310 lines
58 KiB
XML
310 lines
58 KiB
XML
<document id="2B70EDFC46D3B6E368FB06F32068576F" ID-DOI="10.11646/zootaxa.5476.1.26" ID-ISSN="1175-5326" ID-Zenodo-Dep="12681830" ID-ZooBank="84C66278-7194-4F3E-8145-3918E1659289" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720375444750" checkinUser="plazi" docAuthor="Ng, Peter K. L., Prema, Mani & Ravichandran, Samuthirapandian" docDate="2024" docId="03C54B01FF8C375DFFF1FC1595D9FD15" docLanguage="en" docName="zootaxa.5476.1.26.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.26" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acanthophrys keeae Ng, Prema & Ravichandran, 2024, n. sp." docType="treatment" docVersion="3" lastPageNumber="329" masterDocId="FFFC3379FF8E3759FF66FF8A9728FFBC" masterDocTitle="Descriptions of new and rare crabs associated with coral rubble and sponges from Kanyakumari, southeastern Arabian Sea, southern India (Decapoda: Brachyura: Epialtidae and Pilumnidae)" masterLastPageNumber="341" masterPageNumber="325" pageNumber="327" updateTime="1720379182836" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D7E0D214CAD843B9658E9EDFF870F4C7">Descriptions of new and rare crabs associated with coral rubble and sponges from Kanyakumari, southeastern Arabian Sea, southern India (Decapoda: Brachyura: Epialtidae and Pilumnidae)</mods:title>
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<mods:name id="16FA49EC9076D840F0CAE854DC016475" type="personal">
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<mods:roleTerm id="EF0E378B50018F029A866F7F49569430">Author</mods:roleTerm>
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<mods:namePart id="15F3659BEA1E047E4A922318A6559FC0">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="A906325088CD6F9715C8AC8586426F75">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore.</mods:affiliation>
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<mods:name id="D6805090447F7BD604A57A33E022F35F" type="personal">
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<mods:roleTerm id="F4A655E8A4851D315C8967A01EB9B62B">Author</mods:roleTerm>
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<mods:namePart id="6721BD09A1D0CC91C28881A6B4ACA6DD">Prema, Mani</mods:namePart>
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<mods:affiliation id="906E17A897FD9EE7B920C4B717DFF24F">Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Annamalai University, Parangipettai- 608 502, India.</mods:affiliation>
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<mods:namePart id="009C3E62B6B1BBEBBC7782E4E624E261">Ravichandran, Samuthirapandian</mods:namePart>
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<mods:affiliation id="17D9D7179F9E4EECAD756828C048FF12">Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Annamalai University, Parangipettai- 608 502, India. & Department of Zoology, Government Arts & Science College, Nagercoil - 629 004, India.</mods:affiliation>
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<mods:title id="5504AAFB934376608BBAE8DCC7F1E433">Zootaxa</mods:title>
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<mods:date id="29D629D495D9691F065A0E5CB8560188">2024</mods:date>
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<mods:number id="6E91F138899116479714DB5C30F07BAC">2024-07-04</mods:number>
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<mods:number id="0B3BCA4A406461ED29C264EDA380551E">5476</mods:number>
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<treatment id="03C54B01FF8C375DFFF1FC1595D9FD15" ID-DOI="http://doi.org/10.5281/zenodo.12681396" ID-Zenodo-Dep="12681396" LSID="urn:lsid:plazi:treatment:03C54B01FF8C375DFFF1FC1595D9FD15" httpUri="http://treatment.plazi.org/id/03C54B01FF8C375DFFF1FC1595D9FD15" lastPageId="4" lastPageNumber="329" pageId="2" pageNumber="327">
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<subSubSection id="C376A99CFF8C375BFFF1FC1596FEFC06" box="[151,470,927,954]" pageId="2" pageNumber="327" type="nomenclature">
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<paragraph id="8BD3FA17FF8C375BFFF1FC1596FEFC06" blockId="2.[151,470,927,990]" box="[151,470,927,954]" pageId="2" pageNumber="327">
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<heading id="D09B4D7BFF8C375BFFF1FC1596FEFC06" bold="true" box="[151,470,927,954]" fontSize="11" level="1" pageId="2" pageNumber="327" reason="1">
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<emphasis id="B9182605FF8C375BFFF1FC1596FEFC06" bold="true" box="[151,470,927,954]" pageId="2" pageNumber="327">
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<taxonomicName id="4C6C8194FF8C375BFFF1FC1596A4FC06" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[151,396,927,954]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="keeae" status="sp. nov.">
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<emphasis id="B9182605FF8C375BFFF1FC1596A4FC06" bold="true" box="[151,396,927,954]" italics="true" pageId="2" pageNumber="327">Acanthophrys keeae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22B9B7EFF8C375BFEF2FC2A96FEFC06" box="[404,470,928,954]" pageId="2" pageNumber="327" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C376A99CFF8C375BFFF1FC49963AFC62" box="[151,274,963,990]" pageId="2" pageNumber="327" type="description">
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<paragraph id="8BD3FA17FF8C375BFFF1FC49963AFC62" blockId="2.[151,470,927,990]" box="[151,274,963,990]" pageId="2" pageNumber="327">
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(
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<figureCitation id="1357E692FF8C375BFFF9FC499622FC62" box="[159,266,963,990]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="3.[151,250,1908,1932]" captionStartId-1="4.[151,250,1429,1453]" captionStartId-2="5.[151,250,1845,1869]" captionTargetBox-0="[191,1396,181,1885]" captionTargetBox-1="[151,1411,705,1405]" captionTargetBox-2="[165,1423,532,1812]" captionTargetId-0="figure-18@3.[191,1396,181,1885]" captionTargetId-1="figure-292@4.[151,1435,705,1405]" captionTargetId-2="figure-187@5.[151,1435,525,1822]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." captionText-1="FIGURE 3. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) anterior thoracic sternum and pleon; (B) left G1 (ventral view); (C) left G1 (subdorsal view); (D) left G1 (dorsal view); (E) left G2. B–E, photographed to same magnification." captionText-2="FIGURE 4. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) left third maxilliped; (B) outer view of left chela; (C) merus of left cheliped; (D) left P5; (E) pleon; (F) left G1 (ventral view); (G) left G1 (dorsal view); (H) distal part of left G1 (ventral view); (I) distal part of left G1 (ventral view); (J) distal part of left G1 (subdorsal view); (K), (L) distal part of left G1 (dorsal views); (M) left G2. A–E, H–L, denuded. Scales: A–E = 0.5 mm; F–M = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12681839" figureDoi-1="http://doi.org/10.5281/zenodo.12681841" figureDoi-2="http://doi.org/10.5281/zenodo.12681843" httpUri-0="https://zenodo.org/record/12681839/files/figure.png" httpUri-1="https://zenodo.org/record/12681841/files/figure.png" httpUri-2="https://zenodo.org/record/12681843/files/figure.png" pageId="2" pageNumber="327">Figs. 2–4</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C376A99CFF8C375BFFF1FB869417FBD2" pageId="2" pageNumber="327" type="materials_examined">
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<paragraph id="8BD3FA17FF8C375BFFF1FB869417FBD2" blockId="2.[151,1437,1035,2034]" pageId="2" pageNumber="327">
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<emphasis id="B9182605FF8C375BFFF1FB869669FB99" bold="true" box="[151,321,1035,1062]" pageId="2" pageNumber="327">Type material</emphasis>
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||
.
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<materialsCitation id="3B04F04AFF8C375BFE34FB819417FBD2" collectingDate="2021-11-05" collectorName="M. Prema" country="India" county="Southeastern Arabian Sea" elevation="55" latitude="8.091611" location="Manakudy" longLatPrecision="1" longitude="77.48414" municipality="Kanyakumari district" pageId="2" pageNumber="327" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="holotype">
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<emphasis id="B9182605FF8C375BFE34FB819691FB99" box="[338,441,1035,1061]" italics="true" pageId="2" pageNumber="327">
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<typeStatus id="54D744B5FF8C375BFE34FB819691FB99" box="[338,441,1035,1061]" pageId="2" pageNumber="327" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
: male (cw 7.6, cl 13.4, pcl
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||
<quantity id="4C9457F2FF8C375BFD99FB869474FB9A" box="[767,860,1036,1062]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="2" pageNumber="327" unit="mm" value="9.9">9.9 mm</quantity>
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), CASAU-CR-2021-1022,
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<location id="8EB3ACCCFF8C375BFBF5FB819223FB99" LSID="urn:lsid:plazi:treatment:03C54B01FF8C375DFFF1FC1595D9FD15:8EB3ACCCFF8C375BFBF5FB819223FB99" box="[1171,1291,1035,1061]" country="India" county="Southeastern Arabian Sea" latitude="8.091611" longLatPrecision="1" longitude="77.48414" municipality="Kanyakumari district" name="Manakudy" pageId="2" pageNumber="327" stateProvince="Tamil Nadu">Manakudy</location>
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||
fish landing centre,
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||
<collectingMunicipality id="6BB7606DFF8C375BFF8DFBA596F5FBF5" box="[235,477,1071,1097]" pageId="2" pageNumber="327">Kanyakumari district</collectingMunicipality>
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,
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<collectingRegion id="49A834F5FF8C375BFE8EFBA59559FBF5" box="[488,625,1071,1097]" country="India" name="Tamil Nadu" pageId="2" pageNumber="327">Tamil Nadu</collectingRegion>
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, southern
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<collectingCountry id="F37BBA87FF8C375BFD81FBA5940BFBF5" box="[743,803,1071,1097]" name="India" pageId="2" pageNumber="327">India</collectingCountry>
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,
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<collectingCounty id="62B2829BFF8C375BFC49FBA5937DFBF6" box="[815,1109,1071,1098]" pageId="2" pageNumber="327">Southeastern Arabian Sea</collectingCounty>
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,
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<geoCoordinate id="EE589CD0FF8C375BFB07FBBB93D9FBF0" box="[1121,1265,1073,1100]" degrees="8" direction="north" minutes="05" orientation="latitude" pageId="2" pageNumber="327" precision="1" seconds="29.8" value="8.091611">8°05′29.8′′N</geoCoordinate>
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,
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<geoCoordinate id="EE589CD0FF8C375BFB98FBBB92B0FBF0" box="[1278,1432,1073,1100]" degrees="77" direction="east" minutes="29" orientation="longitude" pageId="2" pageNumber="327" precision="1" seconds="02.9" value="77.48414">77°29′02.9′′E</geoCoordinate>
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,
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<quantity id="4C9457F2FF8C375BFFF1FBD997D4FBD2" box="[151,252,1107,1134]" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="2" pageNumber="327" unit="m" value="55.0" valueMax="60.0" valueMin="50.0">
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<elevation id="00411D24FF8C375BFFF1FBD997D4FBD2" box="[151,252,1107,1134]" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="2" pageNumber="327" unit="m" value="55.0" valueMax="60.0" valueMin="50.0">50–60 m</elevation>
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</quantity>
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, in coral rubble, coll.
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<collectorName id="26999FC1FF8C375BFE97FBDE954AFBD2" box="[497,610,1108,1134]" pageId="2" pageNumber="327">M. Prema</collectorName>
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,
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<date id="FFD2DCD7FF8C375BFD08FBD99413FBD2" box="[622,827,1107,1134]" pageId="2" pageNumber="327" value="2021-11-05">
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||
<collectingDate id="EF96253FFF8C375BFD08FBD99413FBD2" box="[622,827,1107,1134]" pageId="2" pageNumber="327" value="2021-11-05">5 November 2021</collectingDate>
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</date>
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||
.
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||
</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376A99CFF8C375BFFA1FBF29695FB0A" pageId="2" pageNumber="327" type="etymology">
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<paragraph id="8BD3FA17FF8C375BFFA1FBF29695FB0A" blockId="2.[151,1437,1035,2034]" pageId="2" pageNumber="327">
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||
<emphasis id="B9182605FF8C375BFFA1FBF29666FB2E" bold="true" box="[199,334,1144,1170]" pageId="2" pageNumber="327">Etymology.</emphasis>
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The species is dedicated to late Dr. Ng Ngan Kee from the National University of
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<collectingCountry id="F37BBA87FF8C375BFB8AFBFD9248FB2D" box="[1260,1376,1143,1169]" name="Singapore" pageId="2" pageNumber="327">Singapore</collectingCountry>
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, who passed away in
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<date id="FFD2DCD7FF8C375BFE2EFB119691FB0A" box="[328,441,1179,1206]" pageId="2" pageNumber="327" value="2022-07">July 2022</date>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376A99CFF8C375BFFA1FB3596C3F96D" pageId="2" pageNumber="327" type="diagnosis">
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<paragraph id="8BD3FA17FF8C375BFFA1FB3594A7F912" blockId="2.[151,1437,1035,2034]" pageId="2" pageNumber="327">
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||
<emphasis id="B9182605FF8C375BFFA1FB359612FB65" bold="true" box="[199,314,1215,1241]" pageId="2" pageNumber="327">Diagnosis</emphasis>
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. Carapace pyriform; dorsal surface covered by large low, rounded tubercles on anterior half of carapace, without sharp tubercles or spines; branchial region with low tubercles and granules and larger submedian tubercle; median gastric regions with 3 low tubercles, no spines; cardiac region elevated but without spine; intestinal region raised with median tubercle; hepatic region with small median tubercle (
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<figureCitation id="1357E692FF8C375BFB43FAA19389FAFA" box="[1061,1185,1323,1350]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="2" pageNumber="327">Fig. 2A–E</figureCitation>
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); pseudorostral spines long, divergent (
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<figureCitation id="1357E692FF8C375BFE28FAC596E0FAD6" box="[334,456,1359,1386]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="2" pageNumber="327">Fig. 2A–C</figureCitation>
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); proepistome distinctly deflexed, margins carinate (
|
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<figureCitation id="1357E692FF8C375BFB68FAC59340FAD6" box="[1038,1128,1359,1386]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="2" pageNumber="327">Fig. 2G</figureCitation>
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); supraorbital eave laterally expanded, distinctly bilobed, antorbital and preorbital lobes subequal in size and length, tips of lobes rounded (
|
||
<figureCitation id="1357E692FF8C375BFA09FAF997F7FA0E" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="2" pageNumber="327">Fig. 2B–D</figureCitation>
|
||
); postorbital spine proportionately longer, extending well beyond margin of supraorbital eave, inner margin with prominent median tubercle (
|
||
<figureCitation id="1357E692FF8C375BFD7BFA3195B6FA6A" box="[541,670,1467,1494]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="2" pageNumber="327">Fig. 2B, C</figureCitation>
|
||
); outer margin of basal antennal article sinuous with median part concave (
|
||
<figureCitation id="1357E692FF8C375BFE64FA559652FA46" box="[258,378,1503,1530]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="2" pageNumber="327">Fig. 2F–H</figureCitation>
|
||
); pterygostomial region with 2 large tubercles (
|
||
<figureCitation id="1357E692FF8C375BFCE1FA5594F3FA46" box="[903,987,1503,1530]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="2" pageNumber="327">Fig. 2F</figureCitation>
|
||
); exopod of third maxilliped wide (
|
||
<figureCitation id="1357E692FF8C375BFA02FA55979DF9A2" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="2" pageNumber="327">Figs. 2F</figureCitation>
|
||
,
|
||
<figureCitation id="1357E692FF8C375BFFA7F98E97CFF9A2" box="[193,231,1540,1566]" captionStart="FIGURE 4" captionStartId="5.[151,250,1845,1869]" captionTargetBox="[165,1423,532,1812]" captionTargetId="figure-187@5.[151,1435,525,1822]" captionTargetPageId="5" captionText="FIGURE 4. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) left third maxilliped; (B) outer view of left chela; (C) merus of left cheliped; (D) left P5; (E) pleon; (F) left G1 (ventral view); (G) left G1 (dorsal view); (H) distal part of left G1 (ventral view); (I) distal part of left G1 (ventral view); (J) distal part of left G1 (subdorsal view); (K), (L) distal part of left G1 (dorsal views); (M) left G2. A–E, H–L, denuded. Scales: A–E = 0.5 mm; F–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12681843" httpUri="https://zenodo.org/record/12681843/files/figure.png" pageId="2" pageNumber="327">4A</figureCitation>
|
||
); margins of merus of cheliped smooth, unarmed (
|
||
<figureCitation id="1357E692FF8C375BFC47F9899452F9A2" box="[801,890,1539,1566]" captionStart="FIGURE 4" captionStartId="5.[151,250,1845,1869]" captionTargetBox="[165,1423,532,1812]" captionTargetId="figure-187@5.[151,1435,525,1822]" captionTargetPageId="5" captionText="FIGURE 4. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) left third maxilliped; (B) outer view of left chela; (C) merus of left cheliped; (D) left P5; (E) pleon; (F) left G1 (ventral view); (G) left G1 (dorsal view); (H) distal part of left G1 (ventral view); (I) distal part of left G1 (ventral view); (J) distal part of left G1 (subdorsal view); (K), (L) distal part of left G1 (dorsal views); (M) left G2. A–E, H–L, denuded. Scales: A–E = 0.5 mm; F–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12681843" httpUri="https://zenodo.org/record/12681843/files/figure.png" pageId="2" pageNumber="327">Fig. 4C</figureCitation>
|
||
); ambulatory legs relatively short, flexor margin of dactylus with 8 or 9 short stout spines (
|
||
<figureCitation id="1357E692FF8C375BFD03F9AD9597F9FE" box="[613,703,1575,1602]" captionStart="FIGURE 4" captionStartId="5.[151,250,1845,1869]" captionTargetBox="[165,1423,532,1812]" captionTargetId="figure-187@5.[151,1435,525,1822]" captionTargetPageId="5" captionText="FIGURE 4. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) left third maxilliped; (B) outer view of left chela; (C) merus of left cheliped; (D) left P5; (E) pleon; (F) left G1 (ventral view); (G) left G1 (dorsal view); (H) distal part of left G1 (ventral view); (I) distal part of left G1 (ventral view); (J) distal part of left G1 (subdorsal view); (K), (L) distal part of left G1 (dorsal views); (M) left G2. A–E, H–L, denuded. Scales: A–E = 0.5 mm; F–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12681843" httpUri="https://zenodo.org/record/12681843/files/figure.png" pageId="2" pageNumber="327">Fig. 4D</figureCitation>
|
||
); male anterior thoracic sternum distinctly elongate, with sternites 3 and 4 constricted to form neck-like structure (
|
||
<figureCitation id="1357E692FF8C375BFDCCF9C1942BF9DA" box="[682,771,1611,1638]" captionStart="FIGURE 3" captionStartId="4.[151,250,1429,1453]" captionTargetBox="[151,1411,705,1405]" captionTargetId="figure-292@4.[151,1435,705,1405]" captionTargetPageId="4" captionText="FIGURE 3. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) anterior thoracic sternum and pleon; (B) left G1 (ventral view); (C) left G1 (subdorsal view); (D) left G1 (dorsal view); (E) left G2. B–E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681841" httpUri="https://zenodo.org/record/12681841/files/figure.png" pageId="2" pageNumber="327">Fig. 3A</figureCitation>
|
||
); male pleon distinctly triangular in shape, pleomeres 3 and 4 extremely wide, telson lingulate, elongate (
|
||
<figureCitation id="1357E692FF8C375BFDF3F9E595D1F936" box="[661,761,1647,1674]" captionStart="FIGURE 3" captionStartId="4.[151,250,1429,1453]" captionTargetBox="[151,1411,705,1405]" captionTargetId="figure-292@4.[151,1435,705,1405]" captionTargetPageId="4" captionText="FIGURE 3. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) anterior thoracic sternum and pleon; (B) left G1 (ventral view); (C) left G1 (subdorsal view); (D) left G1 (dorsal view); (E) left G2. B–E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681841" httpUri="https://zenodo.org/record/12681841/files/figure.png" pageId="2" pageNumber="327">Figs. 3A</figureCitation>
|
||
,
|
||
<figureCitation id="1357E692FF8C375BFC60F9FA9401F936" box="[774,809,1648,1674]" captionStart="FIGURE 4" captionStartId="5.[151,250,1845,1869]" captionTargetBox="[165,1423,532,1812]" captionTargetId="figure-187@5.[151,1435,525,1822]" captionTargetPageId="5" captionText="FIGURE 4. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) left third maxilliped; (B) outer view of left chela; (C) merus of left cheliped; (D) left P5; (E) pleon; (F) left G1 (ventral view); (G) left G1 (dorsal view); (H) distal part of left G1 (ventral view); (I) distal part of left G1 (ventral view); (J) distal part of left G1 (subdorsal view); (K), (L) distal part of left G1 (dorsal views); (M) left G2. A–E, H–L, denuded. Scales: A–E = 0.5 mm; F–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12681843" httpUri="https://zenodo.org/record/12681843/files/figure.png" pageId="2" pageNumber="327">4E</figureCitation>
|
||
); G1 relatively straight, slender, distal two-thirds of G1 almost straight with distal part clearly tapering (
|
||
<figureCitation id="1357E692FF8C375BFDD7F919941EF912" box="[689,822,1683,1710]" captionStart="FIGURE 3" captionStartId="4.[151,250,1429,1453]" captionTargetBox="[151,1411,705,1405]" captionTargetId="figure-292@4.[151,1435,705,1405]" captionTargetPageId="4" captionText="FIGURE 3. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) anterior thoracic sternum and pleon; (B) left G1 (ventral view); (C) left G1 (subdorsal view); (D) left G1 (dorsal view); (E) left G2. B–E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681841" httpUri="https://zenodo.org/record/12681841/files/figure.png" pageId="2" pageNumber="327">Figs. 3B–D</figureCitation>
|
||
,
|
||
<figureCitation id="1357E692FF8C375BFC27F91E94AAF912" box="[833,898,1684,1710]" captionStart="FIGURE 4" captionStartId="5.[151,250,1845,1869]" captionTargetBox="[165,1423,532,1812]" captionTargetId="figure-187@5.[151,1435,525,1822]" captionTargetPageId="5" captionText="FIGURE 4. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) left third maxilliped; (B) outer view of left chela; (C) merus of left cheliped; (D) left P5; (E) pleon; (F) left G1 (ventral view); (G) left G1 (dorsal view); (H) distal part of left G1 (ventral view); (I) distal part of left G1 (ventral view); (J) distal part of left G1 (subdorsal view); (K), (L) distal part of left G1 (dorsal views); (M) left G2. A–E, H–L, denuded. Scales: A–E = 0.5 mm; F–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12681843" httpUri="https://zenodo.org/record/12681843/files/figure.png" pageId="2" pageNumber="327">4F–L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD3FA17FF8C375BFFA1F93D96C3F96D" blockId="2.[151,1437,1035,2034]" box="[199,491,1719,1746]" pageId="2" pageNumber="327">
|
||
<emphasis id="B9182605FF8C375BFFA1F93D9670F96D" bold="true" box="[199,344,1719,1745]" pageId="2" pageNumber="327">Colouration</emphasis>
|
||
. Not known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376A99CFF8C375DFFA1F95695BBFD39" lastPageId="4" lastPageNumber="329" pageId="2" pageNumber="327" type="discussion">
|
||
<paragraph id="8BD3FA17FF8C375DFFA1F95695BBFD39" blockId="2.[151,1437,1035,2034]" lastBlockId="4.[151,1437,151,681]" lastPageId="4" lastPageNumber="329" pageId="2" pageNumber="327">
|
||
<emphasis id="B9182605FF8C375BFFA1F956961CF94A" bold="true" box="[199,308,1756,1782]" pageId="2" pageNumber="327">Remarks</emphasis>
|
||
. The most distinctive character of
|
||
<taxonomicName id="4C6C8194FF8C375BFDA3F9519481F949" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[709,937,1755,1781]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="keeae" status="sp. nov.">
|
||
<emphasis id="B9182605FF8C375BFDA3F9519481F949" box="[709,937,1755,1781]" italics="true" pageId="2" pageNumber="327">Acanthophrys keeae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF8C375BFCD4F95694DBF94A" bold="true" box="[946,1011,1756,1782]" pageId="2" pageNumber="327">
|
||
<taxonomicNameLabel id="A22B9B7EFF8C375BFCD4F95694DBF94A" box="[946,1011,1756,1782]" pageId="2" pageNumber="327" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the bilobed supraorbital eave with the antorbital and preorbital lobes subequal in size and length, a character it shares with
|
||
<taxonomicName id="4C6C8194FF8C375BFB14F88A93C5F8A5" authorityName="Griffin & Tranter" authorityYear="1986" box="[1138,1261,1792,1818]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="B9182605FF8C375BFB14F88A93C5F8A5" box="[1138,1261,1792,1818]" italics="true" pageId="2" pageNumber="327">A. costatus</emphasis>
|
||
</taxonomicName>
|
||
described from Lord Howe Island, eastern
|
||
<collectingCountry id="F37BBA87FF8C375BFEA0F8A99518F881" box="[454,560,1827,1853]" name="Australia" pageId="2" pageNumber="327">Australia</collectingCountry>
|
||
. The other
|
||
<taxonomicName id="4C6C8194FF8C375BFDCBF8A99488F881" box="[685,928,1827,1853]" pageId="2" pageNumber="327">
|
||
<emphasis id="B9182605FF8C375BFDCBF8A99461F881" box="[685,841,1827,1853]" italics="true" pageId="2" pageNumber="327">Acanthophrys</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
have the antorbital lobe much larger than the preorbital lobe.
|
||
<taxonomicName id="4C6C8194FF8C375BFE2DF8CD9507F8DD" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[331,559,1863,1889]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="keeae" status="sp. nov.">
|
||
<emphasis id="B9182605FF8C375BFE2DF8CD9507F8DD" box="[331,559,1863,1889]" italics="true" pageId="2" pageNumber="327">Acanthophrys keeae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF8C375BFD50F8C2955EF8DE" bold="true" box="[566,630,1864,1890]" pageId="2" pageNumber="327">
|
||
<taxonomicNameLabel id="A22B9B7EFF8C375BFD50F8C2955EF8DE" box="[566,630,1864,1890]" pageId="2" pageNumber="327" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, however, can be separated from
|
||
<taxonomicName id="4C6C8194FF8C375BFC93F8C29359F8DD" authorityName="Griffin & Tranter" authorityYear="1986" box="[1013,1137,1864,1890]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="B9182605FF8C375BFC93F8C29359F8DD" box="[1013,1137,1864,1890]" italics="true" pageId="2" pageNumber="327">A. costatus</emphasis>
|
||
</taxonomicName>
|
||
by the tips of the ant- and preorbital lobes of the supraorbital eave being distinctly rounded (
|
||
<figureCitation id="1357E692FF8C375BFC1AF8E194DDF83A" box="[892,1013,1899,1926]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="2" pageNumber="327">Fig. 2B–D</figureCitation>
|
||
) (vs. tips sharp in
|
||
<taxonomicName id="4C6C8194FF8C375BFBA3F8E69268F839" authorityName="Griffin & Tranter" authorityYear="1986" box="[1221,1344,1900,1926]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="B9182605FF8C375BFBA3F8E69268F839" box="[1221,1344,1900,1926]" italics="true" pageId="2" pageNumber="327">A. costatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF8C375BFA29F8E19666F815" author="Griffin, D. J. G. & Tranter, H. A." pageId="2" pageNumber="327" pagination="1 - 335" refId="ref9546" refString="Griffin, D. J. G. & Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie, XXXIX, 1 - 335, figs. 1 - 112, pls. 1 - 22, 3 tabs. https: // doi. org / 10.1163 / 9789004626461 _ 002" type="book chapter" year="1986">Griffin & Tranter, 1986</bibRefCitation>
|
||
: fig. 31h); the postorbital spine is proportionately longer, extending well beyond the margin of the supraorbital eave (
|
||
<figureCitation id="1357E692FF8C375BFE02F83996F1F872" box="[356,473,1971,1998]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="2" pageNumber="327">Fig. 2B, C</figureCitation>
|
||
) (vs. postorbital spine shorter, reaching just beyond the margin of the supraorbital eave in
|
||
<taxonomicName id="4C6C8194FF8C375BFFD2F8529618F84D" authorityName="Griffin & Tranter" authorityYear="1986" box="[180,304,2008,2034]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="B9182605FF8C375BFFD2F8529618F84D" box="[180,304,2008,2034]" italics="true" pageId="2" pageNumber="327">A. costatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF8C375BFE59F85D9562F84D" author="Griffin, D. J. G. & Tranter, H. A." box="[319,586,2007,2034]" pageId="2" pageNumber="327" pagination="1 - 335" refId="ref9546" refString="Griffin, D. J. G. & Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie, XXXIX, 1 - 335, figs. 1 - 112, pls. 1 - 22, 3 tabs. https: // doi. org / 10.1163 / 9789004626461 _ 002" type="book chapter" year="1986">Griffin & Tranter, 1986</bibRefCitation>
|
||
: fig. 31h); the gastric regions are covered with many low rounded tubercles (
|
||
<figureCitation id="1357E692FF8A375DFFF9FF1D963DFF0D" box="[159,277,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="4" pageNumber="329">Fig. 2B–E</figureCitation>
|
||
) (vs. gastric regions smoother in
|
||
<taxonomicName id="4C6C8194FF8A375DFDE9FF1D9422FF0C" authorityName="Griffin & Tranter" authorityYear="1986" box="[655,778,151,177]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="B9182605FF8A375DFDE9FF1D9422FF0C" box="[655,778,151,177]" italics="true" pageId="4" pageNumber="329">A. costatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF8A375DFC7FFF1D930EFF0D" author="Griffin, D. J. G. & Tranter, H. A." box="[793,1062,151,177]" pageId="4" pageNumber="329" pagination="1 - 335" refId="ref9546" refString="Griffin, D. J. G. & Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie, XXXIX, 1 - 335, figs. 1 - 112, pls. 1 - 22, 3 tabs. https: // doi. org / 10.1163 / 9789004626461 _ 002" type="book chapter" year="1986">Griffin & Tranter, 1986</bibRefCitation>
|
||
: fig. 31h); the branchial region is less prominently inflated and narrower, with only low tubercles on the surface (
|
||
<figureCitation id="1357E692FF8A375DFB43FF319388FF69" box="[1061,1184,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="4" pageNumber="329">Fig. 2A, B</figureCitation>
|
||
) (vs. branchial region more inflated, wider, with low sharp spines and tubercles in
|
||
<taxonomicName id="4C6C8194FF8A375DFC53FF559498FF44" authorityName="Griffin & Tranter" authorityYear="1986" box="[821,944,223,249]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="B9182605FF8A375DFC53FF559498FF44" box="[821,944,223,249]" italics="true" pageId="4" pageNumber="329">A. costatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF8A375DFCD8FF5593EAFF45" author="Griffin, D. J. G. & Tranter, H. A." box="[958,1218,223,249]" pageId="4" pageNumber="329" pagination="1 - 335" refId="ref9546" refString="Griffin, D. J. G. & Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie, XXXIX, 1 - 335, figs. 1 - 112, pls. 1 - 22, 3 tabs. https: // doi. org / 10.1163 / 9789004626461 _ 002" type="book chapter" year="1986">Griffin & Tranter 1986</bibRefCitation>
|
||
: fig. 31h); the outer margin of the basal antennal article is sinuous with the median part concave (
|
||
<figureCitation id="1357E692FF8A375DFC89FE89934DFEA1" box="[1007,1125,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="4" pageNumber="329">Fig. 2F–H</figureCitation>
|
||
) (vs. margin almost straight in
|
||
<taxonomicName id="4C6C8194FF8A375DFFD2FEAD9618FEFC" authorityName="Griffin & Tranter" authorityYear="1986" box="[180,304,295,321]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="B9182605FF8A375DFFD2FEAD9618FEFC" box="[180,304,295,321]" italics="true" pageId="4" pageNumber="329">A. costatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF8A375DFE59FEAD956DFEFD" author="Griffin, D. J. G. & Tranter, H. A." box="[319,581,295,321]" pageId="4" pageNumber="329" pagination="1 - 335" refId="ref9546" refString="Griffin, D. J. G. & Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie, XXXIX, 1 - 335, figs. 1 - 112, pls. 1 - 22, 3 tabs. https: // doi. org / 10.1163 / 9789004626461 _ 002" type="book chapter" year="1986">Griffin & Tranter 1986</bibRefCitation>
|
||
: fig. 31a); the exopod of the third maxilliped is proportionately wider (
|
||
<figureCitation id="1357E692FF8A375DFA02FEAD979DFED9" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[191,1396,181,1885]" captionTargetId="figure-18@3.[191,1396,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae." figureDoi="http://doi.org/10.5281/zenodo.12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="4" pageNumber="329">Figs. 2F</figureCitation>
|
||
,
|
||
<figureCitation id="1357E692FF8A375DFFD9FEC197CBFED9" box="[191,227,331,357]" captionStart="FIGURE 4" captionStartId="5.[151,250,1845,1869]" captionTargetBox="[165,1423,532,1812]" captionTargetId="figure-187@5.[151,1435,525,1822]" captionTargetPageId="5" captionText="FIGURE 4. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) left third maxilliped; (B) outer view of left chela; (C) merus of left cheliped; (D) left P5; (E) pleon; (F) left G1 (ventral view); (G) left G1 (dorsal view); (H) distal part of left G1 (ventral view); (I) distal part of left G1 (ventral view); (J) distal part of left G1 (subdorsal view); (K), (L) distal part of left G1 (dorsal views); (M) left G2. A–E, H–L, denuded. Scales: A–E = 0.5 mm; F–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12681843" httpUri="https://zenodo.org/record/12681843/files/figure.png" pageId="4" pageNumber="329">4A</figureCitation>
|
||
) (vs. exopod distinctly narrower in
|
||
<taxonomicName id="4C6C8194FF8A375DFD08FEC195C0FED8" authorityName="Griffin & Tranter" authorityYear="1986" box="[622,744,331,357]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="B9182605FF8A375DFD08FEC195C0FED8" box="[622,744,331,357]" italics="true" pageId="4" pageNumber="329">A. costatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF8A375DFD93FEC194DFFED9" author="Griffin, D. J. G. & Tranter, H. A." box="[757,1015,331,357]" pageId="4" pageNumber="329" pagination="1 - 335" refId="ref9546" refString="Griffin, D. J. G. & Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie, XXXIX, 1 - 335, figs. 1 - 112, pls. 1 - 22, 3 tabs. https: // doi. org / 10.1163 / 9789004626461 _ 002" type="book chapter" year="1986">Griffin & Tranter 1986</bibRefCitation>
|
||
: fig. 31f); the almost smooth merus of the cheliped (
|
||
<figureCitation id="1357E692FF8A375DFE4BFEE596AAFE35" box="[301,386,367,393]" captionStart="FIGURE 4" captionStartId="5.[151,250,1845,1869]" captionTargetBox="[165,1423,532,1812]" captionTargetId="figure-187@5.[151,1435,525,1822]" captionTargetPageId="5" captionText="FIGURE 4. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) left third maxilliped; (B) outer view of left chela; (C) merus of left cheliped; (D) left P5; (E) pleon; (F) left G1 (ventral view); (G) left G1 (dorsal view); (H) distal part of left G1 (ventral view); (I) distal part of left G1 (ventral view); (J) distal part of left G1 (subdorsal view); (K), (L) distal part of left G1 (dorsal views); (M) left G2. A–E, H–L, denuded. Scales: A–E = 0.5 mm; F–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12681843" httpUri="https://zenodo.org/record/12681843/files/figure.png" pageId="4" pageNumber="329">Fig. 4C</figureCitation>
|
||
) (vs. dorsal margin distinctly tuberculated in
|
||
<taxonomicName id="4C6C8194FF8A375DFC1FFEE594DBFE34" authorityName="Griffin & Tranter" authorityYear="1986" box="[889,1011,367,393]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="B9182605FF8A375DFC1FFEE594DBFE34" box="[889,1011,367,393]" italics="true" pageId="4" pageNumber="329">A. costatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF8A375DFB67FEE5922BFE35" author="Griffin, D. J. G. & Tranter, H. A." box="[1025,1283,367,393]" pageId="4" pageNumber="329" pagination="1 - 335" refId="ref9546" refString="Griffin, D. J. G. & Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie, XXXIX, 1 - 335, figs. 1 - 112, pls. 1 - 22, 3 tabs. https: // doi. org / 10.1163 / 9789004626461 _ 002" type="book chapter" year="1986">Griffin & Tranter 1986</bibRefCitation>
|
||
: fig. 31d); the male anterior thoracic sternum is distinctly elongate, with sternites 3 and 4 constricted to form neck-like structure (
|
||
<figureCitation id="1357E692FF8A375DFFF9FE3D97D2FE6D" box="[159,250,439,465]" captionStart="FIGURE 3" captionStartId="4.[151,250,1429,1453]" captionTargetBox="[151,1411,705,1405]" captionTargetId="figure-292@4.[151,1435,705,1405]" captionTargetPageId="4" captionText="FIGURE 3. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) anterior thoracic sternum and pleon; (B) left G1 (ventral view); (C) left G1 (subdorsal view); (D) left G1 (dorsal view); (E) left G2. B–E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681841" httpUri="https://zenodo.org/record/12681841/files/figure.png" pageId="4" pageNumber="329">Fig. 3A</figureCitation>
|
||
) (vs. male anterior thoracic sternum relatively shorter, sternites 3 and 4 not distinctly constricted in
|
||
<taxonomicName id="4C6C8194FF8A375DFAE2FE3D97DBFE48" authorityName="Griffin & Tranter" authorityYear="1986" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="B9182605FF8A375DFAE2FE3D97DBFE48" italics="true" pageId="4" pageNumber="329">A. costatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF8A375DFE67FE51952DFE49" author="Griffin, D. J. G. & Tranter, H. A." box="[257,517,475,501]" pageId="4" pageNumber="329" pagination="1 - 335" refId="ref9546" refString="Griffin, D. J. G. & Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie, XXXIX, 1 - 335, figs. 1 - 112, pls. 1 - 22, 3 tabs. https: // doi. org / 10.1163 / 9789004626461 _ 002" type="book chapter" year="1986">Griffin & Tranter 1986</bibRefCitation>
|
||
: fig. 31c); the male pleon is more triangular in shape, with pleonal somites 3 and 4 distinctly wider with the telson lingulate and elongate (
|
||
<figureCitation id="1357E692FF8A375DFD99FE75944AFDA5" box="[767,866,511,537]" captionStart="FIGURE 3" captionStartId="4.[151,250,1429,1453]" captionTargetBox="[151,1411,705,1405]" captionTargetId="figure-292@4.[151,1435,705,1405]" captionTargetPageId="4" captionText="FIGURE 3. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) anterior thoracic sternum and pleon; (B) left G1 (ventral view); (C) left G1 (subdorsal view); (D) left G1 (dorsal view); (E) left G2. B–E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681841" httpUri="https://zenodo.org/record/12681841/files/figure.png" pageId="4" pageNumber="329">Figs. 3A</figureCitation>
|
||
,
|
||
<figureCitation id="1357E692FF8A375DFC08FE7594A7FDA5" box="[878,911,511,537]" captionStart="FIGURE 4" captionStartId="5.[151,250,1845,1869]" captionTargetBox="[165,1423,532,1812]" captionTargetId="figure-187@5.[151,1435,525,1822]" captionTargetPageId="5" captionText="FIGURE 4. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) left third maxilliped; (B) outer view of left chela; (C) merus of left cheliped; (D) left P5; (E) pleon; (F) left G1 (ventral view); (G) left G1 (dorsal view); (H) distal part of left G1 (ventral view); (I) distal part of left G1 (ventral view); (J) distal part of left G1 (subdorsal view); (K), (L) distal part of left G1 (dorsal views); (M) left G2. A–E, H–L, denuded. Scales: A–E = 0.5 mm; F–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12681843" httpUri="https://zenodo.org/record/12681843/files/figure.png" pageId="4" pageNumber="329">4E</figureCitation>
|
||
) (vs. pleonal somites proportionately narrower with telson triangular and shorter in
|
||
<taxonomicName id="4C6C8194FF8A375DFD52FDA99598FD80" authorityName="Griffin & Tranter" authorityYear="1986" box="[564,688,547,573]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="B9182605FF8A375DFD52FDA99598FD80" box="[564,688,547,573]" italics="true" pageId="4" pageNumber="329">A. costatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF8A375DFDD9FDA994EEFD81" author="Griffin, D. J. G. & Tranter, H. A." box="[703,966,547,573]" pageId="4" pageNumber="329" pagination="1 - 335" refId="ref9546" refString="Griffin, D. J. G. & Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie, XXXIX, 1 - 335, figs. 1 - 112, pls. 1 - 22, 3 tabs. https: // doi. org / 10.1163 / 9789004626461 _ 002" type="book chapter" year="1986">Griffin & Tranter 1986</bibRefCitation>
|
||
: fig. 31c); and the distal two-thirds of the G1 is almost straight with the distal part tapering (
|
||
<figureCitation id="1357E692FF8A375DFDADFDCD9478FDDD" box="[715,848,583,609]" captionStart="FIGURE 3" captionStartId="4.[151,250,1429,1453]" captionTargetBox="[151,1411,705,1405]" captionTargetId="figure-292@4.[151,1435,705,1405]" captionTargetPageId="4" captionText="FIGURE 3. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) anterior thoracic sternum and pleon; (B) left G1 (ventral view); (C) left G1 (subdorsal view); (D) left G1 (dorsal view); (E) left G2. B–E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681841" httpUri="https://zenodo.org/record/12681841/files/figure.png" pageId="4" pageNumber="329">Figs. 3B–D</figureCitation>
|
||
,
|
||
<figureCitation id="1357E692FF8A375DFC3DFDCD94B2FDDD" box="[859,922,583,609]" captionStart="FIGURE 4" captionStartId="5.[151,250,1845,1869]" captionTargetBox="[165,1423,532,1812]" captionTargetId="figure-187@5.[151,1435,525,1822]" captionTargetPageId="5" captionText="FIGURE 4. Acanthophrys keeae n. sp., holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) left third maxilliped; (B) outer view of left chela; (C) merus of left cheliped; (D) left P5; (E) pleon; (F) left G1 (ventral view); (G) left G1 (dorsal view); (H) distal part of left G1 (ventral view); (I) distal part of left G1 (ventral view); (J) distal part of left G1 (subdorsal view); (K), (L) distal part of left G1 (dorsal views); (M) left G2. A–E, H–L, denuded. Scales: A–E = 0.5 mm; F–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12681843" httpUri="https://zenodo.org/record/12681843/files/figure.png" pageId="4" pageNumber="329">4F–L</figureCitation>
|
||
) (vs. G1 distally bent with a rounded tip in
|
||
<taxonomicName id="4C6C8194FF8A375DFAE2FDCD97DBFD38" authorityName="Griffin & Tranter" authorityYear="1986" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="B9182605FF8A375DFAE2FDCD97DBFD38" italics="true" pageId="4" pageNumber="329">A. costatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF8A375DFE64FDE1952FFD39" author="Griffin, D. J. G. & Tranter, H. A." box="[258,519,619,645]" pageId="4" pageNumber="329" pagination="1 - 335" refId="ref9546" refString="Griffin, D. J. G. & Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie, XXXIX, 1 - 335, figs. 1 - 112, pls. 1 - 22, 3 tabs. https: // doi. org / 10.1163 / 9789004626461 _ 002" type="book chapter" year="1986">Griffin & Tranter 1986</bibRefCitation>
|
||
: fig. 32a, b).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF13AA9FFF8D375AFFF1F8FE9592F844" ID-DOI="http://doi.org/10.5281/zenodo.12681839" ID-Zenodo-Dep="12681839" httpUri="https://zenodo.org/record/12681839/files/figure.png" pageId="3" pageNumber="328" startId="3.[151,250,1908,1932]" targetBox="[191,1396,181,1885]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BD3FA17FF8D375AFFF1F8FE9592F844" blockId="3.[151,1436,1908,2041]" pageId="3" pageNumber="328">
|
||
<emphasis id="B9182605FF8D375AFFF1F8FE963BF831" bold="true" box="[151,275,1908,1933]" pageId="3" pageNumber="328">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C6C8194FF8D375AFE7CF8FF96C0F831" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[282,488,1909,1933]" class="Malacostraca" family="Epialtidae" genus="Acanthophrys" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="keeae" status="sp. nov.">
|
||
<emphasis id="B9182605FF8D375AFE7CF8FF96C0F831" box="[282,488,1909,1933]" italics="true" pageId="3" pageNumber="328">Acanthophrys keeae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF8D375AFE88F8FF950FF831" bold="true" box="[494,551,1909,1933]" pageId="3" pageNumber="328">
|
||
<taxonomicNameLabel id="A22B9B7EFF8D375AFE88F8FF950FF831" box="[494,551,1909,1933]" pageId="3" pageNumber="328" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (pcl 9.9 mm) (CASAU-CR-2021-1022), India: (A) overall habitus; (B) dorsal view of carapace; (C) anterior part of carapace; (D) angled subfrontal view of anterior part of carapace; (E) lateral view of right side of cephalothorax; (F) antennae, epistome, orbits, eyes, buccal cavity and third maxilliped; (G) proepistome, antennules and antennae; (H) epistome, antennules and antennae.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C376A99CFF8A375DFFA1FD0495D9FD15" box="[199,753,654,681]" pageId="4" pageNumber="329" type="distribution">
|
||
<paragraph id="8BD3FA17FF8A375DFFA1FD0495D9FD15" blockId="4.[151,1437,151,681]" box="[199,753,654,681]" pageId="4" pageNumber="329">
|
||
<emphasis id="B9182605FF8A375DFFA1FD049671FD14" bold="true" box="[199,345,654,680]" pageId="4" pageNumber="329">Distribution</emphasis>
|
||
. Known only from the
|
||
<typeStatus id="54D744B5FF8A375DFD06FD0595B8FD15" box="[608,656,655,681]" pageId="4" pageNumber="329">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |