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<document id="5CC739AAD044DB43254034C401D30F01" ID-CLB-Dataset="29039" ID-DOI="10.11646/zootaxa.4459.1.1" ID-GBIF-Dataset="9acf71bc-6414-4878-8af0-a64d3ec1fea6" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458302" ID-ZooBank="633A9498-FCC6-4439-BE33-A45B9E51B550" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539287574730" checkinUser="plazi" docAuthor="Grandinete, Yuri Campanholo, Noll, Fernando Barbosa &amp; Carpenter, James" docDate="2018" docId="03C5878FFFD2F323FF30952FD7E9FE57" docLanguage="en" docName="zootaxa.4459.1.1.pdf" docOrigin="Zootaxa 4459 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eumenes filiformis de Saussure 1855" docType="treatment" docVersion="9" lastPageNumber="36" masterDocId="FFFCFFF7FFF3F300FFA79155D339FFDF" masterDocTitle="Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World" masterLastPageNumber="52" masterPageNumber="1" pageNumber="34" updateTime="1698577660185" updateUser="ExternalLinkService">
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<mods:title id="964656DBAD94FA55D87C00B4A4E4A40F">Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World</mods:title>
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<mods:namePart id="0AD0366436F9EA3CC01B20D2C895D2B1">Grandinete, Yuri Campanholo</mods:namePart>
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<mods:namePart id="DF43AB26D2C5E7B901D51BF28D4195C4">Noll, Fernando Barbosa</mods:namePart>
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<mods:namePart id="A0AAA1470084DD1DC1841F7463650C15">Carpenter, James</mods:namePart>
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<mods:title id="B45AF74B763D64C6E6E57DFD7B83E36F">Zootaxa</mods:title>
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<mods:date id="9F6F796E9B0EC1C9B2CD97A9136F4512">2018</mods:date>
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<mods:number id="205F0F3977F2A40FC2A3F0BBDB42B0A8">2018-08-14</mods:number>
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<mods:number id="268BCE430065BC7058B7DFAC9C06F884">4459</mods:number>
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<mods:identifier id="43846F25F7066C4440469C2CA2F2DF8B" type="DOI">10.11646/zootaxa.4459.1.1</mods:identifier>
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<treatment id="03C5878FFFD2F323FF30952FD7E9FE57" ID-DOI="http://doi.org/10.5281/zenodo.5986396" ID-GBIF-Taxon="148692492" ID-Zenodo-Dep="5986396" LSID="urn:lsid:plazi:treatment:03C5878FFFD2F323FF30952FD7E9FE57" httpUri="http://treatment.plazi.org/id/03C5878FFFD2F323FF30952FD7E9FE57" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
<subSubSection id="C3766512FFD2F321FF30952FD140FB08" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BD33699FFD2F321FF30952FD15DFB4A" blockId="33.[151,612,1146,1173]" box="[151,612,1146,1173]" pageId="33" pageNumber="34">
<heading id="D09B81F5FFD2F321FF30952FD15DFB4A" bold="true" box="[151,612,1146,1173]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<taxonomicName id="4C6C4D1AFFD2F321FF30952FD15DFB4A" ID-CoL="3CDGZ" authority="de Saussure, 1855" authorityName="de Saussure" authorityYear="1855" box="[151,612,1146,1173]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD2F321FF30952FD15DFB4A" bold="true" box="[151,612,1146,1173]" pageId="33" pageNumber="34">
<emphasis id="B918EA8BFFD2F321FF30952FD246FB4B" bold="true" box="[151,383,1146,1172]" italics="true" pageId="33" pageNumber="34">Eumenes filiformis</emphasis>
de Saussure, 1855
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD33699FFD2F321FF309597D140FB08" blockId="33.[151,1436,1217,2036]" box="[151,633,1217,1239]" pageId="33" pageNumber="34">
<taxonomicName id="4C6C4D1AFFD2F321FF309597D11CFB08" authority="de Saussure, 1855" authorityName="de Saussure" authorityYear="1855" box="[151,549,1217,1239]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD2F321FF309597D11CFB08" bold="true" box="[151,549,1217,1239]" pageId="33" pageNumber="34">
<emphasis id="B918EA8BFFD2F321FF309597D266FB08" bold="true" box="[151,351,1217,1239]" italics="true" pageId="33" pageNumber="34">Eumenes filiformis</emphasis>
de Saussure, 1855
</emphasis>
</taxonomicName>
, 3: 146.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD2F321FF6095B5D15EF97C" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8BD33699FFD2F321FF6095B5D12EFB29" blockId="33.[151,1436,1217,2036]" box="[199,535,1248,1270]" pageId="33" pageNumber="34">
<emphasis id="B918EA8BFFD2F321FF6095B5D244FB29" bold="true" box="[199,381,1248,1270]" pageId="33" pageNumber="34">
<typeStatus id="54D7883BFFD2F321FF6095B5D3C5FB29" box="[199,252,1248,1270]" pageId="33" pageNumber="34">Type</typeStatus>
data: Lecto
</emphasis>
<typeStatus id="54D7883BFFD2F321FEDA95B4D290FB29" box="[381,425,1249,1270]" pageId="33" pageNumber="34">type</typeStatus>
at
<collectionCode id="ED7DAE5CFFD2F321FE6F95B5D12AFB29" box="[456,531,1248,1270]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15721" name="Museo Regionale di Scienze Naturali" pageId="33" pageNumber="34">MRSN</collectionCode>
.
</paragraph>
<paragraph id="8BD33699FFD2F321FF6095ABD2E2FACB" blockId="33.[151,1436,1217,2036]" box="[199,475,1278,1300]" pageId="33" pageNumber="34">
<materialsCitation id="3B043CC4FFD2F321FF6095ABD2E2FACB" ID-GBIF-Occurrence="1917355161" box="[199,475,1278,1300]" country="Brazil" location="Bresil" pageId="33" pageNumber="34" specimenCount="1" typeStatus="holotype">
<emphasis id="B918EA8BFFD2F321FF6095ABD263FACB" bold="true" box="[199,346,1278,1300]" pageId="33" pageNumber="34">
<typeStatus id="54D7883BFFD2F321FF6095ABD3C5FACB" box="[199,252,1278,1300]" pageId="33" pageNumber="34">Type</typeStatus>
locality:
</emphasis>
“Le
<collectingCountry id="F37B7609FFD2F321FE2B95ABD2F4FACB" box="[396,461,1278,1300]" name="Brazil" pageId="33" pageNumber="34">Bresil</collectingCountry>
”.
</materialsCitation>
</paragraph>
<paragraph id="8BD33699FFD2F321FF609448D2FDFAEC" blockId="33.[151,1436,1217,2036]" box="[199,452,1309,1331]" pageId="33" pageNumber="34">Smith, 1857, 5: 30 (cat.).</paragraph>
<paragraph id="8BD33699FFD2F321FF609469D2D2FA8D" blockId="33.[151,1436,1217,2036]" box="[199,491,1340,1362]" pageId="33" pageNumber="34">de Saussure, 1875, 254: 109.</paragraph>
<paragraph id="8BD33699FFD2F321FF60940FD2C5FAAF" blockId="33.[151,1436,1217,2036]" box="[199,508,1370,1392]" pageId="33" pageNumber="34">Dalla Torre, 1894, 9: 24 (cat.).</paragraph>
<paragraph id="8BD33699FFD2F321FF60942FD29EFA50" blockId="33.[151,1436,1217,2036]" box="[199,423,1401,1423]" pageId="33" pageNumber="34">Fox, 1899, 1899: 457.</paragraph>
<paragraph id="8BD33699FFD2F321FF6094CDD131FA71" blockId="33.[151,1436,1217,2036]" box="[199,520,1432,1454]" pageId="33" pageNumber="34">Dalla Torre, 1904, 19: 22 (cat.).</paragraph>
<paragraph id="8BD33699FFD2F321FF6094E3D2DCFA13" blockId="33.[151,1436,1217,2036]" box="[199,485,1462,1484]" pageId="33" pageNumber="34">Bertoni, 1912, 22: 106, 108.</paragraph>
<paragraph id="8BD33699FFD2F321FF609480D2C1FA34" blockId="33.[151,1436,1217,2036]" box="[199,504,1493,1515]" pageId="33" pageNumber="34">
<bibRefCitation id="EFFD4B68FFD2F321FF609480D25DFA34" box="[199,356,1493,1515]" pageId="33" pageNumber="34" refId="ref44216" refString="Zavattari, E. (1912) Materialien fur eine Monographie der neotropischen Eumeniden. Archiv fur Naturgeschichte, 78 A (4), 1 - 272." type="journal article">Zavattari, 1912</bibRefCitation>
, 78A (4): 129.
</paragraph>
<paragraph id="8BD33699FFD2F321FF6094A1D2CDF9D5" blockId="33.[151,1436,1217,2036]" box="[199,500,1524,1546]" pageId="33" pageNumber="34">Giordani Soika, 1961, 1: 241;</paragraph>
<paragraph id="8BD33699FFD2F321FF609747D03EF9F7" blockId="33.[151,1436,1217,2036]" box="[199,775,1554,1576]" pageId="33" pageNumber="34">
<bibRefCitation id="EFFD4B68FFD2F321FF609747D29BF9F7" box="[199,418,1554,1576]" pageId="33" pageNumber="34" refId="ref43151" refString="Giordani Soika, A. (1978) Revisione degli Eumenidi neotropicali appartenenti ai generi Eumenes Latr., Omicron (Sauss.), Pararhaphidoglossa Schulth. ed afinni. Bolletino della Museo Civico di Storia Naturale di Uenezia, 29, 1 - 420." type="journal article">Giordani Soika, 1978</bibRefCitation>
, 29: 43 (designation of
<typeStatus id="54D7883BFFD2F321FD359746D1C5F9F7" box="[658,764,1555,1576]" pageId="33" pageNumber="34" type="lectotype">Lectotype</typeStatus>
).
</paragraph>
<paragraph id="8BD33699FFD2F321FF609764D060F998" blockId="33.[151,1436,1217,2036]" box="[199,857,1585,1607]" pageId="33" pageNumber="34">
Carpenter &amp; van der
<bibRefCitation id="EFFD4B68FFD2F321FE3B9764D121F998" box="[412,536,1585,1607]" pageId="33" pageNumber="34" refId="ref42766" refString="Carpenter, J. M. &amp; van der Vecht, J. (1991) A study of the Vespidae described by William J. Fox (Insecta: Hymenoptera), with assessment of taxonomic implications. Annals of the Carnegie Museum of Natural History, 60 (3), 211 - 241." type="journal article">Vecht, 1991</bibRefCitation>
, 60: 211, 231 (distinct species).
</paragraph>
<paragraph id="8BD33699FFD2F321FF609705D129F9B9" blockId="33.[151,1436,1217,2036]" box="[199,528,1616,1638]" pageId="33" pageNumber="34">
West-Eberhard
<emphasis id="B918EA8BFFD2F321FEC39704D2AEF9B9" box="[356,407,1616,1638]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
, 1995: 573.
</paragraph>
<paragraph id="8BD33699FFD2F321FF60973AD153F95B" blockId="33.[151,1436,1217,2036]" box="[199,618,1646,1668]" pageId="33" pageNumber="34">Rasmussen &amp; Asenjo, 2009, 15: 38 (list).</paragraph>
<paragraph id="8BD33699FFD2F321FF3097DBD15EF97C" blockId="33.[151,1436,1217,2036]" box="[151,615,1677,1699]" pageId="33" pageNumber="34">
<taxonomicName id="4C6C4D1AFFD2F321FF3097DBD249F97C" box="[151,368,1677,1699]" class="Insecta" family="Vespidae" genus="Zeteumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis id="B918EA8BFFD2F321FF3097DBD249F97C" box="[151,368,1677,1699]" italics="true" pageId="33" pageNumber="34">Zeteumenes filicornis</emphasis>
</taxonomicName>
[!]; Bertoni, 1921, 1: 11
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD2F321FF3097F8D027F920" pageId="33" pageNumber="34" type="description">
<paragraph id="8BD33699FFD2F321FF3097F8D025F91D" blockId="33.[151,1436,1217,2036]" box="[151,796,1708,1730]" pageId="33" pageNumber="34">
<taxonomicName id="4C6C4D1AFFD2F321FF3097F8D139F91D" box="[151,512,1708,1730]" class="Insecta" family="Vespidae" genus="Zeteumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="filicornis" subGenus="Beteumenes">
<emphasis id="B918EA8BFFD2F321FF3097F8D236F91D" box="[151,271,1709,1730]" italics="true" pageId="33" pageNumber="34">Zeteumenes</emphasis>
(
<emphasis id="B918EA8BFFD2F321FEBA97F8D2AFF91D" box="[285,406,1709,1730]" italics="true" pageId="33" pageNumber="34">Beteumenes</emphasis>
)
<emphasis id="B918EA8BFFD2F321FE0397F9D139F91D" box="[420,512,1708,1730]" italics="true" pageId="33" pageNumber="34">filicornis</emphasis>
</taxonomicName>
; Bertoni, 1934, 3: 111, 115.
</paragraph>
<paragraph id="8BD33699FFD2F321FF60979FD1FFF93F" blockId="33.[151,1436,1217,2036]" box="[199,710,1738,1760]" pageId="33" pageNumber="34">Bertoni, 1934, Rev. Soc. Cient. Parag. 3: 111, 115.</paragraph>
<paragraph id="8BD33699FFD2F321FF3097BFD027F920" blockId="33.[151,1436,1217,2036]" box="[151,798,1769,1791]" pageId="33" pageNumber="34">
<taxonomicName id="4C6C4D1AFFD2F321FF3097BFD23AF920" authorityName="Bertoni" authorityYear="1925" box="[151,259,1770,1791]" class="Insecta" family="Vespidae" genus="Zetamenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFD2F321FF3097BFD23AF920" box="[151,259,1770,1791]" italics="true" pageId="33" pageNumber="34">Zetamenes</emphasis>
</taxonomicName>
[!]
<taxonomicName id="4C6C4D1AFFD2F321FE8E97BCD1E3F920" authority="Bertoni, 1925" authorityName="Bertoni" authorityYear="1925" box="[297,730,1769,1791]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="variety" species="filiformis" variety="costarricensis">
<emphasis id="B918EA8BFFD2F321FE8E97BCD2BFF920" box="[297,390,1769,1791]" italics="true" pageId="33" pageNumber="34">filiformis</emphasis>
var.
<emphasis id="B918EA8BFFD2F321FE1F97BFD17FF920" box="[440,582,1770,1791]" italics="true" pageId="33" pageNumber="34">costarricensis</emphasis>
Bertoni, 1925
</taxonomicName>
, 2: 75.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD2F321FF60965DD1DDF884" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8BD33699FFD2F321FF60965DD120F8C1" blockId="33.[151,1436,1217,2036]" box="[199,537,1800,1822]" pageId="33" pageNumber="34">
<emphasis id="B918EA8BFFD2F321FF60965DD203F8C1" bold="true" box="[199,314,1800,1822]" pageId="33" pageNumber="34">Type data:</emphasis>
Type male at IBNPY.
</paragraph>
<paragraph id="8BD33699FFD2F321FF609673D2D4F8E3" blockId="33.[151,1436,1217,2036]" box="[199,493,1830,1852]" pageId="33" pageNumber="34">
<materialsCitation id="3B043CC4FFD2F321FF609673D2D4F8E3" ID-GBIF-Occurrence="1917355183" box="[199,493,1830,1852]" country="Costa Rica" location="Costa Rica" pageId="33" pageNumber="34" specimenCount="1" typeStatus="holotype">
<emphasis id="B918EA8BFFD2F321FF609673D254F8E3" bold="true" box="[199,365,1830,1852]" pageId="33" pageNumber="34">
<typeStatus id="54D7883BFFD2F321FF609673D3C5F8E3" box="[199,252,1830,1852]" pageId="33" pageNumber="34">Type</typeStatus>
locality: “
</emphasis>
<collectingCountry id="F37B7609FFD2F321FECB9673D2E6F8E3" box="[364,479,1830,1852]" name="Costa Rica" pageId="33" pageNumber="34">Costa Rica</collectingCountry>
”.
</materialsCitation>
</paragraph>
<paragraph id="8BD33699FFD2F321FF309610D1DDF884" blockId="33.[151,1436,1217,2036]" box="[151,740,1861,1883]" pageId="33" pageNumber="34">
<taxonomicName id="4C6C4D1AFFD2F321FF309610D2ABF884" box="[151,402,1861,1883]" class="Insecta" genus="Zeteumenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD2F321FF309610D2ABF884" box="[151,402,1861,1883]" italics="true" pageId="33" pageNumber="34">Zeteumenoides filiformis</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFFD4B68FFD2F321FE389610D143F884" box="[415,634,1861,1883]" pageId="33" pageNumber="34" refId="ref43124" refString="Giordani Soika, A. (1972) Notulae vespidologicae XXXII Nuovi Eumenidi Indomalesi. Bolletino dela Societa Entomologica Italiana, 104, 99 - 110." type="journal article">Giordani Soika, 1972</bibRefCitation>
, 104: 110.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD2F321FF309630D092F8A5" box="[151,939,1892,1914]" pageId="33" pageNumber="34" type="description">
<paragraph id="8BD33699FFD2F321FF309630D092F8A5" blockId="33.[151,1436,1217,2036]" box="[151,939,1892,1914]" pageId="33" pageNumber="34">
<taxonomicName id="4C6C4D1AFFD2F321FF309630D148F8A5" box="[151,625,1892,1914]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="versicolor" subGenus="Zeteumenoides" subSpecies="filiformis">
<emphasis id="B918EA8BFFD2F321FF309630D3CAF8A5" box="[151,243,1893,1914]" italics="true" pageId="33" pageNumber="34">Eumenes</emphasis>
(
<emphasis id="B918EA8BFFD2F321FEA69631D2A0F8A5" box="[257,409,1892,1914]" italics="true" pageId="33" pageNumber="34">Zeteumenoides</emphasis>
)
<emphasis id="B918EA8BFFD2F321FE009631D148F8A5" box="[423,625,1892,1914]" italics="true" pageId="33" pageNumber="34">versicolor filiformis</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFFD4B68FFD2F321FDD89631D063F8A5" box="[639,858,1892,1914]" pageId="33" pageNumber="34" refId="ref43151" refString="Giordani Soika, A. (1978) Revisione degli Eumenidi neotropicali appartenenti ai generi Eumenes Latr., Omicron (Sauss.), Pararhaphidoglossa Schulth. ed afinni. Bolletino della Museo Civico di Storia Naturale di Uenezia, 29, 1 - 420." type="journal article">Giordani Soika, 1978</bibRefCitation>
, 29: 43.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD2F321FF6096D7D2CAF82B" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8BD33699FFD2F321FF6096D7D1D2F809" blockId="33.[151,1436,1217,2036]" pageId="33" pageNumber="34">
<emphasis id="B918EA8BFFD2F321FF6096D7D26AF847" bold="true" box="[199,339,1922,1944]" pageId="33" pageNumber="34">Distribution:</emphasis>
<collectingCountry id="F37B7609FFD2F321FEC796D7D2A7F847" box="[352,414,1922,1944]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
(AM, AP*, PA*, RO);
<collectingCountry id="F37B7609FFD2F321FD0A96D7D1C3F847" box="[685,762,1922,1944]" name="Bolivia" pageId="33" pageNumber="34">Bolivia</collectingCountry>
* (Coroico);
<collectingCountry id="F37B7609FFD2F321FC2F96D7D0D5F847" box="[904,1004,1922,1944]" name="Colombia" pageId="33" pageNumber="34">Colombia</collectingCountry>
(
<collectingRegion id="49A8F87BFFD2F321FBA796D7D748F847" box="[1024,1137,1922,1944]" country="Peru" name="Amazonas" pageId="33" pageNumber="34">Amazonas</collectingRegion>
*,
<collectingRegion id="49A8F87BFFD2F321FB2B96D7D7E3F847" box="[1164,1242,1922,1944]" country="Colombia" name="Arauca" pageId="33" pageNumber="34">Arauca</collectingRegion>
*,
<collectingRegion id="49A8F87BFFD2F321FB5296D7D67FF847" box="[1269,1350,1922,1944]" country="Colombia" name="Boyaca" pageId="33" pageNumber="34">Boyaca</collectingRegion>
*,
<collectingRegion id="49A8F87BFFD2F321FAC696D6D6A1F847" box="[1377,1432,1923,1944]" country="Colombia" name="Meta" pageId="33" pageNumber="34">Meta</collectingRegion>
,
<collectingRegion id="49A8F87BFFD2F321FF6096F4D20AF868" box="[199,307,1953,1975]" country="Colombia" name="Putumayo" pageId="33" pageNumber="34">Putumayo</collectingRegion>
*,
<collectingRegion id="49A8F87BFFD2F321FEE096F4D2AEF868" box="[327,407,1953,1975]" country="Colombia" name="Vaupes" pageId="33" pageNumber="34">Vaupés</collectingRegion>
*);
<collectingCountry id="F37B7609FFD2F321FE1396F4D11CF868" box="[436,549,1953,1975]" name="Costa Rica" pageId="33" pageNumber="34">Costa Rica</collectingCountry>
* (
<collectingRegion id="49A8F87BFFD2F321FD9996F4D1A5F868" box="[574,668,1953,1975]" country="Costa Rica" name="San Jose" pageId="33" pageNumber="34">San José</collectingRegion>
); French
<collectingCountry id="F37B7609FFD2F321FD5C96F4D073F868" box="[763,842,1953,1975]" name="Guyana" pageId="33" pageNumber="34">Guyana</collectingCountry>
(St. Laurent du Maroni);
<collectingRegion id="49A8F87BFFD2F321FBF496F4D79FF868" box="[1107,1190,1953,1975]" country="Panama" name="Panama" pageId="33" pageNumber="34">
<collectingCountry id="F37B7609FFD2F321FBF496F4D79FF868" box="[1107,1190,1953,1975]" name="Panama" pageId="33" pageNumber="34">Panama</collectingCountry>
</collectingRegion>
* (
<collectingRegion id="49A8F87BFFD2F321FB1A96F4D62EF868" box="[1213,1303,1953,1975]" country="Panama" name="Chiriqui" pageId="33" pageNumber="34">Chiriqui</collectingRegion>
);
<collectingRegion id="49A8F87BFFD2F321FA8D96F4D6B7F868" box="[1322,1422,1953,1975]" country="Paraguay" name="Asuncion" pageId="33" pageNumber="34">
<collectingCountry id="F37B7609FFD2F321FA8D96F4D6B7F868" box="[1322,1422,1953,1975]" name="Paraguay" pageId="33" pageNumber="34">Paraguay</collectingCountry>
</collectingRegion>
*;
<collectingCountry id="F37B7609FFD2F321FF609694D3CCF809" box="[199,245,1985,2006]" name="Peru" pageId="33" pageNumber="34">Peru</collectingCountry>
(Chamchamayo,
<collectingRegion id="49A8F87BFFD2F321FE0F9695D2C8F809" box="[424,497,1984,2006]" country="Peru" name="Loreto" pageId="33" pageNumber="34">Loreto</collectingRegion>
*,
<collectingRegion id="49A8F87BFFD2F321FDA29695D163F809" box="[517,602,1984,2006]" country="Peru" name="Ucayali" pageId="33" pageNumber="34">Ucayali</collectingRegion>
*);
<collectingCountry id="F37B7609FFD2F321FDD19695D1E4F809" box="[630,733,1984,2006]" name="Suriname" pageId="33" pageNumber="34">Suriname</collectingCountry>
*.
</paragraph>
<paragraph id="8BD33699FFD2F321FF60968BD2CAF82B" blockId="33.[151,1436,1217,2036]" box="[199,499,2014,2036]" pageId="33" pageNumber="34">
<emphasis id="B918EA8BFFD2F321FF60968BD3EDF82B" bold="true" box="[199,212,2014,2036]" pageId="33" pageNumber="34">*</emphasis>
Records from the literature.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD1F322FF3091C2D2AFFCE7" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BD33699FFD1F322FF3091C2D784FD5B" blockId="34.[151,1437,151,2013]" pageId="34" pageNumber="35">
<emphasis id="B918EA8BFFD1F322FF3091C2D2EEFF6F" bold="true" box="[151,471,151,176]" pageId="34" pageNumber="35">Comments and diagnosis.</emphasis>
<bibRefCitation id="EFFD4B68FFD1F322FE4291C2D1D6FF6F" box="[485,751,151,176]" pageId="34" pageNumber="35" refId="ref43151" refString="Giordani Soika, A. (1978) Revisione degli Eumenidi neotropicali appartenenti ai generi Eumenes Latr., Omicron (Sauss.), Pararhaphidoglossa Schulth. ed afinni. Bolletino della Museo Civico di Storia Naturale di Uenezia, 29, 1 - 420." type="journal article">Giordani Soika (1978)</bibRefCitation>
treated
<taxonomicName id="4C6C4D1AFFD1F322FCFF91CDD086FF6F" box="[856,959,152,176]" class="Insecta" genus="Zeteumenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD1F322FCFF91CDD086FF6F" box="[856,959,152,176]" italics="true" pageId="34" pageNumber="35">filiformis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D1AFFD1F322FBA491CDD79CFF6F" authorityName="Fox" authorityYear="1899" box="[1027,1189,152,176]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="rufomaculatus">
<emphasis id="B918EA8BFFD1F322FBA491CDD79CFF6F" box="[1027,1189,152,176]" italics="true" pageId="34" pageNumber="35">rufomaculatus</emphasis>
</taxonomicName>
as subspecies of
<taxonomicName id="4C6C4D1AFFD1F322FA2391CCD230FF0B" authorityName="de Saussure" authorityYear="1852" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B918EA8BFFD1F322FA2391CCD230FF0B" italics="true" pageId="34" pageNumber="35">E. versicolor</emphasis>
</taxonomicName>
that differed basically by coloration. Even though he illustrated some differences in the digitus (p. 40 and figs 35, 37, 39), he judged them as not appreciable. Later,
<bibRefCitation id="EFFD4B68FFD1F322FC8A91B5D76AFF27" box="[813,1107,223,248]" pageId="34" pageNumber="35" refId="ref42766" refString="Carpenter, J. M. &amp; van der Vecht, J. (1991) A study of the Vespidae described by William J. Fox (Insecta: Hymenoptera), with assessment of taxonomic implications. Annals of the Carnegie Museum of Natural History, 60 (3), 211 - 241." type="journal article">Carpenter &amp; Vecth (1991)</bibRefCitation>
studied the specimenss from CMNH and proposed to treat them as different species based on morphological characters. We confirm this treatment herein and sustain
<taxonomicName id="4C6C4D1AFFD1F322FE54907CD146FE9F" box="[499,639,296,320]" class="Insecta" genus="Zeteumenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD1F322FE54907CD146FE9F" box="[499,639,296,320]" italics="true" pageId="34" pageNumber="35">E. filiformis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D1AFFD1F322FD33907CD062FE9F" authorityName="Fox" authorityYear="1899" box="[660,859,296,320]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="rufomaculatus">
<emphasis id="B918EA8BFFD1F322FD33907CD062FE9F" box="[660,859,296,320]" italics="true" pageId="34" pageNumber="35">E. rufomaculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D1AFFD1F322FC39907CD70FFE9F" authorityName="de Saussure" authorityYear="1852" box="[926,1078,296,320]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B918EA8BFFD1F322FC39907CD70FFE9F" box="[926,1078,296,320]" italics="true" pageId="34" pageNumber="35">E. versicolor</emphasis>
</taxonomicName>
as separate species.
<taxonomicName id="4C6C4D1AFFD1F322FA90907CD3C7FEBB" authorityName="de Saussure" authorityYear="1855" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD1F322FA90907CD3C7FEBB" italics="true" pageId="34" pageNumber="35">Eumenes filiformis</emphasis>
</taxonomicName>
has the clypeus somewhat narrow (fig. 74), with the punctation coarse and evident on the basal half and reduced on the apical half, while
<taxonomicName id="4C6C4D1AFFD1F322FDBA9024D1D8FE57" authorityName="Fox" authorityYear="1899" box="[541,737,368,392]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="rufomaculatus">
<emphasis id="B918EA8BFFD1F322FDBA9024D1D8FE57" box="[541,737,368,392]" italics="true" pageId="34" pageNumber="35">E. rufomaculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D1AFFD1F322FCBA9024D088FE57" authorityName="de Saussure" authorityYear="1852" box="[797,945,368,392]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B918EA8BFFD1F322FCBA9024D088FE57" box="[797,945,368,392]" italics="true" pageId="34" pageNumber="35">E. versicolor</emphasis>
</taxonomicName>
have the clypeus broader, with punctation uniformly distributed (fig. 75, 76). Besides that, the humeral region is less projecting, best observed in oblique view. We examined the male genitalia from the three species and found appreciable characters that confirm them as separate taxa: the aedeagus of each taxon is similar, but the ventral margin in
<taxonomicName id="4C6C4D1AFFD1F322FC5B9088D7BBFE2B" authorityName="de Saussure" authorityYear="1855" box="[1020,1154,476,500]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD1F322FC5B9088D7BBFE2B" box="[1020,1154,476,500]" italics="true" pageId="34" pageNumber="35">E. filiformis</emphasis>
</taxonomicName>
is less projecting than
<taxonomicName id="4C6C4D1AFFD1F322FA239088D200FDC7" authorityName="Fox" authorityYear="1899" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="rufomaculatus">
<emphasis id="B918EA8BFFD1F322FA239088D200FDC7" italics="true" pageId="34" pageNumber="35">E. rufomaculatus</emphasis>
</taxonomicName>
. The digitus of
<taxonomicName id="4C6C4D1AFFD1F322FE489354D14FFDC7" authorityName="de Saussure" authorityYear="1855" box="[495,630,512,536]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD1F322FE489354D14FFDC7" box="[495,630,512,536]" italics="true" pageId="34" pageNumber="35">E. filiformis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D1AFFD1F322FD099354D006FDC7" authorityName="de Saussure" authorityYear="1852" box="[686,831,512,536]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="B918EA8BFFD1F322FD099354D006FDC7" box="[686,831,512,536]" italics="true" pageId="34" pageNumber="35">E. versicolor</emphasis>
</taxonomicName>
are flattened, but the dorsal margin in
<taxonomicName id="4C6C4D1AFFD1F322FB5D9354D6BBFDC7" authorityName="de Saussure" authorityYear="1855" box="[1274,1410,512,536]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD1F322FB5D9354D6BBFDC7" box="[1274,1410,512,536]" italics="true" pageId="34" pageNumber="35">E. filiformis</emphasis>
</taxonomicName>
is angled with evident and longer bristles, mostly on the ventral margin (
<figureCitation id="13572A1CFFD1F322FC0E9371D732FDE3" box="[937,1035,548,573]" captionStart="FIGURES 110–125" captionStartId="51.[151,264,1899,1921]" captionTargetBox="[163,1425,209,1857]" captionTargetId="figure@51.[150,1437,193,1878]" captionTargetPageId="51" captionText="FIGURES 110–125. Male genitalia: digitus. 110) Eumenes americanus; 111) E. bequaerti; 112) E. bollii; 113) E. consobrinus; 114) E. coyotae; 115) E. crucifera; 116) E. cubensis; 117) E. flavitinctus; 118) E. fraternus; 119) E. pictus; 120) E. sculleni; 121) E. smithii; 122) E. verticalis; 123) E. filiformis; 124) E. rufomaculatus; 125) E. versicolor." httpUri="https://zenodo.org/record/1458324/files/figure.png" pageId="34" pageNumber="35">Fig. 123</figureCitation>
), while in the other two species it is rounded with slightly shorter and less evident bristles (fig. 124, 125). The digitus of
<taxonomicName id="4C6C4D1AFFD1F322FBD5931CD60FFDBF" authorityName="Fox" authorityYear="1899" box="[1138,1334,584,608]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="rufomaculatus">
<emphasis id="B918EA8BFFD1F322FBD5931CD60FFDBF" box="[1138,1334,584,608]" italics="true" pageId="34" pageNumber="35">E. rufomaculatus</emphasis>
</taxonomicName>
is broad laterally, with the dorsal margin angled and with less evident bristles compared to
<taxonomicName id="4C6C4D1AFFD1F322FB979338D78FFD5B" authorityName="de Saussure" authorityYear="1855" box="[1072,1206,620,644]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD1F322FB979338D78FFD5B" box="[1072,1206,620,644]" italics="true" pageId="34" pageNumber="35">E. filiformis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD33699FFD1F322FF6093DAD7BAFD2F" blockId="34.[151,1437,151,2013]" pageId="34" pageNumber="35">
Besides the features described above,
<taxonomicName id="4C6C4D1AFFD1F322FDD293C4D1C5FD77" authorityName="de Saussure" authorityYear="1855" box="[629,764,656,680]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD1F322FDD293C4D1C5FD77" box="[629,764,656,680]" italics="true" pageId="34" pageNumber="35">E. filiformis</emphasis>
</taxonomicName>
may be identified by having the body predominantly black (
<figureCitation id="13572A1CFFD1F322FF3893E1D3CEFD13" box="[159,247,692,717]" captionStart="FIGURES 13–26" captionStartId="42.[151,264,1649,1671]" captionTargetBox="[152,1436,193,1628]" captionTargetId="figure@42.[151,1436,193,1628]" captionTargetPageId="42" captionText="FIGURES 13–26. Eumenes species, lateral view. 13) E. cubensis Cresson, 1865 (female); 14) E. flavitinctus Bohart, 1950 (Holotype, female); 15) E. fervens Say, 1824 (Lectotype, female); 16) E. pictus (Smith, 1857) (female); 17) E. sculleni Bohart, 1950 (Holotype, male); 18) E. smithii de Saussure, 1852 (female); 19) E. belfragei Cresson, 1872 (Lectotype, female); 20) E. verticalis Say, 1824 (female); 21) Eumenes verticalis coloradensis Cresson, 1875 (lectotype, female); 22) E. verticalis neoboreus Bequaert, 1944 (Holotype, male); 23) E. verticalis tricinctus Isely, 1917 (Holotype, female); 24) E. filiformis de Saussure, 1855 (famele); 25) E. rufomaculata Fox, 1899 (Lectotype, female); 26) Eumenes versicolor de Saussure, 1852 (Lectotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458306/files/figure.png" pageId="34" pageNumber="35">Fig. 24</figureCitation>
), including the pterostigma, costal, medial, and submedial cells of the fore wings; T1 very long and narrow; and the apical region from the ventral surface of T1 ventrally projecting (fig. 61).
</paragraph>
<paragraph id="8BD33699FFD1F322FF6093A8D2AFFCE7" blockId="34.[151,1437,151,2013]" pageId="34" pageNumber="35">
We did not examine the
<typeStatus id="54D7883BFFD1F322FE4B93A8D125FCCA" box="[492,540,765,789]" pageId="34" pageNumber="35">type</typeStatus>
specimen of
<taxonomicName id="4C6C4D1AFFD1F322FD1893A8D070FCCB" authorityName="de Saussure" authorityYear="1855" box="[703,841,764,788]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD1F322FD1893A8D070FCCB" box="[703,841,764,788]" italics="true" pageId="34" pageNumber="35">E. filiformis</emphasis>
</taxonomicName>
but we based the redescription on
<specimenCount id="9D6AFD10FFD1F322FB4B93A9D643FCCA" box="[1260,1402,764,789]" pageId="34" pageNumber="35" type="generic">8 specimens</specimenCount>
(
<specimenCount id="9D6AFD10FFD1F322FA2A93A9D3D7FCE7" pageId="34" pageNumber="35" type="female">7 females</specimenCount>
and
<specimenCount id="9D6AFD10FFD1F322FE839275D2B3FCE7" box="[292,394,799,824]" pageId="34" pageNumber="35" type="female">1 female</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD1F323FF609211D7E9FE57" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="description">
<paragraph id="8BD33699FFD1F322FF609211D75FFBEA" blockId="34.[151,1437,151,2013]" pageId="34" pageNumber="35">
<emphasis id="B918EA8BFFD1F322FF609211D227FC82" bold="true" box="[199,286,836,861]" pageId="34" pageNumber="35">Female</emphasis>
(
<figureCitation id="13572A1CFFD1F322FE899211D2B9FC83" box="[302,384,836,861]" captionStart="FIGURES 13–26" captionStartId="42.[151,264,1649,1671]" captionTargetBox="[152,1436,193,1628]" captionTargetId="figure@42.[151,1436,193,1628]" captionTargetPageId="42" captionText="FIGURES 13–26. Eumenes species, lateral view. 13) E. cubensis Cresson, 1865 (female); 14) E. flavitinctus Bohart, 1950 (Holotype, female); 15) E. fervens Say, 1824 (Lectotype, female); 16) E. pictus (Smith, 1857) (female); 17) E. sculleni Bohart, 1950 (Holotype, male); 18) E. smithii de Saussure, 1852 (female); 19) E. belfragei Cresson, 1872 (Lectotype, female); 20) E. verticalis Say, 1824 (female); 21) Eumenes verticalis coloradensis Cresson, 1875 (lectotype, female); 22) E. verticalis neoboreus Bequaert, 1944 (Holotype, male); 23) E. verticalis tricinctus Isely, 1917 (Holotype, female); 24) E. filiformis de Saussure, 1855 (famele); 25) E. rufomaculata Fox, 1899 (Lectotype, female); 26) Eumenes versicolor de Saussure, 1852 (Lectotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458306/files/figure.png" pageId="34" pageNumber="35">Fig. 24</figureCitation>
).
<emphasis id="B918EA8BFFD1F322FE309211D113FC82" bold="true" box="[407,554,836,861]" pageId="34" pageNumber="35">Color, head:</emphasis>
Black, with yellow marks as follows: longitudinal broad band on each side of clypeus; a band on inter-antennal region; narrow stripe on upper half of gena, adjacent to the compound eye. Part of pedicel and flagellum yellow-orange.
<emphasis id="B918EA8BFFD1F322FDED92D9D1E8FC7A" bold="true" box="[586,721,908,933]" pageId="34" pageNumber="35">Mesosoma:</emphasis>
Black, with yellow marks as follows: small mark on humeral region of pronotum; a narrow stripe along the posterior edge of pronotum; parategula. Small marks on coxae, distal region of femora and longitudinal bands on tibiae.
<emphasis id="B918EA8BFFD1F322FD6B9281D068FC32" bold="true" box="[716,849,980,1005]" pageId="34" pageNumber="35">Metasoma:</emphasis>
Black, with yellow marks: a narrow transverse band on apex of T1, T2, and S2–S5. Apex of T3–T6 and S6 with brownish bands on apex.
<emphasis id="B918EA8BFFD1F322FBF892A2D78DFBCF" bold="true" box="[1119,1204,1015,1040]" pageId="34" pageNumber="35">Wings:</emphasis>
Apical half hyaline; basal half darker, mainly costal, medial, submedial cells and pterostigma. Veins darker.
</paragraph>
<paragraph id="8BD33699FFD1F322FF60956AD70FFA8B" blockId="34.[151,1437,151,2013]" pageId="34" pageNumber="35">
<emphasis id="B918EA8BFFD1F322FF60956AD29AFB87" bold="true" box="[199,419,1087,1112]" pageId="34" pageNumber="35">Pubescence, head:</emphasis>
Clypeus with short whitish (better observed in oblique view) pubescence. Frons, vertex and occipital region with long and golden pubescence, while on gena it is short (shorter than on clypeus). Scape with golden pubescence as short as on clypeus.
<emphasis id="B918EA8BFFD1F322FD0B95D2D00AFB7F" bold="true" box="[684,819,1159,1184]" pageId="34" pageNumber="35">Mesosoma:</emphasis>
covered with golden pubescence. Short pubescence on pronotum, mesoscutum and lateral surface of mesosoma and longer (as long as on frons and vertex) on scutellum, metanotum and posterior face of propodeum. Trochanter and basal half of femur of the fore leg with erect golden bristles. External surface of tibiae also with golden and erect bristles, longer than on fore femur.
<emphasis id="B918EA8BFFD1F322FAB195A1D6A5FAD2" bold="true" box="[1302,1436,1268,1293]" pageId="34" pageNumber="35">Metasoma:</emphasis>
Covered with golden pubescence. Short pubescence on dorsal face of T1 and other segments. Ventral surface of T1 with scattered long bristles. Apex of the other segments with long bristles on apex.
</paragraph>
<paragraph id="8BD33699FFD1F322FF60940AD6A4FA1F" blockId="34.[151,1437,151,2013]" pageId="34" pageNumber="35">
<emphasis id="B918EA8BFFD1F322FF60940AD11CFAA7" bold="true" box="[199,549,1375,1400]" pageId="34" pageNumber="35">Surface of integument, head:</emphasis>
Clypeus with weak punctation, more evident on basal half; frons with coarse and moderately dense punctation; reduced punctation behind the ocelli and on gena.
<emphasis id="B918EA8BFFD1F322FBD794D1D7CEFA42" bold="true" box="[1136,1271,1412,1437]" pageId="34" pageNumber="35">Mesosoma:</emphasis>
Covered with coarse and moderately dense punctation as on frons, except on basal half of mesepisternum where it is sparser.
</paragraph>
<paragraph id="8BD33699FFD1F322FF309499D2F2F98F" blockId="34.[151,1437,151,2013]" pageId="34" pageNumber="35">
<emphasis id="B918EA8BFFD1F322FF309499D224FA3A" bold="true" box="[151,285,1484,1509]" pageId="34" pageNumber="35">Metasoma:</emphasis>
Dorsal surface of T1 with coarser and moderately dense punctation, becoming denser on apex. Lateral surface of T1 with scattered punctures, while ventral surface without evident punctation. Dorsal surface of T2 with coarse and dense punctation, becoming slightly weaker and sparser towards lateral surface. T3–T6 and S2–S6 without evident punctation.
</paragraph>
<paragraph id="8BD33699FFD1F322FF609709D611F866" blockId="34.[151,1437,151,2013]" pageId="34" pageNumber="35">
<emphasis id="B918EA8BFFD1F322FF609709D2B3F9AA" bold="true" box="[199,394,1628,1653]" pageId="34" pageNumber="35">Structure, head:</emphasis>
Clypeus convex, longer than wide, with the apex concave forming two rounded projections with weak carinae on apex (
<figureCitation id="13572A1CFFD1F322FE77972AD11CF947" box="[464,549,1663,1688]" captionStart="FIGURES 66–76" captionStartId="47.[151,264,1772,1794]" captionTargetBox="[151,1436,193,1750]" captionTargetId="figure@47.[151,1436,193,1750]" captionTargetPageId="47" captionText="FIGURES 66–76. Clypeus. 66) Eumenes americanus (female); 67) E. smithii (female); 68) E. crucifera (female); 69) E. crucifera (male); 70) E. verticalis (female); 71) E. fraternus (female); 72) E. fraternus (male); 73) E. sculleni (female); 74) E. filiformis (female); 75) E. rufomaculatus (female); 76) E. versicolor (female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458316/files/figure.png" pageId="34" pageNumber="35">Fig. 74</figureCitation>
); inter-antennal wider than the antennal socket, with a center tubercle on center region; lateral ocelli closer to the compound eye than to each other; occipital carina angled on middle region of gena and close to the compound eyes next to the mandible.
<emphasis id="B918EA8BFFD1F322FCE89792D0EFF93F" bold="true" box="[847,982,1735,1760]" pageId="34" pageNumber="35">Mesosoma:</emphasis>
pronotal fovea present; pronotal carina present dorsally and and more developed laterally; humeral region weakly projecting; mesepimeron slightly elevated regarding the mesepisternum; posterior projection of tegula weakle developed and rounded; parategula lamelliform; posterior face of propodeum strongly concave on basal half. Basal region of the external surface of fore coxa with a short and weak carina.
<emphasis id="B918EA8BFFD1F322FDF19602D1E5F8AF" bold="true" box="[598,732,1879,1904]" pageId="34" pageNumber="35">Metasoma:</emphasis>
T1 very long, more than five times longer than wide (
<figureCitation id="13572A1CFFD1F322FA9D9602D6B6F8AF" box="[1338,1423,1879,1904]" captionStart="FIGURES 27–42" captionStartId="43.[151,264,1902,1924]" captionTargetBox="[151,1435,193,1881]" captionTargetId="figure@43.[151,1436,193,1881]" captionTargetPageId="43" captionText="FIGURES 27–42. First tergum, dorsal view. 27) Eumenes americanus; 28) E. bequaerti; 29) E. bollii; 30) E. consobrinus; 31) E. coyotae; 32) E. crucifera; 33) E. cubensis; 34) E. flavitinctus; 35) E. fraternus; 36) E. pictus; 37) E. sculleni; 38) E. smithii; 39) E. verticalis; 40) E. filiformis; 41) E. rufomaculactus; 42) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458308/files/figure.png" pageId="34" pageNumber="35">Fig. 40</figureCitation>
); T2 longer than wide, with the lateral margins weakly concave (
<figureCitation id="13572A1CFFD1F322FCCC9629D0F8F84A" box="[875,961,1916,1941]" captionStart="FIGURES 43–58" captionStartId="44.[151,264,1946,1968]" captionTargetBox="[202,1388,195,1934]" captionTargetId="figure@44.[201,1388,193,1934]" captionTargetPageId="44" captionText="FIGURES 43–58. Second tergum, dorsal view. 43) Eumenes americanus; 44) E. bequaerti; 45) E. bollii; 46) E. consobrinus; 47) E. coyotae; 48) E. crucifera; 49) E. cubensis; 50) E. flavitinctus; 51) E. fraternus; 52) E. pictus; 53) E. sculleni; 54) E. smithii; 55) E. verticalis; 56) E. filiformis; 57) E. rufomaculactus; 58) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458310/files/figure.png" pageId="34" pageNumber="35">Fig. 56</figureCitation>
); apical lamella on apex of T2, becoming reduced towards lateral margins. S2 with a weak longitudinal elevation, better observed in oblique view.
</paragraph>
<paragraph id="8BD33699FFD1F323FF609691D7A7FF0A" blockId="34.[151,1437,151,2013]" lastBlockId="35.[151,1437,151,392]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<emphasis id="B918EA8BFFD1F322FF609691D234F802" bold="true" box="[199,269,1988,2013]" pageId="34" pageNumber="35">Male:</emphasis>
Clypeus narrower than females; F11 long, without longitudinal carina, apex pointed surpassing the apical edge of F8 (
<figureCitation id="13572A1CFFD0F323FED191C2D2F7FF6F" box="[374,462,151,176]" captionStart="FIGURES 77–92" captionStartId="48.[151,264,1946,1968]" captionTargetBox="[151,1436,193,1925]" captionTargetId="figure@48.[151,1436,193,1925]" captionTargetPageId="48" captionText="FIGURES 77–92. First tergum, dorsal view. 77) Eumenes americanus; 78) E. bequaerti; 79) E. bollii; 80) E. consobrinus; 81) E. coyotae; 82) E. crucifera; 83) E. cubensis; 84) E. flavitinctus; 85) E. fraternus; 86) E. pictus; 87) E. sculleni; 88) E. smithii; 89) E. verticalis; 90) E. filiformis; 91) E. rufomaculactus; 92) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458318/files/figure.png" pageId="35" pageNumber="36">Fig. 90</figureCitation>
). Ventral surface without microscopis erect bristles. S7 flattened apically with erect bristles on apex. Color, pubescence, surface of integument and other structure as in females.
</paragraph>
<paragraph id="8BD33699FFD0F323FF60918AD0A4FEBA" blockId="35.[151,1437,151,392]" pageId="35" pageNumber="36">
<emphasis id="B918EA8BFFD0F323FF60918AD243FF27" bold="true" box="[199,378,223,248]" pageId="35" pageNumber="36">Male genitalia:</emphasis>
Aedeagus as in figure 106 a, b. Paramere (
<figureCitation id="13572A1CFFD0F323FCF1918AD0F9FF27" box="[854,960,223,248]" captionStart="FIGURES 102–109" captionStartId="50.[151,264,1926,1948]" captionTargetBox="[151,1436,193,1903]" captionTargetId="figure@50.[151,1436,193,1904]" captionTargetPageId="50" captionText="FIGURES 102–109. Male genitalia: a, b) aedeagus (lateral and ventral view, respectively); c) paramere, internal view. 102) E. pictus; 103) E. sculleni; 104) E. smithii; 105) E. verticalis; 106) E. filiformis; 107) E. rufomaculatus; 108) E. versicolor; 109) General scheme of the apical projection of the aedeagus: a) weakly concave/truncate; b) strongly concave. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458322/files/figure.png" pageId="35" pageNumber="36">Fig. 106c</figureCitation>
) with bristles on middle area of gonostyle; digitus broad in basal half, becoming slender toward apex (fig. 123), with short and evident bristles on the base and slightly shorter toward apex; cuspis with short and erect bristles; volsella with moderately long and scattered bristles (longer then on digitus); distal lobe truncate, without bristles.
</paragraph>
<paragraph id="8BD33699FFD0F323FF60903AD7E9FE57" blockId="35.[151,1437,151,392]" box="[199,1232,367,392]" pageId="35" pageNumber="36">
<emphasis id="B918EA8BFFD0F323FF60903AD278FE57" bold="true" box="[199,321,367,392]" pageId="35" pageNumber="36">Variation:</emphasis>
Yellow marks on clypeus reduced; band on apex of T1 and T
<date id="FFD21059FFD0F323FC559025D70EFE57" box="[1010,1079,368,392]" pageId="35" pageNumber="36">2 may</date>
be brownish.
</paragraph>
</subSubSection>
</treatment>
</document>
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