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<document id="BB79EEA265CB3390D49251A14D1B0F05" ID-CLB-Dataset="29039" ID-DOI="10.11646/zootaxa.4459.1.1" ID-GBIF-Dataset="9acf71bc-6414-4878-8af0-a64d3ec1fea6" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458302" ID-ZooBank="633A9498-FCC6-4439-BE33-A45B9E51B550" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539287574730" checkinUser="plazi" docAuthor="Grandinete, Yuri Campanholo, Noll, Fernando Barbosa &amp; Carpenter, James" docDate="2018" docId="03C5878FFFFCF313FF309792D674F9AB" docLanguage="en" docName="zootaxa.4459.1.1.pdf" docOrigin="Zootaxa 4459 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eumenes crucifera Provancher 1888" docType="treatment" docVersion="9" lastPageNumber="20" masterDocId="FFFCFFF7FFF3F300FFA79155D339FFDF" masterDocTitle="Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World" masterLastPageNumber="52" masterPageNumber="1" pageNumber="16" updateTime="1698577660185" updateUser="ExternalLinkService">
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<mods:title id="91CE543E99E0B3A196D49DF55893138F">Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World</mods:title>
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<treatment id="03C5878FFFFCF313FF309792D674F9AB" ID-DOI="http://doi.org/10.5281/zenodo.5986374" ID-GBIF-Taxon="148692478" ID-Zenodo-Dep="5986374" LSID="urn:lsid:plazi:treatment:03C5878FFFFCF313FF309792D674F9AB" httpUri="http://treatment.plazi.org/id/03C5878FFFFCF313FF309792D674F9AB" lastPageId="19" lastPageNumber="20" pageId="15" pageNumber="16">
<subSubSection id="C3766512FFFCF30FFF309792D212F89C" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BD33699FFFCF30FFF309792D164F93D" blockId="15.[151,605,1735,1762]" box="[151,605,1735,1762]" pageId="15" pageNumber="16">
<heading id="D09B81F5FFFCF30FFF309792D164F93D" bold="true" box="[151,605,1735,1762]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<taxonomicName id="4C6C4D1AFFFCF30FFF309792D164F93D" ID-CoL="3CDFT" authority="Provancher, 1888" authorityName="Provancher" authorityYear="1888" box="[151,605,1735,1762]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFFCF30FFF309792D164F93D" bold="true" box="[151,605,1735,1762]" pageId="15" pageNumber="16">
<emphasis id="B918EA8BFFFCF30FFF309792D244F93E" bold="true" box="[151,381,1735,1761]" italics="true" pageId="15" pageNumber="16">Eumenes crucifera</emphasis>
Provancher, 1888
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD33699FFFCF30FFF30965AD212F89C" blockId="15.[151,1435,1806,2012]" pageId="15" pageNumber="16">
<taxonomicName id="4C6C4D1AFFFCF30FFF30965AD17CF8FB" authority="de Saussure, 1855" authorityName="de Saussure" authorityYear="1855" box="[151,581,1806,1828]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="globulosus">
<emphasis id="B918EA8BFFFCF30FFF30965AD17CF8FB" bold="true" box="[151,581,1806,1828]" pageId="15" pageNumber="16">
<emphasis id="B918EA8BFFFCF30FFF30965AD24AF8FB" bold="true" box="[151,371,1806,1828]" italics="true" pageId="15" pageNumber="16">Eumenes globulosus</emphasis>
de Saussure, 1855
</emphasis>
</taxonomicName>
, 3: 139 [junior secondary homonym of
<emphasis id="B918EA8BFFFCF30FFC539645D7A7F8FB" box="[1012,1182,1806,1829]" italics="true" pageId="15" pageNumber="16">Uespa globulosa</emphasis>
Fourcroy, 1785].
<emphasis id="B918EA8BFFFCF30FFAF8965BD21FF89C" bold="true" pageId="15" pageNumber="16">NEW STATUS</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFFCF310FF609619D2E0FED5" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BD33699FFFCF30FFF609619D0CCF8BD" blockId="15.[151,1435,1806,2012]" box="[199,1013,1868,1890]" pageId="15" pageNumber="16">
<emphasis id="B918EA8BFFFCF30FFF609619D203F8BD" bold="true" box="[199,314,1868,1890]" pageId="15" pageNumber="16">Type data:</emphasis>
Holotype female at MHNG
<emphasis id="B918EA8BFFFCF30FFDFB9619D1EFF8BD" bold="true" box="[604,726,1868,1890]" pageId="15" pageNumber="16">(examined)</emphasis>
. Paratype female at USNM.
</paragraph>
<paragraph id="8BD33699FFFCF30FFF60963FD081F85F" blockId="15.[151,1435,1806,2012]" box="[199,952,1898,1920]" pageId="15" pageNumber="16">
<materialsCitation id="3B043CC4FFFCF30FFF60963FD1FAF85F" ID-GBIF-Occurrence="1917355186" box="[199,707,1898,1920]" location="Nord" pageId="15" pageNumber="16" specimenCount="2" stateProvince="Nord" typeStatus="holotype">
<emphasis id="B918EA8BFFFCF30FFF60963FD263F85F" bold="true" box="[199,346,1898,1920]" pageId="15" pageNumber="16">
<typeStatus id="54D7883BFFFCF30FFF60963FD3C5F85F" box="[199,252,1898,1920]" pageId="15" pageNumber="16">Type</typeStatus>
locality:
</emphasis>
“LAmérique du
<collectingRegion id="49A8F87BFFFCF30FFDAC963FD178F85F" box="[523,577,1898,1920]" country="Haiti" name="Nord" pageId="15" pageNumber="16">Nord</collectingRegion>
” (
<typeStatus id="54D7883BFFFCF30FFDFF963FD185F85F" box="[600,700,1898,1920]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
)
</materialsCitation>
.
<materialsCitation id="3B043CC4FFFCF30FFD6A963FD08DF85F" ID-GBIF-Occurrence="1917355188" box="[717,948,1898,1920]" location="Colorado" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Colorado" typeStatus="paratype">
<collectingRegion id="49A8F87BFFFCF30FFD71963FD00EF85F" box="[726,823,1898,1920]" country="United States of America" name="Colorado" pageId="15" pageNumber="16">Colorado</collectingRegion>
” (
<typeStatus id="54D7883BFFFCF30FFCE8963FD097F85F" box="[847,942,1898,1920]" pageId="15" pageNumber="16" type="paratype">Paratype</typeStatus>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD33699FFFCF30FFF6096DCD2D2F840" blockId="15.[151,1435,1806,2012]" box="[199,491,1929,1951]" pageId="15" pageNumber="16">de Saussure, 1875, 254: 101.</paragraph>
<paragraph id="8BD33699FFFCF30FFF6096FDD2F4F861" blockId="15.[151,1435,1806,2012]" box="[199,461,1960,1982]" pageId="15" pageNumber="16">Cresson, 1887: 287 (cat.).</paragraph>
<paragraph id="8BD33699FFFCF30FFF609693D2C2F803" blockId="15.[151,1435,1806,2012]" box="[199,507,1990,2012]" pageId="15" pageNumber="16">Dalla Torre, 1894, 9: 25 (cat.).</paragraph>
<paragraph id="8BD33699FFE3F310FF6091CDD131FF71" blockId="16.[151,1436,152,2014]" box="[199,520,152,174]" pageId="16" pageNumber="17">Dalla Torre, 1904, 19: 23 (cat.).</paragraph>
<paragraph id="8BD33699FFE3F310FF6091E2D2E2FF13" blockId="16.[151,1436,152,2014]" box="[199,475,182,204]" pageId="16" pageNumber="17">Bequaert, 1928, 101: 1004.</paragraph>
<paragraph id="8BD33699FFE3F310FF609183D12EFF34" blockId="16.[151,1436,152,2014]" box="[199,535,213,235]" pageId="16" pageNumber="17">
<bibRefCitation id="EFFD4B68FFE3F310FF609183D25DFF34" box="[199,356,213,235]" pageId="16" pageNumber="17" refId="ref42258" refString="Bequaert, J. (1938) The three Eumenes of Canada and the Northeastern United States; with notes on other North American species (Hymenoptera; Vespidae). Bulletin of the Brooklyn Entomological Society, 33 (2), 59 - 70." type="journal article">Bequaert, 1938</bibRefCitation>
, 33: 62 (key), 63.
</paragraph>
<paragraph id="8BD33699FFE3F310FF6091A0D2E0FED5" blockId="16.[151,1436,152,2014]" box="[199,473,244,266]" pageId="16" pageNumber="17">Bequaert, 1943, 38 (2): 43.</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE3F310FF309046D160FEF7" box="[151,601,274,296]" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BD33699FFE3F310FF309046D160FEF7" blockId="16.[151,1436,152,2014]" box="[151,601,274,296]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF309046D16CFEF7" ID-CoL="3CDFT" authority="Provancher, 1888: 421" authorityName="Provancher" authorityPageNumber="421" authorityYear="1888" box="[151,597,274,296]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE3F310FF309046D126FEF7" bold="true" box="[151,543,274,296]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF309046D267FEF7" bold="true" box="[151,350,274,296]" italics="true" pageId="16" pageNumber="17">Eumenes crucifera</emphasis>
Provancher, 1888
</emphasis>
: 421
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE3F310FF609064D072FCCC" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BD33699FFE3F310FF609064D27FFEB9" blockId="16.[151,1436,152,2014]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF609064D204FE98" bold="true" box="[199,317,305,327]" pageId="16" pageNumber="17">
<typeStatus id="54D7883BFFE3F310FF609064D3C5FE98" box="[199,252,305,327]" pageId="16" pageNumber="17">Type</typeStatus>
data:
</emphasis>
<typeStatus id="54D7883BFFE3F310FEE19064D29DFE98" box="[326,420,305,327]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
female at
<collectionCode id="ED7DAE5CFFE3F310FDB09064D159FE98" box="[535,608,305,327]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17">USNM</collectionCode>
(n° 1977)
<emphasis id="B918EA8BFFE3F310FD749064D075FE98" bold="true" box="[723,844,305,327]" pageId="16" pageNumber="17">(examined)</emphasis>
. Three
<typeStatus id="54D7883BFFE3F310FC079067D738FE98" box="[928,1025,306,327]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
(two males and one female) at
<collectionCode id="ED7DAE5CFFE3F310FAF49064D6A5FE98" box="[1363,1436,305,327]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17">USNM</collectionCode>
<emphasis id="B918EA8BFFE3F310FF609005D279FEB9" bold="true" box="[199,320,336,358]" pageId="16" pageNumber="17">(examined)</emphasis>
.
</paragraph>
<paragraph id="8BD33699FFE3F310FF60903BD7CBFE5B" blockId="16.[151,1436,152,2014]" box="[199,1266,366,388]" pageId="16" pageNumber="17">
<materialsCitation id="3B043CC4FFE3F310FF60903BD7CBFE5B" ID-GBIF-Occurrence="1917355165" box="[199,1266,366,388]" county="Los Angeles Co." location="Two" pageId="16" pageNumber="17" specimenCount="2" stateProvince="California" typeStatus="holotype">
<emphasis id="B918EA8BFFE3F310FF60903BD263FE5B" bold="true" box="[199,346,366,388]" pageId="16" pageNumber="17">
<typeStatus id="54D7883BFFE3F310FF60903BD3C5FE5B" box="[199,252,366,388]" pageId="16" pageNumber="17">Type</typeStatus>
locality:
</emphasis>
“Los Angeles” (
<typeStatus id="54D7883BFFE3F310FDA4903BD15EFE5B" box="[515,615,366,388]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
).
<location id="8EB36042FFE3F310FDD0903AD19AFE5B" LSID="urn:lsid:plazi:treatment:03C5878FFFFCF313FF309792D674F9AB:8EB36042FFE3F310FDD0903AD19AFE5B" box="[631,675,367,388]" county="Los Angeles Co." name="Two" pageId="16" pageNumber="17" stateProvince="California">Two</location>
males and one female from
<collectingRegion id="49A8F87BFFE3F310FC63903BD712FE5B" box="[964,1067,366,388]" country="United States of America" name="California" pageId="16" pageNumber="17">California</collectingRegion>
(
<collectingCounty id="62B24E15FFE3F310FB9F903BD7DFFE5B" box="[1080,1254,366,388]" pageId="16" pageNumber="17">Los Angeles Co.</collectingCounty>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD33699FFE3F310FF6090D8D2C2FE7C" blockId="16.[151,1436,152,2014]" box="[199,507,397,419]" pageId="16" pageNumber="17">Dalla Torre, 1894, 9: 22 (cat.).</paragraph>
<paragraph id="8BD33699FFE3F310FF6090F9D131FE1D" blockId="16.[151,1436,152,2014]" box="[199,520,428,450]" pageId="16" pageNumber="17">Dalla Torre, 1904, 19: 22 (cat.).</paragraph>
<paragraph id="8BD33699FFE3F310FF60909ED170FE3F" blockId="16.[151,1436,152,2014]" box="[199,585,458,480]" pageId="16" pageNumber="17">
<bibRefCitation id="EFFD4B68FFE3F310FF60909ED25DFE3F" box="[199,356,458,480]" pageId="16" pageNumber="17" refId="ref42258" refString="Bequaert, J. (1938) The three Eumenes of Canada and the Northeastern United States; with notes on other North American species (Hymenoptera; Vespidae). Bulletin of the Brooklyn Entomological Society, 33 (2), 59 - 70." type="journal article">Bequaert, 1938</bibRefCitation>
, 33: 69 (note on type).
</paragraph>
<paragraph id="8BD33699FFE3F310FF6090BCD16AFE20" blockId="16.[151,1436,152,2014]" box="[199,595,489,511]" pageId="16" pageNumber="17">
<bibRefCitation id="EFFD4B68FFE3F310FF6090BCD2BEFE20" box="[199,391,489,511]" pageId="16" pageNumber="17" refId="ref43608" refString="MacLachlan, W. B. (1980) A key to and notes on the Eumenes of America North of Mexico (Hymenoptera: Eumenidae). Journal of the Kansas Entomological Society, 53 (3), 617 - 621." type="journal article">MacLachlan, 1980</bibRefCitation>
, 53: 618, 620 (key).
</paragraph>
<paragraph id="8BD33699FFE3F310FF60935DD110FDC1" blockId="16.[151,1436,152,2014]" box="[199,553,520,542]" pageId="16" pageNumber="17">
Buck
<emphasis id="B918EA8BFFE3F310FEA4935CD20FFDC1" box="[259,310,520,542]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
, 2008, 5: 44 (key), 155.
</paragraph>
<paragraph id="8BD33699FFE3F310FF309372D1CAFDE3" blockId="16.[151,1436,152,2014]" box="[151,755,550,572]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF309372D1ADFDE3" authority="Viereck, 1907" authorityName="Viereck" authorityYear="1907" box="[151,660,550,572]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="marginilineatus" subGenus="Alpha">
<emphasis id="B918EA8BFFE3F310FF309372D1ADFDE3" bold="true" box="[151,660,550,572]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF309372D3C1FDE3" bold="true" box="[151,248,551,572]" italics="true" pageId="16" pageNumber="17">Eumenes</emphasis>
(
<emphasis id="B918EA8BFFE3F310FEA19373D27CFDE3" bold="true" box="[262,325,550,572]" italics="true" pageId="16" pageNumber="17">Alpha</emphasis>
)
<emphasis id="B918EA8BFFE3F310FEF39373D2C3FDE3" bold="true" box="[340,506,550,572]" italics="true" pageId="16" pageNumber="17">marginilineatus</emphasis>
Viereck, 1907
</emphasis>
</taxonomicName>
, 33: 381.
</paragraph>
<paragraph id="8BD33699FFE3F310FF609310D197FD84" blockId="16.[151,1436,152,2014]" box="[199,686,581,603]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF609310D203FD84" bold="true" box="[199,314,581,603]" pageId="16" pageNumber="17">
<typeStatus id="54D7883BFFE3F310FF609310D3C5FD84" box="[199,252,581,603]" pageId="16" pageNumber="17">Type</typeStatus>
data:
</emphasis>
<typeStatus id="54D7883BFFE3F310FEE69313D24AFD84" box="[321,371,582,603]" pageId="16" pageNumber="17">Type</typeStatus>
male at
<collectionCode id="ED7DAE5CFFE3F310FE6E9313D110FD84" box="[457,553,582,603]" pageId="16" pageNumber="17">KUNHM</collectionCode>
<emphasis id="B918EA8BFFE3F310FD889310D191FD84" bold="true" box="[559,680,581,603]" pageId="16" pageNumber="17">(examined)</emphasis>
.
</paragraph>
<paragraph id="8BD33699FFE3F310FF609331D003FDA5" blockId="16.[151,1436,152,2014]" box="[199,826,612,634]" pageId="16" pageNumber="17">
<materialsCitation id="3B043CC4FFE3F310FF609331D003FDA5" ID-GBIF-Occurrence="1917355214" box="[199,826,612,634]" country="United States of America" county="Larimer County" location="Este's Park" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Colorado" typeStatus="holotype">
<emphasis id="B918EA8BFFE3F310FF609331D263FDA5" bold="true" box="[199,346,612,634]" pageId="16" pageNumber="17">
<typeStatus id="54D7883BFFE3F310FF609331D3C5FDA5" box="[199,252,612,634]" pageId="16" pageNumber="17">Type</typeStatus>
locality:
</emphasis>
<location id="8EB36042FFE3F310FEC79331D2EAFDA5" LSID="urn:lsid:plazi:treatment:03C5878FFFFCF313FF309792D674F9AB:8EB36042FFE3F310FEC79331D2EAFDA5" box="[352,467,612,634]" country="United States of America" county="Larimer County" name="Este's Park" pageId="16" pageNumber="17" stateProvince="Colorado">Estes Park</location>
,
<collectingCounty id="62B24E15FFE3F310FE7A9331D1B9FDA5" box="[477,640,612,634]" pageId="16" pageNumber="17">Larimer County</collectingCounty>
,
<collectingRegion id="49A8F87BFFE3F310FD2C9331D1D0FDA5" box="[651,745,612,634]" country="United States of America" name="Colorado" pageId="16" pageNumber="17">Colorado</collectingRegion>
(
<collectingCountry id="F37B7609FFE3F310FD509331D016FDA5" box="[759,815,612,634]" name="United States of America" pageId="16" pageNumber="17">USA</collectingCountry>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD33699FFE3F310FF6093D7D1AFFD47" blockId="16.[151,1436,152,2014]" box="[199,662,642,664]" pageId="16" pageNumber="17">
Viereck, 1908, 33: 388 (
<emphasis id="B918EA8BFFE3F310FE1A93D7D151FD47" box="[445,616,642,664]" italics="true" pageId="16" pageNumber="17">emarginilineatus</emphasis>
[!]).
</paragraph>
<paragraph id="8BD33699FFE3F310FF6093F7D170FD68" blockId="16.[151,1436,152,2014]" box="[199,585,673,695]" pageId="16" pageNumber="17">
<bibRefCitation id="EFFD4B68FFE3F310FF6093F7D25DFD68" box="[199,356,673,695]" pageId="16" pageNumber="17" refId="ref42258" refString="Bequaert, J. (1938) The three Eumenes of Canada and the Northeastern United States; with notes on other North American species (Hymenoptera; Vespidae). Bulletin of the Brooklyn Entomological Society, 33 (2), 59 - 70." type="journal article">Bequaert, 1938</bibRefCitation>
, 33: 69 (note on type).
</paragraph>
<paragraph id="8BD33699FFE3F310FF309395D089FD09" blockId="16.[151,1436,152,2014]" box="[151,944,703,726]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF309395D19AFD09" authority="Viereck, 1908" authorityName="Viereck" authorityYear="1908" box="[151,675,703,726]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bolliformis" subGenus="Pachymenes">
<emphasis id="B918EA8BFFE3F310FF309395D19AFD09" bold="true" box="[151,675,703,726]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF309395D3C1FD0A" bold="true" box="[151,248,704,725]" italics="true" pageId="16" pageNumber="17">Eumenes</emphasis>
(
<emphasis id="B918EA8BFFE3F310FEA193EAD2B0FD0A" bold="true" box="[262,393,703,725]" italics="true" pageId="16" pageNumber="17">Pachymenes</emphasis>
)
<emphasis id="B918EA8BFFE3F310FE3F93EAD133FD0A" bold="true" box="[408,522,703,725]" italics="true" pageId="16" pageNumber="17">bolliformis</emphasis>
Viereck, 1908
</emphasis>
</taxonomicName>
, 33: 387.
<emphasis id="B918EA8BFFE3F310FCAF9395D089FD09" bold="true" box="[776,944,704,726]" pageId="16" pageNumber="17">NEW STATUS.</emphasis>
</paragraph>
<paragraph id="8BD33699FFE3F310FF60938BD710FD2B" blockId="16.[151,1436,152,2014]" box="[199,1065,734,756]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF60938BD203FD2B" bold="true" box="[199,314,734,756]" pageId="16" pageNumber="17">
<typeStatus id="54D7883BFFE3F310FF60938BD3C5FD2B" box="[199,252,734,756]" pageId="16" pageNumber="17">Type</typeStatus>
data:
</emphasis>
<typeStatus id="54D7883BFFE3F310FEE6938AD290FD2B" box="[321,425,735,756]" pageId="16" pageNumber="17" type="lectotype">Lectotype</typeStatus>
, by present designation, female and
<typeStatus id="54D7883BFFE3F310FCBE938BD09AFD2B" box="[793,931,734,756]" pageId="16" pageNumber="17" type="paralectotype">Paralectotype</typeStatus>
at
<collectionCode id="ED7DAE5CFFE3F310FC66938AD71DFD2B" box="[961,1060,735,756]" pageId="16" pageNumber="17">KUNHM</collectionCode>
.
</paragraph>
<paragraph id="8BD33699FFE3F310FF6093A8D072FCCC" blockId="16.[151,1436,152,2014]" box="[199,843,765,787]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF6093A8D263FCCC" bold="true" box="[199,346,765,787]" pageId="16" pageNumber="17">Type locality:</emphasis>
“S. Arizona” (both lectotype and paralectotype).
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE3F310FF309249D072FCED" box="[151,843,795,818]" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BD33699FFE3F310FF309249D072FCED" blockId="16.[151,1436,152,2014]" box="[151,843,795,818]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF309249D2F7FCED" ID-CoL="6GWPJ" authority="Isely, 1917" authorityName="Isely" authorityYear="1917" box="[151,462,795,818]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B918EA8BFFE3F310FF309249D2F7FCED" bold="true" box="[151,462,795,818]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF309249D261FCEE" bold="true" box="[151,344,795,817]" italics="true" pageId="16" pageNumber="17">Eumenes robustus</emphasis>
<bibRefCitation id="EFFD4B68FFE3F310FEF99249D2F7FCED" box="[350,462,796,818]" pageId="16" pageNumber="17" refId="ref43377" refString="Isely, D. (1917) A synopsis of the petiolate wasps of the family Eumenidae (Hymenoptera), found in America North of Mexico. Annals of the Entomological Society of America, 10 (4), 345 - 366. https: // doi. org / 10.1093 / aesa / 10.4.345" type="journal article">Isely, 1917</bibRefCitation>
</emphasis>
</taxonomicName>
, 10: 347 (key), 360.
<emphasis id="B918EA8BFFE3F310FD049249D072FCED" bold="true" box="[675,843,796,818]" pageId="16" pageNumber="17">NEW STATUS.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE3F310FF60926FD2BFFC14" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BD33699FFE3F310FF60926FD1F3FC8F" blockId="16.[151,1436,152,2014]" box="[199,714,826,848]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF60926FD203FC8F" bold="true" box="[199,314,826,848]" pageId="16" pageNumber="17">Type data:</emphasis>
Holotype female at USNM (n° 21382).
</paragraph>
<paragraph id="8BD33699FFE3F310FF60920CD195FCB0" blockId="16.[151,1436,152,2014]" box="[199,684,857,879]" pageId="16" pageNumber="17">
<materialsCitation id="3B043CC4FFE3F310FF60920CD195FCB0" ID-GBIF-Occurrence="1917355185" box="[199,684,857,879]" location="Beulah" pageId="16" pageNumber="17" specimenCount="1" stateProvince="San Miguel" typeStatus="holotype">
<emphasis id="B918EA8BFFE3F310FF60920CD263FCB0" bold="true" box="[199,346,857,879]" pageId="16" pageNumber="17">
<typeStatus id="54D7883BFFE3F310FF60920CD3C5FCB0" box="[199,252,857,879]" pageId="16" pageNumber="17">Type</typeStatus>
locality:
</emphasis>
<location id="8EB36042FFE3F310FECD920CD28CFCB0" LSID="urn:lsid:plazi:treatment:03C5878FFFFCF313FF309792D674F9AB:8EB36042FFE3F310FECD920CD28CFCB0" box="[362,437,857,879]" name="Beulah" pageId="16" pageNumber="17" stateProvince="San Miguel">Beulah</location>
,
<collectingRegion id="49A8F87BFFE3F310FE18920CD10DFCB0" box="[447,564,857,879]" country="El Salvador" name="San Miguel" pageId="16" pageNumber="17">San Miguel</collectingRegion>
Co., N.M.”
</materialsCitation>
</paragraph>
<paragraph id="8BD33699FFE3F310FF60922CD16AFC51" blockId="16.[151,1436,152,2014]" box="[199,595,888,910]" pageId="16" pageNumber="17">
<bibRefCitation id="EFFD4B68FFE3F310FF60922CD25DFC51" box="[199,356,888,910]" pageId="16" pageNumber="17" refId="ref42258" refString="Bequaert, J. (1938) The three Eumenes of Canada and the Northeastern United States; with notes on other North American species (Hymenoptera; Vespidae). Bulletin of the Brooklyn Entomological Society, 33 (2), 59 - 70." type="journal article">Bequaert, 1938</bibRefCitation>
, 33: 70 (note on types).
</paragraph>
<paragraph id="8BD33699FFE3F310FF6092C2D28CFC73" blockId="16.[151,1436,152,2014]" box="[199,437,918,940]" pageId="16" pageNumber="17">
<bibRefCitation id="EFFD4B68FFE3F310FF6092C2D25DFC73" box="[199,356,918,940]" pageId="16" pageNumber="17" refId="ref42355" refString="Bequaert, J. (1944) The genus Eumenes in the Dominion of Canada. Le Naturaliste Canadien, 71, 75 - 88." type="journal article">Bequaert, 1944</bibRefCitation>
, 71: 81.
</paragraph>
<paragraph id="8BD33699FFE3F310FF6092E0D2BFFC14" blockId="16.[151,1436,152,2014]" box="[199,390,949,971]" pageId="16" pageNumber="17">Bohart, 1951: 884.</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE3F310FF309281D117FC35" box="[151,558,979,1002]" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BD33699FFE3F310FF309281D117FC35" blockId="16.[151,1436,152,2014]" box="[151,558,979,1002]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF309281D2F7FC35" ID-CoL="3CDPF" authority="Isely, 1917" authorityName="Isely" authorityYear="1917" box="[151,462,979,1002]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sternalis">
<emphasis id="B918EA8BFFE3F310FF309281D2F7FC35" bold="true" box="[151,462,979,1002]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF309281D26EFC36" bold="true" box="[151,343,979,1001]" italics="true" pageId="16" pageNumber="17">Eumenes sternalis</emphasis>
<bibRefCitation id="EFFD4B68FFE3F310FEFA9281D2F7FC35" box="[349,462,980,1002]" pageId="16" pageNumber="17" refId="ref43377" refString="Isely, D. (1917) A synopsis of the petiolate wasps of the family Eumenidae (Hymenoptera), found in America North of Mexico. Annals of the Entomological Society of America, 10 (4), 345 - 366. https: // doi. org / 10.1093 / aesa / 10.4.345" type="journal article">Isely, 1917</bibRefCitation>
</emphasis>
</taxonomicName>
, 10: 353.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE3F310FF6092A7D16AFB99" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BD33699FFE3F310FF6092A7D070FBD7" blockId="16.[151,1436,152,2014]" box="[199,841,1010,1032]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF6092A7D203FBD7" bold="true" box="[199,314,1010,1032]" pageId="16" pageNumber="17">Type data:</emphasis>
Holotype female at USNM (n° 21380)
<emphasis id="B918EA8BFFE3F310FD6D92A7D07AFBD7" bold="true" box="[714,835,1010,1032]" pageId="16" pageNumber="17">(examined)</emphasis>
.
</paragraph>
<paragraph id="8BD33699FFE3F310FF609544D1A7FBF8" blockId="16.[151,1436,152,2014]" box="[199,670,1041,1063]" pageId="16" pageNumber="17">
<materialsCitation id="3B043CC4FFE3F310FF609544D1A7FBF8" ID-GBIF-Occurrence="1917355136" box="[199,670,1041,1063]" country="United States of America" location="Beaver Canyon" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Utah" typeStatus="holotype">
<emphasis id="B918EA8BFFE3F310FF609544D263FBF8" bold="true" box="[199,346,1041,1063]" pageId="16" pageNumber="17">
<typeStatus id="54D7883BFFE3F310FF609544D3C5FBF8" box="[199,252,1041,1063]" pageId="16" pageNumber="17">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F37B7609FFE3F310FEC79544D29FFBF8" box="[352,422,1041,1063]" name="United States of America" pageId="16" pageNumber="17">U.S.A.</collectingCountry>
(
<location id="8EB36042FFE3F310FE159544D16AFBF8" LSID="urn:lsid:plazi:treatment:03C5878FFFFCF313FF309792D674F9AB:8EB36042FFE3F310FE159544D16AFBF8" box="[434,595,1041,1063]" country="United States of America" name="Beaver Canyon" pageId="16" pageNumber="17" stateProvince="Utah">Beaver Canyon</location>
,
<collectingRegion id="49A8F87BFFE3F310FDFA9544D1AAFBF8" box="[605,659,1041,1063]" country="United States of America" name="Utah" pageId="16" pageNumber="17">Utah</collectingRegion>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD33699FFE3F310FF609564D16AFB99" blockId="16.[151,1436,152,2014]" box="[199,595,1072,1094]" pageId="16" pageNumber="17">
<bibRefCitation id="EFFD4B68FFE3F310FF609564D25DFB99" box="[199,356,1072,1094]" pageId="16" pageNumber="17" refId="ref42258" refString="Bequaert, J. (1938) The three Eumenes of Canada and the Northeastern United States; with notes on other North American species (Hymenoptera; Vespidae). Bulletin of the Brooklyn Entomological Society, 33 (2), 59 - 70." type="journal article">Bequaert, 1938</bibRefCitation>
, 33: 70 (note on types).
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE3F310FF30951AD167FBBB" box="[151,606,1102,1124]" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BD33699FFE3F310FF30951AD167FBBB" blockId="16.[151,1436,152,2014]" box="[151,606,1102,1124]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF30951AD2C4FBBB" ID-CoL="3CDQG" authority="Isely, 1917" authorityName="Isely" authorityYear="1917" box="[151,509,1102,1124]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="xanthogaster">
<emphasis id="B918EA8BFFE3F310FF30951AD2C4FBBB" bold="true" box="[151,509,1102,1124]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF30951AD2BEFBBB" bold="true" box="[151,391,1102,1124]" italics="true" pageId="16" pageNumber="17">Eumenes xanthogaster</emphasis>
<bibRefCitation id="EFFD4B68FFE3F310FE2B951BD2C4FBBB" box="[396,509,1102,1124]" pageId="16" pageNumber="17" refId="ref43377" refString="Isely, D. (1917) A synopsis of the petiolate wasps of the family Eumenidae (Hymenoptera), found in America North of Mexico. Annals of the Entomological Society of America, 10 (4), 345 - 366. https: // doi. org / 10.1093 / aesa / 10.4.345" type="journal article">Isely, 1917</bibRefCitation>
</emphasis>
</taxonomicName>
, 10: 359.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE3F310FF609538D14DFAC3" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BD33699FFE3F310FF609538D00CFB5C" blockId="16.[151,1436,152,2014]" box="[199,821,1133,1155]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF609538D203FB5C" bold="true" box="[199,314,1133,1155]" pageId="16" pageNumber="17">Type data:</emphasis>
Holotype male at USNM (n° 21381)
<emphasis id="B918EA8BFFE3F310FD119538D016FB5C" bold="true" box="[694,815,1133,1155]" pageId="16" pageNumber="17">(examined)</emphasis>
.
</paragraph>
<paragraph id="8BD33699FFE3F310FF6095D9D18FFB7D" blockId="16.[151,1436,152,2014]" box="[199,694,1164,1186]" pageId="16" pageNumber="17">
<materialsCitation id="3B043CC4FFE3F310FF6095D9D18FFB7D" ID-GBIF-Occurrence="1917355182" box="[199,694,1164,1186]" country="United States of America" location="Los Angeles" pageId="16" pageNumber="17" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="B918EA8BFFE3F310FF6095D9D263FB7D" bold="true" box="[199,346,1164,1186]" pageId="16" pageNumber="17">
<typeStatus id="54D7883BFFE3F310FF6095D9D3C5FB7D" box="[199,252,1164,1186]" pageId="16" pageNumber="17">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F37B7609FFE3F310FEC795D9D29FFB7D" box="[352,422,1164,1186]" name="United States of America" pageId="16" pageNumber="17">U.S.A.</collectingCountry>
(
<location id="8EB36042FFE3F310FE1495D9D10CFB7D" LSID="urn:lsid:plazi:treatment:03C5878FFFFCF313FF309792D674F9AB:8EB36042FFE3F310FE1495D9D10CFB7D" box="[435,565,1164,1186]" country="United States of America" name="Los Angeles" pageId="16" pageNumber="17" stateProvince="California">Los Angeles</location>
,
<collectingRegion id="49A8F87BFFE3F310FDE795D9D195FB7D" box="[576,684,1164,1186]" country="United States of America" name="California" pageId="16" pageNumber="17">California</collectingRegion>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD33699FFE3F310FF6095FED170FB1F" blockId="16.[151,1436,152,2014]" box="[199,585,1194,1216]" pageId="16" pageNumber="17">
<bibRefCitation id="EFFD4B68FFE3F310FF6095FED25DFB1F" box="[199,356,1194,1216]" pageId="16" pageNumber="17" refId="ref42258" refString="Bequaert, J. (1938) The three Eumenes of Canada and the Northeastern United States; with notes on other North American species (Hymenoptera; Vespidae). Bulletin of the Brooklyn Entomological Society, 33 (2), 59 - 70." type="journal article">Bequaert, 1938</bibRefCitation>
, 33: 70 (note on type).
</paragraph>
<paragraph id="8BD33699FFE3F310FF30959FD18AFB00" blockId="16.[151,1436,152,2014]" box="[151,691,1225,1247]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF30959FD25EFB00" authorityName="Viereck" authorityYear="1908" box="[151,359,1225,1247]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bolliformis">
<emphasis id="B918EA8BFFE3F310FF30959FD25EFB00" box="[151,359,1225,1247]" italics="true" pageId="16" pageNumber="17">Eumenes bolliformis</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFFD4B68FFE3F310FED2959CD2DFFB00" box="[373,486,1225,1247]" pageId="16" pageNumber="17" refId="ref43377" refString="Isely, D. (1917) A synopsis of the petiolate wasps of the family Eumenidae (Hymenoptera), found in America North of Mexico. Annals of the Entomological Society of America, 10 (4), 345 - 366. https: // doi. org / 10.1093 / aesa / 10.4.345" type="journal article">Isely, 1917</bibRefCitation>
, 10: 347 (key), 352.
</paragraph>
<paragraph id="8BD33699FFE3F310FF6095BCD170FB21" blockId="16.[151,1436,152,2014]" box="[199,585,1256,1278]" pageId="16" pageNumber="17">
<bibRefCitation id="EFFD4B68FFE3F310FF6095BCD25DFB21" box="[199,356,1256,1278]" pageId="16" pageNumber="17" refId="ref42258" refString="Bequaert, J. (1938) The three Eumenes of Canada and the Northeastern United States; with notes on other North American species (Hymenoptera; Vespidae). Bulletin of the Brooklyn Entomological Society, 33 (2), 59 - 70." type="journal article">Bequaert, 1938</bibRefCitation>
, 33: 68 (note on type).
</paragraph>
<paragraph id="8BD33699FFE3F310FF30945DD14DFAC3" blockId="16.[151,1436,152,2014]" box="[151,628,1286,1309]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF30945DD2AEFAC3" authorityName="Viereck" authorityYear="1907" box="[151,407,1286,1309]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="marginilineatus">
<emphasis id="B918EA8BFFE3F310FF30945DD2AEFAC3" box="[151,407,1286,1309]" italics="true" pageId="16" pageNumber="17">Eumenes marginilineatus</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFFD4B68FFE3F310FE029453D12FFAC3" box="[421,534,1286,1308]" pageId="16" pageNumber="17" refId="ref43377" refString="Isely, D. (1917) A synopsis of the petiolate wasps of the family Eumenidae (Hymenoptera), found in America North of Mexico. Annals of the Entomological Society of America, 10 (4), 345 - 366. https: // doi. org / 10.1093 / aesa / 10.4.345" type="journal article">Isely, 1917</bibRefCitation>
, 10: 363.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE3F310FF609473D09CFA48" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BD33699FFE3F310FF609473D17AFA85" blockId="16.[151,1436,152,2014]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF609473D297FAE4" authorityName="Cresson" authorityYear="1875" box="[199,430,1317,1339]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="coloradensis">
<emphasis id="B918EA8BFFE3F310FF609473D297FAE4" box="[199,430,1317,1339]" italics="true" pageId="16" pageNumber="17">Eumenes coloradensis</emphasis>
</taxonomicName>
var. (or subsp.)
<taxonomicName id="4C6C4D1AFFE3F310FDC79473D04BFAE4" ID-CoL="5H8JK" authority="Bequaert, 1944" authorityName="Bequaert" authorityYear="1944" box="[608,882,1317,1339]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="nearcticus">
<emphasis id="B918EA8BFFE3F310FDC79473D1F0FAE4" box="[608,713,1318,1339]" italics="true" pageId="16" pageNumber="17">nearcticus</emphasis>
<bibRefCitation id="EFFD4B68FFE3F310FD759473D04BFAE4" box="[722,882,1317,1339]" pageId="16" pageNumber="17" refId="ref42355" refString="Bequaert, J. (1944) The genus Eumenes in the Dominion of Canada. Le Naturaliste Canadien, 71, 75 - 88." type="journal article">Bequaert, 1944</bibRefCitation>
</taxonomicName>
, 71: 76, 82 [replacement name for
<taxonomicName id="4C6C4D1AFFE3F310FB4A9473D25BFA85" ID-CoL="3CDHS" authority="de Saussure, 1855" authorityName="de Saussure" authorityYear="1855" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="globulosus">
<emphasis id="B918EA8BFFE3F310FB4A9473D643FAE4" box="[1261,1402,1317,1339]" italics="true" pageId="16" pageNumber="17">E. globulosus</emphasis>
de Saussure, 1855
</taxonomicName>
,
<emphasis id="B918EA8BFFE3F310FECA9410D2AAFA85" box="[365,403,1349,1370]" italics="true" pageId="16" pageNumber="17">non</emphasis>
Fourcroy, 1785].
</paragraph>
<paragraph id="8BD33699FFE3F310FF309436D028FAA7" blockId="16.[151,1436,152,2014]" box="[151,785,1378,1400]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF309436D150FAA7" ID-CoL="5H8JL" authority="Bequaert, 1944" authorityName="Bequaert" authorityYear="1944" box="[151,617,1378,1400]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="crucifera" status="stat. nov." subSpecies="stricklandi">
<emphasis id="B918EA8BFFE3F310FF309436D2F0FAA7" box="[151,457,1378,1400]" italics="true" pageId="16" pageNumber="17">Eumenes crucifera stricklandi</emphasis>
<bibRefCitation id="EFFD4B68FFE3F310FE699436D150FAA7" box="[462,617,1378,1400]" pageId="16" pageNumber="17" refId="ref42355" refString="Bequaert, J. (1944) The genus Eumenes in the Dominion of Canada. Le Naturaliste Canadien, 71, 75 - 88." type="journal article">Bequaert, 1944</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A22B57F0FFE3F310FDC89436D034FAA7" box="[623,781,1378,1400]" pageId="16" pageNumber="17">NEW STATUS</taxonomicNameLabel>
.
</paragraph>
<paragraph id="8BD33699FFE3F310FF3094D7D09CFA48" blockId="16.[151,1436,152,2014]" box="[151,933,1409,1431]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF3094D7D243FA48" authorityName="Cresson" authorityYear="1875" box="[151,378,1409,1431]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="coloradensis">
<emphasis id="B918EA8BFFE3F310FF3094D7D243FA48" box="[151,378,1409,1431]" italics="true" pageId="16" pageNumber="17">Eumenes coloradensis</emphasis>
</taxonomicName>
var. (or subsp.)
<taxonomicName id="4C6C4D1AFFE3F310FDB994D4D017FA48" ID-CoL="5H8JL" authority="Bequaert, 1944" authorityName="Bequaert" authorityYear="1944" box="[542,814,1409,1431]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="stricklandi">
<emphasis id="B918EA8BFFE3F310FDB994D4D1B3FA48" box="[542,650,1409,1431]" italics="true" pageId="16" pageNumber="17">stricklandi</emphasis>
<bibRefCitation id="EFFD4B68FFE3F310FD3694D7D017FA48" box="[657,814,1409,1431]" pageId="16" pageNumber="17" refId="ref42355" refString="Bequaert, J. (1944) The genus Eumenes in the Dominion of Canada. Le Naturaliste Canadien, 71, 75 - 88." type="journal article">Bequaert, 1944</bibRefCitation>
</taxonomicName>
, 71: 76, 84.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE3F310FF6094F5D2DCF801" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BD33699FFE3F310FF6094F5D00FFA69" blockId="16.[151,1436,152,2014]" box="[199,822,1440,1462]" pageId="16" pageNumber="17">
<emphasis id="B918EA8BFFE3F310FF6094F5D203FA69" bold="true" box="[199,314,1440,1462]" pageId="16" pageNumber="17">Type data:</emphasis>
Holotype female at MCZ (n° 33184)
<emphasis id="B918EA8BFFE3F310FD1094F5D00FFA69" bold="true" box="[695,822,1440,1462]" pageId="16" pageNumber="17">(examined).</emphasis>
</paragraph>
<paragraph id="8BD33699FFE3F310FF6094EBD1FBFA0B" blockId="16.[151,1436,152,2014]" box="[199,706,1470,1492]" pageId="16" pageNumber="17">
<materialsCitation id="3B043CC4FFE3F310FF6094EBD1FBFA0B" ID-GBIF-Occurrence="1917355197" box="[199,706,1470,1492]" country="Canada" location="Cypress Hills" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Alberta" typeStatus="holotype">
<emphasis id="B918EA8BFFE3F310FF6094EBD263FA0B" bold="true" box="[199,346,1470,1492]" pageId="16" pageNumber="17">
<typeStatus id="54D7883BFFE3F310FF6094EBD3C5FA0B" box="[199,252,1470,1492]" pageId="16" pageNumber="17">Type</typeStatus>
locality:
</emphasis>
<location id="8EB36042FFE3F310FECE94EBD2CEFA0B" LSID="urn:lsid:plazi:treatment:03C5878FFFFCF313FF309792D674F9AB:8EB36042FFE3F310FECE94EBD2CEFA0B" box="[361,503,1470,1492]" country="Canada" name="Cypress Hills" pageId="16" pageNumber="17" stateProvince="Alberta">Cypress Hills</location>
,
<collectingRegion id="49A8F87BFFE3F310FDA694EBD176FA0B" box="[513,591,1470,1492]" country="Canada" name="Alberta" pageId="16" pageNumber="17">Alberta</collectingRegion>
” (
<collectingCountry id="F37B7609FFE3F310FDC294EBD18EFA0B" box="[613,695,1470,1492]" name="Canada" pageId="16" pageNumber="17">Canada</collectingCountry>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD33699FFE3F310FF30948BD1F7FA2C" blockId="16.[151,1436,152,2014]" box="[151,718,1501,1523]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF30948BD2F1FA2C" authorityName="Bequaert" authorityYear="1944" box="[151,456,1501,1523]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="stricklandi">
<emphasis id="B918EA8BFFE3F310FF30948BD2F1FA2C" box="[151,456,1501,1523]" italics="true" pageId="16" pageNumber="17">Eumenes crucifera stricklandi</emphasis>
</taxonomicName>
; Bohart, 1951: 884 (cat.).
</paragraph>
<paragraph id="8BD33699FFE3F310FF6094A9D2D7F9CD" blockId="16.[151,1436,152,2014]" box="[199,494,1532,1554]" pageId="16" pageNumber="17">Krombein, 1979: 1507 (cat.).</paragraph>
<paragraph id="8BD33699FFE3F310FF30974ED1E8F9EF" blockId="16.[151,1436,152,2014]" box="[151,721,1562,1584]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF30974ED2F2F9EF" authorityName="Viereck" authorityYear="1908" box="[151,459,1562,1584]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="bolliformis">
<emphasis id="B918EA8BFFE3F310FF30974ED2F2F9EF" box="[151,459,1562,1584]" italics="true" pageId="16" pageNumber="17">Eumenes crucifera bolliformis</emphasis>
</taxonomicName>
; Bohart, 1951: 884 (cat.).
</paragraph>
<paragraph id="8BD33699FFE3F310FF60976CD288F990" blockId="16.[151,1436,152,2014]" box="[199,433,1593,1615]" pageId="16" pageNumber="17">Linsley, 1962, 55: 153.</paragraph>
<paragraph id="8BD33699FFE3F310FF60970DD29EF9B1" blockId="16.[151,1436,152,2014]" box="[199,423,1624,1646]" pageId="16" pageNumber="17">Krombein, 1967: 376.</paragraph>
<paragraph id="8BD33699FFE3F310FF609723D2CDF953" blockId="16.[151,1436,152,2014]" box="[199,500,1654,1676]" pageId="16" pageNumber="17">
<bibRefCitation id="EFFD4B68FFE3F310FF609723D29BF953" box="[199,418,1654,1676]" pageId="16" pageNumber="17" refId="ref43151" refString="Giordani Soika, A. (1978) Revisione degli Eumenidi neotropicali appartenenti ai generi Eumenes Latr., Omicron (Sauss.), Pararhaphidoglossa Schulth. ed afinni. Bolletino della Museo Civico di Storia Naturale di Uenezia, 29, 1 - 420." type="journal article">Giordani Soika, 1978</bibRefCitation>
, 29: 28.
</paragraph>
<paragraph id="8BD33699FFE3F310FF6097C0D2D7F974" blockId="16.[151,1436,152,2014]" box="[199,494,1685,1707]" pageId="16" pageNumber="17">Krombein, 1979: 1507 (cat.).</paragraph>
<paragraph id="8BD33699FFE3F310FF6097E1D16AF915" blockId="16.[151,1436,152,2014]" box="[199,595,1716,1738]" pageId="16" pageNumber="17">
<bibRefCitation id="EFFD4B68FFE3F310FF6097E1D2BEF915" box="[199,391,1716,1738]" pageId="16" pageNumber="17" refId="ref43608" refString="MacLachlan, W. B. (1980) A key to and notes on the Eumenes of America North of Mexico (Hymenoptera: Eumenidae). Journal of the Kansas Entomological Society, 53 (3), 617 - 621." type="journal article">MacLachlan, 1980</bibRefCitation>
, 53: 618, 621 (key).
</paragraph>
<paragraph id="8BD33699FFE3F310FF609787D12FF937" blockId="16.[151,1436,152,2014]" box="[199,534,1746,1768]" pageId="16" pageNumber="17">
Ruiz C.
<emphasis id="B918EA8BFFE3F310FEBC9786D277F937" box="[283,334,1746,1768]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
, 1993, 88: 86 (list).
</paragraph>
<paragraph id="8BD33699FFE3F310FF6097A4D10FF8D8" blockId="16.[151,1436,152,2014]" box="[199,566,1777,1799]" pageId="16" pageNumber="17">Rodríguez-Palafox, 1996: 479 (list).</paragraph>
<paragraph id="8BD33699FFE3F310FF309644D186F8F9" blockId="16.[151,1436,152,2014]" box="[151,703,1808,1830]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF309644D280F8F9" authority="crucifera" box="[151,441,1808,1830]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="crucifera">
<emphasis id="B918EA8BFFE3F310FF309644D280F8F9" box="[151,441,1808,1830]" italics="true" pageId="16" pageNumber="17">Eumenes crucifera crucifera</emphasis>
</taxonomicName>
; Bohart, 1951: 884 (cat.).
</paragraph>
<paragraph id="8BD33699FFE3F310FF60967BD2D7F89B" blockId="16.[151,1436,152,2014]" box="[199,494,1838,1860]" pageId="16" pageNumber="17">Krombein, 1979: 1507 (cat.).</paragraph>
<paragraph id="8BD33699FFE3F310FF609618D2DCF8BC" blockId="16.[151,1436,152,2014]" box="[199,485,1869,1891]" pageId="16" pageNumber="17">
<bibRefCitation id="EFFD4B68FFE3F310FF609618D2BEF8BC" box="[199,391,1869,1891]" pageId="16" pageNumber="17" refId="ref43608" refString="MacLachlan, W. B. (1980) A key to and notes on the Eumenes of America North of Mexico (Hymenoptera: Eumenidae). Journal of the Kansas Entomological Society, 53 (3), 617 - 621." type="journal article">MacLachlan, 1980</bibRefCitation>
, 53: 618.
</paragraph>
<paragraph id="8BD33699FFE3F310FF309638D1F2F85D" blockId="16.[151,1436,152,2014]" box="[151,715,1900,1922]" pageId="16" pageNumber="17">
<taxonomicName id="4C6C4D1AFFE3F310FF309638D2FCF85D" authorityName="Bequaert" authorityYear="1944" box="[151,453,1900,1922]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="nearcticus">
<emphasis id="B918EA8BFFE3F310FF309638D2FCF85D" box="[151,453,1900,1922]" italics="true" pageId="16" pageNumber="17">Eumenes crucifera nearcticus</emphasis>
</taxonomicName>
; Bohart, 1951: 884 (cat.).
</paragraph>
<paragraph id="8BD33699FFE3F310FF6096DFD29EF87F" blockId="16.[151,1436,152,2014]" box="[199,423,1930,1952]" pageId="16" pageNumber="17">Krombein, 1958: 163.</paragraph>
<paragraph id="8BD33699FFE3F310FF6096FCD2D7F860" blockId="16.[151,1436,152,2014]" box="[199,494,1961,1983]" pageId="16" pageNumber="17">Krombein, 1979: 1507 (cat.).</paragraph>
<paragraph id="8BD33699FFE3F310FF60969DD2DCF801" blockId="16.[151,1436,152,2014]" box="[199,485,1992,2014]" pageId="16" pageNumber="17">
<bibRefCitation id="EFFD4B68FFE3F310FF60969DD2BEF801" box="[199,391,1992,2014]" pageId="16" pageNumber="17" refId="ref43608" refString="MacLachlan, W. B. (1980) A key to and notes on the Eumenes of America North of Mexico (Hymenoptera: Eumenidae). Journal of the Kansas Entomological Society, 53 (3), 617 - 621." type="journal article">MacLachlan, 1980</bibRefCitation>
, 53: 618.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE2F311FF6091CDD2CAFED5" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BD33699FFE2F311FF6091CDD0B4FF34" blockId="17.[199,1436,152,266]" pageId="17" pageNumber="18">
<emphasis id="B918EA8BFFE2F311FF6091CDD26AFF71" bold="true" box="[199,339,152,174]" pageId="17" pageNumber="18">Distribution:</emphasis>
<collectingRegion id="49A8F87BFFE2F311FEFA91CDD293FF71" box="[349,426,152,174]" country="Mexico" name="Mexico" pageId="17" pageNumber="18">
<collectingCountry id="F37B7609FFE2F311FEFA91CDD293FF71" box="[349,426,152,174]" name="Mexico" pageId="17" pageNumber="18">Mexico</collectingCountry>
</collectingRegion>
(
<collectingRegion id="49A8F87BFFE2F311FE1C91CDD116FF71" box="[443,559,152,174]" country="Mexico" name="Chihuahua" pageId="17" pageNumber="18">Chihuahua</collectingRegion>
);
<collectingCountry id="F37B7609FFE2F311FDE491CDD1B6FF71" box="[579,655,152,174]" name="Canada" pageId="17" pageNumber="18">Canada</collectingCountry>
(
<collectingRegion id="49A8F87BFFE2F311FD0791CDD1C9FF71" box="[672,752,152,174]" country="Canada" name="Alberta" pageId="17" pageNumber="18">Alberta</collectingRegion>
,
<collectingRegion id="49A8F87BFFE2F311FD5991CDD08FFF71" box="[766,950,152,174]" country="Canada" name="British Columbia" pageId="17" pageNumber="18">British Columbia</collectingRegion>
*;
<collectingRegion id="49A8F87BFFE2F311FC6991CDD70CFF71" box="[974,1077,152,174]" country="Canada" name="Manitoba" pageId="17" pageNumber="18">Manitoba</collectingRegion>
*,
<collectingRegion id="49A8F87BFFE2F311FBEC91CCD7C1FF71" box="[1099,1272,152,174]" country="Canada" name="New Brunswick" pageId="17" pageNumber="18">New Brunswick</collectingRegion>
*, New Scotia*, Northwest Territory*,
<collectingRegion id="49A8F87BFFE2F311FE0E91E3D2C2FF13" box="[425,507,182,204]" country="Canada" name="Quebec" pageId="17" pageNumber="18">Quebec</collectingRegion>
*,
<collectingRegion id="49A8F87BFFE2F311FDA891E3D158FF13" box="[527,609,182,204]" country="Canada" name="Ontario" pageId="17" pageNumber="18">Ontario</collectingRegion>
);
<collectingCountry id="F37B7609FFE2F311FDD591E3D19CFF13" box="[626,677,182,204]" name="United States of America" pageId="17" pageNumber="18">USA</collectingCountry>
(AZ, CA, CO, CT*, ID, IL*, IN, KY*, MA*, ME, MI, MN, MO, ND, NH, NJ, NM, NV, NY, OH*, OR*, PE, SC, SD, UT, VT*, WA*, WI, WY).
</paragraph>
<paragraph id="8BD33699FFE2F311FF6091A1D2CAFED5" blockId="17.[199,1436,152,266]" box="[199,499,244,266]" pageId="17" pageNumber="18">* Records from the literature.</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE2F311FF309063D1D3FB9C" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BD33699FFE2F311FF309063D128FDDC" blockId="17.[151,1437,310,2027]" pageId="17" pageNumber="18">
<emphasis id="B918EA8BFFE2F311FF309063D2E8FE90" bold="true" box="[151,465,310,335]" pageId="17" pageNumber="18">Comments and diagnosis:</emphasis>
<taxonomicName id="4C6C4D1AFFE2F311FE7B9062D0BDFE90" authority="Provancher, 1888" authorityName="Provancher" authorityYear="1888" box="[476,900,310,335]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE2F311FE7B9062D18BFE90" box="[476,690,311,335]" italics="true" pageId="17" pageNumber="18">Eumenes crucifera</emphasis>
Provancher, 1888
</taxonomicName>
(
<figureCitation id="13572A1CFFE2F311FC319063D0E5FE90" box="[918,988,310,335]" captionStart="FIGURES 112" captionStartId="41.[151,264,1386,1408]" captionTargetBox="[151,1436,193,1341]" captionTargetId="figure@41.[151,1436,193,1341]" captionTargetPageId="41" captionText="FIGURES 112. Eumenes species, lateral view. 1) Eumenes americanus de Saussure (female); 2) E. belfragei aureus Isely, 1917 (Holotype, female); 3) E. bollii Cresson, 1872 (Lectotype, female); 4) E. bollii oregonensis Bequaert, 1938 (Holotype, female); 5); E. bequaerti Isely, 1917 (Holotype, male); 6) E. iturbide (= E. consobrinus; Holotype, female); 7) E. coyotae Bohart, 1948 (Holotype, male); 8) E. crucifera Provancher, 1888 (Holotype, female). 9) E. bolliformis Viereck, 1908 (Lectotype, female); 10) E. crucifera var. nearcticus Bequaert, 1944 (female); 11) E. crucifera stricklandi Bequaert, 1944 (Holotype, female); 12) E. cruciferoides Giordani Soika, 1978 (Holotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458304/files/figure.png" pageId="17" pageNumber="18">Fig. 8</figureCitation>
) can be identified by having long and golden pubescence along the body; punctation on the clypeus; black integument with yellow marks varying in size (see
<emphasis id="B918EA8BFFE2F311FF69902AD20CFE49" box="[206,309,383,406]" italics="true" pageId="17" pageNumber="18">variation</emphasis>
section below); T2 wide, with the lateral margins strongly concave (
<figureCitation id="13572A1CFFE2F311FBFD902BD78BFE48" box="[1114,1202,382,407]" captionStart="FIGURES 4358" captionStartId="44.[151,264,1946,1968]" captionTargetBox="[202,1388,195,1934]" captionTargetId="figure@44.[201,1388,193,1934]" captionTargetPageId="44" captionText="FIGURES 4358. Second tergum, dorsal view. 43) Eumenes americanus; 44) E. bequaerti; 45) E. bollii; 46) E. consobrinus; 47) E. coyotae; 48) E. crucifera; 49) E. cubensis; 50) E. flavitinctus; 51) E. fraternus; 52) E. pictus; 53) E. sculleni; 54) E. smithii; 55) E. verticalis; 56) E. filiformis; 57) E. rufomaculactus; 58) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458310/files/figure.png" pageId="17" pageNumber="18">Fig. 48</figureCitation>
); and mainly by the posterior face of the propodeum weakly concave. The species has a wide distribution across the
<collectingCountry id="F37B7609FFE2F311FB7590F6D633FE64" box="[1234,1290,419,443]" name="United States of America" pageId="17" pageNumber="18">USA</collectingCountry>
and
<collectingCountry id="F37B7609FFE2F311FAE690F7D6A1FE64" box="[1345,1432,418,443]" name="Canada" pageId="17" pageNumber="18">Canada</collectingCountry>
, and consequently we observed a range of variation mainly in coloration, which previous authors used to justify the description of various subspecies.
</paragraph>
<paragraph id="8BD33699FFE2F311FF60935AD0B9FD4C" blockId="17.[151,1437,310,2027]" pageId="17" pageNumber="18">
We had access to all the
<typeStatus id="54D7883BFFE2F311FE79935AD137FDF8" box="[478,526,527,551]" pageId="17" pageNumber="18">type</typeStatus>
specimens of the typical
<taxonomicName id="4C6C4D1AFFE2F311FC88935AD08FFDF8" authorityName="Provancher" authorityYear="1888" box="[815,950,527,551]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE2F311FC88935AD08FFDF8" box="[815,950,527,551]" italics="true" pageId="17" pageNumber="18">E. crucifera</emphasis>
</taxonomicName>
and its subspecies, except for the
<typeStatus id="54D7883BFFE2F311FA9D935BD6A5FDF8" box="[1338,1436,526,551]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C6C4D1AFFE2F311FF119366D2F9FD94" authorityName="Bohart" authorityYear="1950" box="[182,448,563,587]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="flavitinctus">
<emphasis id="B918EA8BFFE2F311FF119366D2F9FD94" box="[182,448,563,587]" italics="true" pageId="17" pageNumber="18">E. crucifera flavitinctus</emphasis>
</taxonomicName>
, of which we examined four
<typeStatus id="54D7883BFFE2F311FCAA9366D040FD94" box="[781,889,563,587]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
(females and males). It is important to highlight that one specimen deposited at USNM (n° 21382), labeled as
<typeStatus id="54D7883BFFE2F311FCD39302D0EDFDB0" box="[884,980,599,623]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C6C4D1AFFE2F311FBA59302D7B5FDB0" authorityName="Provancher" authorityYear="1888" box="[1026,1164,599,623]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE2F311FBA59302D7B5FDB0" box="[1026,1164,599,623]" italics="true" pageId="17" pageNumber="18">E. crucifera</emphasis>
</taxonomicName>
, is actually a non-type specimen, as Provancher did not mention a
<typeStatus id="54D7883BFFE2F311FDD8932ED1E6FD4C" box="[639,735,635,659]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
from
<collectingRegion id="49A8F87BFFE2F311FC83932ED045FD4C" box="[804,892,635,659]" country="United States of America" name="Oregon" pageId="17" pageNumber="18">Oregon</collectingRegion>
.
</paragraph>
<paragraph id="8BD33699FFE2F311FF6093CAD1D3FB9C" blockId="17.[151,1437,310,2027]" pageId="17" pageNumber="18">
A total of
<specimenCount id="9D6AFD10FFE2F311FE9893CBD2DEFD68" box="[319,487,670,695]" pageId="17" pageNumber="18" type="generic">361 specimens</specimenCount>
were examined, females and males, and the genitalia from all the subspecies were extracted and examined. As a result, we observed that the same range of variation that occurs in
<taxonomicName id="4C6C4D1AFFE2F311FB799396D65FFD04" authorityName="Say" authorityYear="1824" box="[1246,1382,707,731]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFE2F311FB799396D65FFD04" box="[1246,1382,707,731]" italics="true" pageId="17" pageNumber="18">E. verticalis</emphasis>
</taxonomicName>
also exists for
<taxonomicName id="4C6C4D1AFFE2F311FEA193B2D2B4FD20" authorityName="Provancher" authorityYear="1888" box="[262,397,743,767]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE2F311FEA193B2D2B4FD20" box="[262,397,743,767]" italics="true" pageId="17" pageNumber="18">E. crucifera</emphasis>
</taxonomicName>
, which has an extensive geographical distribution that causes intra-specific variation among the specimens, including some overlapping morphological characters. All the variation examined was found in the
<taxonomicName id="4C6C4D1AFFE2F311FF30927BD22BFC99" authorityName="Viereck" authorityYear="1908" box="[151,274,814,838]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bolliformis">
<emphasis id="B918EA8BFFE2F311FF30927BD22BFC99" box="[151,274,814,838]" italics="true" pageId="17" pageNumber="18">bolliformis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D1AFFE2F311FE87927AD2ADFC99" authorityName="Bequaert" authorityYear="1944" box="[288,404,815,838]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="nearcticus">
<emphasis id="B918EA8BFFE2F311FE87927AD2ADFC99" box="[288,404,815,838]" italics="true" pageId="17" pageNumber="18">nearcticus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D1AFFE2F311FE6C927BD17AFC99" authorityName="Bequaert" authorityYear="1944" box="[459,579,814,838]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="stricklandi">
<emphasis id="B918EA8BFFE2F311FE6C927BD17AFC99" box="[459,579,814,838]" italics="true" pageId="17" pageNumber="18">stricklandi</emphasis>
</taxonomicName>
subspecies (the particulars are discussed in the
<emphasis id="B918EA8BFFE2F311FBF9927AD7FCFC99" box="[1118,1221,815,838]" italics="true" pageId="17" pageNumber="18">variation</emphasis>
section) in various combinations; the male genitalia were compared in each form and presented no significant differences. Therefore, to avoid treating those names as a subspecific category based just on color differences, which is known to be unreliable (
<bibRefCitation id="EFFD4B68FFE2F311FEBF92CED12CFC6C" box="[280,533,923,947]" pageId="17" pageNumber="18" refId="ref43652" refString="MacLean, B. K., Chandler, L. &amp; MacLean, D. B. (1978) Phenotypic expression in the paper wasp Polistes fuscatus (Hymenoptera: Vespidae). Great Lakes Entomologist, 11, 105 - 116." type="journal article">
MacLean
<emphasis id="B918EA8BFFE2F311FE2992CED2F0FC6C" box="[398,457,923,947]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
, 1978
</bibRefCitation>
; Carpenter, 1988; Carpenter and van der
<bibRefCitation id="EFFD4B68FFE2F311FC5192CFD7BBFC6C" box="[1014,1154,922,947]" pageId="17" pageNumber="18" refId="ref42766" refString="Carpenter, J. M. &amp; van der Vecht, J. (1991) A study of the Vespidae described by William J. Fox (Insecta: Hymenoptera), with assessment of taxonomic implications. Annals of the Carnegie Museum of Natural History, 60 (3), 211 - 241." type="journal article">Vecht, 1991</bibRefCitation>
; Carpenter and Garcete- Barrett, 2003), we propose the synonymy of
<taxonomicName id="4C6C4D1AFFE2F311FD0B92EAD053FC09" authorityName="Viereck" authorityYear="1908" box="[684,874,958,982]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="bolliformis">
<emphasis id="B918EA8BFFE2F311FD0B92EAD053FC09" box="[684,874,958,982]" italics="true" pageId="17" pageNumber="18">E. c. bolliformis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D1AFFE2F311FCDB92EAD70AFC09" authorityName="Bequaert" authorityYear="1944" box="[892,1075,959,982]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="nearcticus">
<emphasis id="B918EA8BFFE2F311FCDB92EAD70AFC09" box="[892,1075,959,982]" italics="true" pageId="17" pageNumber="18">E. c. nearcticus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D1AFFE2F311FBD592EAD614FC09" authorityName="Bequaert" authorityYear="1944" box="[1138,1325,958,982]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="stricklandi">
<emphasis id="B918EA8BFFE2F311FBD592EAD614FC09" box="[1138,1325,958,982]" italics="true" pageId="17" pageNumber="18">E. c. stricklandi</emphasis>
</taxonomicName>
under
<taxonomicName id="4C6C4D1AFFE2F311FA2392EAD3C7FC24" authorityName="Provancher" authorityYear="1888" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE2F311FA2392EAD3C7FC24" italics="true" pageId="17" pageNumber="18">E. crucifera</emphasis>
</taxonomicName>
and treat these names as variations of the typical species as a consequence of the great geographical distribution of this species. Finally, we elevated the subspecies
<taxonomicName id="4C6C4D1AFFE2F311FCC79552D72EFBC1" authorityName="Bohart" authorityYear="1950" box="[864,1047,1030,1054]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="flavitinctus">
<emphasis id="B918EA8BFFE2F311FCC79552D72EFBC1" box="[864,1047,1030,1054]" italics="true" pageId="17" pageNumber="18">E. c. flavitinctus</emphasis>
</taxonomicName>
to specific level based on features that are discussed in the redescription of that species.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE2F313FF60951BD6AAFC33" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="description">
<paragraph id="8BD33699FFE2F311FF60951BD0C5FBB8" blockId="17.[151,1437,310,2027]" box="[199,1020,1102,1127]" pageId="17" pageNumber="18">
<emphasis id="B918EA8BFFE2F311FF60951BD098FBB8" bold="true" box="[199,929,1102,1127]" pageId="17" pageNumber="18">
Redescription. Holotype
<taxonomicName id="4C6C4D1AFFE2F311FE48951AD098FBB8" authority="Provancher, 1888" authorityName="Provancher" authorityYear="1888" box="[495,929,1102,1127]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE2F311FE48951AD1F2FBB8" bold="true" box="[495,715,1102,1127]" italics="true" pageId="17" pageNumber="18">Eumenes crucifera</emphasis>
Provancher, 1888
</taxonomicName>
</emphasis>
(
<figureCitation id="13572A1CFFE2F311FC17951BD0CAFBB8" box="[944,1011,1102,1127]" captionStart="FIGURES 112" captionStartId="41.[151,264,1386,1408]" captionTargetBox="[151,1436,193,1341]" captionTargetId="figure@41.[151,1436,193,1341]" captionTargetPageId="41" captionText="FIGURES 112. Eumenes species, lateral view. 1) Eumenes americanus de Saussure (female); 2) E. belfragei aureus Isely, 1917 (Holotype, female); 3) E. bollii Cresson, 1872 (Lectotype, female); 4) E. bollii oregonensis Bequaert, 1938 (Holotype, female); 5); E. bequaerti Isely, 1917 (Holotype, male); 6) E. iturbide (= E. consobrinus; Holotype, female); 7) E. coyotae Bohart, 1948 (Holotype, male); 8) E. crucifera Provancher, 1888 (Holotype, female). 9) E. bolliformis Viereck, 1908 (Lectotype, female); 10) E. crucifera var. nearcticus Bequaert, 1944 (female); 11) E. crucifera stricklandi Bequaert, 1944 (Holotype, female); 12) E. cruciferoides Giordani Soika, 1978 (Holotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458304/files/figure.png" pageId="17" pageNumber="18">Fig. 8</figureCitation>
)
</paragraph>
<paragraph id="8BD33699FFE2F311FF609527D621F9C8" blockId="17.[151,1437,310,2027]" pageId="17" pageNumber="18">
<emphasis id="B918EA8BFFE2F311FF609527D2FEFB54" bold="true" box="[199,455,1138,1163]" pageId="17" pageNumber="18">Female. Color, head:</emphasis>
Black, with apex the tooth of mandibles brown. Yellow marks as follows: apex of labrum; clypeus, except for a blackish spot on the center region; inter-antennal region; short and narrow stripe on upper half of the gena, adjacent to the compound eye; broad longitudinal stripe on scape.
<emphasis id="B918EA8BFFE2F311FB3695EFD620FB0C" bold="true" box="[1169,1305,1210,1235]" pageId="17" pageNumber="18">Mesosoma:</emphasis>
Black with yellow marks as follows: anterior portion of the dorsal face of pronotum; wide spot on upper half of the mesepisternum, adjacent to the mesepimeron; tegula (with a translucent spot on center of it); a triangular shaped spot on each side of mesoscutum and a mark on the center region of it; metanotum, except for a narrow stripe on the basal edge; wide spot on each side of upper half of the propodeum; a small spot on mid coxa; distal half of femur, with some brownish marks; entirely tibiae and tarsi, which becomes darker towards apex.
<emphasis id="B918EA8BFFE2F311FB49943BD64DFA58" bold="true" box="[1262,1396,1390,1415]" pageId="17" pageNumber="18">Metasoma:</emphasis>
T1 black with two big yellow spots on center region and a transverse stripe on apex of T1 and S1. T2 yellow with a black “T” on dorsal face of it. T3T6 entirely yellow with a transverse narrow brownish stripe on apex. S2 with basal half blackish and narrow stripes on the lateral margins almost reaching the apex; S3S6 with black marks on basal half.
<emphasis id="B918EA8BFFE2F311FEB694ABD25CF9C8" bold="true" box="[273,357,1534,1559]" pageId="17" pageNumber="18">Wings:</emphasis>
Hyaline with costal, medial and submedial yellow. Pterostigma and veins brownish.
</paragraph>
<paragraph id="8BD33699FFE2F311FF609777D7C4F8E9" blockId="17.[151,1437,310,2027]" pageId="17" pageNumber="18">
<emphasis id="B918EA8BFFE2F311FF609777D29BF9E4" bold="true" box="[199,418,1570,1595]" pageId="17" pageNumber="18">Pubescence, head:</emphasis>
Frons, vertex, gena, scape and occipital region with long and golden pubescence (longer on frons and occipital region). The pubescence on scape is the shortest regarding those other regions. Clypeus with long and slightly whitish (better observed in oblique view) pubescence mostly on basal and lateral regions, while on apex it is shorter.
<emphasis id="B918EA8BFFE2F311FE2197DBD137F978" bold="true" box="[390,526,1678,1703]" pageId="17" pageNumber="18">Mesosoma:</emphasis>
All covered with golden and long (as on frons) pubescence. Anterior surface of coxae with long and golden pubescence, longer on fore coxa and shorter on hind coxa. Scattered erect golden bristles on trochanter and basal half of the fore femur.
<emphasis id="B918EA8BFFE2F311FCA39783D0B3F930" bold="true" box="[772,906,1750,1775]" pageId="17" pageNumber="18">Metasoma:</emphasis>
T1 covered with golden and long pubescence, longer on basal half. T2 covered with golden pubescence, longer on basal third, becoming shorter towards apex. T3T6 and S2S6 covered with short and golden pubescence, except for some long bristles on apex.
</paragraph>
<paragraph id="8BD33699FFE2F312FF609617D083FE9F" blockId="17.[151,1437,310,2027]" lastBlockId="18.[151,1437,151,2013]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="B918EA8BFFE2F311FF609617D127F884" bold="true" box="[199,542,1858,1883]" pageId="17" pageNumber="18">Surface of integument, head:</emphasis>
Clypeus with and sparse punctation mainly on basal half. Frons and vertex with coarse punctation, moderately dense, becoming weaker and sparser behind the ocelli.
<emphasis id="B918EA8BFFE2F311FBCD9633D7C8F8A0" bold="true" box="[1130,1265,1894,1919]" pageId="17" pageNumber="18">Mesosoma:</emphasis>
Dorsal face of pronotum, mesoscutum, scutelum and metanotum with coarse and moderately dense punctation, as in frons. Mesepisternum with coarse punctation as mesosucutum but slightly sparser. Posterior face of propodeum with coarse and denser punctation, forming carinae between the punctures. Posterior half from the lateral face of propodeum with coarse but sparser punctation, while on the anterior half the punctation is reduced.
<emphasis id="B918EA8BFFE1F312FAB191C2D6A5FF6F" bold="true" box="[1302,1436,151,176]" pageId="18" pageNumber="19">Metasoma:</emphasis>
Dorsal and ventral face of T1 with coarse and moderately dense (sparser than mesoscutum) punctation mainly on apical two-third; dorsal face of T2 with weaker and sparser punctation regardless T1, and the punctation becomes and sparser towards the lateral face. Basal half of S2 with (more than in dorsal face of T2) scattered punctation. T3T6 with punctation as in S2, and S3S6 with no evident punctation.
</paragraph>
<paragraph id="8BD33699FFE1F312FF609019D6A4FE0F" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
<emphasis id="B918EA8BFFE1F312FF609019D2B4FEBA" bold="true" box="[199,397,332,357]" pageId="18" pageNumber="19">Structure, head:</emphasis>
Clypeus weakly convex, slightly longer than wide with the apex concave, forming two rounded projections with a weak carina on apex of each projection (
<figureCitation id="13572A1CFFE1F312FC38903AD0CDFE57" box="[927,1012,367,392]" captionStart="FIGURES 6676" captionStartId="47.[151,264,1772,1794]" captionTargetBox="[151,1436,193,1750]" captionTargetId="figure@47.[151,1436,193,1750]" captionTargetPageId="47" captionText="FIGURES 6676. Clypeus. 66) Eumenes americanus (female); 67) E. smithii (female); 68) E. crucifera (female); 69) E. crucifera (male); 70) E. verticalis (female); 71) E. fraternus (female); 72) E. fraternus (male); 73) E. sculleni (female); 74) E. filiformis (female); 75) E. rufomaculatus (female); 76) E. versicolor (female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458316/files/figure.png" pageId="18" pageNumber="19">Fig. 68</figureCitation>
). Inter-antennal region longitudinally cariniform, as wide as the diameter of antennal socket. Lateral ocelli closer to the compound eyes more than to each other. Occipital carina well developed along all its extension, being slightly angled on middle region of gena.
</paragraph>
<paragraph id="8BD33699FFE1F312FF309089D6A4FDBF" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
<emphasis id="B918EA8BFFE1F312FF309089D227FE2A" bold="true" box="[151,286,476,501]" pageId="18" pageNumber="19">Mesosoma:</emphasis>
Pronotal carina present, weakly developed on dorsal face and stronger on lateral face; pronotal fovea present; mesepimeron slightly elevated when compared to the mesepisternum level; tegula with posterior emargination weakly developed, rounded; parategula lamelliform. Posterior face of propodeum moderately concave on basal half. Fore coxa with a longitudinal carina; posterior face of the propodeum concave on lower half.
</paragraph>
<paragraph id="8BD33699FFE1F312FF309339D249FCCB" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
<emphasis id="B918EA8BFFE1F312FF309339D224FD5A" bold="true" box="[151,285,620,645]" pageId="18" pageNumber="19">Metasoma:</emphasis>
T1 about two times longer than wide, strongly swollen in dorsal view (
<figureCitation id="13572A1CFFE1F312FBDE9339D7E8FD5B" box="[1145,1233,620,645]" captionStart="FIGURES 2742" captionStartId="43.[151,264,1902,1924]" captionTargetBox="[151,1435,193,1881]" captionTargetId="figure@43.[151,1436,193,1881]" captionTargetPageId="43" captionText="FIGURES 2742. First tergum, dorsal view. 27) Eumenes americanus; 28) E. bequaerti; 29) E. bollii; 30) E. consobrinus; 31) E. coyotae; 32) E. crucifera; 33) E. cubensis; 34) E. flavitinctus; 35) E. fraternus; 36) E. pictus; 37) E. sculleni; 38) E. smithii; 39) E. verticalis; 40) E. filiformis; 41) E. rufomaculactus; 42) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458308/files/figure.png" pageId="18" pageNumber="19">Fig. 32</figureCitation>
), with the lateral margins subparallel on the distal half, except on the apex which is slightly divergent. Base of T2, in lateral view, rising up abruptly; apical third of T2, in lateral view, with a weak sinuosity before the apex; T2 as long as wide in dorsal view; in dorsal view (
<figureCitation id="13572A1CFFE1F312FE7D9382D117FD2F" box="[474,558,727,752]" captionStart="FIGURES 4358" captionStartId="44.[151,264,1946,1968]" captionTargetBox="[202,1388,195,1934]" captionTargetId="figure@44.[201,1388,193,1934]" captionTargetPageId="44" captionText="FIGURES 4358. Second tergum, dorsal view. 43) Eumenes americanus; 44) E. bequaerti; 45) E. bollii; 46) E. consobrinus; 47) E. coyotae; 48) E. crucifera; 49) E. cubensis; 50) E. flavitinctus; 51) E. fraternus; 52) E. pictus; 53) E. sculleni; 54) E. smithii; 55) E. verticalis; 56) E. filiformis; 57) E. rufomaculactus; 58) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458310/files/figure.png" pageId="18" pageNumber="19">Fig. 48</figureCitation>
), the lateral margins of T2 are strongly concave on middle region; lamella on apex of T2 present.
</paragraph>
<paragraph id="8BD33699FFE1F312FF60924AD7E1FBCF" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
<emphasis id="B918EA8BFFE1F312FF60924AD234FCE7" bold="true" box="[199,269,799,824]" pageId="18" pageNumber="19">Male:</emphasis>
Clypeus narrower than females (
<figureCitation id="13572A1CFFE1F312FDD8924AD1EAFCE7" box="[639,723,799,824]" captionStart="FIGURES 6676" captionStartId="47.[151,264,1772,1794]" captionTargetBox="[151,1436,193,1750]" captionTargetId="figure@47.[151,1436,193,1750]" captionTargetPageId="47" captionText="FIGURES 6676. Clypeus. 66) Eumenes americanus (female); 67) E. smithii (female); 68) E. crucifera (female); 69) E. crucifera (male); 70) E. verticalis (female); 71) E. fraternus (female); 72) E. fraternus (male); 73) E. sculleni (female); 74) E. filiformis (female); 75) E. rufomaculatus (female); 76) E. versicolor (female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458316/files/figure.png" pageId="18" pageNumber="19">Fig. 69</figureCitation>
). Sometimes the males of
<taxonomicName id="4C6C4D1AFFE1F312FC5E9274D746FCE7" authorityName="Provancher" authorityYear="1888" box="[1017,1151,800,824]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE1F312FC5E9274D746FCE7" box="[1017,1151,800,824]" italics="true" pageId="18" pageNumber="19">E. crucifera</emphasis>
</taxonomicName>
may confused with those from
<taxonomicName id="4C6C4D1AFFE1F312FF719210D266FC83" authorityName="Say" authorityYear="1824" box="[214,351,836,860]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFE1F312FF719210D266FC83" box="[214,351,836,860]" italics="true" pageId="18" pageNumber="19">E. verticalis</emphasis>
</taxonomicName>
, but they can be rightly separated from them by the absence of the microscopic bristles on the ventral surface of F11. Furthermore, males of
<taxonomicName id="4C6C4D1AFFE1F312FD1E923CD07AFC5F" authorityName="Provancher" authorityYear="1888" box="[697,835,872,896]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE1F312FD1E923CD07AFC5F" box="[697,835,872,896]" italics="true" pageId="18" pageNumber="19">E. crucifera</emphasis>
</taxonomicName>
present the F11 long, broad basally, with the apex pointed, surpassing the apical edge of F8, sometimes reaching the basal edge of it (in
<taxonomicName id="4C6C4D1AFFE1F312FBC192D8D7D7FC7B" authorityName="Say" authorityYear="1824" box="[1126,1262,908,932]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFE1F312FBC192D8D7D7FC7B" box="[1126,1262,908,932]" italics="true" pageId="18" pageNumber="19">E. verticalis</emphasis>
</taxonomicName>
it reaches only the apical edge) (fig. 82). Dorsal surface with a longitudinal carina. Ventral surface without microscopic erect bristles, different from
<taxonomicName id="4C6C4D1AFFE1F312FE3A9280D11CFC33" authorityName="Say" authorityYear="1824" box="[413,549,980,1004]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFE1F312FE3A9280D11CFC33" box="[413,549,980,1004]" italics="true" pageId="18" pageNumber="19">E. verticalis</emphasis>
</taxonomicName>
, which have them. The lateral margins of T2 usually are not so concave as on females. S7 flattened apically, with some moderately long bristles scattered on apex (also on T7).
</paragraph>
<paragraph id="8BD33699FFE1F312FF609549D2A0FB37" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
<emphasis id="B918EA8BFFE1F312FF609549D243FBEA" bold="true" box="[199,378,1052,1077]" pageId="18" pageNumber="19">Male genitalia:</emphasis>
Aedeagus as in figure 98a, b. The dorsal margim may vary between less to strongly angled, as we observed in
<taxonomicName id="4C6C4D1AFFE1F312FEF09514D13CFB87" authorityName="de Saussure" authorityYear="1855" box="[343,517,1088,1112]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B918EA8BFFE1F312FEF09514D13CFB87" box="[343,517,1088,1112]" italics="true" pageId="18" pageNumber="19">E. consobrinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D1AFFE1F312FDE29514D1EBFB87" authorityName="Say" authorityYear="1824" box="[581,722,1088,1112]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFE1F312FDE29514D1EBFB87" box="[581,722,1088,1112]" italics="true" pageId="18" pageNumber="19">E. verticalis</emphasis>
</taxonomicName>
. Paramere (fig. 98c) with long bristles on middle region of gonostyle; digitus long, surpassing the middle region of the gonostyle, with short bristles on apex, becoming moderately longer towards base (
<figureCitation id="13572A1CFFE1F312FDAA95D2D156FB7F" box="[525,623,1159,1184]" captionStart="FIGURES 110125" captionStartId="51.[151,264,1899,1921]" captionTargetBox="[163,1425,209,1857]" captionTargetId="figure@51.[150,1437,193,1878]" captionTargetPageId="51" captionText="FIGURES 110125. Male genitalia: digitus. 110) Eumenes americanus; 111) E. bequaerti; 112) E. bollii; 113) E. consobrinus; 114) E. coyotae; 115) E. crucifera; 116) E. cubensis; 117) E. flavitinctus; 118) E. fraternus; 119) E. pictus; 120) E. sculleni; 121) E. smithii; 122) E. verticalis; 123) E. filiformis; 124) E. rufomaculatus; 125) E. versicolor." httpUri="https://zenodo.org/record/1458324/files/figure.png" pageId="18" pageNumber="19">Fig. 115</figureCitation>
). Distal lobe truncate, without evident bristles. Cuspis with long (longer than on digitus) and erect bristles. Volsella with long bristles (longer than on cuspis) on the ventral edge and scattered short bristles.
</paragraph>
<paragraph id="8BD33699FFE1F312FF6095A1D2DAF8F7" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
<emphasis id="B918EA8BFFE1F312FF6095A1D2B5FAD2" bold="true" box="[199,396,1268,1293]" pageId="18" pageNumber="19">Variation, color:</emphasis>
The color variation listed was observed also in specimens from the same localities. Clypeus entirely black, except for small marks on each side on the basal half and sometimes on apical half (NM, NV, UT), or entirely yellow; yellow marks on scape, pronotum, mesepisternum, scutellum, T
<date id="FFD21059FFE1F312FB9A9469D747FA8A" box="[1085,1150,1340,1365]" pageId="18" pageNumber="19">1 may</date>
be reduced or sometimes absent; yellow mark on central area of scutellum may be absent; black mark on T
<date id="FFD21059FFE1F312FB9E9435D747FAA6" box="[1081,1150,1376,1401]" pageId="18" pageNumber="19">2 may</date>
be covering all the basal half, with just a broad yellow band on each lateral margin; or black mark on S
<date id="FFD21059FFE1F312FBA094D1D774FA42" box="[1031,1101,1412,1437]" pageId="18" pageNumber="19">2 may</date>
be covering almost the entire sclerite.
<emphasis id="B918EA8BFFE1F312FF5294F2D2B1FA1F" bold="true" box="[245,392,1447,1472]" pageId="18" pageNumber="19">Pubescence:</emphasis>
Pubescence on apical half of T
<date id="FFD21059FFE1F312FD4094FDD015FA1E" box="[743,812,1448,1473]" pageId="18" pageNumber="19">2 may</date>
be slightly longer (as long as on basal half).
<emphasis id="B918EA8BFFE1F312FA8594F2D21EFA3A" bold="true" pageId="18" pageNumber="19">Surface of integument:</emphasis>
punctation on clypeus may be more evident (specimens from the same localities varied between less and more evident punctures); punctation of T
<date id="FFD21059FFE1F312FD0694A5D1D1F9D6" box="[673,744,1520,1545]" pageId="18" pageNumber="19">2 may</date>
be stronger, although some specimens outside of
<collectingRegion id="49A8F87BFFE1F312FA8C94BAD6A4F9D7" box="[1323,1437,1519,1544]" country="United States of America" name="California" pageId="18" pageNumber="19">California</collectingRegion>
presented punctures as in the typical pattern (NV, UT, ID, WY).
<emphasis id="B918EA8BFFE1F312FCCC9741D0DCF9F2" bold="true" box="[875,997,1556,1581]" pageId="18" pageNumber="19">Structure:</emphasis>
posterior face of propodeum may vary from weakly to strongly concave; generally concave in males. T
<date id="FFD21059FFE1F312FCD4976DD08FF98E" box="[883,950,1592,1617]" pageId="18" pageNumber="19">1 may</date>
be shorter and more developed on apex or longer; lateral margins of T1 more concave; some specimens presented an elevation on the middle region of T2 (better observed in lateral view), which produces a weak sinuosity on the apical third (not so evident as on
<taxonomicName id="4C6C4D1AFFE1F312FA2397D4D219F963" authorityName="de Saussure" authorityYear="1855" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B918EA8BFFE1F312FA2397D4D219F963" italics="true" pageId="18" pageNumber="19">E. consobrinus</emphasis>
</taxonomicName>
). We observed in two specimens that the sinuosity was strong, but this was clearly caused by a malformation of the sclerite that produced the impression of a strong sinuosity; the lateral margins of T
<date id="FFD21059FFE1F312FA94979DD640F93E" box="[1331,1401,1736,1761]" pageId="18" pageNumber="19">2 may</date>
be less concave, mainly in males; and some specimens may present the basal angle of T2 rising up weakly, as in the
<typeStatus id="54D7883BFFE1F312FF309644D3FEF8F6" box="[151,199,1809,1833]" pageId="18" pageNumber="19">type</typeStatus>
specimen of
<taxonomicName id="4C6C4D1AFFE1F312FEFA9644D2E5F8F7" authorityName="Isely" authorityYear="1917" box="[349,476,1808,1832]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="sternalis">
<emphasis id="B918EA8BFFE1F312FEFA9644D2E5F8F7" box="[349,476,1808,1832]" italics="true" pageId="18" pageNumber="19">E. sternalis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD33699FFE1F312FF609661D637F84B" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
Viereck (1908) described
<taxonomicName id="4C6C4D1AFFE1F312FE579660D1B4F893" authorityName="Viereck" authorityYear="1908" box="[496,653,1844,1868]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bolliformis">
<emphasis id="B918EA8BFFE1F312FE579660D1B4F893" box="[496,653,1844,1868]" italics="true" pageId="18" pageNumber="19">E. bolliformis</emphasis>
</taxonomicName>
(
<typeStatus id="54D7883BFFE1F312FD399661D03EF892" box="[670,775,1844,1869]" pageId="18" pageNumber="19" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54D7883BFFE1F312FCE69661D0E1F892" box="[833,984,1844,1869]" pageId="18" pageNumber="19" type="paralectotype">paralectotype</typeStatus>
designated herein) (
<figureCitation id="13572A1CFFE1F312FB639661D632F893" box="[1220,1291,1844,1869]" captionStart="FIGURES 112" captionStartId="41.[151,264,1386,1408]" captionTargetBox="[151,1436,193,1341]" captionTargetId="figure@41.[151,1436,193,1341]" captionTargetPageId="41" captionText="FIGURES 112. Eumenes species, lateral view. 1) Eumenes americanus de Saussure (female); 2) E. belfragei aureus Isely, 1917 (Holotype, female); 3) E. bollii Cresson, 1872 (Lectotype, female); 4) E. bollii oregonensis Bequaert, 1938 (Holotype, female); 5); E. bequaerti Isely, 1917 (Holotype, male); 6) E. iturbide (= E. consobrinus; Holotype, female); 7) E. coyotae Bohart, 1948 (Holotype, male); 8) E. crucifera Provancher, 1888 (Holotype, female). 9) E. bolliformis Viereck, 1908 (Lectotype, female); 10) E. crucifera var. nearcticus Bequaert, 1944 (female); 11) E. crucifera stricklandi Bequaert, 1944 (Holotype, female); 12) E. cruciferoides Giordani Soika, 1978 (Holotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458304/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
), which was considered by Bohart (1951) as a subspecies of
<taxonomicName id="4C6C4D1AFFE1F312FD65960CD073F8AF" authorityName="Provancher" authorityYear="1888" box="[706,842,1880,1904]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE1F312FD65960CD073F8AF" box="[706,842,1880,1904]" italics="true" pageId="18" pageNumber="19">E. crucifera</emphasis>
</taxonomicName>
, identified by its ferrugineous color adjacent to the yellow marks along the body (mainly on the mesosoma, T1, and T2), and the coarse punctation of T2.
</paragraph>
<paragraph id="8BD33699FFE1F313FF6096F5D112FDE3" blockId="18.[151,1437,151,2013]" lastBlockId="19.[151,1437,151,1653]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
The
<typeStatus id="54D7883BFFE1F312FF5C96F4D212F866" box="[251,299,1953,1977]" pageId="18" pageNumber="19">type</typeStatus>
of
<taxonomicName id="4C6C4D1AFFE1F312FEF396F4D2E5F867" authorityName="Provancher" authorityYear="1888" box="[340,476,1952,1976]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE1F312FEF396F4D2E5F867" box="[340,476,1952,1976]" italics="true" pageId="18" pageNumber="19">E. crucifera</emphasis>
</taxonomicName>
is from
<collectingRegion id="49A8F87BFFE1F312FD9896F5D188F866" box="[575,689,1952,1977]" country="United States of America" name="California" pageId="18" pageNumber="19">California</collectingRegion>
and shows a pattern of strongly developed yellow marks and the punctation of T2 often weaker. As the distribution goes to the central-east regions, the punctation becomes stronger and the yellow marks often are less developed, although this variation was not a pattern for some specimens from those regions (see
<emphasis id="B918EA8BFFE0F313FECE91E8D2E9FF0B" box="[361,464,189,212]" italics="true" pageId="19" pageNumber="20">variation</emphasis>
section). Besides the ferrugineous marks, we also observed for
<taxonomicName id="4C6C4D1AFFE0F313FB0291E8D679FF0B" authorityName="Viereck" authorityYear="1908" box="[1189,1344,188,212]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bolliformis">
<emphasis id="B918EA8BFFE0F313FB0291E8D679FF0B" box="[1189,1344,188,212]" italics="true" pageId="19" pageNumber="20">E. bolliformis</emphasis>
</taxonomicName>
that the specimens often present the lateral margins of T1 more concave and the posterior face of propodeum also with the concavity more developed. However, this pattern could not justify a subspecies rank because we observed typical
<taxonomicName id="4C6C4D1AFFE0F313FF30907DD3C7FE9F" authorityName="Provancher" authorityYear="1888" box="[151,254,296,320]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE0F313FF30907DD3C7FE9F" box="[151,254,296,320]" italics="true" pageId="19" pageNumber="20">crucifera</emphasis>
</taxonomicName>
mainly from
<collectingRegion id="49A8F87BFFE0F313FE059072D2E1FE9F" box="[418,472,295,320]" country="United States of America" name="Utah" pageId="19" pageNumber="20">Utah</collectingRegion>
and
<collectingRegion id="49A8F87BFFE0F313FDB19072D147FE9F" box="[534,638,295,320]" country="United States of America" name="Colorado" pageId="19" pageNumber="20">Colorado</collectingRegion>
that also showed these features, including some ferrugineous marks adjacent to the yellow marks, apparently like a “transitional species”. As southern specimens were examined, the ferrugineous marks became more developed and the concavity of the posterior face of the propodeum stronger, although it was variable. One specimen identified as
<taxonomicName id="4C6C4D1AFFE0F313FD5390C1D056FE73" authorityName="Viereck" authorityYear="1908" box="[756,879,404,428]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bolliformis">
<emphasis id="B918EA8BFFE0F313FD5390C1D056FE73" box="[756,879,404,428]" italics="true" pageId="19" pageNumber="20">bolliformis</emphasis>
</taxonomicName>
was recorded from
<collectingRegion id="49A8F87BFFE0F313FBF290C1D7FEFE72" box="[1109,1223,404,429]" country="United States of America" name="California" pageId="19" pageNumber="20">California</collectingRegion>
and as expected it showed more developed yellow marks (with ferrugineous), but the punctation of T2 is weaker. Finally, we compared the genitalia from males of typical
<taxonomicName id="4C6C4D1AFFE0F313FD0C9089D02BFE2B" authorityName="Provancher" authorityYear="1888" box="[683,786,476,500]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE0F313FD0C9089D02BFE2B" box="[683,786,476,500]" italics="true" pageId="19" pageNumber="20">crucifera</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D1AFFE0F313FCE99089D0F0FE2B" authorityName="Viereck" authorityYear="1908" box="[846,969,476,500]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bolliformis">
<emphasis id="B918EA8BFFE0F313FCE99089D0F0FE2B" box="[846,969,476,500]" italics="true" pageId="19" pageNumber="20">bolliformis</emphasis>
</taxonomicName>
and we did not find any difference that would be enough to separate the subspecies into different species. Therefore, we propose to treat
<taxonomicName id="4C6C4D1AFFE0F313FB769354D652FDC7" authorityName="Viereck" authorityYear="1908" box="[1233,1387,512,536]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bolliformis">
<emphasis id="B918EA8BFFE0F313FB769354D652FDC7" box="[1233,1387,512,536]" italics="true" pageId="19" pageNumber="20">E. bolliformis</emphasis>
</taxonomicName>
as a synonymous variant of
<taxonomicName id="4C6C4D1AFFE0F313FE399370D11DFDE3" authorityName="Provancher" authorityYear="1888" box="[414,548,548,572]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE0F313FE399370D11DFDE3" box="[414,548,548,572]" italics="true" pageId="19" pageNumber="20">E. crucifera</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD33699FFE0F313FF60931DD7F5FCA0" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="20">
We examined the
<typeStatus id="54D7883BFFE0F313FE369312D2CAFDBF" box="[401,499,583,608]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C6C4D1AFFE0F313FDBE931CD188FDBF" authorityName="de Saussure" authorityYear="1855" box="[537,689,584,608]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="globulosus">
<emphasis id="B918EA8BFFE0F313FDBE931CD188FDBF" box="[537,689,584,608]" italics="true" pageId="19" pageNumber="20">E. globulosus</emphasis>
</taxonomicName>
(
<figureCitation id="13572A1CFFE0F313FD669312D02AFDBF" box="[705,787,583,608]" captionStart="FIGURES 112" captionStartId="41.[151,264,1386,1408]" captionTargetBox="[151,1436,193,1341]" captionTargetId="figure@41.[151,1436,193,1341]" captionTargetPageId="41" captionText="FIGURES 112. Eumenes species, lateral view. 1) Eumenes americanus de Saussure (female); 2) E. belfragei aureus Isely, 1917 (Holotype, female); 3) E. bollii Cresson, 1872 (Lectotype, female); 4) E. bollii oregonensis Bequaert, 1938 (Holotype, female); 5); E. bequaerti Isely, 1917 (Holotype, male); 6) E. iturbide (= E. consobrinus; Holotype, female); 7) E. coyotae Bohart, 1948 (Holotype, male); 8) E. crucifera Provancher, 1888 (Holotype, female). 9) E. bolliformis Viereck, 1908 (Lectotype, female); 10) E. crucifera var. nearcticus Bequaert, 1944 (female); 11) E. crucifera stricklandi Bequaert, 1944 (Holotype, female); 12) E. cruciferoides Giordani Soika, 1978 (Holotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458304/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
) and, as we observed in
<taxonomicName id="4C6C4D1AFFE0F313FB8D931CD788FDBF" authorityName="Say" authorityYear="1824" box="[1066,1201,584,608]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFE0F313FB8D931CD788FDBF" box="[1066,1201,584,608]" italics="true" pageId="19" pageNumber="20">E. verticalis</emphasis>
</taxonomicName>
, the specimens from
<collectingCountry id="F37B7609FFE0F313FF309339D3D2FD5A" box="[151,235,620,645]" name="Canada" pageId="19" pageNumber="20">Canada</collectingCountry>
and northern to central-west regions of
<collectingCountry id="F37B7609FFE0F313FD129339D1D4FD5B" box="[693,749,620,644]" name="United States of America" pageId="19" pageNumber="20">USA</collectingCountry>
presented a pattern with reduced yellow marks on the body. All the variation, especially on T1, the posterior face of propodeum, and punctation of T2, was observed on the specimens identified as
<taxonomicName id="4C6C4D1AFFE0F313FE0593E0D19BFD13" authorityName="Bequaert" authorityYear="1944" box="[418,674,692,716]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="nearcticus">
<emphasis id="B918EA8BFFE0F313FE0593E0D19BFD13" box="[418,674,692,716]" italics="true" pageId="19" pageNumber="20">E. crucifera nearcticus</emphasis>
</taxonomicName>
. As we did not find any pattern to treat it as a different species from the typical species, we propose to treat
<taxonomicName id="4C6C4D1AFFE0F313FDCF938DD1E5FD30" authorityName="Bequaert" authorityYear="1944" box="[616,732,728,751]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="nearcticus">
<emphasis id="B918EA8BFFE0F313FDCF938DD1E5FD30" box="[616,732,728,751]" italics="true" pageId="19" pageNumber="20">nearcticus</emphasis>
</taxonomicName>
as a variant of
<emphasis id="B918EA8BFFE0F313FC3D938CD712FD2F" box="[922,1067,728,752]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="4C6C4D1AFFE0F313FC3D938CD71FFD2F" authorityName="Provancher" authorityYear="1888" box="[922,1062,728,752]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="crucifera">E. crucifera</taxonomicName>
.
</emphasis>
Furthermore, the differences in yellow marks are more visible in females, while males often have the yellow marks reduced and less variable. Some females presented the clypeus with coarse punctation, which may lead them to be confused with
<taxonomicName id="4C6C4D1AFFE0F313FAB29274D6A5FCE7" authorityName="Say" authorityYear="1824" box="[1301,1436,800,824]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFE0F313FAB29274D6A5FCE7" box="[1301,1436,800,824]" italics="true" pageId="19" pageNumber="20">E. verticalis</emphasis>
</taxonomicName>
(mostly the
<taxonomicName id="4C6C4D1AFFE0F313FE819211D2A3FC83" authorityName="Bequaert" authorityYear="1944" box="[294,410,836,860]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="neoboreus">
<emphasis id="B918EA8BFFE0F313FE819211D2A3FC83" box="[294,410,836,860]" italics="true" pageId="19" pageNumber="20">neoboreus</emphasis>
</taxonomicName>
variation), but the latter species still presents the clypeus with punctures clearly more evident, T1 is usually less swollen in dorsal view and the lateral margins of T2 are less concave.
</paragraph>
<paragraph id="8BD33699FFE0F313FF6092D8D6AAFC33" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="20">
<taxonomicName id="4C6C4D1AFFE0F313FF6092D8D120FC7B" authorityName="Bequaert" authorityYear="1944" box="[199,537,908,932]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="stricklandi">
<emphasis id="B918EA8BFFE0F313FF6092D8D120FC7B" box="[199,537,908,932]" italics="true" pageId="19" pageNumber="20">Eumenes crucifera stricklandi</emphasis>
</taxonomicName>
(
<figureCitation id="13572A1CFFE0F313FD8E92D9D140FC7B" box="[553,633,908,933]" captionStart="FIGURES 112" captionStartId="41.[151,264,1386,1408]" captionTargetBox="[151,1436,193,1341]" captionTargetId="figure@41.[151,1436,193,1341]" captionTargetPageId="41" captionText="FIGURES 112. Eumenes species, lateral view. 1) Eumenes americanus de Saussure (female); 2) E. belfragei aureus Isely, 1917 (Holotype, female); 3) E. bollii Cresson, 1872 (Lectotype, female); 4) E. bollii oregonensis Bequaert, 1938 (Holotype, female); 5); E. bequaerti Isely, 1917 (Holotype, male); 6) E. iturbide (= E. consobrinus; Holotype, female); 7) E. coyotae Bohart, 1948 (Holotype, male); 8) E. crucifera Provancher, 1888 (Holotype, female). 9) E. bolliformis Viereck, 1908 (Lectotype, female); 10) E. crucifera var. nearcticus Bequaert, 1944 (female); 11) E. crucifera stricklandi Bequaert, 1944 (Holotype, female); 12) E. cruciferoides Giordani Soika, 1978 (Holotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458304/files/figure.png" pageId="19" pageNumber="20">Fig. 11</figureCitation>
) is known only from the
<typeStatus id="54D7883BFFE0F313FC3F92D8D0F1FC7A" box="[920,968,909,933]" pageId="19" pageNumber="20">type</typeStatus>
specimen and besides the yellow-creamy marks on metasoma, there is no difference in pubescence, punctation, and structure compared to typical
<taxonomicName id="4C6C4D1AFFE0F313FA2392E4D3C7FC33" authorityName="Provancher" authorityYear="1888" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE0F313FA2392E4D3C7FC33" italics="true" pageId="19" pageNumber="20">E. crucifera</emphasis>
</taxonomicName>
. We did not examine any males of the subspecies, but propose to treat it also as a variant of
<taxonomicName id="4C6C4D1AFFE0F313FAA09280D6B5FC33" authorityName="Provancher" authorityYear="1888" box="[1287,1420,980,1004]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE0F313FAA09280D6B5FC33" box="[1287,1420,980,1004]" italics="true" pageId="19" pageNumber="20">E. crucifera</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE0F313FF6092A2D674F9AB" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8BD33699FFE0F313FF6092A2D1A8FAD3" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="20">
<emphasis id="B918EA8BFFE0F313FF6092A2D240FBCF" bold="true" box="[199,377,1015,1040]" pageId="19" pageNumber="20">Type material:</emphasis>
The holotype of
<taxonomicName id="4C6C4D1AFFE0F313FDE592ACD02EFBCF" authorityName="Provancher" authorityYear="1888" box="[578,791,1016,1040]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE0F313FDE592ACD02EFBCF" box="[578,791,1016,1040]" italics="true" pageId="19" pageNumber="20">Eumenes crucifera</emphasis>
</taxonomicName>
(female) is in good condition and bears the labels: Los Angeles \ Co. CAL. (C) 178 [red label] Type No \ 1977 \ USNM [red label] Type \ No. 1977 \ U.S.N.M. [partially handwritten label]
<emphasis id="B918EA8BFFE0F313FE4D9514D125FB87" box="[490,540,1089,1112]" italics="true" pageId="19" pageNumber="20">E. c.</emphasis>
\
<taxonomicName id="4C6C4D1AFFE0F313FD959515D1A0FB87" authorityName="Provancher" authorityYear="1888" box="[562,665,1088,1112]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE0F313FD959515D1A0FB87" box="[562,665,1088,1112]" italics="true" pageId="19" pageNumber="20">crucifera</emphasis>
</taxonomicName>
\ Prov. DO NOT REMOVE \ SI DB Reference \ Not a property tag \ T. Schultz. NMHN \\ USNM ENT \ 0 0 537193 [bar code] Loan from \ USNMNH \ 2073492. The paratypes at USNM are in good condition and bear the labels: Los Angeles \ Co. CAL. (C) [Red Label] Paratype \ No. 21381 \ U.S.N.M.
<taxonomicName id="4C6C4D1AFFE0F313FE9395F8D144FB1B" authority="crucifera" box="[308,637,1196,1220]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="crucifera" subSpecies="crucifera">
<emphasis id="B918EA8BFFE0F313FE9395F8D144FB1B" box="[308,637,1196,1220]" italics="true" pageId="19" pageNumber="20">Eumenes crucifera crucifera</emphasis>
</taxonomicName>
\ R. Bohart`44 Prov; Los Angeles \ Co. CAL. (C) [Red Label] Paratype \ No. 21381 \ U.S.N.M.; 178 Los Angeles \ Co., Cal. Collection \ Coquillett [Red Label] Type No \ 1977 \ USNM
<taxonomicName id="4C6C4D1AFFE0F313FEE895A0D28DFAD3" box="[335,436,1269,1292]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFE0F313FEE895A0D28DFAD3" box="[335,436,1269,1292]" italics="true" pageId="19" pageNumber="20">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFE0F313FE6D95A1D108FAD3" authorityName="Provancher" authorityYear="1888" box="[458,561,1268,1292]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B918EA8BFFE0F313FE6D95A1D108FAD3" box="[458,561,1268,1292]" italics="true" pageId="19" pageNumber="20">crucifera</emphasis>
</taxonomicName>
\ Prov..
</paragraph>
<paragraph id="8BD33699FFE0F313FF609442D2EAFA8B" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="20">
The
<typeStatus id="54D7883BFFE0F313FF5E9442D262FAEF" box="[249,347,1303,1328]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C6C4D1AFFE0F313FED9944CD145FAEF" authorityName="Isely" authorityYear="1917" box="[382,636,1304,1328]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="xanthogaster">
<emphasis id="B918EA8BFFE0F313FED9944CD145FAEF" box="[382,636,1304,1328]" italics="true" pageId="19" pageNumber="20">Eumenes xanthogaster</emphasis>
</taxonomicName>
(male) is in bad condition with the metasoma broken (glued on a label), and lacking the left F2F11.
</paragraph>
<paragraph id="8BD33699FFE0F313FF60940AD151F9D7" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="20">
The
<typeStatus id="54D7883BFFE0F313FF5C940AD264FAA7" box="[251,349,1375,1400]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
female of
<taxonomicName id="4C6C4D1AFFE0F313FE7F9434D186FAA7" authorityName="de Saussure" authorityYear="1855" box="[472,703,1376,1400]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="globulosus">
<emphasis id="B918EA8BFFE0F313FE7F9434D186FAA7" box="[472,703,1376,1400]" italics="true" pageId="19" pageNumber="20">Eumenes globulosus</emphasis>
</taxonomicName>
is in bad condition, missing the apical tarsi of the right fore leg, entirely mid right leg, all tarsi of the right hind leg and the flagellum of the both antenna. Bears the labels: Amer. Sept. [Red Label] Holo- \
<collectionCode id="ED7DAE5CFFE0F313FE7494FDD120FA60" box="[467,537,1448,1471]" pageId="19" pageNumber="20">
<typeStatus id="54D7883BFFE0F313FE7494FDD120FA60" box="[467,537,1448,1471]" pageId="19" pageNumber="20">TYPE</typeStatus>
</collectionCode>
\
<specimenCount id="9D6AFD10FFE0F313FD9794FDD17CFA60" box="[560,581,1448,1471]" pageId="19" pageNumber="20" type="female"></specimenCount>
<taxonomicName id="4C6C4D1AFFE0F313FDEA94FDD1FFFA1F" authorityName="de Saussure" authorityYear="1855" box="[589,710,1448,1472]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="globulosus">
<emphasis id="B918EA8BFFE0F313FDEA94FDD1FFFA1F" box="[589,710,1448,1472]" italics="true" pageId="19" pageNumber="20">globulosus</emphasis>
</taxonomicName>
\ Saussure
<collectionCode id="ED7DAE5CFFE0F313FCF094FDD08BFA1F" box="[855,946,1448,1472]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="19" pageNumber="20">MHNG</collectionCode>
\
<collectionCode id="ED7DAE5CFFE0F313FC6D94FDD72DFA1F" box="[970,1044,1448,1472]" pageId="19" pageNumber="20">ENTO</collectionCode>
\ 00008850. The
<typeStatus id="54D7883BFFE0F313FB4094FDD67EFA1F" box="[1255,1351,1448,1472]" pageId="19" pageNumber="20" type="paratype">paratype</typeStatus>
female (
<collectionCode id="ED7DAE5CFFE0F313FF399499D3CBFA3B" box="[158,242,1484,1508]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="20">USNM</collectionCode>
) is in good condition and bears the labels: Ft. Collins \ Col. 1047
<taxonomicName id="4C6C4D1AFFE0F313FC539498D760FA3B" box="[1012,1113,1485,1508]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFE0F313FC539498D760FA3B" box="[1012,1113,1485,1508]" italics="true" pageId="19" pageNumber="20">Eumenes</emphasis>
</taxonomicName>
<specimenCount id="9D6AFD10FFE0F313FBC79499D74CFA3A" box="[1120,1141,1484,1509]" pageId="19" pageNumber="20" type="female">
<emphasis id="B918EA8BFFE0F313FBC79499D74CFA3A" bold="true" box="[1120,1141,1484,1509]" pageId="19" pageNumber="20"></emphasis>
</specimenCount>
\
<taxonomicName id="4C6C4D1AFFE0F313FB239499D669FA3B" authority="Sauss." authorityName="de Saussure" authorityYear="1855" box="[1156,1360,1484,1508]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="globulosus">
<emphasis id="B918EA8BFFE0F313FB239499D7C4FA3B" box="[1156,1277,1484,1508]" italics="true" pageId="19" pageNumber="20">globulosus</emphasis>
Sauss.
</taxonomicName>
[Red Label]
<typeStatus id="54D7883BFFE0F313FF4094A5D270F9D7" box="[231,329,1520,1544]" pageId="19" pageNumber="20" type="paratype">Paratype</typeStatus>
\
<collectingRegion id="49A8F87BFFE0F313FEF894A5D2BDF9D8" box="[351,388,1520,1543]" country="Haiti" name="Nord" pageId="19" pageNumber="20">No</collectingRegion>
. 21382 \
<collectingCountry id="F37B7609FFE0F313FE4C94A5D124F9D7" box="[491,541,1520,1544]" name="United States of America" pageId="19" pageNumber="20">U.S.</collectingCountry>
N.M..
</paragraph>
<paragraph id="8BD33699FFE0F313FF609741D674F9AB" blockId="19.[151,1437,151,1653]" pageId="19" pageNumber="20">
The
<typeStatus id="54D7883BFFE0F313FF5C9741D264F9F2" box="[251,349,1556,1581]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
female of
<taxonomicName id="4C6C4D1AFFE0F313FE7E9740D06CF9F3" box="[473,853,1556,1580]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="coloradensis" subSpecies="stricklandi">
<emphasis id="B918EA8BFFE0F313FE7E9740D06CF9F3" box="[473,853,1556,1580]" italics="true" pageId="19" pageNumber="20">Eumenes coloradensis stricklandi</emphasis>
</taxonomicName>
is in good condition and bears the labels: Cypress Hills, Alta. \
<date id="FFD21059FFE0F313FE92976DD2E9F98F" box="[309,464,1592,1616]" pageId="19" pageNumber="20" value="1939-08-10">10 VIII 1939</date>
\ E. H. Stricklandi [Red Label] M.C.Z. \
<typeStatus id="54D7883BFFE0F313FC719762D707F98F" box="[982,1086,1591,1616]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
\
<taxonomicName id="4C6C4D1AFFE0F313FBFA9762D7EAF98F" box="[1117,1235,1591,1616]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="coloradensis" subSpecies="stricklandi">stricklandi</taxonomicName>
<emphasis id="B918EA8BFFE0F313FB799762D7DFF98F" bold="true" box="[1246,1254,1591,1616]" pageId="19" pageNumber="20">\</emphasis>
<specimenCount id="9D6AFD10FFE0F313FB56976DD631F98F" box="[1265,1288,1592,1616]" pageId="19" pageNumber="20" type="female"></specimenCount>
<taxonomicName id="4C6C4D1AFFE0F313FA83976CD6B0F98F" box="[1316,1417,1593,1616]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFE0F313FA83976CD6B0F98F" box="[1316,1417,1593,1616]" italics="true" pageId="19" pageNumber="20">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFE0F313FF309709D21FF9AB" authorityName="Cresson" authorityYear="1875" box="[151,294,1628,1652]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="coloradensis">
<emphasis id="B918EA8BFFE0F313FF309709D21FF9AB" box="[151,294,1628,1652]" italics="true" pageId="19" pageNumber="20">coloradensis</emphasis>
</taxonomicName>
\ var. \
<taxonomicName id="4C6C4D1AFFE0F313FEDC9709D2CAF9AB" box="[379,499,1628,1652]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="coloradensis" subSpecies="stricklandi">
<emphasis id="B918EA8BFFE0F313FEDC9709D2CAF9AB" box="[379,499,1628,1652]" italics="true" pageId="19" pageNumber="20">stricklandi</emphasis>
</taxonomicName>
\ J. Beq [Red Label] M.C.Z. \
<typeStatus id="54D7883BFFE0F313FCFA9709D0FCF9AA" box="[861,965,1628,1653]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
\ 33184 MCZ-ENT \ 00033184.
</paragraph>
</subSubSection>
</treatment>
</document>