465 lines
54 KiB
XML
465 lines
54 KiB
XML
<document id="F845BC9B9A5E71A6EFACBAC52144FE48" ID-DOI="10.5852/ejt.2021.758.1423" ID-GBIF-Dataset="de9c0c06-0e7d-44a6-a582-d41a346233f6" ID-ISSN="2118-9773" ID-Zenodo-Dep="5075824" ID-ZooBank="01BBC12C-E715-4393-A9F6-6EA85CB1289F" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1625579995989" checkinUser="felipe" docAuthor="Wesener, Thomas & Sagorny, Christina" docDate="2021" docId="03BF4B62FFAFE0271C0CFEF38716F81C" docLanguage="en" docName="ejt-758_1-48.pdf" docOrigin="European Journal of Taxonomy 758 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.758.1423" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zoosphaerium coquerelianum" docType="key" docVersion="10" lastPageNumber="20" masterDocId="FF86331AFFBEE0341E06FFFB8261FFB5" masterDocTitle="Seven new giant pill-millipede species and numerous new records of the genus Zoosphaerium from Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="18" updateTime="1733759775900" updateUser="felipe" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CB1109306B7730DC133C4C4FA7D52EA7">Seven new giant pill-millipede species and numerous new records of the genus Zoosphaerium from Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae)</mods:title>
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<mods:namePart id="AB089BD13C894B65E83CC1021717C5F3">Wesener, Thomas</mods:namePart>
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<mods:affiliation id="44F8B6719354112BF582B87C2AD99A5C">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="141B1A0D939A768A15664ED5228F16A8" type="email">t.wesener@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:namePart id="5724A0EB740B75FF6308EA3FDEFC2AF8">Sagorny, Christina</mods:namePart>
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<mods:affiliation id="2C271D220987DCF2F257A7E85BFA297F">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany. & University of Bonn, Institute of Evolutionary Biology and Ecology, D- 53121 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="2B587D57B19DC647B1903D67B75EE588" type="email">csagorny@evolution.uni-bonn.de</mods:nameIdentifier>
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<mods:title id="DB655CFB8C46DFA5E795AB56578DAE54">European Journal of Taxonomy</mods:title>
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<mods:date id="3D6B083CA3CF9F5A2AE5DD9A585E9D75">2021</mods:date>
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<treatment id="03BF4B62FFAFE0271C0CFEF38716F81C" ID-DOI="http://doi.org/10.5281/zenodo.5076198" ID-GBIF-Taxon="183392509" ID-Zenodo-Dep="5076198" LSID="urn:lsid:plazi:treatment:03BF4B62FFAFE0271C0CFEF38716F81C" httpUri="http://treatment.plazi.org/id/03BF4B62FFAFE0271C0CFEF38716F81C" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" scope_class="Diplopoda" scope_family="Arthrosphaeridae" scope_order="Sphaerotheriida">
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<subSubSection id="C30CA9FFFFAFE0251C0CFEF3864BFE97" box="[522,1066,264,291]" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="8BA9FA74FFAFE0251C0CFEF3864BFE97" blockId="17.[522,1066,264,291]" box="[522,1066,264,291]" pageId="17" pageNumber="18">
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<heading id="D0E14D18FFAFE0251C0CFEF3864BFE97" box="[522,1066,264,291]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
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<emphasis id="B9622666FFAFE0251C0CFEF3864BFE97" bold="true" box="[522,1066,264,291]" pageId="17" pageNumber="18">
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<taxonomicName id="4C1681F7FFAFE0251C0CFEF38119FE96" ID-CoL="5D7GQ" authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[522,888,264,291]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="coquerelianum">
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<emphasis id="B9622666FFAFE0251C0CFEF38119FE96" bold="true" box="[522,888,264,291]" italics="true" pageId="17" pageNumber="18">Zoosphaerium coquerelianum</emphasis>
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</taxonomicName>
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species-group
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C30CA9FFFFAFE0251EBBFEA281D1FCB9" pageId="17" pageNumber="18" type="discussion">
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<paragraph id="8BA9FA74FFAFE0251EBBFEA2834EFEC6" blockId="17.[189,303,345,371]" box="[189,303,345,371]" pageId="17" pageNumber="18">
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<heading id="D0E14D18FFAFE0251EBBFEA2834EFEC6" bold="true" box="[189,303,345,371]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
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<emphasis id="B9622666FFAFE0251EBBFEA2834EFEC6" bold="true" box="[189,303,345,371]" pageId="17" pageNumber="18">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA9FA74FFAFE0251EBBFE7C8702FD0E" blockId="17.[189,1399,391,700]" pageId="17" pageNumber="18">
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Species can be identified as members of the
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<taxonomicName id="4C1681F7FFAFE0251D0FFE7C8610FE14" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[777,1137,391,417]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="coquerelianum">
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<emphasis id="B9622666FFAFE0251D0FFE7C8610FE14" box="[777,1137,391,417]" italics="true" pageId="17" pageNumber="18">Zoosphaerium coquerelianum</emphasis>
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</taxonomicName>
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species-group based on morphological characters of the posterior telopods (
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<bibRefCitation id="EF878785FFAFE0251D6CFE5086C6FE70" author="Wesener T. & Wagele J. W." box="[874,1191,427,453]" pageId="17" pageNumber="18" pagination="5 - 82" refId="ref28537" refString="Wesener T. & Wagele J. W. 2008. The giant pill-millipedes of Madagascar: revision of the genus Zoosphaerium (Myriapoda, Diplopoda, Sphaerotheriida). Zoosystema 30 (1): 5 - 82." type="journal article" year="2008">Wesener & Wägele 2008</bibRefCitation>
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; Wesener 2009). Telopoditomere 3 only carries small sclerotized teeth and is always slightly longer and>1.5 times as wide as the immovable finger. Similar telopod characters occur in some members of the
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<taxonomicName id="4C1681F7FFAFE0251AD5FE098717FDBE" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1902" box="[1235,1398,497,523]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="platylabum">
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<emphasis id="B9622666FFAFE0251AD5FE098717FDBE" box="[1235,1398,497,523]" italics="true" pageId="17" pageNumber="18">Z. platylabum</emphasis>
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</taxonomicName>
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species-group, but the locking carinae in this group are located close to the last pleurites, while they are positioned close to the anal shield margin in species of the
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<taxonomicName id="4C1681F7FFAFE0251DB3FDC386E4FDE4" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[949,1157,567,593]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="coquerelianum">
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<emphasis id="B9622666FFAFE0251DB3FDC386E4FDE4" box="[949,1157,567,593]" italics="true" pageId="17" pageNumber="18">Z. coquerelianum</emphasis>
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</taxonomicName>
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group. The speciesgroup now contains 23 described species distributed in humid to subhumid forests all over
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<collectingCountry id="F301BAE4FFAFE0251AE5FDA08712FDC0" box="[1251,1395,603,629]" name="Madagascar" pageId="17" pageNumber="18">Madagascar</collectingCountry>
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. Species of this group are conspicuously absent from the spiny forest ecosystem and the western dry forest (Wesener 2009;
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<bibRefCitation id="EF878785FFAFE0251FC0FD5980A0FD0E" author="Wesener T. & Bespalova I. & Sierwald P." box="[454,705,673,700]" pageId="17" pageNumber="18" pagination="133 - 161" refId="ref28826" refString="Wesener T., Bespalova I. & Sierwald P. 2010 b. Madagascar's living giants: discovery of five new species of endemic giant pill-millipedes from Madagascar (Diplopoda: Sphaerotheriida: Arthrosphaeridae: Zoosphaerium). African Invertebrates 51 (1): 133 - 161." type="journal article" year="2010">
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Wesener
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<emphasis id="B9622666FFAFE0251C34FD59800DFD0E" box="[562,620,673,699]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
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2010b
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</bibRefCitation>
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;
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||
<bibRefCitation id="EF878785FFAFE0251CCAFD5A819CFD0E" author="Sagorny C. & Wesener T." box="[716,1021,673,700]" pageId="17" pageNumber="18" pagination="273 - 294" refId="ref27608" refString="Sagorny C. & Wesener T. 2017. Two new giant pill-millipede species of the genus Zoosphaerium endemic to the Bemanevika area in northern Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae). Zootaxa 4263 (2): 273 - 294. https: // doi. org / 10.11646 / zootaxa. 4263.2.4" type="journal article" year="2017">Sagorny & Wesener 2017</bibRefCitation>
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;
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||
<bibRefCitation id="EF878785FFAFE0251A0EFD598736FD0E" author="Wesener T. & Anilkumar P. A." box="[1032,1367,673,700]" pageId="17" pageNumber="18" pagination="3 - 35" refId="ref28188" refString="Wesener T. & Anilkumar P. A. 2020. The millipedes collected by the Museum " La Specola " on Madagascar 1989 / 1991, with the description of three new species of giant pill-millipedes (Diplopoda, Sphaerotheriida, Arthrosphaeridae). In: Korsos Z. & Danyi L. (eds) Proceedings of the 18 th International Congress of Myriapodology, Budapest, Hungary. ZooKeys 930: 3 - 35. https: // doi. org / 10.3897 / zookeys. 930.47620" type="journal article" year="2020">Wesener & Anilkumar 2020</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BA9FA74FFAFE0251EBBFD0981D1FCB9" blockId="17.[189,944,753,780]" box="[189,944,753,780]" pageId="17" pageNumber="18">
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<heading id="D0E14D18FFAFE0251EBBFD0981D1FCB9" bold="true" box="[189,944,753,780]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
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<emphasis id="B9622666FFAFE0251EBBFD0981D1FCB9" bold="true" box="[189,944,753,780]" pageId="17" pageNumber="18">
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Key to the species of the
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<taxonomicName id="4C1681F7FFAFE0251FF7FD0A813EFCB9" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[497,863,753,780]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="coquerelianum">
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||
<emphasis id="B9622666FFAFE0251FF7FD0A813EFCB9" bold="true" box="[497,863,753,780]" italics="true" pageId="17" pageNumber="18">Zoosphaerium coquerelianum</emphasis>
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</taxonomicName>
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group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30CA9FFFFAFE0271EBBFCDB8716F81C" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="key">
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<key id="FF87AD45FFAFE0271EBBFCDB8716F81C" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
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<keyStep id="30E2E2D1FFAFE0251EBBFCDB8717FCEB" pageId="17" pageNumber="18">
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<paragraph id="8BA9FA74FFAFE0251EBBFCDB8716FC8F" blockId="17.[189,1400,800,1969]" box="[189,1399,800,827]" pageId="17" pageNumber="18">
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<keyLead id="30E75941FFAFE0251EBBFCDB8716FC8F" box="[189,1399,800,827]" pageId="17" pageNumber="18">1. Male harp with two to three stridulation ribs..................................................................................2</keyLead>
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</paragraph>
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<paragraph id="8BA9FA74FFAFE0251EC7FCBF8717FCEB" blockId="17.[189,1400,800,1969]" box="[193,1398,835,862]" pageId="17" pageNumber="18">
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<keyLead id="30E75941FFAFE0251EC7FCBF8717FCEB" box="[193,1398,835,862]" pageId="17" pageNumber="18">– Male harp with a single stridulation rib........................................................................................11</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E2E2D1FFAFE0251EBBFC768717FBE2" pageId="17" pageNumber="18">
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<paragraph id="8BA9FA74FFAFE0251EBBFC768716FC58" blockId="17.[189,1400,800,1969]" pageId="17" pageNumber="18">
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<keyLead id="30E75941FFAFE0251EBBFC768716FC58" pageId="17" pageNumber="18">
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2. Tergites glossy, anteriorly shiny light green, posteriorly dark green. Specimen more than
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<quantity id="4CEE5791FFAFE0251B27FC778716FC12" box="[1313,1399,908,935]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="mm" value="50.0">50 mm</quantity>
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long. Anal shield not well-rounded, with steep slope. Movable finger of posterior telopods 1.6 times as long as wide, apically with groove ......................................................
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||
<taxonomicName id="4C1681F7FFAFE0251A3BFC288716FC58" ID-CoL="5D7GB" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1085,1399,978,1005]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="amabile">
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||
<emphasis id="B9622666FFAFE0251A3BFC2886DEFC58" bold="true" box="[1085,1215,978,1005]" italics="true" pageId="17" pageNumber="18">Z. amabile</emphasis>
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Wesener, 2009
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</taxonomicName>
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</keyLead>
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||
</paragraph>
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<paragraph id="8BA9FA74FFAFE0251EC7FC0C8717FBE2" blockId="17.[189,1400,800,1969]" pageId="17" pageNumber="18">
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<keyLead id="30E75941FFAFE0251EC7FC0C8717FBE2" pageId="17" pageNumber="18">
|
||
– Coloration pattern different, or if similar then specimens less than
|
||
<quantity id="4CEE5791FFAFE0251A0EFC0D863DFBA4" box="[1032,1116,1014,1041]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="17" pageNumber="18" unit="mm" value="35.0">35 mm</quantity>
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||
long. Anal shield wellrounded or bell-shaped. Movable finger of posterior telopods 1.9–2.5 times as long as wide, without apical groove.........................................................................................................................3
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</keyLead>
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||
</paragraph>
|
||
</keyStep>
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||
<keyStep id="30E2E2D1FFAFE0251EBBFB7D8717FB76" pageId="17" pageNumber="18">
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<paragraph id="8BA9FA74FFAFE0251EBBFB7D8716FB15" blockId="17.[189,1400,800,1969]" box="[189,1399,1157,1184]" pageId="17" pageNumber="18">
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||
<keyLead id="30E75941FFAFE0251EBBFB7D8716FB15" box="[189,1399,1157,1184]" pageId="17" pageNumber="18">3. Antenna with four apical cones.......................................................................................................4</keyLead>
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</paragraph>
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<paragraph id="8BA9FA74FFAFE0251EC7FB528717FB76" blockId="17.[189,1400,800,1969]" box="[193,1398,1192,1219]" pageId="17" pageNumber="18">
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||
<keyLead id="30E75941FFAFE0251EC7FB528717FB76" box="[193,1398,1192,1219]" pageId="17" pageNumber="18">– Antenna with>10 apical cones......................................................................................................5</keyLead>
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||
</paragraph>
|
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</keyStep>
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<keyStep id="30E2E2D1FFAFE0251EBBFB098719FAC0" pageId="17" pageNumber="18">
|
||
<paragraph id="8BA9FA74FFAFE0251EBBFB098719FA9A" blockId="17.[189,1400,800,1969]" pageId="17" pageNumber="18">
|
||
<keyLead id="30E75941FFAFE0251EBBFB098719FA9A" pageId="17" pageNumber="18">
|
||
4. Anal shield surface covered with short hairs. Posterior locking carina of anal shield 2.5–3 times as long as anterior carina .............................................
|
||
<taxonomicName id="4C1681F7FFAFE0251D6FFAEE8719FA9A" ID-CoL="7GH2H" authority="(de Saussure & Zehntner, 1902)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1902" box="[873,1400,1300,1327]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="alluaudi">
|
||
<emphasis id="B9622666FFAFE0251D6FFAEE8191FA9A" bold="true" box="[873,1008,1300,1327]" italics="true" pageId="17" pageNumber="18">Z. alluaudi</emphasis>
|
||
(de
|
||
<bibRefCitation id="EF878785FFAFE0251A2CFAEE8711FA9A" author="Saussure H. de & Zehntner L." box="[1066,1392,1301,1327]" pageId="17" pageNumber="18" pagination="1 - 356" refId="ref27698" refString="Saussure H. de & Zehntner L. 1902. Myriapodes de Madagascar. In: Grandidier A. (ed.) Histoire physique, naturelle et politique de Madagascar 27 (53): i - viii, 1 - 356." type="journal article" year="1902">Saussure & Zehntner, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFAFE0251EC7FAC28719FAC0" blockId="17.[189,1400,800,1969]" pageId="17" pageNumber="18">
|
||
<keyLead id="30E75941FFAFE0251EC7FAC28719FAC0" pageId="17" pageNumber="18">
|
||
– Anal shield surface glabrous. Posterior locking carina of anal shield more than 3.5 times as long as anterior carina .............................................
|
||
<taxonomicName id="4C1681F7FFAFE0251D0DFAA78719FAC0" ID-CoL="5D7JF" authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[779,1400,1371,1398]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="voeltzkowianum">
|
||
<emphasis id="B9622666FFAFE0251D0DFAA78190FAC3" bold="true" box="[779,1009,1371,1398]" italics="true" pageId="17" pageNumber="18">Z. voeltzkowianum</emphasis>
|
||
(de
|
||
<bibRefCitation id="EF878785FFAFE0251A2CFAA08711FAC0" author="Saussure H. de & Zehntner L." box="[1066,1392,1371,1397]" pageId="17" pageNumber="18" pagination="1 - 12" refId="ref27664" refString="Saussure H. de & Zehntner L. 1897. Atlas de l'histoire naturelle des Myriapodes. Histoire physique, naturelle et politique de Madagascar 27 (53): 1 - 12." type="journal article" year="1897">Saussure & Zehntner, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFAFE0251EBBFA5F8717F9DB" pageId="17" pageNumber="18">
|
||
<paragraph id="8BA9FA74FFAFE0251EBBFA5F8716F99D" blockId="17.[189,1400,800,1969]" pageId="17" pageNumber="18">
|
||
<keyLead id="30E75941FFAFE0251EBBFA5F8716F99D" pageId="17" pageNumber="18">
|
||
5. Male harp with three stridulation ribs. Endotergum with three rows of marginal bristles, bristles slightly extending beyond tergite. Sclerotized teeth present on antennomeres 1–5. Movable finger of posterior telopods 2 times as long as wide. Posterior locking carina of anal shield four times as long as anterior carina. Male anal shield bell-shaped .......
|
||
<taxonomicName id="4C1681F7FFAFE0251A16F9F58716F99D" ID-CoL="5D7JB" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1040,1399,1549,1576]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="trichordum">
|
||
<emphasis id="B9622666FFAFE0251A16F9F586DDF99D" bold="true" box="[1040,1212,1549,1576]" italics="true" pageId="17" pageNumber="18">Z. trichordum</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFAFE0251EC7F9C98717F9DB" blockId="17.[189,1400,800,1969]" pageId="17" pageNumber="18">
|
||
<keyLead id="30E75941FFAFE0251EC7F9C98717F9DB" pageId="17" pageNumber="18">– Male harp with two stridulation ribs. Endotergum with one up to two rows of marginal bristles, bristles not reaching tergite margin. Sclerotized teeth only present up to antennomere 3 or 4......6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFAFE0251EBBF9668717F96E" pageId="17" pageNumber="18">
|
||
<paragraph id="8BA9FA74FFAFE0251EBBF9668716F90D" blockId="17.[189,1400,800,1969]" box="[189,1399,1693,1720]" pageId="17" pageNumber="18">
|
||
<keyLead id="30E75941FFAFE0251EBBF9668716F90D" box="[189,1399,1693,1720]" pageId="17" pageNumber="18">6. Only up to eight ventral spines on walking legs.............................................................................7</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFAFE0251EC7F93A8717F96E" blockId="17.[189,1400,800,1969]" box="[193,1398,1728,1755]" pageId="17" pageNumber="18">
|
||
<keyLead id="30E75941FFAFE0251EC7F93A8717F96E" box="[193,1398,1728,1755]" pageId="17" pageNumber="18">– At least nine ventral spines on walking legs.................................................................................8</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFAFE0251EBBF8F18716F805" pageId="17" pageNumber="18">
|
||
<paragraph id="8BA9FA74FFAFE0251EBBF8F18716F8F2" blockId="17.[189,1400,800,1969]" pageId="17" pageNumber="18">
|
||
<keyLead id="30E75941FFAFE0251EBBF8F18716F8F2" pageId="17" pageNumber="18">
|
||
7. Movable finger of posterior telopod thick, 2.5 times as long as wide. Leg 3 with an apical spine. On anal shield, second locking carina 2–3 times as long as first ............
|
||
<taxonomicName id="4C1681F7FFAFE0251A5EF8D6876CF8F2" ID-CoL="9BDGP" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[1112,1293,1836,1863]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ambatovaky" status="sp. nov.">
|
||
<emphasis id="B9622666FFAFE0251A5EF8D6876CF8F2" bold="true" box="[1112,1293,1836,1863]" italics="true" pageId="17" pageNumber="18">Z. ambatovaky</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFAFE0251B1FF8D68716F8F2" box="[1305,1399,1837,1863]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFAFE0251EC7F8AB8716F805" blockId="17.[189,1400,800,1969]" pageId="17" pageNumber="18">
|
||
<keyLead id="30E75941FFAFE0251EC7F8AB8716F805" pageId="17" pageNumber="18">
|
||
– Movable finger of posterior telopod very thick, 2 times as long as wide. Leg 3 without an apical spine. On anal shield, second locking carina 4–5 times as long as first .............................................. .........................................................................................
|
||
<taxonomicName id="4C1681F7FFAFE0251D71F86C8716F805" ID-CoL="5D7HK" authority="Wesener & Bespalova, 2010" authorityName="Wesener & Bespalova" authorityYear="2010" box="[887,1399,1942,1969]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="muscorum">
|
||
<emphasis id="B9622666FFAFE0251D71F86C867BF804" bold="true" box="[887,1050,1943,1969]" italics="true" pageId="17" pageNumber="18">Z. muscorum</emphasis>
|
||
<bibRefCitation id="EF878785FFAFE0251A25F86C8716F805" author="Wesener T. & Bespalova I. & Sierwald P." box="[1059,1399,1942,1969]" pageId="17" pageNumber="18" pagination="133 - 161" refId="ref28826" refString="Wesener T., Bespalova I. & Sierwald P. 2010 b. Madagascar's living giants: discovery of five new species of endemic giant pill-millipedes from Madagascar (Diplopoda: Sphaerotheriida: Arthrosphaeridae: Zoosphaerium). African Invertebrates 51 (1): 133 - 161." type="journal article" year="2010">Wesener & Bespalova, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFACE0261EBBFEF28716FE05" pageId="18" pageNumber="19">
|
||
<paragraph id="8BA9FA74FFACE0261EBBFEF28716FEDC" blockId="18.[189,1401,265,2002]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EBBFEF28716FEDC" pageId="18" pageNumber="19">
|
||
8. Male anal shield well-rounded, with slight bell shape. Posterior locking carina of anal shield six times as long as anterior carina. Up to 10 ventral spines on walking legs .............................. ....................................................................................
|
||
<taxonomicName id="4C1681F7FFACE0261D54FEAB8716FEDC" ID-CoL="5D7GK" authority="Wesener & Bespalova, 2010" authorityName="Wesener & Bespalova" authorityYear="2010" box="[850,1399,335,362]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bambusoides">
|
||
<emphasis id="B9622666FFACE0261D54FEAB8672FEDF" bold="true" box="[850,1043,335,362]" italics="true" pageId="18" pageNumber="19">Z. bambusoides</emphasis>
|
||
<bibRefCitation id="EF878785FFACE0261A18FEAB8716FEDC" author="Wesener T. & Bespalova I. & Sierwald P." box="[1054,1399,335,362]" pageId="18" pageNumber="19" pagination="133 - 161" refId="ref28826" refString="Wesener T., Bespalova I. & Sierwald P. 2010 b. Madagascar's living giants: discovery of five new species of endemic giant pill-millipedes from Madagascar (Diplopoda: Sphaerotheriida: Arthrosphaeridae: Zoosphaerium). African Invertebrates 51 (1): 133 - 161." type="journal article" year="2010">Wesener & Bespalova, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFACE0261EC7FE888716FE05" blockId="18.[189,1401,265,2002]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EC7FE888716FE05" pageId="18" pageNumber="19">– Male anal shield well-rounded. Posterior locking carina of anal shield three to four times as long as anterior carina. Usually more than 10 ventral spines on walking legs.............................9</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFACE0261EBBFE248716FD79" pageId="18" pageNumber="19">
|
||
<paragraph id="8BA9FA74FFACE0261EBBFE248716FDD7" blockId="18.[189,1401,265,2002]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EBBFE248716FDD7" pageId="18" pageNumber="19">
|
||
9. Leg 3 without an apical spine. Specimens only slightly more than
|
||
<quantity id="4CEE5791FFACE0261A2EFE24861EFE4C" box="[1064,1151,479,505]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="19" unit="mm" value="30.0">30 mm</quantity>
|
||
long. Tergites jadelike green with thin brown posterior margin, texture glossy. Operculum of vulva deeply notched, strongly extending beyond coxa. Apical disc with ca 14 apical cones ............................................... ..........................................................................................................
|
||
<taxonomicName id="4C1681F7FFACE0261DF2FDB28716FDD7" ID-CoL="5D7J2" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1012,1399,584,611]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="smaragdinum">
|
||
<emphasis id="B9622666FFACE0261DF2FDB286DFFDD6" bold="true" box="[1012,1214,584,611]" italics="true" pageId="18" pageNumber="19">Z. smaragdinum</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFACE0261EC7FD978716FD79" blockId="18.[189,1401,265,2002]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EC7FD978716FD79" pageId="18" pageNumber="19">
|
||
– Leg 3 with an apical spine. Specimens more than
|
||
<quantity id="4CEE5791FFACE0261D31FD9081EBFD33" box="[823,906,619,646]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="mm" value="50.0">50 mm</quantity>
|
||
long. Tergites never jade-like, not glossy. Operculum of vulva not protruding above coxal margin. Apical disc with more than 20 apical cones................................................................................................................................................10
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFACE0261EBBFD008716FC70" pageId="18" pageNumber="19">
|
||
<paragraph id="8BA9FA74FFACE0261EBBFD008716FCE9" blockId="18.[189,1401,265,2002]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EBBFD008716FCE9" pageId="18" pageNumber="19">
|
||
10. Specimens more than
|
||
<quantity id="4CEE5791FFACE0261C0DFD008006FCA0" box="[523,615,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="mm" value="60.0">60 mm</quantity>
|
||
long. Tergites dark-green, covered with numerous large pits and short setae. Sclerotized teeth present on antennomeres 1–3, disc with ca 45 apical cones. Endotergum with two rows of marginal bristles .........
|
||
<taxonomicName id="4C1681F7FFACE0261D8AFCB98716FCE9" ID-CoL="5D7JD" authority="Wesener & Sierwald, 2005" authorityName="Wesener & Sierwald" authorityYear="2005" box="[908,1399,833,860]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="villosum">
|
||
<emphasis id="B9622666FFACE0261D8AFCB9867BFCE9" bold="true" box="[908,1050,833,860]" italics="true" pageId="18" pageNumber="19">Z. villosum</emphasis>
|
||
Wesener & Sierwald, 2005
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFACE0261EC7FC9E8716FC70" blockId="18.[189,1401,265,2002]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EC7FC9E8716FC70" pageId="18" pageNumber="19">
|
||
– Specimens less than
|
||
<quantity id="4CEE5791FFACE0261FE9FC9F8029FCCA" box="[495,584,868,895]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="mm" value="60.0">60 mm</quantity>
|
||
long. Tergites light-brown, glabrous. Sclerotized teeth present on antennomeres 1–4, disc with 20–30 apical cones. Endotergum with a single row of marginal bristles ..............................................................................
|
||
<taxonomicName id="4C1681F7FFACE0261D82FC508716FC70" ID-CoL="84F5T" authority="Sagorny & Wesener, 2017" authorityName="Sagorny & Wesener" authorityYear="2017" box="[900,1399,938,965]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bemanevika">
|
||
<emphasis id="B9622666FFACE0261D82FC508656FC70" bold="true" box="[900,1079,938,965]" italics="true" pageId="18" pageNumber="19">Z. bemanevika</emphasis>
|
||
<bibRefCitation id="EF878785FFACE0261A39FC508716FC70" author="Sagorny C. & Wesener T." box="[1087,1399,939,965]" pageId="18" pageNumber="19" pagination="273 - 294" refId="ref27608" refString="Sagorny C. & Wesener T. 2017. Two new giant pill-millipede species of the genus Zoosphaerium endemic to the Bemanevika area in northern Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae). Zootaxa 4263 (2): 273 - 294. https: // doi. org / 10.11646 / zootaxa. 4263.2.4" type="journal article" year="2017">Sagorny & Wesener, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFACE0261EBBFC0F8716FB84" pageId="18" pageNumber="19">
|
||
<paragraph id="8BA9FA74FFACE0261EBBFC0F8716FBBB" blockId="18.[189,1401,265,2002]" box="[189,1399,1011,1038]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EBBFC0F8716FBBB" box="[189,1399,1011,1038]" pageId="18" pageNumber="19">
|
||
11. Movable finger 3 times as long as wide ................................................
|
||
<taxonomicName id="4C1681F7FFACE0261A2FFC0F8716FBBB" ID-CoL="5D7GC" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1065,1399,1011,1038]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ambrense">
|
||
<emphasis id="B9622666FFACE0261A2FFC0F86A1FBBB" bold="true" box="[1065,1216,1011,1038]" italics="true" pageId="18" pageNumber="19">Z. ambrense</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFACE0261EC7FBE38716FB84" blockId="18.[189,1401,265,2002]" box="[193,1399,1047,1074]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EC7FBE38716FB84" box="[193,1399,1047,1074]" pageId="18" pageNumber="19">– Movable finger <2.5 times as long as wide....................................................................................12</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFACE0261EBBFB9B8716FB51" pageId="18" pageNumber="19">
|
||
<paragraph id="8BA9FA74FFACE0261EBBFB9B8716FB28" blockId="18.[189,1401,265,2002]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EBBFB9B8716FB28" pageId="18" pageNumber="19">12. Antenna with four apical cones. Movable finger of posterior telopods at inner margin with two membranous lobes...........................................................................................................................13</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFACE0261EC7FB5C8716FB51" blockId="18.[189,1401,265,2002]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EC7FB5C8716FB51" pageId="18" pageNumber="19">– Antenna with>10 apical cones. Movable finger of posterior telopods at inner margin with a single membranous lobe............................................................................................................................17</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFACE0261EBBFAE88716FA0F" pageId="18" pageNumber="19">
|
||
<paragraph id="8BA9FA74FFACE0261EBBFAE88716FA23" blockId="18.[189,1401,265,2002]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EBBFAE88716FA23" pageId="18" pageNumber="19">
|
||
13. Process of second podomere of anterior telopod not visible in anterior view. Fifth antennomere with a field of sensilla basiconica. Collum with isolated, long setae. Endotergum with one row of large cuticular impressions and a second row of much smaller impressions; bristles long, strongly protruding above tergite ...........................................................
|
||
<taxonomicName id="4C1681F7FFACE0261A32FA868716FA23" ID-CoL="5D7GL" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1076,1399,1404,1431]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bilobum">
|
||
<emphasis id="B9622666FFACE0261A32FA8686DAFA22" bold="true" box="[1076,1211,1404,1431]" italics="true" pageId="18" pageNumber="19">Z. bilobum</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFACE0261EC7FA5B8716FA0F" blockId="18.[189,1401,265,2002]" box="[193,1399,1439,1466]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EC7FA5B8716FA0F" box="[193,1399,1439,1466]" pageId="18" pageNumber="19">– Process of second podomere of anterior telopod visible in anterior view.................................14</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFACE0261EBBFA128716F993" pageId="18" pageNumber="19">
|
||
<paragraph id="8BA9FA74FFACE0261EBBFA128717F9B6" blockId="18.[189,1401,265,2002]" box="[189,1398,1512,1539]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EBBFA128717F9B6" box="[189,1398,1512,1539]" pageId="18" pageNumber="19">14. Podomere 3 of anterior telopod with crenulated teeth...................................................................15</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFACE0261EC7F9F78716F993" blockId="18.[189,1401,265,2002]" box="[193,1399,1548,1574]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EC7F9F78716F993" box="[193,1399,1548,1574]" pageId="18" pageNumber="19">– Podomere 3 of anterior telopod without crenulated teeth.............................................................16</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFACE0261EBBF9AE8716F8F7" pageId="18" pageNumber="19">
|
||
<paragraph id="8BA9FA74FFACE0261EBBF9AE8716F96D" blockId="18.[189,1401,265,2002]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EBBF9AE8716F96D" pageId="18" pageNumber="19">
|
||
15. Collum glabrous, with a few setae at corners on either side of head. Endotergum with two rows of regularly distributed circular impressions, marginal bristles strongly protruding above tergite. Sensilla basiconica present on antennomeres 1 and 2. Anal shield weakly bellshaped ................................................................
|
||
<taxonomicName id="4C1681F7FFACE0261D24F9448716F96D" authority="Wesener & Bespalova, 2010" authorityName="Wesener & Bespalova" authorityYear="2010" box="[802,1399,1726,1753]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="trigrioculatum">
|
||
<emphasis id="B9622666FFACE0261D24F944819DF96C" bold="true" box="[802,1020,1726,1753]" italics="true" pageId="18" pageNumber="19">Z. trigrioculatum</emphasis>
|
||
<bibRefCitation id="EF878785FFACE0261A0BF9448716F96D" author="Wesener T. & Bespalova I. & Sierwald P." box="[1037,1399,1726,1753]" pageId="18" pageNumber="19" pagination="133 - 161" refId="ref28826" refString="Wesener T., Bespalova I. & Sierwald P. 2010 b. Madagascar's living giants: discovery of five new species of endemic giant pill-millipedes from Madagascar (Diplopoda: Sphaerotheriida: Arthrosphaeridae: Zoosphaerium). African Invertebrates 51 (1): 133 - 161." type="journal article" year="2010">Wesener & Bespalova, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFACE0261EC7F9198716F8F7" blockId="18.[189,1401,265,2002]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EC7F9198716F8F7" pageId="18" pageNumber="19">
|
||
– Collum glabrous, anterior margin with two rows of setae. Endotergum with a single row of elliptical cuticular impression, marginal bristles protruding to margin. Sensilla basiconica absent. Anal shield well rounded ..................................
|
||
<taxonomicName id="4C1681F7FFACE0261D7CF8D38716F8F7" authority="Anilkumar & Wesener, 2020" authorityName="Anilkumar & Wesener" authorityYear="2020" box="[890,1399,1831,1858]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bartolozzi">
|
||
<emphasis id="B9622666FFACE0261D7CF8D38673F8F7" bold="true" box="[890,1042,1831,1858]" italics="true" pageId="18" pageNumber="19">Z. bartolozzi</emphasis>
|
||
Anilkumar & Wesener, 2020
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFACE0271EBBF88A8716FEDC" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
|
||
<paragraph id="8BA9FA74FFACE0261EBBF88A8716F864" blockId="18.[189,1401,265,2002]" pageId="18" pageNumber="19">
|
||
<keyLead id="30E75941FFACE0261EBBF88A8716F864" pageId="18" pageNumber="19">
|
||
16. Collum glabrous. Endotergum with a single row of large cuticular impressions, marginal bristles slightly protruding above tergite margin. Second leg with 4 or 5 ventral spines. Anal shield tapering ........................................................................................................................
|
||
<taxonomicName id="4C1681F7FFACE0261A5CF8438716F864" ID-CoL="5D7HC" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1114,1399,1975,2002]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="isalo">
|
||
<emphasis id="B9622666FFACE0261A5CF84386D8F867" bold="true" box="[1114,1209,1975,2002]" italics="true" pageId="18" pageNumber="19">Z. isalo</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFADE0271EC7FEF28716FEDC" blockId="19.[189,1399,265,1961]" pageId="19" pageNumber="20">
|
||
<keyLead id="30E75941FFADE0271EC7FEF28716FEDC" pageId="19" pageNumber="20">
|
||
– Collum glabrous. Endotergum with a single row of slightly rounded elliptical cuticular impressions, marginal bristles protruding to margin. Second leg with 6 or 7 ventral spines. Anal shield well rounded ..................................................................
|
||
<taxonomicName id="4C1681F7FFADE0271DC2FEAB8716FEDC" authority="Anilkumar & Wesener, 2020" authorityName="Anilkumar & Wesener" authorityYear="2020" box="[964,1399,335,362]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="taiti">
|
||
<emphasis id="B9622666FFADE0271DC2FEAB8674FEDC" bold="true" box="[964,1045,335,362]" italics="true" pageId="19" pageNumber="20">Z. taiti</emphasis>
|
||
Anilkumar & Wesener, 2020
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFADE0271EBBFE638716FE63" pageId="19" pageNumber="20">
|
||
<paragraph id="8BA9FA74FFADE0271EBBFE638716FE07" blockId="19.[189,1399,265,1961]" box="[189,1399,408,435]" pageId="19" pageNumber="20">
|
||
<keyLead id="30E75941FFADE0271EBBFE638716FE07" box="[189,1399,408,435]" pageId="19" pageNumber="20">17. First pair of stigmatic plates greatly enlarged, with a well-rounded tip......................................18</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFADE0271EC7FE478716FE63" blockId="19.[189,1399,265,1961]" box="[193,1399,443,470]" pageId="19" pageNumber="20">
|
||
<keyLead id="30E75941FFADE0271EC7FE478716FE63" box="[193,1399,443,470]" pageId="19" pageNumber="20">– First pair of stigmatic plates triangular, tapering towards tip.......................................................19</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFADE0271EBBFDFE8716FCEE" pageId="19" pageNumber="20">
|
||
<paragraph id="8BA9FA74FFADE0271EBBFDFE8716FD1E" blockId="19.[189,1399,265,1961]" pageId="19" pageNumber="20">
|
||
<keyLead id="30E75941FFADE0271EBBFDFE8716FD1E" pageId="19" pageNumber="20">
|
||
18. Operculum of vulva ending clearly before coxal margin, deeply notched, divided into two large, well-rounded, similar-sized lobes. Third podomere of anterior telopod laterally with ca five large sclerotized teeth. Sensilla basiconica on first antennomere absent. Posterior locking carina of anal shield two times as long as anterior carina.Endotergal marginal bristles protruding up to tergite margin. Movable finger of posterior telopods thick, 1.9 times as long as wide ......
|
||
<taxonomicName id="4C1681F7FFADE0271A47FD698716FD1E" ID-CoL="5D7J8" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1089,1399,657,684]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tampolo">
|
||
<emphasis id="B9622666FFADE0271A47FD6986A2FD19" bold="true" box="[1089,1219,657,684]" italics="true" pageId="19" pageNumber="20">Z. tampolo</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFADE0271EC7FD4E8716FCEE" blockId="19.[189,1399,265,1961]" pageId="19" pageNumber="20">
|
||
<keyLead id="30E75941FFADE0271EC7FD4E8716FCEE" pageId="19" pageNumber="20">
|
||
– Operculum of vulva ending before coxal margin, elongated into a single median lobe, lobe only shallowly notched at tip. Third podomere of anterior telopod without any large sclerotized teeth. Sensilla basiconica on first antennomere present. Locking carinae of anal shield of equal length. Endotergal marginal bristles very short, ending far before tergite margin. Movable finger of posterior telopods thick, 2.1 times as long as wide ...............
|
||
<taxonomicName id="4C1681F7FFADE0271A33FCB98716FCEE" ID-CoL="5D7H9" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1077,1399,833,860]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ignotum">
|
||
<emphasis id="B9622666FFADE0271A33FCB986DCFCEE" bold="true" box="[1077,1213,833,860]" italics="true" pageId="19" pageNumber="20">Z. ignotum</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFADE0271EBBFC718717FBE1" pageId="19" pageNumber="20">
|
||
<paragraph id="8BA9FA74FFADE0271EBBFC718717FC5E" blockId="19.[189,1399,265,1961]" pageId="19" pageNumber="20">
|
||
<keyLead id="30E75941FFADE0271EBBFC718717FC5E" pageId="19" pageNumber="20">
|
||
19. Tergite texture orange-peel like. Collum covered with isolated, long hairs. Body size of adult female>
|
||
<quantity id="4CEE5791FFADE0271F5BFC5683DBFC7D" box="[349,442,941,968]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="20" unit="mm" value="50.0">50 mm</quantity>
|
||
. Endotergum with two rows of marginal bristles and a single row of large circular cuticular impressions........................................................................................................20
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFADE0271EC7FC0F8717FBE1" blockId="19.[189,1399,265,1961]" pageId="19" pageNumber="20">
|
||
<keyLead id="30E75941FFADE0271EC7FC0F8717FBE1" pageId="19" pageNumber="20">
|
||
– Tergite texture glossy or smooth. Collum glabrous. Body size of adults <
|
||
<quantity id="4CEE5791FFADE0271A7DFC0F86B7FBBB" box="[1147,1238,1012,1039]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="19" pageNumber="20" unit="mm" value="35.0">35 mm</quantity>
|
||
. Endotergum with a single row of marginal bristles and either without or with two rows of cuticular impressions.........................................................................................................................................22
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFADE0271EBBFB788717FAF8" pageId="19" pageNumber="20">
|
||
<paragraph id="8BA9FA74FFADE0271EBBFB788717FB51" blockId="19.[189,1399,265,1961]" pageId="19" pageNumber="20">
|
||
<keyLead id="30E75941FFADE0271EBBFB788717FB51" pageId="19" pageNumber="20">
|
||
20. Posterior locking carina on anal shield 2.5 times as long as anterior carina. Third podomere of anterior telopods without large sclerotized teeth. Anal shield glabrous ............................................ ...........................................................................
|
||
<taxonomicName id="4C1681F7FFADE0271D16FB318717FB51" ID-CoL="5D7GQ" authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[784,1398,1225,1252]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="coquerelianum">
|
||
<emphasis id="B9622666FFADE0271D16FB31818AFB51" bold="true" box="[784,1003,1225,1252]" italics="true" pageId="19" pageNumber="20">Z. coquerelianum</emphasis>
|
||
(de
|
||
<bibRefCitation id="EF878785FFADE0271A22FB31870FFB51" author="Saussure H. de & Zehntner L." box="[1060,1390,1226,1252]" pageId="19" pageNumber="20" pagination="1 - 12" refId="ref27664" refString="Saussure H. de & Zehntner L. 1897. Atlas de l'histoire naturelle des Myriapodes. Histoire physique, naturelle et politique de Madagascar 27 (53): 1 - 12." type="journal article" year="1897">Saussure & Zehntner, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFADE0271EC7FB168717FAF8" blockId="19.[189,1399,265,1961]" pageId="19" pageNumber="20">
|
||
<keyLead id="30E75941FFADE0271EC7FB168717FAF8" pageId="19" pageNumber="20">– Posterior locking carina on anal shield>4 times as long as anterior carina. Third podomere of anterior telopods with at least three large sclerotized teeth. Anal shield covered with short hair.............................................................................................................................................21</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFADE0271EBBFA878716F9F3" pageId="19" pageNumber="20">
|
||
<paragraph id="8BA9FA74FFADE0271EBBFA878716FA68" blockId="19.[189,1399,265,1961]" pageId="19" pageNumber="20">
|
||
<keyLead id="30E75941FFADE0271EBBFA878716FA68" pageId="19" pageNumber="20">
|
||
21. Third podomere on anterior telopods with 3 or 4 crenulated teeth. Endotergal marginal bristles very short, protruding up to ⅓ of distance to tergite margin. Tips of vulvar operculum widely separated .................................................................................................
|
||
<taxonomicName id="4C1681F7FFADE0271A2DFA388716FA68" ID-CoL="2f382a91-4a4d-4e52-9f40-9a6d4640968d" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[1067,1399,1474,1501]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="mangabe">
|
||
<emphasis id="B9622666FFADE0271A2DFA3886DCFA68" bold="true" box="[1067,1213,1474,1501]" italics="true" pageId="19" pageNumber="20">Z. mangabe</emphasis>
|
||
Wesener, 2020
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFADE0271EC7FA1D8716F9F3" blockId="19.[189,1399,265,1961]" pageId="19" pageNumber="20">
|
||
<keyLead id="30E75941FFADE0271EC7FA1D8716F9F3" pageId="19" pageNumber="20">
|
||
– Third podomere on anterior telopods with ca seven large sclerotized teeth. Endotergal marginal bristles protruding up to tergite margin. Tips of vulvar operculum fused ...................................... .............................................................................................................
|
||
<taxonomicName id="4C1681F7FFADE0271A0CF9D68716F9F3" ID-CoL="5D7J7" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1034,1399,1580,1607]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tainkintana">
|
||
<emphasis id="B9622666FFADE0271A0CF9D686DBF9F2" bold="true" box="[1034,1210,1580,1607]" italics="true" pageId="19" pageNumber="20">Z. tainkintana</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E2E2D1FFADE0271EBBF98E8716F81C" pageId="19" pageNumber="20">
|
||
<paragraph id="8BA9FA74FFADE0271EBBF98E8716F94C" blockId="19.[189,1399,265,1961]" pageId="19" pageNumber="20">
|
||
<keyLead id="30E75941FFADE0271EBBF98E8716F94C" pageId="19" pageNumber="20">
|
||
22. Tergites glossy. Female anal shield bell-shaped, in male with steep slope. Fixed finger of posterior telopods wide, only slightly curved towards movable finger. Movable finger 1.8 times as long as wide, on posterior side with ten small sclerotized teeth. Sclerotized teeth on antenna only present on first antennomere. Endotergum without cuticular impressions .........
|
||
<taxonomicName id="4C1681F7FFADE0271A03F9248716F94C" ID-CoL="7GGQS" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1029,1399,1758,1785]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="viridissimum">
|
||
<emphasis id="B9622666FFADE0271A03F92486A3F94C" bold="true" box="[1029,1218,1758,1785]" italics="true" pageId="19" pageNumber="20">Z. viridissimum</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFADE0271EC7F8F88716F81C" blockId="19.[189,1399,265,1961]" pageId="19" pageNumber="20">
|
||
<keyLead id="30E75941FFADE0271EC7F8F88716F81C" pageId="19" pageNumber="20">
|
||
– Tergites matte. Anal shield well-rounded. Fixed finger of posterior telopods slender, strongly curved towards movable finger. Movable finger 2.4 times as long as wide, on posterior side with ca 25 small sclerotized teeth. Sclerotized teeth present on antennomeres 1–3. Endotergum with a single row of large and a second row of smaller circular cuticular impressions ............... .........................................................................................................
|
||
<taxonomicName id="4C1681F7FFADE0271DEAF8748716F81C" ID-CoL="7GGPZ" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1004,1399,1934,1961]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="denticulatum">
|
||
<emphasis id="B9622666FFADE0271DEAF87486D5F81C" bold="true" box="[1004,1204,1934,1961]" italics="true" pageId="19" pageNumber="20">Z. denticulatum</emphasis>
|
||
Wesener, 2009
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |