594 lines
103 KiB
XML
594 lines
103 KiB
XML
<document id="5EBCD28586699CE1E9BB6242D1CE817B" ID-DOI="10.11646/zootaxa.5476.1.26" ID-ISSN="1175-5326" ID-Zenodo-Dep="12681830" ID-ZooBank="84C66278-7194-4F3E-8145-3918E1659289" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1720375444750" checkinUser="plazi" docAuthor="Ng, Peter K. L., Prema, Mani & Ravichandran, Samuthirapandian" docDate="2024" docId="03C54B01FF843757FFF1FF1D95D9FED9" docLanguage="en" docName="zootaxa.5476.1.26.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.26" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vellumnus intonsus Ng, Prema & Ravichandran, 2024, n. sp." docType="treatment" docVersion="4" lastPageNumber="339" masterDocId="FFFC3379FF8E3759FF66FF8A9728FFBC" masterDocTitle="Descriptions of new and rare crabs associated with coral rubble and sponges from Kanyakumari, southeastern Arabian Sea, southern India (Decapoda: Brachyura: Epialtidae and Pilumnidae)" masterLastPageNumber="341" masterPageNumber="325" pageNumber="335" updateTime="1721068086350" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B7215795ECA372A07C8BC6F6ACA74AF6">Descriptions of new and rare crabs associated with coral rubble and sponges from Kanyakumari, southeastern Arabian Sea, southern India (Decapoda: Brachyura: Epialtidae and Pilumnidae)</mods:title>
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<mods:namePart id="7449F9DD9D5C1C9E79E5C6E2B9C286C2">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="15F8CE85E86CC9DB05007EDE6FAB1801">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore.</mods:affiliation>
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<mods:namePart id="FBF638DA062D52A83C9FD7AC53D99AE6">Prema, Mani</mods:namePart>
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<mods:affiliation id="07DFE8FB5EA6AC1D76280879320ADA5C">Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Annamalai University, Parangipettai- 608 502, India.</mods:affiliation>
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<mods:namePart id="CDFDD0F49CCCC5E09DEC044DD914ED9E">Ravichandran, Samuthirapandian</mods:namePart>
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<mods:affiliation id="11216A5FE9347EE362DDE509ED360235">Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Annamalai University, Parangipettai- 608 502, India. & Department of Zoology, Government Arts & Science College, Nagercoil - 629 004, India.</mods:affiliation>
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<mods:title id="AC745FA46406D73EA0540D0EDE272D09">Zootaxa</mods:title>
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<mods:date id="525D70F3FE403FBB1256C7E87E7F12BC">2024</mods:date>
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<taxonomicName id="4C6C8194FF843753FFF1FF1D96AEFF0D" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[151,390,151,178]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="intonsus" status="sp. nov.">
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<emphasis id="B9182605FF843753FFF1FF1D96AEFF0D" bold="true" box="[151,390,151,178]" italics="true" pageId="10" pageNumber="335">Vellumnus intonsus</emphasis>
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<taxonomicNameLabel id="A22B9B7EFF843753FEEBFF1D96F8FF0D" box="[397,464,151,177]" pageId="10" pageNumber="335" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BD3FA17FF843753FFF1FF319608FF69" blockId="10.[151,464,151,213]" box="[151,288,187,213]" pageId="10" pageNumber="335">
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(
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<figureCitation id="1357E692FF843753FFF9FF319630FF69" box="[159,280,187,213]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStartId-0="11.[151,250,1710,1734]" captionStartId-1="12.[151,250,1552,1576]" captionStartId-2="13.[151,250,1687,1711]" captionTargetBox-0="[151,1436,181,1686]" captionTargetBox-1="[151,1435,381,1529]" captionTargetBox-2="[162,1429,251,1647]" captionTargetId-0="figure-18@11.[151,1436,181,1686]" captionTargetId-1="figure-137@12.[151,1435,381,1529]" captionTargetId-2="figure-43@13.[151,1436,237,1664]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="FIGURE 8. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) overall habitus (carapace gently brushed with right side denuded); (C, D) right side of carapace (denuded); (E) frontal view of cephalothorax; (F) antennules, antenna, orbit and epistome; (B, D) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India." captionText-1="FIGURE 9. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A) anterior thoracic sternum and pleon; (B) dorsal view of left cheliped; (C) dorsal view of right cheliped; (D) outer view of left chela; (E) outer view of right chela; (F) left P5. B, C, and D, E, photographed to same magnification." captionText-2="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12681853" figureDoi-1="http://doi.org/10.5281/zenodo.12681859" figureDoi-2="http://doi.org/10.5281/zenodo.12681861" httpUri-0="https://zenodo.org/record/12681853/files/figure.png" httpUri-1="https://zenodo.org/record/12681859/files/figure.png" httpUri-2="https://zenodo.org/record/12681861/files/figure.png" pageId="10" pageNumber="335">Figs. 8–10</figureCitation>
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)
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</paragraph>
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<taxonomicName id="4C6C8194FF843753FFF1FE8A96ACFEA4" ID-CoL="6VJL4" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[151,388,256,280]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="labyrinthicus">
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<emphasis id="B9182605FF843753FFF1FE8A96ACFEA4" box="[151,388,256,280]" italics="true" pageId="10" pageNumber="335">Pilumnus labyrinthicus</emphasis>
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</taxonomicName>
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.—
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<treatmentCitation id="0ACDDC06FF843753FEC7FE8A95ACFEAB" author="Henderson, J. R." box="[417,644,255,280]" page="365" pageId="10" pageNumber="335" year="1893">
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<bibRefCitation id="EFFD87E6FF843753FEC7FE8A95ACFEAB" author="Henderson, J. R." box="[417,644,255,280]" pageId="10" pageNumber="335" pagination="325 - 458" refId="ref9614" refString="Henderson, J. R. (1893) A contribution to Indian carcinology. Transactions of the Linnaean Society of London Zoology, Series 2, 5, 325 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.1893. tb 00653. x" type="journal article" year="1893">Henderson, 1893: 365</bibRefCitation>
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(Gulf of Manaar);
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<treatmentCitation id="0ACDDC06FF843753FC21FE8A9359FEA4" author="Kathirvel, M. & Gokul, A." box="[839,1137,256,280]" page="27" pageId="10" pageNumber="335" year="2010">
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<bibRefCitation id="EFFD87E6FF843753FC21FE8A9359FEA4" author="Kathirvel, M. & Gokul, A." box="[839,1137,256,280]" pageId="10" pageNumber="335" refId="ref9718" refString="Kathirvel, M. & Gokul, A. (2010) Checklist of brachyuran crabs from the Gulf of Mannar marine biosphere reserve. In: Technical Bulletin. Vol. 4. The Fisheries Technocrats Forum, Chennai, pp. 1 - 45." type="book" year="2010">Kathirvel & Gokul, 2010: 27</bibRefCitation>
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</treatmentCitation>
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. (Not
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<taxonomicName id="4C6C8194FF843753FBD6FE8A9668FE8A" ID-CoL="6VJL4" authority="Miers 1884" authorityName="Miers" authorityYear="1884" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="labyrinthicus">
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||
<emphasis id="B9182605FF843753FBD6FE8A92B5FEA4" box="[1200,1437,256,280]" italics="true" pageId="10" pageNumber="335">Pilumnus labyrinthicus</emphasis>
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<bibRefCitation id="EFFD87E6FF843753FFA1FE949668FE8A" author="Miers, E. J." box="[199,320,286,310]" pageId="10" pageNumber="335" pagination="18 - 32" refId="ref10234" refString="Miers, E. J. (1884) Crustacea. In: Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. Alert 1881 - 1882. Part I. The collections from Melanesia. British Museum (Natural History), London, pp. 178 - 322, pls. 18 - 32." type="book chapter" year="1884">Miers 1884</bibRefCitation>
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</taxonomicName>
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).
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</paragraph>
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<paragraph id="8BD3FA17FF843753FFF1FEB795E3FEE9" blockId="10.[151,1437,255,341]" box="[151,715,317,341]" pageId="10" pageNumber="335">
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<treatmentCitationGroup id="AB7CDD39FF843753FFF1FEB795E3FEE9" box="[151,715,317,341]" pageId="10" pageNumber="335">
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<taxonomicName id="4C6C8194FF843753FFF1FEB796A4FEE9" ID-CoL="7FKC6" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[151,396,317,341]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="labyrinthicus">
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||
<emphasis id="B9182605FF843753FFF1FEB796A4FEE9" box="[151,396,317,341]" italics="true" pageId="10" pageNumber="335">Vellumnus labyrinthicus</emphasis>
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</taxonomicName>
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.—
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<treatmentCitation id="0ACDDC06FF843753FECFFEB795A5FEE9" author="Trivedi, J. N. & Trivedi, D. J. & Vachhrajani, K. D. & Ng, P. K. L." box="[425,653,317,341]" page="61" pageId="10" pageNumber="335" year="2018">
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<bibRefCitation id="EFFD87E6FF843753FECFFEB795A5FEE9" author="Trivedi, J. N. & Trivedi, D. J. & Vachhrajani, K. D. & Ng, P. K. L." box="[425,653,317,341]" pageId="10" pageNumber="335" pagination="1 - 83" refId="ref11685" refString="Trivedi, J. N., Trivedi, D. J., Vachhrajani, K. D. & Ng, P. K. L. (2018) An annotated checklist of marine brachyuran crabs (Crustacea: Decapoda: Brachyura) of India. Zootaxa, 4502 (1), 1 - 83. https: // doi. org / 10.11646 / zootaxa. 4502.1.1" type="journal article" year="2018">
|
||
Trivedi
|
||
<emphasis id="B9182605FF843753FE9DFEB49500FEE9" box="[507,552,317,341]" italics="true" pageId="10" pageNumber="335">et al</emphasis>
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||
. 2018: 61
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||
</bibRefCitation>
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||
</treatmentCitation>
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(list).
|
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<paragraph id="8BD3FA17FF843753FFF1FE099570FDB5" blockId="10.[151,1437,387,1997]" pageId="10" pageNumber="335">
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<emphasis id="B9182605FF843753FFF1FE09966BFE21" bold="true" box="[151,323,387,413]" pageId="10" pageNumber="335">Type material</emphasis>
|
||
.
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||
<materialsCitation id="3B04F04AFF843753FE30FE099404FE59" collectingDate="2021-11-05" collectorName="M. Prema" country="India" county="Southeastern Arabian Sea" elevation="55" latitude="8.091611" location="Mankudy" longLatPrecision="1" longitude="77.48414" municipality="Kanyakumari district" pageId="10" pageNumber="335" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="holotype">
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<emphasis id="B9182605FF843753FE30FE099695FE21" box="[342,445,387,413]" italics="true" pageId="10" pageNumber="335">
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<typeStatus id="54D744B5FF843753FE30FE099695FE21" box="[342,445,387,413]" pageId="10" pageNumber="335" type="holotype">Holotype</typeStatus>
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</emphasis>
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: male (cw
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<quantity id="4C9457F2FF843753FD2FFE09958EFE22" box="[585,678,387,414]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="10" pageNumber="335" unit="mm" value="4.3">4.3 mm</quantity>
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, cl 3.0 mm), CASAU-CR-2021-1024, in coral rubble,
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fish landing centre,
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,
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<collectingRegion id="49A834F5FF843753FDE7FE2D9424FE7D" box="[641,780,423,449]" country="India" name="Tamil Nadu" pageId="10" pageNumber="335">Tamil Nadu</collectingRegion>
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, southern
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<collectingCountry id="F37BBA87FF843753FCE3FE2D94E9FE7D" box="[901,961,423,449]" name="India" pageId="10" pageNumber="335">India</collectingCountry>
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,
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,
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,
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,
|
||
<quantity id="4C9457F2FF843753FE5AFE419689FE5A" box="[316,417,459,486]" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="10" pageNumber="335" unit="m" value="55.0" valueMax="60.0" valueMin="50.0">
|
||
<elevation id="00411D24FF843753FE5AFE419689FE5A" box="[316,417,459,486]" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="10" pageNumber="335" unit="m" value="55.0" valueMax="60.0" valueMin="50.0">50–60 m</elevation>
|
||
</quantity>
|
||
, coll.
|
||
<collectorName id="26999FC1FF843753FE82FE46957DFE5A" box="[484,597,460,486]" pageId="10" pageNumber="335">M. Prema</collectorName>
|
||
,
|
||
<date id="FFD2DCD7FF843753FD06FE419404FE59" box="[608,812,459,486]" pageId="10" pageNumber="335" value="2021-11-05">
|
||
<collectingDate id="EF96253FFF843753FD06FE419404FE59" box="[608,812,459,486]" pageId="10" pageNumber="335" value="2021-11-05">5 November 2021</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B04F04AFF843753FC51FE46957CFDB5" collectingDate="2021-11-05" collectionCode="ZRC" collectorName="M. Prema" country="India" county="Southeastern Arabian Sea" elevation="55" latitude="8.091611" location="Mankudy" longLatPrecision="1" longitude="77.48414" municipality="Kanyakumari district" pageId="10" pageNumber="335" specimenCode="ZRC 2022.0999" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="paratype">
|
||
<emphasis id="B9182605FF843753FC51FE4694B6FE59" box="[823,926,460,485]" italics="true" pageId="10" pageNumber="335">
|
||
<typeStatus id="54D744B5FF843753FC51FE4694B6FE59" box="[823,926,460,485]" pageId="10" pageNumber="335" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
: 1 ovigerous female (cw
|
||
<quantity id="4C9457F2FF843753FBDFFE419239FE5A" box="[1209,1297,459,486]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="10" pageNumber="335" unit="mm" value="4.3">4.3 mm</quantity>
|
||
, cl
|
||
<quantity id="4C9457F2FF843753FA51FE4192B8FE59" box="[1335,1424,459,485]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="10" pageNumber="335" unit="mm" value="3.1">3.1 mm</quantity>
|
||
),
|
||
<specimenCode id="DBCA526CFF843753FFF1FE659667FDB5" box="[151,335,495,521]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="335">ZRC 2022.0999</specimenCode>
|
||
, same data as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376A99CFF843753FFA1FD9997D8FDED" pageId="10" pageNumber="335" type="etymology">
|
||
<paragraph id="8BD3FA17FF843753FFA1FD9997D8FDED" blockId="10.[151,1437,387,1997]" pageId="10" pageNumber="335">
|
||
<emphasis id="B9182605FF843753FFA1FD999660FD91" bold="true" box="[199,328,531,557]" pageId="10" pageNumber="335">Etymology</emphasis>
|
||
. The name is derived from the Latin for bearded and unshaven; alluding to the appearance of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376A99CFF843753FFA1FDD19794FB2D" pageId="10" pageNumber="335" type="diagnosis">
|
||
<paragraph id="8BD3FA17FF843753FFA1FDD19794FB2D" blockId="10.[151,1437,387,1997]" pageId="10" pageNumber="335">
|
||
<emphasis id="B9182605FF843753FFA1FDD19612FDC9" bold="true" box="[199,314,603,629]" pageId="10" pageNumber="335">Diagnosis</emphasis>
|
||
. Carapace hexagonal, wide; surface and pereiopods covered with dense, short, soft pubescence as well as scattered long plumose setae completely obscuring margins and surfaces, longer setae arranged in clumps on carapace regions, forming faint circular patterns on gastric and cardiac regions (
|
||
<figureCitation id="1357E692FF843753FB6CFD2993A9FD01" box="[1034,1153,675,701]" captionStart="FIGURE 8" captionStartId="11.[151,250,1710,1734]" captionTargetBox="[151,1436,181,1686]" captionTargetId="figure-18@11.[151,1436,181,1686]" captionTargetPageId="11" captionText="FIGURE 8. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) overall habitus (carapace gently brushed with right side denuded); (C, D) right side of carapace (denuded); (E) frontal view of cephalothorax; (F) antennules, antenna, orbit and epistome; (B, D) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India." figureDoi="http://doi.org/10.5281/zenodo.12681853" httpUri="https://zenodo.org/record/12681853/files/figure.png" pageId="10" pageNumber="335">Fig. 8A, B</figureCitation>
|
||
); epigastric, mesogastric, postorbital cristae low, gastric and subhepatic regions with scattered low granules, hepatic region with low broad tubercle, may be very low (
|
||
<figureCitation id="1357E692FF843753FEB3FD619576FCBA" box="[469,606,747,774]" captionStart="FIGURE 8" captionStartId="11.[151,250,1710,1734]" captionTargetBox="[151,1436,181,1686]" captionTargetId="figure-18@11.[151,1436,181,1686]" captionTargetPageId="11" captionText="FIGURE 8. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) overall habitus (carapace gently brushed with right side denuded); (C, D) right side of carapace (denuded); (E) frontal view of cephalothorax; (F) antennules, antenna, orbit and epistome; (B, D) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India." figureDoi="http://doi.org/10.5281/zenodo.12681853" httpUri="https://zenodo.org/record/12681853/files/figure.png" pageId="10" pageNumber="335">Figs. 8C, D</figureCitation>
|
||
,
|
||
<figureCitation id="1357E692FF843753FD0AFD6195E9FCB9" box="[620,705,747,773]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="10" pageNumber="335">10A, B</figureCitation>
|
||
); frontal margin distinctly convex, lateral lobule sharp, distinct, separated from frontal lobes by deep V-shaped cleft; supraorbital margin with median fissure, margin convex (
|
||
<figureCitation id="1357E692FF843753FA02FC8597F5FCF2" captionStart="FIGURE 8" captionStartId="11.[151,250,1710,1734]" captionTargetBox="[151,1436,181,1686]" captionTargetId="figure-18@11.[151,1436,181,1686]" captionTargetPageId="11" captionText="FIGURE 8. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) overall habitus (carapace gently brushed with right side denuded); (C, D) right side of carapace (denuded); (E) frontal view of cephalothorax; (F) antennules, antenna, orbit and epistome; (B, D) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India." figureDoi="http://doi.org/10.5281/zenodo.12681853" httpUri="https://zenodo.org/record/12681853/files/figure.png" pageId="10" pageNumber="335">Figs. 8C, D</figureCitation>
|
||
,
|
||
<figureCitation id="1357E692FF843753FF8EFCB99613FCF1" box="[232,315,819,845]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="10" pageNumber="335">10A, B</figureCitation>
|
||
); external orbital tooth triangular, first anterolateral tooth triangular, third tooth smallest; subhepatic tooth low or distinct, may be visible in dorsal view (
|
||
<figureCitation id="1357E692FF843753FD9EFCDD94A9FCCE" box="[760,897,855,882]" captionStart="FIGURE 8" captionStartId="11.[151,250,1710,1734]" captionTargetBox="[151,1436,181,1686]" captionTargetId="figure-18@11.[151,1436,181,1686]" captionTargetPageId="11" captionText="FIGURE 8. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) overall habitus (carapace gently brushed with right side denuded); (C, D) right side of carapace (denuded); (E) frontal view of cephalothorax; (F) antennules, antenna, orbit and epistome; (B, D) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India." figureDoi="http://doi.org/10.5281/zenodo.12681853" httpUri="https://zenodo.org/record/12681853/files/figure.png" pageId="10" pageNumber="335">Figs. 8C, D</figureCitation>
|
||
,
|
||
<figureCitation id="1357E692FF843753FCE9FCDD94CDFCCD" box="[911,997,855,881]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="10" pageNumber="335">10A, B</figureCitation>
|
||
); surface of merus of third maxilliped smooth, anteroexternal angle of rounded, not auriculiform, ischium relatively short (
|
||
<figureCitation id="1357E692FF843753FB03FCF193E7FC29" box="[1125,1231,891,917]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="10" pageNumber="335">Fig. 10C</figureCitation>
|
||
); outer surface of chelae with numerous relatively large round granules, vaguely arranged in longitudinal rows, carpus with numerous round granules, without distinct tooth on inner angle, fingers shorter than palm (
|
||
<figureCitation id="1357E692FF843753FB60FC499355FC61" box="[1030,1149,963,989]" captionStart="FIGURE 9" captionStartId="12.[151,250,1552,1576]" captionTargetBox="[151,1435,381,1529]" captionTargetId="figure-137@12.[151,1435,381,1529]" captionTargetPageId="12" captionText="FIGURE 9. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A) anterior thoracic sternum and pleon; (B) dorsal view of left cheliped; (C) dorsal view of right cheliped; (D) outer view of left chela; (E) outer view of right chela; (F) left P5. B, C, and D, E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681859" httpUri="https://zenodo.org/record/12681859/files/figure.png" pageId="10" pageNumber="335">Fig. 9B–E</figureCitation>
|
||
); ambulatory legs without distinct crests or ridges, unarmed; surfaces completely obscured by numerous long, short setae (
|
||
<figureCitation id="1357E692FF843753FBBFFC6D9207FBBD" box="[1241,1327,999,1025]" captionStart="FIGURE 9" captionStartId="12.[151,250,1552,1576]" captionTargetBox="[151,1435,381,1529]" captionTargetId="figure-137@12.[151,1435,381,1529]" captionTargetPageId="12" captionText="FIGURE 9. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A) anterior thoracic sternum and pleon; (B) dorsal view of left cheliped; (C) dorsal view of right cheliped; (D) outer view of left chela; (E) outer view of right chela; (F) left P5. B, C, and D, E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681859" httpUri="https://zenodo.org/record/12681859/files/figure.png" pageId="10" pageNumber="335">Fig. 9F</figureCitation>
|
||
); anterior thoracic sternum with surfaces relatively smooth, suture between sternites 3 and 4 shallow, medially interrupted (
|
||
<figureCitation id="1357E692FF843753FFF9FBA597D1FBF5" box="[159,249,1071,1097]" captionStart="FIGURE 9" captionStartId="12.[151,250,1552,1576]" captionTargetBox="[151,1435,381,1529]" captionTargetId="figure-137@12.[151,1435,381,1529]" captionTargetPageId="12" captionText="FIGURE 9. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A) anterior thoracic sternum and pleon; (B) dorsal view of left cheliped; (C) dorsal view of right cheliped; (D) outer view of left chela; (E) outer view of right chela; (F) left P5. B, C, and D, E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681859" httpUri="https://zenodo.org/record/12681859/files/figure.png" pageId="10" pageNumber="335">Fig. 9A</figureCitation>
|
||
); male pleon relatively narrow, subrectangular, pleonal somites 1, 3 subequal in width; telson semicircular (
|
||
<figureCitation id="1357E692FF843753FFF9FBD9962DFBD1" box="[159,261,1107,1133]" captionStart="FIGURE 9" captionStartId="12.[151,250,1552,1576]" captionTargetBox="[151,1435,381,1529]" captionTargetId="figure-137@12.[151,1435,381,1529]" captionTargetPageId="12" captionText="FIGURE 9. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A) anterior thoracic sternum and pleon; (B) dorsal view of left cheliped; (C) dorsal view of right cheliped; (D) outer view of left chela; (E) outer view of right chela; (F) left P5. B, C, and D, E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681859" httpUri="https://zenodo.org/record/12681859/files/figure.png" pageId="10" pageNumber="335">Figs. 9A</figureCitation>
|
||
,
|
||
<figureCitation id="1357E692FF843753FE77FBD99668FBD1" box="[273,320,1107,1133]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="10" pageNumber="335">10F</figureCitation>
|
||
); G1 slender, sinuous along entire length, distal part gently bent laterally with sharp tip (
|
||
<figureCitation id="1357E692FF843753FA56FBD99787FB2D" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="10" pageNumber="335">Fig. 10G, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376A99CFF843753FFA1FB1196C3FAD5" pageId="10" pageNumber="335" type="description">
|
||
<paragraph id="8BD3FA17FF843753FFA1FB1194D1FAF9" blockId="10.[151,1437,387,1997]" pageId="10" pageNumber="335">
|
||
<emphasis id="B9182605FF843753FFA1FB119610FB09" bold="true" box="[199,312,1179,1205]" pageId="10" pageNumber="335">Variation</emphasis>
|
||
. In the
|
||
<typeStatus id="54D744B5FF843753FEEDFB1196C5FB09" box="[395,493,1179,1205]" pageId="10" pageNumber="335" type="holotype">holotype</typeStatus>
|
||
male, there is a distinct large but low tubercle behind the first anterolateral tooth and the subhepatic region has a distinct tooth which is clearly visible in dorsal view (
|
||
<figureCitation id="1357E692FF843753FB53FB3593B3FB65" box="[1077,1179,1215,1241]" captionStart="FIGURE 8" captionStartId="11.[151,250,1710,1734]" captionTargetBox="[151,1436,181,1686]" captionTargetId="figure-18@11.[151,1436,181,1686]" captionTargetPageId="11" captionText="FIGURE 8. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) overall habitus (carapace gently brushed with right side denuded); (C, D) right side of carapace (denuded); (E) frontal view of cephalothorax; (F) antennules, antenna, orbit and epistome; (B, D) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India." figureDoi="http://doi.org/10.5281/zenodo.12681853" httpUri="https://zenodo.org/record/12681853/files/figure.png" pageId="10" pageNumber="335">Figs. 8D</figureCitation>
|
||
,
|
||
<figureCitation id="1357E692FF843753FBCEFB3593F4FB65" box="[1192,1244,1215,1241]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="10" pageNumber="335">10A</figureCitation>
|
||
). In the
|
||
<typeStatus id="54D744B5FF843753FA5AFB4A92B4FB66" box="[1340,1436,1216,1242]" pageId="10" pageNumber="335" type="paratype">paratype</typeStatus>
|
||
female, however, the surface behind the first anterolateral tooth is only slightly raised and barely visible and the subhepatic tooth is low and poorly developed, and hardly visible in dorsal view (
|
||
<figureCitation id="1357E692FF843753FB76FA8D9358FA9D" box="[1040,1136,1287,1313]" captionStart="FIGURE 8" captionStartId="11.[151,250,1710,1734]" captionTargetBox="[151,1436,181,1686]" captionTargetId="figure-18@11.[151,1436,181,1686]" captionTargetPageId="11" captionText="FIGURE 8. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) overall habitus (carapace gently brushed with right side denuded); (C, D) right side of carapace (denuded); (E) frontal view of cephalothorax; (F) antennules, antenna, orbit and epistome; (B, D) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India." figureDoi="http://doi.org/10.5281/zenodo.12681853" httpUri="https://zenodo.org/record/12681853/files/figure.png" pageId="10" pageNumber="335">Figs. 8C</figureCitation>
|
||
,
|
||
<figureCitation id="1357E692FF843753FB1DFA8D9385FA9D" box="[1147,1197,1287,1313]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="10" pageNumber="335">10B</figureCitation>
|
||
). The
|
||
<specimenCount id="9D6A319EFF843753FB97FA8292B1FA9D" box="[1265,1433,1287,1314]" count="2" pageId="10" pageNumber="335" type="generic">two specimens</specimenCount>
|
||
, however, agree in all other aspects and we are confident they are conspecific.
|
||
</paragraph>
|
||
<paragraph id="8BD3FA17FF843753FFA1FAC596C3FAD5" blockId="10.[151,1437,387,1997]" box="[199,491,1359,1386]" pageId="10" pageNumber="335">
|
||
<emphasis id="B9182605FF843753FFA1FAC59670FAD5" bold="true" box="[199,344,1359,1385]" pageId="10" pageNumber="335">Colouration</emphasis>
|
||
. Not known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376A99CFF843757FFA1FAF99438FEFD" lastPageId="14" lastPageNumber="339" pageId="10" pageNumber="335" type="discussion">
|
||
<paragraph id="8BD3FA17FF843753FFA1FAF993A2F9D9" blockId="10.[151,1437,387,1997]" pageId="10" pageNumber="335">
|
||
<emphasis id="B9182605FF843753FFA1FAF9961CFA31" bold="true" box="[199,308,1395,1421]" pageId="10" pageNumber="335">Remarks</emphasis>
|
||
.
|
||
<bibRefCitation id="EFFD87E6FF843753FE20FAF995A1FA32" author="Henderson, J. R." box="[326,649,1395,1422]" pageId="10" pageNumber="335" pagination="325 - 458" refId="ref9614" refString="Henderson, J. R. (1893) A contribution to Indian carcinology. Transactions of the Linnaean Society of London Zoology, Series 2, 5, 325 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.1893. tb 00653. x" type="journal article" year="1893">Henderson (1893: 365–366)</bibRefCitation>
|
||
recorded one small specimen cw 5.0 by cl 4.0 mm in size (no sex indicated) as “
|
||
<taxonomicName id="4C6C8194FF843753FE51FA1D9513FA0D" authorityName="Miers" authorityYear="1884" box="[311,571,1431,1457]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="labyrinthicus">
|
||
<emphasis id="B9182605FF843753FE51FA1D9513FA0D" box="[311,571,1431,1457]" italics="true" pageId="10" pageNumber="335">Pilumnus labyrinthicus</emphasis>
|
||
</taxonomicName>
|
||
” from Rameswaram, southern
|
||
<collectingCountry id="F37BBA87FF843753FCF3FA1D94F9FA0D" box="[917,977,1431,1457]" name="India" pageId="10" pageNumber="335">India</collectingCountry>
|
||
, noting that it has “the very characteristic markings or lines on its dorsal surface, which hear some resemblance to a face.”
|
||
<bibRefCitation id="EFFD87E6FF843753FB0EFA3192B4FA69" author="Kathirvel, M. & Gokul, A." box="[1128,1436,1467,1493]" pageId="10" pageNumber="335" refId="ref9718" refString="Kathirvel, M. & Gokul, A. (2010) Checklist of brachyuran crabs from the Gulf of Mannar marine biosphere reserve. In: Technical Bulletin. Vol. 4. The Fisheries Technocrats Forum, Chennai, pp. 1 - 45." type="book" year="2010">Kathirvel & Gokul (2010)</bibRefCitation>
|
||
subsequently listed the species based on this record from the Gulf of Mannar. On the basis of this description and distribution, this record is likely to be conspecific with
|
||
<taxonomicName id="4C6C8194FF843753FC6FF98E94AAF9A1" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[777,898,1540,1565]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="intonsus" status="n. sp.">
|
||
<emphasis id="B9182605FF843753FC6FF98E94AAF9A1" box="[777,898,1540,1565]" italics="true" pageId="10" pageNumber="335">V. intonsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF843753FCECF98994E2F9A1" bold="true" box="[906,970,1539,1565]" pageId="10" pageNumber="335">
|
||
<taxonomicNameLabel id="A22B9B7EFF843753FCECF98994E2F9A1" box="[906,970,1539,1565]" pageId="10" pageNumber="335" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The records of “
|
||
<taxonomicName id="4C6C8194FF843753FBECF98992B8F9A1" authorityName="Miers" authorityYear="1884" box="[1162,1424,1539,1565]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="labyrinthicus">
|
||
<emphasis id="B9182605FF843753FBECF98992B8F9A1" box="[1162,1424,1539,1565]" italics="true" pageId="10" pageNumber="335">Pilumnus labyrinthicus</emphasis>
|
||
</taxonomicName>
|
||
” from the
|
||
<collectingRegion id="49A834F5FF843753FF9BF9AD957AF9FD" box="[253,594,1575,1601]" country="India" name="Andaman and Nicobar Islands" pageId="10" pageNumber="335">Andaman and Nicobar Islands</collectingRegion>
|
||
by
|
||
<bibRefCitation id="EFFD87E6FF843753FD1BF9A294B6F9FD" author="Dev Roy, M. K. & Nandi, N. C." box="[637,926,1575,1602]" pageId="10" pageNumber="335" pagination="185 - 236" refId="ref9019" refString="Dev Roy, M. K. & Nandi, N. C. (2012) Brachyuran crabs (Crustacea). In: Director ZSI, Kolkata (Ed.), Fauna of Andaman and Nicobar Islands. State Fauna Series. Vol. 19. Part 1. Zoological Survey of India, Kolkata, pp. 185 - 236." type="book chapter" year="2012">Dev Roy & Nandi (2012)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFFD87E6FF843753FCB3F9A293A4F9FE" author="Dev Roy, M. K." box="[981,1164,1575,1602]" pageId="10" pageNumber="335" pagination="77 - 98" refId="ref8986" refString="Dev Roy, M. K. (2015) Conservation concerns on crustacean fauna of India. Journal of Environment and Socio-biology, 12 (1), 77 - 98." type="journal article" year="2015">Dev Roy (2015)</bibRefCitation>
|
||
need to be clarified and their specimens examined to confirm if they are
|
||
<taxonomicName id="4C6C8194FF843753FDD0F9C6944DF9D9" authorityName="Miers" authorityYear="1884" box="[694,869,1611,1637]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="labyrinthicus">
|
||
<emphasis id="B9182605FF843753FDD0F9C6944DF9D9" box="[694,869,1611,1637]" italics="true" pageId="10" pageNumber="335">V. labyrinthicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22B9B7EFF843753FC0AF9C6948FF9DA" box="[876,935,1612,1638]" pageId="10" pageNumber="335" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
or
|
||
<taxonomicName id="4C6C8194FF843753FCAAF9C6936CF9D9" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[972,1092,1612,1637]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="intonsus" status="n. sp.">
|
||
<emphasis id="B9182605FF843753FCAAF9C6936CF9D9" box="[972,1092,1612,1637]" italics="true" pageId="10" pageNumber="335">V. intonsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF843753FB2AF9C193A2F9D9" bold="true" box="[1100,1162,1611,1637]" pageId="10" pageNumber="335">
|
||
<taxonomicNameLabel id="A22B9B7EFF843753FB2AF9C193A2F9D9" box="[1100,1162,1611,1637]" pageId="10" pageNumber="335" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BD3FA17FF843753FFA1F9E5967DF871" blockId="10.[151,1437,387,1997]" pageId="10" pageNumber="335">
|
||
The small adult size (cw less than
|
||
<quantity id="4C9457F2FF843753FD3AF9E5958BF935" box="[604,675,1647,1673]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="335" unit="mm" value="8.0">8 mm</quantity>
|
||
), less obvious setal pattern on the carapace, the absence of ridges under the vermiform setal patterns, and relatively wider male thoracic sternum and male pleon allies
|
||
<taxonomicName id="4C6C8194FF843753FA63F91E9256F911" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[1285,1406,1684,1709]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="intonsus" status="n. sp.">
|
||
<emphasis id="B9182605FF843753FA63F91E9256F911" box="[1285,1406,1684,1709]" italics="true" pageId="10" pageNumber="335">V. intonsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF843753FAE0F9199790F96D" bold="true" pageId="10" pageNumber="335">
|
||
<taxonomicNameLabel id="A22B9B7EFF843753FAE0F9199790F96D" pageId="10" pageNumber="335" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
most closely with
|
||
<taxonomicName id="4C6C8194FF843753FEF4F932951AF96D" baseAuthorityName="Gordon" baseAuthorityYear="1930" box="[402,562,1719,1745]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="penicillatus">
|
||
<emphasis id="B9182605FF843753FEF4F932951AF96D" box="[402,562,1719,1745]" italics="true" pageId="10" pageNumber="335">V. penicillatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54D744B5FF843753FD24F93D955CF96D" box="[578,628,1719,1745]" pageId="10" pageNumber="335">type</typeStatus>
|
||
locality
|
||
<collectingCountry id="F37BBA87FF843753FDBFF932944BF96D" box="[729,867,1719,1746]" name="Hong Kong" pageId="10" pageNumber="335">Hong Kong</collectingCountry>
|
||
).
|
||
<taxonomicName id="4C6C8194FF843753FC11F93D9367F96D" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[887,1103,1719,1745]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="intonsus" status="sp. nov.">
|
||
<emphasis id="B9182605FF843753FC11F93D9367F96D" box="[887,1103,1719,1745]" italics="true" pageId="10" pageNumber="335">Vellumnus intonsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF843753FB3EF93D93B0F96D" bold="true" box="[1112,1176,1719,1745]" pageId="10" pageNumber="335">
|
||
<taxonomicNameLabel id="A22B9B7EFF843753FB3EF93D93B0F96D" box="[1112,1176,1719,1745]" pageId="10" pageNumber="335" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be separated from by the lateral lobule of the front being separated from the frontal lobes by a deep cleft (
|
||
<figureCitation id="1357E692FF843753FB0DF95193DAF949" box="[1131,1266,1755,1781]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="10" pageNumber="335">Fig. 10A, B</figureCitation>
|
||
) (vs. separated by a shallow concavity in
|
||
<taxonomicName id="4C6C8194FF843753FEA5F88A954BF8A5" baseAuthorityName="Gordon" baseAuthorityYear="1930" box="[451,611,1791,1817]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="penicillatus">
|
||
<emphasis id="B9182605FF843753FEA5F88A954BF8A5" box="[451,611,1791,1817]" italics="true" pageId="10" pageNumber="335">V. penicillatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF843753FD15F88A9416F8A5" author="Ng, P. K. L. & Clark, P. F." box="[627,830,1791,1818]" pageId="10" pageNumber="335" pagination="1 - 10" refId="ref10753" refString="Ng, P. K. L. & Clark, P. F. (2023) Redescription of Vellumnus penicillatus (Gordon, 1930) from Hong Kong and description of a new species from Macclesfield Bank, South China Sea (Crustacea: Brachyura: Pilumnidae). Nauplius, 31, 1 - 10. https: // doi. org / 10.1590 / 2358 - 2936 e 2023014" type="journal article" year="2023">Ng & Clark 2023</bibRefCitation>
|
||
: fig. 3A–C); the external orbital tooth is triangular in shape (
|
||
<figureCitation id="1357E692FF843753FF80F8A99644F881" box="[230,364,1827,1853]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="10" pageNumber="335">Fig. 10A, B</figureCitation>
|
||
) (vs. truncatiform in
|
||
<taxonomicName id="4C6C8194FF843753FD3AF8AE95D2F881" baseAuthorityName="Gordon" baseAuthorityYear="1930" box="[604,762,1827,1853]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="penicillatus">
|
||
<emphasis id="B9182605FF843753FD3AF8AE95D2F881" box="[604,762,1827,1853]" italics="true" pageId="10" pageNumber="335">V. penicillatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF843753FC6EF8AE94E7F881" author="Ng, P. K. L. & Clark, P. F." box="[776,975,1827,1854]" pageId="10" pageNumber="335" pagination="1 - 10" refId="ref10753" refString="Ng, P. K. L. & Clark, P. F. (2023) Redescription of Vellumnus penicillatus (Gordon, 1930) from Hong Kong and description of a new species from Macclesfield Bank, South China Sea (Crustacea: Brachyura: Pilumnidae). Nauplius, 31, 1 - 10. https: // doi. org / 10.1590 / 2358 - 2936 e 2023014" type="journal article" year="2023">Ng & Clark 2023</bibRefCitation>
|
||
: fig. 3A–C); the granules on the cheliped carpus and chela are proportionately larger and fewer in number (
|
||
<figureCitation id="1357E692FF843753FC10F8CD94C4F8DD" box="[886,1004,1863,1889]" captionStart="FIGURE 9" captionStartId="12.[151,250,1552,1576]" captionTargetBox="[151,1435,381,1529]" captionTargetId="figure-137@12.[151,1435,381,1529]" captionTargetPageId="12" captionText="FIGURE 9. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A) anterior thoracic sternum and pleon; (B) dorsal view of left cheliped; (C) dorsal view of right cheliped; (D) outer view of left chela; (E) outer view of right chela; (F) left P5. B, C, and D, E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681859" httpUri="https://zenodo.org/record/12681859/files/figure.png" pageId="10" pageNumber="335">Fig. 9B–E</figureCitation>
|
||
) (vs. smaller and more numerous in
|
||
<taxonomicName id="4C6C8194FF843753FAEEF8C29632F839" baseAuthorityName="Gordon" baseAuthorityYear="1930" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="penicillatus">
|
||
<emphasis id="B9182605FF843753FAEEF8C29632F839" italics="true" pageId="10" pageNumber="335">V. penicillatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF843753FE4EF8E696C5F839" author="Ng, P. K. L. & Clark, P. F." box="[296,493,1899,1926]" pageId="10" pageNumber="335" pagination="1 - 10" refId="ref10753" refString="Ng, P. K. L. & Clark, P. F. (2023) Redescription of Vellumnus penicillatus (Gordon, 1930) from Hong Kong and description of a new species from Macclesfield Bank, South China Sea (Crustacea: Brachyura: Pilumnidae). Nauplius, 31, 1 - 10. https: // doi. org / 10.1590 / 2358 - 2936 e 2023014" type="journal article" year="2023">Ng & Clark 2023</bibRefCitation>
|
||
: figs. 1A, B, 2A, B); and the distal part of the G1 is distinctly curved downwards and hooked (
|
||
<figureCitation id="1357E692FF843753FF9FF80596A8F815" box="[249,384,1935,1961]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="10" pageNumber="335">Fig. 10G, H</figureCitation>
|
||
) (vs. distal part is curved laterally at approximately right angles in
|
||
<taxonomicName id="4C6C8194FF843753FB0AF81A9223F815" baseAuthorityName="Gordon" baseAuthorityYear="1930" box="[1132,1291,1935,1961]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="penicillatus">
|
||
<emphasis id="B9182605FF843753FB0AF81A9223F815" box="[1132,1291,1935,1961]" italics="true" pageId="10" pageNumber="335">V. penicillatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF843753FA7FF81A97FBF871" author="Ng, P. K. L. & Clark, P. F." pageId="10" pageNumber="335" pagination="1 - 10" refId="ref10753" refString="Ng, P. K. L. & Clark, P. F. (2023) Redescription of Vellumnus penicillatus (Gordon, 1930) from Hong Kong and description of a new species from Macclesfield Bank, South China Sea (Crustacea: Brachyura: Pilumnidae). Nauplius, 31, 1 - 10. https: // doi. org / 10.1590 / 2358 - 2936 e 2023014" type="journal article" year="2023">Ng & Clark 2023</bibRefCitation>
|
||
: fig. 3H, I).
|
||
</paragraph>
|
||
<caption id="DF13AA9FFF853752FFF1F924967BF88E" ID-DOI="http://doi.org/10.5281/zenodo.12681853" ID-Zenodo-Dep="12681853" httpUri="https://zenodo.org/record/12681853/files/figure.png" pageId="11" pageNumber="336" startId="11.[151,250,1710,1734]" targetBox="[151,1436,181,1686]" targetPageId="11" targetType="figure">
|
||
<paragraph id="8BD3FA17FF853752FFF1F924967BF88E" blockId="11.[151,1436,1710,1842]" pageId="11" pageNumber="336">
|
||
<emphasis id="B9182605FF853752FFF1F924963FF97A" bold="true" box="[151,279,1710,1734]" pageId="11" pageNumber="336">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C6C8194FF853752FE44F92496C1F97A" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[290,489,1710,1734]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="336" phylum="Arthropoda" rank="species" species="intonsus" status="sp. nov.">
|
||
<emphasis id="B9182605FF853752FE44F92496C1F97A" box="[290,489,1710,1734]" italics="true" pageId="11" pageNumber="336">Vellumnus intonsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF853752FE93F924951AF97A" bold="true" box="[501,562,1710,1734]" pageId="11" pageNumber="336">
|
||
<taxonomicNameLabel id="A22B9B7EFF853752FE93F924951AF97A" box="[501,562,1710,1734]" pageId="11" pageNumber="336" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) overall habitus (carapace gently brushed with right side denuded); (C, D) right side of carapace (denuded); (E) frontal view of cephalothorax; (F) antennules, antenna, orbit and epistome; (B, D) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD3FA17FF853755FFA1F8EA94C8FED9" blockId="11.[151,1437,1888,2023]" lastBlockId="12.[151,1437,150,357]" lastPageId="12" lastPageNumber="337" pageId="11" pageNumber="336">
|
||
<taxonomicName id="4C6C8194FF853752FFA1F8EA96B7F8C6" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[199,415,1888,1914]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="336" phylum="Arthropoda" rank="species" species="intonsus" status="sp. nov.">
|
||
<emphasis id="B9182605FF853752FFA1F8EA96B7F8C6" box="[199,415,1888,1914]" italics="true" pageId="11" pageNumber="336">Vellumnus intonsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF853752FEC1F8EB96CEF8C7" bold="true" box="[423,486,1889,1915]" pageId="11" pageNumber="336">
|
||
<taxonomicNameLabel id="A22B9B7EFF853752FEC1F8EB96CEF8C7" box="[423,486,1889,1915]" pageId="11" pageNumber="336" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also resembles
|
||
<taxonomicName id="4C6C8194FF853752FDFDF8EB940FF8C6" baseAuthorityName="Takeda" baseAuthorityYear="1977" box="[667,807,1889,1914]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="336" phylum="Arthropoda" rank="species" species="pygmaeus">
|
||
<emphasis id="B9182605FF853752FDFDF8EB940FF8C6" box="[667,807,1889,1914]" italics="true" pageId="11" pageNumber="336">V. pygmaeus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54D744B5FF853752FC50F8EB9440F8C7" box="[822,872,1889,1915]" pageId="11" pageNumber="336">type</typeStatus>
|
||
locality Ogasawara Islands) and
|
||
<taxonomicName id="4C6C8194FF853752FB86F8EB9217F8C6" authorityName="Fahimi & Naderloo" authorityYear="2023" box="[1248,1343,1889,1914]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="336" phylum="Arthropoda" rank="species" species="pictus">
|
||
<emphasis id="B9182605FF853752FB86F8EB9217F8C6" box="[1248,1343,1889,1914]" italics="true" pageId="11" pageNumber="336">V. pictus</emphasis>
|
||
</taxonomicName>
|
||
(known only from
|
||
<specimenCount id="9D6A319EFF853752FE6FF80F96AAF822" box="[265,386,1924,1951]" count="1" pageId="11" pageNumber="336" type="female">one female</specimenCount>
|
||
from the Persian Gulf) and most of the differences between them are shared with
|
||
<taxonomicName id="4C6C8194FF853752FB9FF80F92BEF822" baseAuthorityName="Gordon" baseAuthorityYear="1930" box="[1273,1430,1924,1950]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="336" phylum="Arthropoda" rank="species" species="penicillatus">
|
||
<emphasis id="B9182605FF853752FB9FF80F92BEF822" box="[1273,1430,1924,1950]" italics="true" pageId="11" pageNumber="336">V. penicillatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C6C8194FF853752FFF1F82296A8F87E" baseAuthorityName="Takeda" baseAuthorityYear="1977" box="[151,384,1960,1986]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="336" phylum="Arthropoda" rank="species" species="pygmaeus">
|
||
<emphasis id="B9182605FF853752FFF1F82296A8F87E" box="[151,384,1960,1986]" italics="true" pageId="11" pageNumber="336">Vellumnus pygmaeus</emphasis>
|
||
</taxonomicName>
|
||
can be separated as it has the supraorbital margin prominently more convex (
|
||
<bibRefCitation id="EFFD87E6FF853752FBB4F8229245F87F" author="Takeda, M." box="[1234,1389,1960,1987]" pageId="11" pageNumber="336" pagination="113 - 140" refId="ref11496" refString="Takeda, M. (1977) Crabs of the Ogasawara Islands, V. A collection made by dredging. Memoirs of the National Science Museum, Tokyo, 10, 113 - 140." type="journal article" year="1977">Takeda, 1977</bibRefCitation>
|
||
: fig. 5A) (vs. supraorbital margin gently convex with median fissure in
|
||
<taxonomicName id="4C6C8194FF853752FCEFF847932AF85A" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[905,1026,1997,2022]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="336" phylum="Arthropoda" rank="species" species="intonsus" status="n. sp.">
|
||
<emphasis id="B9182605FF853752FCEFF847932AF85A" box="[905,1026,1997,2022]" italics="true" pageId="11" pageNumber="336">V. intonsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF853752FB6CF8479361F85B" bold="true" box="[1034,1097,1997,2023]" pageId="11" pageNumber="336">
|
||
<taxonomicNameLabel id="A22B9B7EFF853752FB6CF8479361F85B" box="[1034,1097,1997,2023]" pageId="11" pageNumber="336" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<figureCitation id="1357E692FF853752FB3FF84693FCF85B" box="[1113,1236,1996,2023]" captionStart="FIGURE 8" captionStartId="11.[151,250,1710,1734]" captionTargetBox="[151,1436,181,1686]" captionTargetId="figure-18@11.[151,1436,181,1686]" captionTargetPageId="11" captionText="FIGURE 8. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) overall habitus (carapace gently brushed with right side denuded); (C, D) right side of carapace (denuded); (E) frontal view of cephalothorax; (F) antennules, antenna, orbit and epistome; (B, D) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India." figureDoi="http://doi.org/10.5281/zenodo.12681853" httpUri="https://zenodo.org/record/12681853/files/figure.png" pageId="11" pageNumber="336">Fig. 8C, D</figureCitation>
|
||
,
|
||
<figureCitation id="1357E692FF853752FB86F847921BF85B" box="[1248,1331,1997,2023]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="11" pageNumber="336">10A, B</figureCitation>
|
||
); and the G1 is relatively stouter with the distal part curved laterally (
|
||
<bibRefCitation id="EFFD87E6FF823755FC54FF1D94E6FF0D" author="Takeda, M." box="[818,974,151,177]" pageId="12" pageNumber="337" pagination="113 - 140" refId="ref11496" refString="Takeda, M. (1977) Crabs of the Ogasawara Islands, V. A collection made by dredging. Memoirs of the National Science Museum, Tokyo, 10, 113 - 140." type="journal article" year="1977">Takeda, 1977</bibRefCitation>
|
||
: fig. 5B, C) (vs. G1 slender, sinuous with the distal part gently bent laterally, with a sharp tip in
|
||
<taxonomicName id="4C6C8194FF823755FD9EFF319459FF68" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[760,881,187,212]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="intonsus" status="n. sp.">
|
||
<emphasis id="B9182605FF823755FD9EFF319459FF68" box="[760,881,187,212]" italics="true" pageId="12" pageNumber="337">V. intonsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF823755FC1FFF319490FF69" bold="true" box="[889,952,187,213]" pageId="12" pageNumber="337">
|
||
<taxonomicNameLabel id="A22B9B7EFF823755FC1FFF319490FF69" box="[889,952,187,213]" pageId="12" pageNumber="337" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<figureCitation id="1357E692FF823755FCA1FF31937AFF69" box="[967,1106,187,213]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="12" pageNumber="337">Fig. 10G, H</figureCitation>
|
||
). Compared with
|
||
<taxonomicName id="4C6C8194FF823755FA7DFF319251FF68" authorityName="Fahimi & Naderloo" authorityYear="2023" box="[1307,1401,187,212]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="pictus">
|
||
<emphasis id="B9182605FF823755FA7DFF319251FF68" box="[1307,1401,187,212]" italics="true" pageId="12" pageNumber="337">V. pictus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C8194FF823755FAEEFF3197DCFF44" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="intonsus" status="n. sp.">
|
||
<emphasis id="B9182605FF823755FAEEFF3197DCFF44" italics="true" pageId="12" pageNumber="337">V. intonsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF823755FF9AFF559614FF45" bold="true" box="[252,316,223,249]" pageId="12" pageNumber="337">
|
||
<taxonomicNameLabel id="A22B9B7EFF823755FF9AFF559614FF45" box="[252,316,223,249]" pageId="12" pageNumber="337" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also has the merus of the third maxilliped smooth with the anterolateral angle gently rounded (
|
||
<figureCitation id="1357E692FF823755FA09FF5597EFFEA1" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="12" pageNumber="337">Fig. 10C</figureCitation>
|
||
) (vs. surface with granules and the anterolateral angle is auriculiform in
|
||
<taxonomicName id="4C6C8194FF823755FB67FE899349FEA0" authorityName="Fahimi & Naderloo" authorityYear="2023" box="[1025,1121,259,284]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="pictus">
|
||
<emphasis id="B9182605FF823755FB67FE899349FEA0" box="[1025,1121,259,284]" italics="true" pageId="12" pageNumber="337">V. pictus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF823755FB16FE8992B0FEA1" author="Fahimi, N. & Naderloo, R." box="[1136,1432,259,285]" pageId="12" pageNumber="337" pagination="537 - 545" refId="ref9084" refString="Fahimi, N. & Naderloo, R. (2023) A new species of pilumnid crab of the genus Vellumnus Ng, 2010 (Brachyura, Pilumnoidea) from the Persian Gulf. Crustaceana, 96 (6), 537 - 545. https: // doi. org / 10.1163 / 15685403 - bja 10292" type="journal article" year="2023">Fahimi & Naderloo, 2023</bibRefCitation>
|
||
: fig. 1D); and there are relatively more numerous granules on the outer surface of the chela (
|
||
<figureCitation id="1357E692FF823755FBC3FEAD9233FEFD" box="[1189,1307,295,321]" captionStart="FIGURE 9" captionStartId="12.[151,250,1552,1576]" captionTargetBox="[151,1435,381,1529]" captionTargetId="figure-137@12.[151,1435,381,1529]" captionTargetPageId="12" captionText="FIGURE 9. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A) anterior thoracic sternum and pleon; (B) dorsal view of left cheliped; (C) dorsal view of right cheliped; (D) outer view of left chela; (E) outer view of right chela; (F) left P5. B, C, and D, E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681859" httpUri="https://zenodo.org/record/12681859/files/figure.png" pageId="12" pageNumber="337">Fig. 9B–E</figureCitation>
|
||
) (vs. fewer and less evenly distributed in
|
||
<taxonomicName id="4C6C8194FF823755FE83FEC1956BFED8" authorityName="Fahimi & Naderloo" authorityYear="2023" box="[485,579,331,356]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="pictus">
|
||
<emphasis id="B9182605FF823755FE83FEC1956BFED8" box="[485,579,331,356]" italics="true" pageId="12" pageNumber="337">V. pictus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF823755FD34FEC1945FFED9" author="Fahimi, N. & Naderloo, R." box="[594,887,331,357]" pageId="12" pageNumber="337" pagination="537 - 545" refId="ref9084" refString="Fahimi, N. & Naderloo, R. (2023) A new species of pilumnid crab of the genus Vellumnus Ng, 2010 (Brachyura, Pilumnoidea) from the Persian Gulf. Crustaceana, 96 (6), 537 - 545. https: // doi. org / 10.1163 / 15685403 - bja 10292" type="journal article" year="2023">Fahimi & Naderloo, 2023</bibRefCitation>
|
||
: fig. 1C).
|
||
</paragraph>
|
||
<caption id="DF13AA9FFF823755FFF1F99A9306F9CD" ID-DOI="http://doi.org/10.5281/zenodo.12681859" ID-Zenodo-Dep="12681859" httpUri="https://zenodo.org/record/12681859/files/figure.png" pageId="12" pageNumber="337" startId="12.[151,250,1552,1576]" targetBox="[151,1435,381,1529]" targetPageId="12" targetType="figure">
|
||
<paragraph id="8BD3FA17FF823755FFF1F99A9306F9CD" blockId="12.[151,1437,1552,1649]" pageId="12" pageNumber="337">
|
||
<emphasis id="B9182605FF823755FFF1F99A963CF994" bold="true" box="[151,276,1552,1576]" pageId="12" pageNumber="337">FIGURE 9.</emphasis>
|
||
<taxonomicName id="4C6C8194FF823755FE7AF99B96F7F994" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[284,479,1553,1577]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="intonsus" status="sp. nov.">
|
||
<emphasis id="B9182605FF823755FE7AF99B96F7F994" box="[284,479,1553,1577]" italics="true" pageId="12" pageNumber="337">Vellumnus intonsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF823755FE81F99B950AF995" bold="true" box="[487,546,1553,1577]" pageId="12" pageNumber="337">
|
||
<taxonomicNameLabel id="A22B9B7EFF823755FE81F99B950AF995" box="[487,546,1553,1577]" pageId="12" pageNumber="337" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A) anterior thoracic sternum and pleon; (B) dorsal view of left cheliped; (C) dorsal view of right cheliped; (D) outer view of left chela; (E) outer view of right chela; (F
|
||
<emphasis id="B9182605FF823755FED2F9D29694F9CC" bold="true" box="[436,444,1624,1648]" pageId="12" pageNumber="337">)</emphasis>
|
||
left P5. B, C, and D, E, photographed to same magnification.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD3FA17FF823754FFA1F915964FFF69" blockId="12.[151,1437,1695,2009]" lastBlockId="13.[151,1437,151,213]" lastPageId="13" lastPageNumber="338" pageId="12" pageNumber="337">
|
||
<taxonomicName id="4C6C8194FF823755FFA1F9159689F905" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[199,417,1695,1721]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="intonsus" status="sp. nov.">
|
||
<emphasis id="B9182605FF823755FFA1F9159689F905" box="[199,417,1695,1721]" italics="true" pageId="12" pageNumber="337">Vellumnus intonsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF823755FECCF91596C3F905" bold="true" box="[426,491,1695,1721]" pageId="12" pageNumber="337">
|
||
<taxonomicNameLabel id="A22B9B7EFF823755FECCF91596C3F905" box="[426,491,1695,1721]" pageId="12" pageNumber="337" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also differs from the
|
||
<typeStatus id="54D744B5FF823755FD8FF9159431F905" box="[745,793,1695,1721]" pageId="12" pageNumber="337">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="4C6C8194FF823755FCE2F92A931EF905" authorityName="Miers" authorityYear="1884" box="[900,1078,1695,1721]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="labyrinthicus">
|
||
<emphasis id="B9182605FF823755FCE2F92A931EF905" box="[900,1078,1695,1721]" italics="true" pageId="12" pageNumber="337">V. labyrinthicus</emphasis>
|
||
</taxonomicName>
|
||
, in the vermiform setal pattern being only evident on the gastric and cardiac regions and when denuded, the underlying surface is smooth (
|
||
<figureCitation id="1357E692FF823755FA09F94997F5F8BD" captionStart="FIGURE 8" captionStartId="11.[151,250,1710,1734]" captionTargetBox="[151,1436,181,1686]" captionTargetId="figure-18@11.[151,1436,181,1686]" captionTargetPageId="11" captionText="FIGURE 8. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) overall habitus (carapace gently brushed with right side denuded); (C, D) right side of carapace (denuded); (E) frontal view of cephalothorax; (F) antennules, antenna, orbit and epistome; (B, D) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India." figureDoi="http://doi.org/10.5281/zenodo.12681853" httpUri="https://zenodo.org/record/12681853/files/figure.png" pageId="12" pageNumber="337">Fig. 8A–D</figureCitation>
|
||
) (vermiform pattern more pronounced in
|
||
<taxonomicName id="4C6C8194FF823755FDDCF9629443F8BD" authorityName="Miers" authorityYear="1884" box="[698,875,1767,1793]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="labyrinthicus">
|
||
<emphasis id="B9182605FF823755FDDCF9629443F8BD" box="[698,875,1767,1793]" italics="true" pageId="12" pageNumber="337">V. labyrinthicus</emphasis>
|
||
</taxonomicName>
|
||
with the setae is present on most of the carapace regions, and there is an ridge under these setae when denuded;
|
||
<bibRefCitation id="EFFD87E6FF823755FC0BF88194FDF899" author="Ng, P. K. L." box="[877,981,1803,1829]" pageId="12" pageNumber="337" pagination="33 - 61" refId="ref10515" refString="Ng, P. K. L. (2010) On the Planopilumnidae Serene, 1984 (Crustacea: Brachyura: Pseudozioidea), with diagnoses of two new pilumnoid genera for species previously assigned to Planopilumnus Balss, 1933. Zootaxa, 2392 (1), 33 - 61. https: // doi. org / 10.11646 / zootaxa. 2392.1. 2" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
: figs. 13A, 14A, 15A); the anterolateral teeth are relatively shorter with the last tooth directed obliquely anteriorly (
|
||
<figureCitation id="1357E692FF823755FC86F8A59370F8F5" box="[992,1112,1839,1865]" captionStart="FIGURE 8" captionStartId="11.[151,250,1710,1734]" captionTargetBox="[151,1436,181,1686]" captionTargetId="figure-18@11.[151,1436,181,1686]" captionTargetPageId="11" captionText="FIGURE 8. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) overall habitus (carapace gently brushed with right side denuded); (C, D) right side of carapace (denuded); (E) frontal view of cephalothorax; (F) antennules, antenna, orbit and epistome; (B, D) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India." figureDoi="http://doi.org/10.5281/zenodo.12681853" httpUri="https://zenodo.org/record/12681853/files/figure.png" pageId="12" pageNumber="337">Fig. 8C, D</figureCitation>
|
||
) (anterolateral teeth stronger and longer with the last tooth directed laterally in
|
||
<taxonomicName id="4C6C8194FF823755FDB6F8DE94A9F8D1" authorityName="Miers" authorityYear="1884" box="[720,897,1875,1901]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="labyrinthicus">
|
||
<emphasis id="B9182605FF823755FDB6F8DE94A9F8D1" box="[720,897,1875,1901]" italics="true" pageId="12" pageNumber="337">V. labyrinthicus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF823755FCF6F8D994DFF8D1" author="Ng, P. K. L." box="[912,1015,1875,1901]" pageId="12" pageNumber="337" pagination="33 - 61" refId="ref10515" refString="Ng, P. K. L. (2010) On the Planopilumnidae Serene, 1984 (Crustacea: Brachyura: Pseudozioidea), with diagnoses of two new pilumnoid genera for species previously assigned to Planopilumnus Balss, 1933. Zootaxa, 2392 (1), 33 - 61. https: // doi. org / 10.11646 / zootaxa. 2392.1. 2" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
: fig. 14A); the male anterior thoracic sternum is proportionately wider (
|
||
<figureCitation id="1357E692FF823755FD7DF8FD955CF82D" box="[539,628,1911,1937]" captionStart="FIGURE 9" captionStartId="12.[151,250,1552,1576]" captionTargetBox="[151,1435,381,1529]" captionTargetId="figure-137@12.[151,1435,381,1529]" captionTargetPageId="12" captionText="FIGURE 9. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A) anterior thoracic sternum and pleon; (B) dorsal view of left cheliped; (C) dorsal view of right cheliped; (D) outer view of left chela; (E) outer view of right chela; (F) left P5. B, C, and D, E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681859" httpUri="https://zenodo.org/record/12681859/files/figure.png" pageId="12" pageNumber="337">Fig. 9A</figureCitation>
|
||
) (proportionately narrower in
|
||
<taxonomicName id="4C6C8194FF823755FCA9F8F293A8F82D" authorityName="Miers" authorityYear="1884" box="[975,1152,1911,1937]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="labyrinthicus">
|
||
<emphasis id="B9182605FF823755FCA9F8F293A8F82D" box="[975,1152,1911,1937]" italics="true" pageId="12" pageNumber="337">V. labyrinthicus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF823755FBF6F8FD93DFF82D" author="Ng, P. K. L." box="[1168,1271,1911,1937]" pageId="12" pageNumber="337" pagination="33 - 61" refId="ref10515" refString="Ng, P. K. L. (2010) On the Planopilumnidae Serene, 1984 (Crustacea: Brachyura: Pseudozioidea), with diagnoses of two new pilumnoid genera for species previously assigned to Planopilumnus Balss, 1933. Zootaxa, 2392 (1), 33 - 61. https: // doi. org / 10.11646 / zootaxa. 2392.1. 2" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
: fig. 14B); the male pleon is relatively wider, especially across somites 3 and 4 (
|
||
<figureCitation id="1357E692FF823755FCE7F81194CCF809" box="[897,996,1947,1973]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="12" pageNumber="337">Fig. 10F</figureCitation>
|
||
) (entire pleon distinctly narrower in
|
||
<taxonomicName id="4C6C8194FF823755FAEEF8169603F865" authorityName="Miers" authorityYear="1884" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="labyrinthicus">
|
||
<emphasis id="B9182605FF823755FAEEF8169603F865" italics="true" pageId="12" pageNumber="337">V. labyrinthicus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF823755FE5EF83596B4F865" author="Ng, P. K. L." box="[312,412,1983,2009]" pageId="12" pageNumber="337" pagination="33 - 61" refId="ref10515" refString="Ng, P. K. L. (2010) On the Planopilumnidae Serene, 1984 (Crustacea: Brachyura: Pseudozioidea), with diagnoses of two new pilumnoid genera for species previously assigned to Planopilumnus Balss, 1933. Zootaxa, 2392 (1), 33 - 61. https: // doi. org / 10.11646 / zootaxa. 2392.1. 2" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
: fig. 16D); and the G1 is more distinctly sinuous with the proximal half strongly curved (
|
||
<figureCitation id="1357E692FF823754FA09F83597C3FF0D" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" lastPageId="13" lastPageNumber="338" pageId="12" pageNumber="337">Fig. 10G, H</figureCitation>
|
||
) (the proximal half is only gently curved with the G1 relatively less sinuous in
|
||
<taxonomicName id="4C6C8194FF833754FB14FF1D920BFF0D" authorityName="Miers" authorityYear="1884" box="[1138,1315,151,177]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="labyrinthicus">
|
||
<emphasis id="B9182605FF833754FB14FF1D920BFF0D" box="[1138,1315,151,177]" italics="true" pageId="13" pageNumber="338">V. labyrinthicus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFFD87E6FF833754FA54FF1D92B0FF0D" author="Ng, P. K. L." box="[1330,1432,151,177]" pageId="13" pageNumber="338" pagination="33 - 61" refId="ref10515" refString="Ng, P. K. L. (2010) On the Planopilumnidae Serene, 1984 (Crustacea: Brachyura: Pseudozioidea), with diagnoses of two new pilumnoid genera for species previously assigned to Planopilumnus Balss, 1933. Zootaxa, 2392 (1), 33 - 61. https: // doi. org / 10.11646 / zootaxa. 2392.1. 2" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
: figs. 15C, 16E, F).
|
||
</paragraph>
|
||
<caption id="DF13AA9FFF833754FFF1F91D94BCF883" ID-DOI="http://doi.org/10.5281/zenodo.12681861" ID-Zenodo-Dep="12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="13" pageNumber="338" startId="13.[151,250,1687,1711]" targetBox="[162,1429,251,1647]" targetPageId="13" targetType="figure">
|
||
<paragraph id="8BD3FA17FF833754FFF1F91D94BCF883" blockId="13.[151,1436,1687,1855]" pageId="13" pageNumber="338">
|
||
<emphasis id="B9182605FF833754FFF1F91D9608F913" bold="true" box="[151,288,1687,1711]" pageId="13" pageNumber="338">FIGURE 10.</emphasis>
|
||
<taxonomicName id="4C6C8194FF833754FE41F91D96C2F913" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[295,490,1687,1711]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="intonsus" status="sp. nov.">
|
||
<emphasis id="B9182605FF833754FE41F91D96C2F913" box="[295,490,1687,1711]" italics="true" pageId="13" pageNumber="338">Vellumnus intonsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF833754FE97F91D9503F913" bold="true" box="[497,555,1687,1711]" pageId="13" pageNumber="338">
|
||
<taxonomicNameLabel id="A22B9B7EFF833754FE97F91D9503F913" box="[497,555,1687,1711]" pageId="13" pageNumber="338" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD3FA17FF833757FFA1F8E79438FEFD" blockId="13.[151,1437,1901,2036]" lastBlockId="14.[151,1437,151,357]" lastPageId="14" lastPageNumber="339" pageId="13" pageNumber="338">
|
||
<taxonomicName id="4C6C8194FF833754FFA1F8E79689F83B" authorityName="Ng & Prema & Ravichandran" authorityYear="2024" box="[199,417,1901,1927]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="intonsus" status="sp. nov.">
|
||
<emphasis id="B9182605FF833754FFA1F8E79689F83B" box="[199,417,1901,1927]" italics="true" pageId="13" pageNumber="338">Vellumnus intonsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9182605FF833754FECDF8E496C5F834" bold="true" box="[427,493,1902,1928]" pageId="13" pageNumber="338">
|
||
<taxonomicNameLabel id="A22B9B7EFF833754FECDF8E496C5F834" box="[427,493,1902,1928]" pageId="13" pageNumber="338" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be separated from
|
||
<taxonomicName id="4C6C8194FF833754FD99F8E494B4F83B" baseAuthorityName="Sakai T" baseAuthorityYear="1969" box="[767,924,1901,1927]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="minabensis">
|
||
<emphasis id="B9182605FF833754FD99F8E494B4F83B" box="[767,924,1901,1927]" italics="true" pageId="13" pageNumber="338">V. minabensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C8194FF833754FCBEF8E4933AF83B" authorityName="Ng & Clark" authorityYear="2023" box="[984,1042,1901,1927]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="tki">
|
||
<emphasis id="B9182605FF833754FCBEF8E4933AF83B" box="[984,1042,1901,1927]" italics="true" pageId="13" pageNumber="338">V. tki</emphasis>
|
||
</taxonomicName>
|
||
by its proportionately wider male anterior thoracic sternum (
|
||
<figureCitation id="1357E692FF833754FEA7F81B9531F810" box="[449,537,1937,1964]" captionStart="FIGURE 9" captionStartId="12.[151,250,1552,1576]" captionTargetBox="[151,1435,381,1529]" captionTargetId="figure-137@12.[151,1435,381,1529]" captionTargetPageId="12" captionText="FIGURE 9. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A) anterior thoracic sternum and pleon; (B) dorsal view of left cheliped; (C) dorsal view of right cheliped; (D) outer view of left chela; (E) outer view of right chela; (F) left P5. B, C, and D, E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681859" httpUri="https://zenodo.org/record/12681859/files/figure.png" pageId="13" pageNumber="338">Fig. 9A</figureCitation>
|
||
) (vs. anterior thoracic sternum proportionately narrower in
|
||
<taxonomicName id="4C6C8194FF833754FBD1F8189279F817" baseAuthorityName="Sakai T" baseAuthorityYear="1969" box="[1207,1361,1937,1963]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="minabensis">
|
||
<emphasis id="B9182605FF833754FBD1F8189279F817" box="[1207,1361,1937,1963]" italics="true" pageId="13" pageNumber="338">V. minabensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C8194FF833754FAEEF818979BF873" authorityName="Ng & Clark" authorityYear="2023" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="tki">
|
||
<emphasis id="B9182605FF833754FAEEF818979BF873" italics="true" pageId="13" pageNumber="338">V. tki</emphasis>
|
||
</taxonomicName>
|
||
, e.g.,
|
||
<bibRefCitation id="EFFD87E6FF833754FF9EF83C96E8F86C" author="Ng, P. K. L. & Clark, P. F." box="[248,448,1973,2000]" pageId="13" pageNumber="338" pagination="1 - 10" refId="ref10753" refString="Ng, P. K. L. & Clark, P. F. (2023) Redescription of Vellumnus penicillatus (Gordon, 1930) from Hong Kong and description of a new species from Macclesfield Bank, South China Sea (Crustacea: Brachyura: Pilumnidae). Nauplius, 31, 1 - 10. https: // doi. org / 10.1590 / 2358 - 2936 e 2023014" type="journal article" year="2023">Ng & Clark 2023</bibRefCitation>
|
||
: fig. 6E); the median lobe of the posterior margin of the epistome is distinctly triangular in shape (
|
||
<figureCitation id="1357E692FF833754FE6FF853965AF848" box="[265,370,2009,2036]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="13" pageNumber="338">Fig. 10D</figureCitation>
|
||
) (vs. median lobe is lower and more lobiform in
|
||
<taxonomicName id="4C6C8194FF833754FCD3F8509379F84F" baseAuthorityName="Sakai T" baseAuthorityYear="1969" box="[949,1105,2009,2035]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="minabensis">
|
||
<emphasis id="B9182605FF833754FCD3F8509379F84F" box="[949,1105,2009,2035]" italics="true" pageId="13" pageNumber="338">V. minabensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C8194FF833754FBE8F85093E0F84F" authorityName="Ng & Clark" authorityYear="2023" box="[1166,1224,2009,2035]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="tki">
|
||
<emphasis id="B9182605FF833754FBE8F85093E0F84F" box="[1166,1224,2009,2035]" italics="true" pageId="13" pageNumber="338">V. tki</emphasis>
|
||
</taxonomicName>
|
||
, e.g.,
|
||
<bibRefCitation id="EFFD87E6FF833757FA74F85097FBFF0D" author="Ng, P. K. L. & Clark, P. F." lastPageId="14" lastPageNumber="339" pageId="13" pageNumber="338" pagination="1 - 10" refId="ref10753" refString="Ng, P. K. L. & Clark, P. F. (2023) Redescription of Vellumnus penicillatus (Gordon, 1930) from Hong Kong and description of a new species from Macclesfield Bank, South China Sea (Crustacea: Brachyura: Pilumnidae). Nauplius, 31, 1 - 10. https: // doi. org / 10.1590 / 2358 - 2936 e 2023014" type="journal article" year="2023">Ng & Clark 2023</bibRefCitation>
|
||
: fig. 6B); the ischium of the third maxilliped is relatively shorter (
|
||
<figureCitation id="1357E692FF803757FCA6FF1D930EFF0D" box="[960,1062,151,177]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="14" pageNumber="339">Fig. 10C</figureCitation>
|
||
) (vs. ischium distinctly longer in
|
||
<taxonomicName id="4C6C8194FF803757FFF1FF319619FF69" baseAuthorityName="Sakai T" baseAuthorityYear="1969" box="[151,305,187,213]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="339" phylum="Arthropoda" rank="species" species="minabensis">
|
||
<emphasis id="B9182605FF803757FFF1FF319619FF69" box="[151,305,187,213]" italics="true" pageId="14" pageNumber="339">V. minabensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C8194FF803757FE01FF3196B6FF69" authorityName="Ng & Clark" authorityYear="2023" box="[359,414,187,213]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="339" phylum="Arthropoda" rank="species" species="tki">
|
||
<emphasis id="B9182605FF803757FE01FF3196B6FF69" box="[359,414,187,213]" italics="true" pageId="14" pageNumber="339">V. tki</emphasis>
|
||
</taxonomicName>
|
||
, e.g.,
|
||
<bibRefCitation id="EFFD87E6FF803757FE85FF319581FF69" author="Ng, P. K. L. & Clark, P. F." box="[483,681,187,213]" pageId="14" pageNumber="339" pagination="1 - 10" refId="ref10753" refString="Ng, P. K. L. & Clark, P. F. (2023) Redescription of Vellumnus penicillatus (Gordon, 1930) from Hong Kong and description of a new species from Macclesfield Bank, South China Sea (Crustacea: Brachyura: Pilumnidae). Nauplius, 31, 1 - 10. https: // doi. org / 10.1590 / 2358 - 2936 e 2023014" type="journal article" year="2023">Ng & Clark 2023</bibRefCitation>
|
||
: fig. 6C); there are fewer but larger granules on the outer surface of the cheliped (
|
||
<figureCitation id="1357E692FF803757FE56FF55968EFF45" box="[304,422,223,249]" captionStart="FIGURE 9" captionStartId="12.[151,250,1552,1576]" captionTargetBox="[151,1435,381,1529]" captionTargetId="figure-137@12.[151,1435,381,1529]" captionTargetPageId="12" captionText="FIGURE 9. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A) anterior thoracic sternum and pleon; (B) dorsal view of left cheliped; (C) dorsal view of right cheliped; (D) outer view of left chela; (E) outer view of right chela; (F) left P5. B, C, and D, E, photographed to same magnification." figureDoi="http://doi.org/10.5281/zenodo.12681859" httpUri="https://zenodo.org/record/12681859/files/figure.png" pageId="14" pageNumber="339">Fig. 9B–E</figureCitation>
|
||
) (vs. more numerous smaller granules in
|
||
<taxonomicName id="4C6C8194FF803757FC18FF559330FF45" baseAuthorityName="Sakai T" baseAuthorityYear="1969" box="[894,1048,223,249]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="339" phylum="Arthropoda" rank="species" species="minabensis">
|
||
<emphasis id="B9182605FF803757FC18FF559330FF45" box="[894,1048,223,249]" italics="true" pageId="14" pageNumber="339">V. minabensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C8194FF803757FB36FF5593A1FF45" authorityName="Ng & Clark" authorityYear="2023" box="[1104,1161,223,249]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="339" phylum="Arthropoda" rank="species" species="tki">
|
||
<emphasis id="B9182605FF803757FB36FF5593A1FF45" box="[1104,1161,223,249]" italics="true" pageId="14" pageNumber="339">V. tki</emphasis>
|
||
</taxonomicName>
|
||
, e.g.,
|
||
<bibRefCitation id="EFFD87E6FF803757FBA9FF5592B0FF45" author="Ng, P. K. L. & Clark, P. F." box="[1231,1432,223,249]" pageId="14" pageNumber="339" pagination="1 - 10" refId="ref10753" refString="Ng, P. K. L. & Clark, P. F. (2023) Redescription of Vellumnus penicillatus (Gordon, 1930) from Hong Kong and description of a new species from Macclesfield Bank, South China Sea (Crustacea: Brachyura: Pilumnidae). Nauplius, 31, 1 - 10. https: // doi. org / 10.1590 / 2358 - 2936 e 2023014" type="journal article" year="2023">Ng & Clark 2023</bibRefCitation>
|
||
: fig. 5B, C); and the distal part of the G1 is distinctly more elongate (
|
||
<figureCitation id="1357E692FF803757FCC3FE899307FEA1" box="[933,1071,259,285]" captionStart="FIGURE 10" captionStartId="13.[151,250,1687,1711]" captionTargetBox="[162,1429,251,1647]" captionTargetId="figure-43@13.[151,1436,237,1664]" captionTargetPageId="13" captionText="FIGURE 10. Vellumnus intonsus n. sp., holotype male (cw 4.3 mm, cl 3.0 mm) (CASAU-CR-2021-1024), India: (A, B) right side of carapace (setae denuded); (C) left third maxilliped (denuded); (D) posterior margin of epistome; (E) left P5 dactylopropodal lock; (F) pleon; (G) left G1 (ventral view); (H) left G1 (dorsal view); (I) = left G2. Paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. (I) right side of carapace (setae denuded); B) paratype ovigerous female (cw 4.3 mm, cl 3.1 mm) (ZRC 2022.0999), India. Scales: A–F = 0.5 mm; G–I = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12681861" httpUri="https://zenodo.org/record/12681861/files/figure.png" pageId="14" pageNumber="339">Fig. 10G, H</figureCitation>
|
||
) (vs. G1 distal part shorter in
|
||
<taxonomicName id="4C6C8194FF803757FAEEFE89963DFEFD" baseAuthorityName="Sakai T" baseAuthorityYear="1969" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="339" phylum="Arthropoda" rank="species" species="minabensis">
|
||
<emphasis id="B9182605FF803757FAEEFE89963DFEFD" italics="true" pageId="14" pageNumber="339">V. minabensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C8194FF803757FE2AFEAD96ABFEFD" authorityName="Ng & Clark" authorityYear="2023" box="[332,387,295,321]" class="Malacostraca" family="Pilumnidae" genus="Vellumnus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="339" phylum="Arthropoda" rank="species" species="tki">
|
||
<emphasis id="B9182605FF803757FE2AFEAD96ABFEFD" box="[332,387,295,321]" italics="true" pageId="14" pageNumber="339">V. tki</emphasis>
|
||
</taxonomicName>
|
||
, e.g.,
|
||
<bibRefCitation id="EFFD87E6FF803757FEAEFEAD95A7FEFD" author="Ng, P. K. L. & Clark, P. F." box="[456,655,295,321]" pageId="14" pageNumber="339" pagination="1 - 10" refId="ref10753" refString="Ng, P. K. L. & Clark, P. F. (2023) Redescription of Vellumnus penicillatus (Gordon, 1930) from Hong Kong and description of a new species from Macclesfield Bank, South China Sea (Crustacea: Brachyura: Pilumnidae). Nauplius, 31, 1 - 10. https: // doi. org / 10.1590 / 2358 - 2936 e 2023014" type="journal article" year="2023">Ng & Clark 2023</bibRefCitation>
|
||
: fig. 6H, I).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376A99CFF803757FFA1FEC095D9FED9" box="[199,753,330,357]" pageId="14" pageNumber="339" type="distribution">
|
||
<paragraph id="8BD3FA17FF803757FFA1FEC095D9FED9" blockId="14.[151,1437,151,357]" box="[199,753,330,357]" pageId="14" pageNumber="339">
|
||
<emphasis id="B9182605FF803757FFA1FEC09671FED8" bold="true" box="[199,345,330,356]" pageId="14" pageNumber="339">Distribution</emphasis>
|
||
. Only known from the
|
||
<typeStatus id="54D744B5FF803757FD06FEC195B8FED9" box="[608,656,331,357]" pageId="14" pageNumber="339">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |