325 lines
39 KiB
XML
325 lines
39 KiB
XML
<document id="AE8E50697B0520343811BF3599AA344B" ID-DOI="10.1080/00222933.2018.1435833" ID-ISSN="1464-5262" ID-Zenodo-Dep="5174451" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628558511936" checkinUser="felipe" docAuthor="Wagner, Philipp, Greenbaum, Eli, Bauer, Aaron M., Kusamba, Chifundera & Leaché, Adam D." docDate="2018" docId="03BFD058FFBEFF83143C8BC0BDBEFE41" docLanguage="en" docName="JNatHist.52.13-16.771-817.pdf" docOrigin="Journal of Natural History 52 (13 - 16)" docSource="http://dx.doi.org/10.1080/00222933.2018.1435833" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Acanthocercus minutus" docType="treatment" docVersion="4" lastPageNumber="810" masterDocId="FF86A820FF9BFFAB154A8901BE4DFF9E" masterDocTitle="Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae)" masterLastPageNumber="817" masterPageNumber="771" pageNumber="807" updateTime="1699046512504" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="02FCDE296510B0DD22E66303E33EA996">Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae)</mods:title>
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<mods:namePart id="A05015EAA0EC3B6D1AD10A24D5663427">Wagner, Philipp</mods:namePart>
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<mods:namePart id="960082A968610A7B1ADD7CA294BA6A9B">Greenbaum, Eli</mods:namePart>
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<mods:namePart id="C330B6C7458ABAD715098667F245FB66">Bauer, Aaron M.</mods:namePart>
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<mods:namePart id="7550DB6D7C67F62E9D746696FB1E8334">Kusamba, Chifundera</mods:namePart>
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<mods:namePart id="EB67130BC835EE5B18499829F15A2B61">Leaché, Adam D.</mods:namePart>
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<mods:title id="465C44DBCDC9E9BD24A493984A0F6964">Journal of Natural History</mods:title>
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<mods:date id="09BEE67BFAE7DAE8BD1E649A8910AAFA">2018</mods:date>
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<mods:number id="6640D160C2E76086C480097930485F45">2018-03-19</mods:number>
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<mods:number id="0412AD074B6F739E132AA92420F8155A">52</mods:number>
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<mods:number id="3499AFEECAA32B0FC19F161F0D025D10">13 - 16</mods:number>
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<mods:identifier id="761F9A730F31C235B82E21E62EAEF99F" type="DOI">10.1080/00222933.2018.1435833</mods:identifier>
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<treatment id="03BFD058FFBEFF83143C8BC0BDBEFE41" ID-DOI="http://doi.org/10.5281/zenodo.5187234" ID-Zenodo-Dep="5187234" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41" httpUri="http://treatment.plazi.org/id/03BFD058FFBEFF83143C8BC0BDBEFE41" lastPageId="40" lastPageNumber="810" pageId="37" pageNumber="807">
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<subSubSection id="C30C32C5FFBEFF8E143C8BC0BDE1FD45" box="[374,940,705,731]" pageId="37" pageNumber="807" type="nomenclature">
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<paragraph id="8BA9614EFFBEFF8E143C8BC0BDE1FD45" blockId="37.[374,940,705,766]" box="[374,940,705,731]" pageId="37" pageNumber="807">
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<heading id="D0E1D622FFBEFF8E143C8BC0BDE1FD45" box="[374,940,705,731]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="807" reason="5">
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<treatmentCitationGroup id="AB064660FFBEFF8E143C8BC0BDE1FD45" box="[374,940,705,731]" pageId="37" pageNumber="807">
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<treatmentCitation id="0AB7475FFFBEFF8E143C8BC0BDE1FD45" author="Klausewitz W" box="[374,940,705,731]" page="170" pageId="37" pageNumber="807" year="1957">
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<taxonomicName id="4C161ACDFFBEFF8E143C8BC0BDE1FD45" authority="(Klausewitz, 1957: 170)" baseAuthorityName="Klausewitz" baseAuthorityPageNumber="170" baseAuthorityYear="1957" box="[374,940,705,731]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="807" phylum="Chordata" rank="species" species="minutus">
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<emphasis id="B962BD5CFFBEFF8E143C8BC0BCD8FD45" bold="true" box="[374,661,705,731]" italics="true" pageId="37" pageNumber="807">Acanthocercus minutus</emphasis>
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(
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<bibRefCitation id="EF871CBFFFBEFF8E17EE8BC3BDEAFD45" author="Klausewitz W" box="[676,935,706,731]" pageId="37" pageNumber="807" pagination="157 - 174" refId="ref20422" refString="Klausewitz W. 1957. Eidonomische Untersuchungen uber die Rassenkreise Agama cyanogaster und Agama atricollis. Die Unterarten von Agama atricollis [Eidonomic studies about the species complex of Agama cyanogaster and Agama atricollis. The subspecies of Agama atricollis]. Senckenb Biol. 38: 157 - 174." type="journal article" year="1957">Klausewitz, 1957: 170</bibRefCitation>
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)
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||
</taxonomicName>
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||
</treatmentCitation>
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||
</treatmentCitationGroup>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30C32C5FFBEFF8E171B8BE4BCCDFC18" pageId="37" pageNumber="807" type="description">
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<paragraph id="8BA9614EFFBEFF8E171B8BE4BC9FFD60" blockId="37.[374,940,705,766]" box="[593,722,741,766]" pageId="37" pageNumber="807">
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||
(
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<figureCitation id="132D7DCBFFBEFF8E17138BE4BC87FD60" box="[601,714,741,766]" captionStart="Figure 17" captionStartId="39.[160,225,1418,1441]" captionTargetBox="[178,1138,142,1387]" captionTargetId="figure-60@39.[178,1138,142,1387]" captionTargetPageId="39" captionText="Figure 17. Holotype (ZMB 29089) of Acanthocercus minutus, from Dscheffedenza, Ethiopia." figureDoi="http://doi.org/10.5281/zenodo.5174487" httpUri="https://zenodo.org/record/5174487/files/figure.png" pageId="37" pageNumber="807">Figure 17</figureCitation>
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)
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</paragraph>
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<paragraph id="8BA9614EFFBEFF8E15EA8A28BCCDFC18" blockId="37.[160,1156,809,902]" pageId="37" pageNumber="807">
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1957
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<taxonomicName id="4C161ACDFFBEFF8E15AA8A28BD95FCDF" authority="Klausewitz, Eidonomische Untersuchungen" authorityName="Klausewitz, Eidonomische Untersuchungen" authorityYear="1957" box="[224,984,809,833]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="807" phylum="Chordata" rank="subSpecies" species="atricollis" subSpecies="minuta">
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<emphasis id="B962BD5CFFBEFF8E15AA8A28BFA5FCDF" box="[224,488,809,833]" italics="true" pageId="37" pageNumber="807">Agama atricollis minuta</emphasis>
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Klausewitz, Eidonomische Untersuchungen
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</taxonomicName>
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über die Rassenkreise
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<taxonomicName id="4C161ACDFFBEFF8E14088A4ABC5FFCFD" authorityName="Ruppell" authorityYear="1835" box="[322,530,843,867]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="807" phylum="Chordata" rank="species" species="cyanogaster">
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<emphasis id="B962BD5CFFBEFF8E14088A4ABC5FFCFD" box="[322,530,843,867]" italics="true" pageId="37" pageNumber="807">Agama cyanogaster</emphasis>
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</taxonomicName>
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und
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<taxonomicName id="4C161ACDFFBEFF8E170E8A4ABCA7FCFD" authorityName="Smith, Illustrations" authorityYear="1849" box="[580,746,843,867]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="807" phylum="Chordata" rank="species" species="atricollis">
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<emphasis id="B962BD5CFFBEFF8E170E8A4ABCA7FCFD" box="[580,746,843,867]" italics="true" pageId="37" pageNumber="807">Agama atricollis</emphasis>
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</taxonomicName>
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. 2. Die Unterarten von
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<taxonomicName id="4C161ACDFFBEFF8E16928A4ABA33FCFD" authorityName="Smith, Illustrations" authorityYear="1849" box="[984,1150,843,867]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="807" phylum="Chordata" rank="species" species="atricollis">
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<emphasis id="B962BD5CFFBEFF8E16928A4ABA33FCFD" box="[984,1150,843,867]" italics="true" pageId="37" pageNumber="807">Agama atricollis</emphasis>
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</taxonomicName>
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. Senckenbergiana biologica 38: 157–174.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30C32C5FFBEFF8E15EA8AC1BF7DFB81" pageId="37" pageNumber="807" type="materials_examined">
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||
<paragraph id="8BA9614EFFBEFF8E15EA8AC1BF46FC47" blockId="37.[160,1155,960,1055]" box="[160,267,960,985]" pageId="37" pageNumber="807">
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||
<emphasis id="B962BD5CFFBEFF8E15EA8AC1BF46FC47" bold="true" box="[160,267,960,985]" italics="true" pageId="37" pageNumber="807">
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<typeStatus id="54ADDFECFFBEFF8E15EA8AC1BF46FC47" box="[160,267,960,985]" pageId="37" pageNumber="807" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="8BA9614EFFBEFF8E15EA8AE5BF7DFB81" blockId="37.[160,1155,960,1055]" pageId="37" pageNumber="807">
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<materialsCitation id="3B7E6B13FFBEFF8E15EA8AE5BF7DFB81" accessionNumber="ZMB 29089" collectionCode="ZMB" country="Ethiopia" location="Dscheffedenza" municipality="Shoa'" pageId="37" pageNumber="807" specimenCount="1" typeStatus="holotype">
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||
<collectionCode id="ED07F98BFFBEFF8E15EA8AE5BE9EFC62" box="[160,211,996,1020]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="37" pageNumber="807" type="Museum">ZMB</collectionCode>
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29089, from ‘
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<location id="8EC93795FFBEFF8E14CF8AE5BC67FC62" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBEFF8E14CF8AE5BC67FC62" box="[389,554,996,1020]" country="Ethiopia" municipality="Shoa'" name="Dscheffedenza" pageId="37" pageNumber="807">Dscheffedenza</location>
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[=Ciaffedenza],
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<collectingMunicipality id="6BCDFB34FFBEFF8E17BA8AE5BD61FC62" box="[752,812,996,1020]" pageId="37" pageNumber="807">Shoa’</collectingMunicipality>
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,
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<collectingCountry id="F30121DEFFBEFF8E16088AE5BDD3FC62" box="[834,926,996,1020]" name="Ethiopia" pageId="37" pageNumber="807">Ethiopia</collectingCountry>
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(for comment see distribution).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C30C32C5FFBEFF8D15EA8D58BA0BF919" lastPageId="38" lastPageNumber="808" pageId="37" pageNumber="807" type="description">
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<paragraph id="8BA9614EFFBEFF8E15EA8D58BF6BFBEC" blockId="37.[160,1156,1113,1727]" box="[160,294,1113,1138]" pageId="37" pageNumber="807">
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<emphasis id="B962BD5CFFBEFF8E15EA8D58BF6BFBEC" bold="true" box="[160,294,1113,1138]" italics="true" pageId="37" pageNumber="807">Description</emphasis>
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</paragraph>
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<paragraph id="8BA9614EFFBEFF8E15EA8D7CBF6CF921" blockId="37.[160,1156,1113,1727]" pageId="37" pageNumber="807">
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A small species of the genus with a total length below
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<quantity id="4CEECCABFFBEFF8E16088D7CBDE9FB0B" box="[834,932,1149,1173]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" pageId="37" pageNumber="807" unit="mm" value="290.0">290 mm</quantity>
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(SVL:
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<quantity id="4CEECCABFFBEFF8E16BB8D7CBACDFB0B" box="[1009,1152,1148,1173]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.17" metricValueMin="0.93" pageId="37" pageNumber="807" unit="mm" value="105.0" valueMax="117.0" valueMin="93.0">93–117 mm</quantity>
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, x =
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<quantity id="4CEECCABFFBEFF8E15978D9EBF16FB29" box="[221,347,1183,1207]" metricMagnitude="-1" metricUnit="m" metricValue="1.082" pageId="37" pageNumber="807" unit="mm" value="108.2">108.2 mm</quantity>
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, n = 3), tail only one fourth longer than SVL, shorter than in
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<taxonomicName id="4C161ACDFFBEFF8E15EA8DC3BF03FB44" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[160,334,1218,1242]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="807" phylum="Chordata" rank="species" species="ugandaensis">
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<emphasis id="B962BD5CFFBEFF8E15EA8DC3BF03FB44" box="[160,334,1218,1242]" italics="true" pageId="37" pageNumber="807">A. ugandaensis</emphasis>
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</taxonomicName>
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or
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<taxonomicName id="4C161ACDFFBEFF8E14368DC3BC44FB44" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[380,521,1218,1242]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="807" phylum="Chordata" rank="species" species="kiwuensis">
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<emphasis id="B962BD5CFFBEFF8E14368DC3BC44FB44" box="[380,521,1218,1242]" italics="true" pageId="37" pageNumber="807">A. kiwuensis</emphasis>
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</taxonomicName>
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. Both
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<bibRefCitation id="EF871CBFFFBEFF8E17118DC3BD6EFB44" author="Loveridge A" box="[603,803,1218,1242]" pageId="37" pageNumber="807" pagination="131 - 167" refId="ref20689" refString="Loveridge A. 1920. Notes on East African lizards collected 1915 - 1919, with the description of a new genus and species of skink and new subspecies of gecko. Proc Zool Soc London. 1920: 131 - 167." type="journal article" year="1920">Loveridge (1920)</bibRefCitation>
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and
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<bibRefCitation id="EF871CBFFFBEFF8E162F8DC3BA79FB44" author="Klausewitz W" box="[869,1076,1218,1242]" pageId="37" pageNumber="807" pagination="157 - 174" refId="ref20422" refString="Klausewitz W. 1957. Eidonomische Untersuchungen uber die Rassenkreise Agama cyanogaster und Agama atricollis. Die Unterarten von Agama atricollis [Eidonomic studies about the species complex of Agama cyanogaster and Agama atricollis. The subspecies of Agama atricollis]. Senckenb Biol. 38: 157 - 174." type="journal article" year="1957">Klausewitz (1957)</bibRefCitation>
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noted the small size of this species. The head seems to be only moderately broad because of the small size, but in relation to SVL length it is broader than in other species of the genus. Ear openings as large as eyes, with the tympanum visible. Occipital scale lacking. Nostril slightly below the canthus rostralis. No vertebral crest. Scales arranged in 104–114 (x = 109.7, n = 3) rows around midbody and 72–85 (x = 79.7, n = 3) longitudinal rows along the vertebrae. Matrix scales small, rhomboidal and keeled, scattered with enlarged scales. These larger scales are not as large and distinct as in
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<taxonomicName id="4C161ACDFFBEFF8E17578CD6BCD8FA71" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[541,661,1495,1519]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="807" phylum="Chordata" rank="species" species="gregorii">
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<emphasis id="B962BD5CFFBEFF8E17578CD6BCD8FA71" box="[541,661,1495,1519]" italics="true" pageId="37" pageNumber="807">A. gregorii</emphasis>
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</taxonomicName>
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. Vertebral scales keeled, enlarged scales intermixed with matrix scales but distinct to the lateral scalation, not arranged in distinct vertebral bands or framed by scales which are larger than the scales of the centre of the band. Gular scales small, smooth and flat. Ventral scales small and smooth, in 71–75 (x = 73.7, n = 3) longitudinal rows. Males usually with two (sometimes three) continuous rows of 14–17 precloacal pores, which are lacking in females.
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</paragraph>
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<caption id="DF6931C6FFBDFF8D15EA8D36BF00FBF5" ID-DOI="http://doi.org/10.5281/zenodo.5174485" ID-Zenodo-Dep="5174485" httpUri="https://zenodo.org/record/5174485/files/figure.png" pageId="38" pageNumber="808" startId="38.[160,225,1079,1102]" targetBox="[178,1138,142,1048]" targetPageId="38">
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<paragraph id="8BA9614EFFBDFF8D15EA8D36BF00FBF5" blockId="38.[160,1155,1078,1131]" pageId="38" pageNumber="808">
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<emphasis id="B962BD5CFFBDFF8D15EA8D36BF46FBD0" bold="true" box="[160,267,1079,1102]" pageId="38" pageNumber="808">Figure 16.</emphasis>
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Distribution of
|
||
<taxonomicName id="4C161ACDFFBDFF8D14E58D37BCD2FBD3" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[431,671,1078,1101]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="808" phylum="Chordata" rank="species" species="kiwuensis">
|
||
<emphasis id="B962BD5CFFBDFF8D14E58D37BCD2FBD3" box="[431,671,1078,1101]" italics="true" pageId="38" pageNumber="808">Acanthocercus kiwuensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C161ACDFFBDFF8D17938D37BD21FBD3" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[729,876,1078,1101]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="808" phylum="Chordata" rank="species" species="ugandaensis">
|
||
<emphasis id="B962BD5CFFBDFF8D17938D37BD21FBD3" box="[729,876,1078,1101]" italics="true" pageId="38" pageNumber="808">A. ugandaensis</emphasis>
|
||
</taxonomicName>
|
||
. The numbers refer to the specific localities.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA9614EFFBDFF8D15EA8D79BFB0FA2C" blockId="38.[160,1157,1144,1458]" pageId="38" pageNumber="808">
|
||
<materialsCitation id="3B7E6B13FFBDFF8D15EA8D79BFB4FA2C" county="Okapi Faunal Reserve" latitude="1.52" location="Okapi Faunal Reserve" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" pageId="38" pageNumber="808" specimenCount="1" stateProvince="Kasese">
|
||
1 = without locality [
|
||
<geoCoordinate id="EE220789FFBDFF8D14148D79BFEBFB12" box="[350,422,1144,1164]" degrees="03.21" direction="north" orientation="latitude" pageId="38" pageNumber="808" precision="555" value="3.21">03.21°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE220789FFBDFF8D14FB8D79BFBBFB12" box="[433,502,1144,1164]" degrees="28.52" direction="east" orientation="longitude" pageId="38" pageNumber="808" precision="555" value="28.52">28.52°E</geoCoordinate>
|
||
]; 2 =
|
||
<collectingCounty id="62C819C2FFBDFF8D17618D79BCA0FB12" box="[555,749,1144,1164]" pageId="38" pageNumber="808">Okapi Faunal Reserve</collectingCounty>
|
||
, Kayumaga stream; 3 = Avakubi,
|
||
<collectingMunicipality id="6BCDFB34FFBDFF8D11548D79BACCFB12" box="[1054,1153,1144,1164]" pageId="38" pageNumber="808">Ituri Forest</collectingMunicipality>
|
||
; 4 = Epulu; 5 =
|
||
<location id="8EC93795FFBDFF8D147E8D92BFB4FB39" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D147E8D92BFB4FB39" box="[308,505,1171,1191]" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Okapi Faunal Reserve" pageId="38" pageNumber="808" stateProvince="Kasese">Okapi Faunal Reserve</location>
|
||
; 6 = without locality [
|
||
<geoCoordinate id="EE220789FFBDFF8D17838D92BD4BFB39" box="[713,774,1171,1191]" degrees="1.12" direction="north" orientation="latitude" pageId="38" pageNumber="808" precision="555" value="1.12">1.12°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE220789FFBDFF8D16588D92BD1AFB39" box="[786,855,1171,1191]" degrees="29.25" direction="east" orientation="longitude" pageId="38" pageNumber="808" precision="555" value="29.25">29.25°E</geoCoordinate>
|
||
]; 7 = Bunia; 8 =
|
||
<location id="8EC93795FFBDFF8D16B08D92BACCFB39" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D16B08D92BACCFB39" box="[1018,1153,1171,1191]" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Ndjuma Forest" pageId="38" pageNumber="808" stateProvince="Kasese">Ndjuma Forest</location>
|
||
; 9 = Kisanzi [village near Mt. Teye]; 10 = Hakitengya; 11 = Byumba; 12 =
|
||
<location id="8EC93795FFBDFF8D16118DAFBDF3FB5C" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D16118DAFBDF3FB5C" box="[859,958,1198,1218]" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Fort Portal" pageId="38" pageNumber="808" stateProvince="Kasese">Fort Portal</location>
|
||
; 13 =
|
||
<location id="8EC93795FFBDFF8D114A8DAFBACCFB5C" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D114A8DAFBACCFB5C" box="[1024,1153,1198,1218]" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Semliki Forest" pageId="38" pageNumber="808" stateProvince="Kasese">Semliki Forest</location>
|
||
; 14 = Nyakalengijo; 15 = Kyehundwe; 16 =
|
||
<location id="8EC93795FFBDFF8D176E8DC9BCEFFB42" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D176E8DC9BCEFFB42" box="[548,674,1224,1244]" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Jambo Village" pageId="38" pageNumber="808" stateProvince="Kasese">Jambo Village</location>
|
||
; 17 =
|
||
<collectingRegion id="49D2AFACFFBDFF8D17968DC9BD54FB42" box="[732,793,1224,1244]" country="Uganda" name="Kasese" pageId="38" pageNumber="808">Kasese</collectingRegion>
|
||
; 18 =
|
||
<location id="8EC93795FFBDFF8D161E8DC9BDB9FB42" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D161E8DC9BDB9FB42" box="[852,1012,1224,1244]" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Mount Ruwenzori" pageId="38" pageNumber="808" stateProvince="Kasese">Mount Ruwenzori</location>
|
||
; 19 = Butembo; 20 =
|
||
<location id="8EC93795FFBDFF8D15858DE2BF0CFB69" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D15858DE2BF0CFB69" box="[207,321,1251,1271]" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Lake Edward" pageId="38" pageNumber="808" stateProvince="Kasese">Lake Edward</location>
|
||
; 21 = Kazinga channel; 22 = Kichwamba; 23 =
|
||
<collectingRegion id="49D2AFACFFBDFF8D17AA8DE2BD52FB69" box="[736,799,1251,1271]" country="Uganda" name="Ibanda" pageId="38" pageNumber="808">Ibanda</collectingRegion>
|
||
; 24 = Kabasha Escarpment; 25 =
|
||
<collectingRegion id="49D2AFACFFBDFF8D110E8DE2BACCFB69" box="[1092,1153,1251,1271]" country="Uganda" name="Kasese" pageId="38" pageNumber="808">Kasese</collectingRegion>
|
||
; 26 = Nyakabande; 27 = near Goma; 28 = Gisenyi [Rutshuru]; 29 = Buyumba; 30 = Isale; 31 =
|
||
<location id="8EC93795FFBDFF8D16B48DFFBE83FAB2" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D16B48DFFBE83FAB2" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Upper Mulinga River" pageId="38" pageNumber="808" stateProvince="Kasese">Upper Mulinga River</location>
|
||
,
|
||
<location id="8EC93795FFBDFF8D159C8C19BF70FAB2" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D159C8C19BF70FAB2" box="[214,317,1304,1324]" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Idjwi Island" pageId="38" pageNumber="808" stateProvince="Kasese">Idjwi Island</location>
|
||
; 32 = Kibuye; 33 = Bukavu; 34 = Cyamudongo; 35 =
|
||
<collectingRegion id="49D2AFACFFBDFF8D165C8C19BD34FAB2" box="[790,889,1304,1324]" country="Burundi" name="Bujumbura Mairie" pageId="38" pageNumber="808">Bujumbura</collectingRegion>
|
||
; 36 = Kikonda; 37 =
|
||
<location id="8EC93795FFBDFF8D11798C19BE95FAD9" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D11798C19BE95FAD9" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Budongo Forest" pageId="38" pageNumber="808" stateProvince="Kasese">Budongo Forest</location>
|
||
; 38 = Masinde; 39 = Kabulamuliro; 40 = Entebbe; 41 =
|
||
<collectingRegion id="49D2AFACFFBDFF8D17B48C32BD01FAD9" box="[766,844,1331,1351]" country="Uganda" name="Kampala" pageId="38" pageNumber="808">Kampala</collectingRegion>
|
||
; 42 =
|
||
<location id="8EC93795FFBDFF8D16D88C32BA5CFAD9" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D16D88C32BA5CFAD9" box="[914,1041,1331,1351]" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Mabira Forest" pageId="38" pageNumber="808" stateProvince="Kasese">Mabira Forest</location>
|
||
; 43 =
|
||
<collectingRegion id="49D2AFACFFBDFF8D111D8C32BACCFAD9" box="[1111,1153,1331,1351]" country="Uganda" name="Jinja" pageId="38" pageNumber="808">Jinja</collectingRegion>
|
||
; 44 = Ntotoro; 45 = Osukuru; 46 = Elgonyi; 47 = Kitale; 48 = Kisere; 49 = Kamwega; 50 =
|
||
<collectingRegion id="49D2AFACFFBDFF8D16AF8C4FBA0DFAFC" box="[997,1088,1358,1378]" country="Kenya" name="Kakamega" pageId="38" pageNumber="808">Kakamega</collectingRegion>
|
||
Forest; 51 =
|
||
<location id="8EC93795FFBDFF8D15858C69BF12FAE2" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D15858C69BF12FAE2" box="[207,351,1384,1404]" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Buyangu Village" pageId="38" pageNumber="808" stateProvince="Kasese">Buyangu Village</location>
|
||
; 52 = Kaimosi; 53 =
|
||
<collectingRegion id="49D2AFACFFBDFF8D175E8C69BC09FAE2" box="[532,580,1384,1404]" country="Kenya" name="Siaya" pageId="38" pageNumber="808">Siaya</collectingRegion>
|
||
; 54 = Ngiya; 55 =
|
||
<collectingRegion id="49D2AFACFFBDFF8D17AC8C69BD64FAE2" box="[742,809,1384,1404]" country="Kenya" name="Kisumu" pageId="38" pageNumber="808">Kisumu</collectingRegion>
|
||
; 56 = Lumbwa; 57 =
|
||
<collectingRegion id="49D2AFACFFBDFF8D16AC8C69BA6CFAE2" box="[998,1057,1384,1404]" country="Kenya" name="Migori" pageId="38" pageNumber="808">Migori</collectingRegion>
|
||
; 58 =
|
||
<location id="8EC93795FFBDFF8D11128C69BE98FA09" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D11128C69BE98FA09" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Loita Plains" pageId="38" pageNumber="808" stateProvince="Kasese">Loita Plains</location>
|
||
; 59 =
|
||
<collectingRegion id="49D2AFACFFBDFF8D14448C82BF1DFA09" box="[270,336,1411,1431]" country="Kenya" name="Nakuru" pageId="38" pageNumber="808">Nakuru</collectingRegion>
|
||
; 60 =
|
||
<collectingRegion id="49D2AFACFFBDFF8D14C28C82BF8AFA09" box="[392,455,1411,1431]" country="Kenya" name="Nakuru" pageId="38" pageNumber="808">Nakuru</collectingRegion>
|
||
National Park; 61 =
|
||
<location id="8EC93795FFBDFF8D17348C82BCB3FA09" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBDFF8D17348C82BCB3FA09" box="[638,766,1411,1431]" county="Okapi Faunal Reserve" latitude="1.52" longLatPrecision="785" longitude="28.59" municipality="Ituri Forest" name="Lake Naivasha" pageId="38" pageNumber="808" stateProvince="Kasese">Lake Naivasha</location>
|
||
; 62 = Thika; 63 = Mount
|
||
<collectingRegion id="49D2AFACFFBDFF8D16AA8C82BA5DFA09" box="[992,1040,1411,1431]" country="Kenya" name="Meru" pageId="38" pageNumber="808">Meru</collectingRegion>
|
||
; 64 = Kitosh; 65 = without locality [
|
||
<geoCoordinate id="EE220789FFBDFF8D14238C9FBFEBFA2C" box="[361,422,1438,1458]" degrees="1.52" direction="north" orientation="latitude" pageId="38" pageNumber="808" precision="555" value="1.52">1.52°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE220789FFBDFF8D14FA8C9FBFB8FA2C" box="[432,501,1438,1458]" degrees="28.59" direction="east" orientation="longitude" pageId="38" pageNumber="808" precision="555" value="28.59">28.59°E</geoCoordinate>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA9614EFFBDFF8D158A8F2BBA0BF919" blockId="38.[160,1156,1578,1671]" pageId="38" pageNumber="808">
|
||
There is no information about life coloration available. According to
|
||
<bibRefCitation id="EF871CBFFFBDFF8D168A8F2BBAC9F9DC" author="Klausewitz W" box="[960,1156,1578,1602]" pageId="38" pageNumber="808" pagination="157 - 174" refId="ref20422" refString="Klausewitz W. 1957. Eidonomische Untersuchungen uber die Rassenkreise Agama cyanogaster und Agama atricollis. Die Unterarten von Agama atricollis [Eidonomic studies about the species complex of Agama cyanogaster and Agama atricollis. The subspecies of Agama atricollis]. Senckenb Biol. 38: 157 - 174." type="journal article" year="1957">Klausewitz (1957)</bibRefCitation>
|
||
this species, like
|
||
<taxonomicName id="4C161ACDFFBDFF8D14158F4DBC48F9FA" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[351,517,1612,1636]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="808" phylum="Chordata" rank="species" species="ugandaensis">
|
||
<emphasis id="B962BD5CFFBDFF8D14158F4DBC48F9FA" box="[351,517,1612,1636]" italics="true" pageId="38" pageNumber="808">A. ugandaensis</emphasis>
|
||
</taxonomicName>
|
||
, is uniform brownish with a reticulated blue pattern on the throat. However, this description seems to be based on preserved specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6931C6FFBCFF8C15EA8C8BBA63FA3F" ID-DOI="http://doi.org/10.5281/zenodo.5174487" ID-Zenodo-Dep="5174487" box="[160,1070,1417,1441]" httpUri="https://zenodo.org/record/5174487/files/figure.png" pageId="39" pageNumber="809" startId="39.[160,225,1418,1441]" targetBox="[178,1138,142,1387]" targetPageId="39">
|
||
<paragraph id="8BA9614EFFBCFF8C15EA8C8BBA63FA3F" blockId="39.[160,1070,1417,1441]" box="[160,1070,1417,1441]" pageId="39" pageNumber="809">
|
||
<emphasis id="B962BD5CFFBCFF8C15EA8C8BBF44FA3F" bold="true" box="[160,265,1418,1441]" pageId="39" pageNumber="809">Figure 17.</emphasis>
|
||
Holotype (ZMB 29089) of
|
||
<taxonomicName id="4C161ACDFFBCFF8C175C8C88BCB8FA3E" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[534,757,1417,1440]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="809" phylum="Chordata" rank="species" species="minutus">
|
||
<emphasis id="B962BD5CFFBCFF8C175C8C88BCB8FA3E" box="[534,757,1417,1440]" italics="true" pageId="39" pageNumber="809">Acanthocercus minutus</emphasis>
|
||
</taxonomicName>
|
||
, from Dscheffedenza, Ethiopia.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30C32C5FFBCFF8315EA8CF2BDBEFE41" lastPageId="40" lastPageNumber="810" pageId="39" pageNumber="809" type="distribution">
|
||
<paragraph id="8BA9614EFFBCFF8C15EA8CF2BF61F992" blockId="39.[160,1155,1523,1687]" box="[160,300,1523,1548]" pageId="39" pageNumber="809">
|
||
<emphasis id="B962BD5CFFBCFF8C15EA8CF2BF61F992" bold="true" box="[160,300,1523,1548]" italics="true" pageId="39" pageNumber="809">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA9614EFFBCFF8C15EA8F16BFC0F909" blockId="39.[160,1155,1523,1687]" pageId="39" pageNumber="809">
|
||
<taxonomicName id="4C161ACDFFBCFF8C15EA8F16BF59F9B1" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[160,276,1559,1583]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="809" phylum="Chordata" rank="species" species="minutus">
|
||
<emphasis id="B962BD5CFFBCFF8C15EA8F16BF59F9B1" box="[160,276,1559,1583]" italics="true" pageId="39" pageNumber="809">A. minutus</emphasis>
|
||
</taxonomicName>
|
||
is mainly restricted to
|
||
<collectingCountry id="F30121DEFFBCFF8C175E8F16BC3DF9B1" box="[532,624,1559,1583]" name="Ethiopia" pageId="39" pageNumber="809">Ethiopia</collectingCountry>
|
||
with few records from
|
||
<collectingCountry id="F30121DEFFBCFF8C163C8F16BDF7F9B1" box="[886,954,1559,1583]" name="Kenya" pageId="39" pageNumber="809">Kenya</collectingCountry>
|
||
(e.g. adult female from Nairobi mentioned by
|
||
<bibRefCitation id="EF871CBFFFBCFF8C14AC8F38BCD0F9CF" author="Klausewitz W" box="[486,669,1593,1617]" pageId="39" pageNumber="809" pagination="157 - 174" refId="ref20422" refString="Klausewitz W. 1957. Eidonomische Untersuchungen uber die Rassenkreise Agama cyanogaster und Agama atricollis. Die Unterarten von Agama atricollis [Eidonomic studies about the species complex of Agama cyanogaster and Agama atricollis. The subspecies of Agama atricollis]. Senckenb Biol. 38: 157 - 174." type="journal article" year="1957">Klausewitz 1957</bibRefCitation>
|
||
,
|
||
<figureCitation id="132D7DCBFFBCFF8C17E78F38BD55F9CF" box="[685,792,1593,1617]" captionStart="Figure 14" captionStartId="34.[160,225,1390,1413]" captionTargetBox="[178,1138,195,1358]" captionTargetId="figure-2@34.[178,1138,195,1359]" captionTargetPageId="34" captionText="Figure 14. Distribution of Acanthocercus gregorii and A. minutus. The numbers refer to the specific localities." figureDoi="http://doi.org/10.5281/zenodo.5174481" httpUri="https://zenodo.org/record/5174481/files/figure.png" pageId="39" pageNumber="809">Figure 14</figureCitation>
|
||
).
|
||
<materialsCitation id="3B7E6B13FFBCFF8C167A8F38BFC4F909" county="Addis Ababa" location="Dscheffedenza" municipality="Ciaffedenza'" pageId="39" pageNumber="809" specimenCount="1">
|
||
The published
|
||
<typeStatus id="54ADDFECFFBCFF8C16978F38BA42F9CF" box="[989,1039,1593,1617]" pageId="39" pageNumber="809">type</typeStatus>
|
||
locality is
|
||
<location id="8EC93795FFBCFF8C15EA8F5DBF08F9EA" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBCFF8C15EA8F5DBF08F9EA" box="[160,325,1628,1652]" county="Addis Ababa" municipality="Ciaffedenza'" name="Dscheffedenza" pageId="39" pageNumber="809">Dscheffedenza</location>
|
||
in ‘
|
||
<location id="8EC93795FFBCFF8C143A8F5DBFE0F9EA" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBCFF8C143A8F5DBFE0F9EA" box="[368,429,1628,1652]" county="Addis Ababa" municipality="Ciaffedenza'" name="Shoa'" pageId="39" pageNumber="809">Shoa’</location>
|
||
, which seems to be the
|
||
<location id="8EC93795FFBCFF8C178D8F5DBD6DF9EA" LSID="urn:lsid:plazi:treatment:03BFD058FFBEFF83143C8BC0BDBEFE41:8EC93795FFBCFF8C178D8F5DBD6DF9EA" box="[711,800,1628,1652]" county="Addis Ababa" municipality="Ciaffedenza'" name="German" pageId="39" pageNumber="809">German</location>
|
||
name for ‘
|
||
<collectingMunicipality id="6BCDFB34FFBCFF8C16D48F5DBA6AF9EA" box="[926,1063,1628,1652]" pageId="39" pageNumber="809">Ciaffedenza’</collectingMunicipality>
|
||
, a town east of
|
||
<collectingCounty id="62C819C2FFBCFF8C15BC8F7EBFC4F909" box="[246,393,1663,1687]" pageId="39" pageNumber="809">Addis Ababa</collectingCounty>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA9614EFFB3FF8315EA898FBFD8FF39" blockId="40.[160,1156,142,479]" box="[160,405,142,167]" pageId="40" pageNumber="810">
|
||
<emphasis id="B962BD5CFFB3FF8315EA898FBFD8FF39" bold="true" box="[160,405,142,167]" italics="true" pageId="40" pageNumber="810">Habitat and ecology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA9614EFFB3FF8315EA89B3BDBEFE41" blockId="40.[160,1156,142,479]" pageId="40" pageNumber="810">
|
||
Little information about habitat or ecology is available. Within a species account on
|
||
<taxonomicName id="4C161ACDFFB3FF8315EA89D5BC15FF72" authority=", Largen and Spawls (2010)" authorityName=", Largen and Spawls" authorityYear="2010" box="[160,600,212,236]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="810" phylum="Chordata" rank="species" species="atricollis">
|
||
<emphasis id="B962BD5CFFB3FF8315EA89D5BF57FF72" box="[160,282,212,236]" italics="true" pageId="40" pageNumber="810">A. atricollis</emphasis>
|
||
,
|
||
<bibRefCitation id="EF871CBFFFB3FF83146189D5BC15FF72" author="Largen MJ & Spawls S" box="[299,600,212,236]" pageId="40" pageNumber="810" refId="ref20508" refString="Largen MJ, Spawls S. 2010. Amphibians and reptiles of ethiopia and eritrea. Frankfurt: Edition Chimaira; p. 694." type="journal volume" year="2010">Largen and Spawls (2010)</bibRefCitation>
|
||
</taxonomicName>
|
||
mentioned Ethiopian populations of small, frequently less colourful and ground-living specimens which probably refer to
|
||
<taxonomicName id="4C161ACDFFB3FF83114289F6BA30FE91" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[1032,1149,247,271]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="810" phylum="Chordata" rank="species" species="minutus">
|
||
<emphasis id="B962BD5CFFB3FF83114289F6BA30FE91" box="[1032,1149,247,271]" italics="true" pageId="40" pageNumber="810">A. minutus</emphasis>
|
||
</taxonomicName>
|
||
. These populations are distributed within an altitudinal range of
|
||
<quantity id="4CEECCABFFB3FF8316C1881BBA6BFEAC" box="[907,1062,281,306]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="40" pageNumber="810" unit="m" value="2250.0" valueMax="3000.0" valueMin="1500.0">1500–3000 m</quantity>
|
||
asl and inhabit dry savannah and woodland habitats.
|
||
<bibRefCitation id="EF871CBFFFB3FF831789883DBDA1FECA" author="Largen MJ & Spawls S" box="[707,1004,316,340]" pageId="40" pageNumber="810" pagination="21 - 109" refId="ref20473" refString="Largen MJ, Spawls S. 2006. Lizards of Ethiopia (Reptilia Sauria): an annotated checklist, bibliography, gazetteer and identification. Trop Zool. 19: 21 - 109." type="journal article" year="2006">Largen and Spawls (2006</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF871CBFFFB3FF8316B4883DBA78FECA" author="Largen MJ & Spawls S" box="[1022,1077,316,340]" pageId="40" pageNumber="810" refId="ref20508" refString="Largen MJ, Spawls S. 2010. Amphibians and reptiles of ethiopia and eritrea. Frankfurt: Edition Chimaira; p. 694." type="journal volume" year="2010">2010</bibRefCitation>
|
||
) mentioned that they do not show affinity for trees and that the specimens were most frequently encountered in open environments in holes or beneath rocks. However, they only mentioned that there was uncertainty about which name to apply to these populations, but did not give citations or mention particular taxon names.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |