFD21041BE05292AFCA6FD3DFC7CF5E8" box="[870,971,692,716]" pageId="5" pageNumber="6" value="1996-01-08">
I.1996,
<specimenCount id="9D6AFD08BE05292AFC7DFD3DFC7CF5E8" box="[957,971,692,716]" pageId="5" pageNumber="6" type="male">8</specimenCount>
</date>
males (
<collectionCode id="ED7DAE44BE05292AFBEDFD3DFBC6F5E8" box="[1069,1137,692,716]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="6">INPA</collectionCode>
),
<specimenCount id="9D6AFD08BE05292AFB44FD3DFB6DF5E8" box="[1156,1242,692,716]" pageId="5" pageNumber="6" type="male">4 males</specimenCount>
(
<collectionCode id="ED7DAE44BE05292AFB2AFD3DFAF9F5E8" box="[1258,1358,692,716]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="6">MZUSP</collectionCode>
). [
<collectingCountry id="F37B7611BE05292AFAA7FD3CFF02F5CB" name="Brazil" pageId="5" pageNumber="6">Brasil</collectingCountry>
: Amazonas, Reserva Ducke,
<quantity id="4C949B64BE05292AFDC1FD5EFDFEF5CB" box="[513,585,727,751]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="6" unit="km" value="26.0">26 km</quantity>
NE Manaus,
<date id="FFD21041BE05292AFD26FD5EFCEAF5CB" box="[742,861,727,751]" pageId="5" pageNumber="6" value="1978-02-28">28.II.1978</date>
, J. Arias &amp; A. Penny],
<specimenCount id="9D6AFD08BE05292AFBA8FD5EFB7AF5CB" box="[1128,1229,727,751]" pageId="5" pageNumber="6" type="male">64 males</specimenCount>
(
<collectionCode id="ED7DAE44BE05292AFB1EFD5EFA94F5CB" box="[1246,1315,727,751]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="6">INPA</collectionCode>
),
<specimenCount id="9D6AFD08BE05292AFAF6FD5EFA2CF5CB" box="[1334,1435,727,751]" pageId="5" pageNumber="6" type="male">24 males</specimenCount>
(
<collectionCode id="ED7DAE44BE05292AFF5FFD75FEB4F430" box="[159,259,764,788]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="6">MZUSP</collectionCode>
). [
<collectingCountry id="F37B7611BE05292AFEDCFD75FED1F430" box="[284,358,764,788]" name="Brazil" pageId="5" pageNumber="6">Brasil</collectingCountry>
: Amazonas, AM 0 10, Km 26, Reserva Ducke,
<date id="FFD21041BE05292AFCB6FD75FC68F430" box="[886,991,764,788]" pageId="5" pageNumber="6" value="1978-02-07">7.II.1978</date>
, Jorge Arias coll.]
<specimenCount id="9D6AFD08BE05292AFB74FD75FAAEF430" box="[1204,1305,764,788]" pageId="5" pageNumber="6" type="male">17 males</specimenCount>
(
<collectionCode id="ED7DAE44BE05292AFAE8FD75FADAF430" box="[1320,1389,764,788]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="6">INPA</collectionCode>
),
<specimenCount id="9D6AFD08BE05292AFA40FD75FF6FF413" pageId="5" pageNumber="6" type="male">12 males</specimenCount>
(
<collectionCode id="ED7DAE44BE05292AFF27FC96FEFCF413" box="[231,331,799,823]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="6">MZUSP</collectionCode>
); 14.
<date id="FFD21041BE05292AFEBFFC96FE4AF413" box="[383,509,799,823]" pageId="5" pageNumber="6" value="1978-02-18">
II.1978,
<specimenCount id="9D6AFD08BE05292AFE21FC96FE4AF413" box="[481,509,799,823]" pageId="5" pageNumber="6" type="male">18</specimenCount>
</date>
males (
<collectionCode id="ED7DAE44BE05292AFD93FC96FD2FF413" box="[595,664,799,823]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="6">INPA</collectionCode>
),
<specimenCount id="9D6AFD08BE05292AFD6BFC96FCB6F413" box="[683,769,799,823]" pageId="5" pageNumber="6" type="male">4 males</specimenCount>
(
<collectionCode id="ED7DAE44BE05292AFCD0FC96FCC3F413" box="[784,884,799,823]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="6">MZUSP</collectionCode>
); 21.
<date id="FFD21041BE05292AFC68FC96FBAFF413" box="[936,1048,799,823]" pageId="5" pageNumber="6" value="1978-02-09">
II.1978,
<specimenCount id="9D6AFD08BE05292AFBCAFC96FBAFF413" box="[1034,1048,799,823]" pageId="5" pageNumber="6" type="male">9</specimenCount>
</date>
males (
<collectionCode id="ED7DAE44BE05292AFBAFFC96FB02F413" box="[1135,1205,799,823]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="6">INPA</collectionCode>
).
</paragraph>
<paragraph id="8BD33681BE05292AFF05FCCAFD53F478" blockId="5.[151,1437,151,932]" box="[197,740,835,860]" pageId="5" pageNumber="6">
<emphasis id="B918EA93BE05292AFF05FCCAFEFCF478" bold="true" box="[197,331,835,860]" pageId="5" pageNumber="6">Ocurrence.</emphasis>
<collectingCountry id="F37B7611BE05292AFE92FCCDFE0CF478" box="[338,443,836,860]" name="Brazil" pageId="5" pageNumber="6">BRAZIL</collectingCountry>
. Amazonas state: Manaus.
</paragraph>
</subSubSection>
<subSubSection id="C376650ABE05292AFF05FCE1FBCBF480" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8BD33681BE05292AFF05FCE1FBCBF480" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="6">
<emphasis id="B918EA93BE05292AFF05FCE1FE8EF4A4" bold="true" box="[197,313,872,896]" pageId="5" pageNumber="6">Remarks.</emphasis>
Hundreds of male specimens were collected with malaise trap from November to May, with peak population in February at the height of the rainy season. No females have been collected.
</paragraph>
</subSubSection>
</treatment>
</document>
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