300 lines
43 KiB
XML
300 lines
43 KiB
XML
<document id="19DC90BD7B33EFE6B1AB2F9D3A7961BA" ID-CLB-Dataset="5669" ID-DOI="10.5852/ejt.2021.758.1423" ID-GBIF-Dataset="de9c0c06-0e7d-44a6-a582-d41a346233f6" ID-ISSN="2118-9773" ID-Zenodo-Dep="5075824" ID-ZooBank="01BBC12C-E715-4393-A9F6-6EA85CB1289F" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625579995989" checkinUser="felipe" docAuthor="Wesener, Thomas & Sagorny, Christina" docDate="2021" docId="03BF4B62FFA6E02E1C67FCF18669F852" docLanguage="en" docName="ejt-758_1-48.pdf" docOrigin="European Journal of Taxonomy 758 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.758.1423" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zoosphaerium beanka Wesener & Sagorny 2021, sp. nov." docType="treatment" docUuid="7F9589AA-AD5D-473F-A7C1-FB94F30FF8AD" docUuidSource="ZooBank" docVersion="7" lastPageNumber="27" masterDocId="FF86331AFFBEE0341E06FFFB8261FFB5" masterDocTitle="Seven new giant pill-millipede species and numerous new records of the genus Zoosphaerium from Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="25" updateTime="1699025899143" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A775756170B8F4D1A5CF6780DF4BA730">Seven new giant pill-millipede species and numerous new records of the genus Zoosphaerium from Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae)</mods:title>
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<mods:namePart id="96835A1F2104E4DE41E8ECA0FFB02655">Wesener, Thomas</mods:namePart>
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<mods:affiliation id="5053C2F92DEA41CC1A0E730F5DFBD305">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="7AC170C2C08307278C000EB0AB510F8D" type="email">t.wesener@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:namePart id="75E17C16969EAACAD03DB0DD006515F8">Sagorny, Christina</mods:namePart>
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<mods:affiliation id="A8E3497044651DA1CA6EEFBFFD8AC697">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany. & University of Bonn, Institute of Evolutionary Biology and Ecology, D- 53121 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="338BF3BE81829208FC9D3759B734DD44" type="email">csagorny@evolution.uni-bonn.de</mods:nameIdentifier>
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<mods:title id="E4E69692B41AD77CFED1E0616C2DAAB4">European Journal of Taxonomy</mods:title>
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<mods:date id="0ED4EAAE98A0E0AA194A5C7FE9020DD6">2021</mods:date>
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<mods:number id="6E54C4C31BAAC1AB13E035BE67AD89C5">2021-07-06</mods:number>
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<treatment id="03BF4B62FFA6E02E1C67FCF18669F852" ID-DOI="http://doi.org/10.5281/zenodo.5076204" ID-GBIF-Taxon="183392514" ID-Zenodo-Dep="5076204" LSID="urn:lsid:zoobank.org:act:7F9589AA-AD5D-473F-A7C1-FB94F30FF8AD" httpUri="http://treatment.plazi.org/id/03BF4B62FFA6E02E1C67FCF18669F852" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
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<subSubSection id="C30CA9FFFFA6E02C1C67FCF181B2FC90" box="[609,979,778,805]" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph id="8BA9FA74FFA6E02C1C67FCF181B2FC90" blockId="24.[357,1230,778,875]" box="[609,979,778,805]" pageId="24" pageNumber="25">
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<heading id="D0E14D18FFA6E02C1C67FCF181B2FC90" box="[609,979,778,805]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
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<taxonomicName id="4C1681F7FFA6E02C1C67FCF18110FC90" ID-CoL="5d6cbe1a-70e0-4fa9-b389-adf9019f6119" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[609,881,778,805]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="beanka" status="sp. nov.">
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<emphasis id="B9622666FFA6E02C1C67FCF18110FC90" bold="true" box="[609,881,778,805]" italics="true" pageId="24" pageNumber="25">Zoosphaerium beanka</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2519B1DFFA6E02C1D7EFCF081B2FC90" box="[888,979,779,805]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="C30CA9FFFFA6E02C1F63FCD6811EFCDE" pageId="24" pageNumber="25" type="description">
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<paragraph id="8BA9FA74FFA6E02C1F63FCD686AFFCFD" blockId="24.[357,1230,778,875]" box="[357,1230,813,840]" pageId="24" pageNumber="25">
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<uri id="FF87F676FFA6E02C1F63FCD686AFFCFD" box="[357,1230,813,840]" pageId="24" pageNumber="25">
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urn:lsid:zoobank.org:act:
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<uuid id="FFB0C0A1FFA6E02C1C85FCD686AFFCFD" box="[643,1230,813,840]" pageId="24" pageNumber="25">7F9589AA-AD5D-473F-A7C1-FB94F30FF8AD</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BA9FA74FFA6E02C1CB2FCAB811EFCDE" blockId="24.[357,1230,778,875]" box="[692,895,848,875]" pageId="24" pageNumber="25">
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<figureCitation id="132DE6F1FFA6E02C1CB2FCAB8175FCDE" box="[692,788,848,875]" captionStart="Fig" captionStartId="9.[189,232,1764,1790]" captionTargetBox="[198,1390,265,1713]" captionTargetId="figure-92@9.[189,1398,265,1719]" captionTargetPageId="9" captionText="Fig. 2. Habitus photographs of the seven new species of Zoosphaerium, lateral view. A. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). B. Z. silens sp. nov., holotype (FMNH-INS 3119887). C. Z. ambatovaky sp. nov., holotype (CASENT 9068294/1).D. Z. beanka sp. nov., holotype (FMNH-INS 3119897). E. Z. voahangy sp. nov., holotype (FMNH-INS 3119921). F. Z. masoala sp. nov., holotype (FMNH-INS 3195651). G. Z. spinopiligerum sp. nov., holotype (FMNH-INS 3119885). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5075828" httpUri="https://zenodo.org/record/5075828/files/figure.png" pageId="24" pageNumber="25">Figs 2D</figureCitation>
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,
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<figureCitation id="132DE6F1FFA6E02C1D27FCAA8150FCDE" box="[801,817,849,875]" captionStart="Fig" captionStartId="10.[189,232,1891,1917]" captionTargetBox="[228,1372,266,1870]" captionTargetId="figure-87@10.[202,1385,239,1881]" captionTargetPageId="10" captionText="Fig. 3. Distribution map of the newly described species of Zoosphaerium. Vegetation color marks modified from Moat & Smith 2007. 1 = Z. nigrum sp. nov.; 2 = Z. silens sp. nov.; 3 = Z. ambatovaky sp. nov.; 4 = Z. beanka sp. nov.; 5 = Z. voahangy sp. nov.; 6 = Z. masoala sp. nov.; 7 = Z. spinopiligerum sp. nov. Numbers followed by ‘a’ mark additional, non-type localities." figureDoi="http://doi.org/10.5281/zenodo.5075830" httpUri="https://zenodo.org/record/5075830/files/figure.png" pageId="24" pageNumber="25">3</figureCitation>
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,
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<figureCitation id="132DE6F1FFA6E02C1D38FCAB8105FCDE" box="[830,868,848,875]" captionStart="Fig" captionStartId="12.[189,232,1891,1917]" captionTargetBox="[270,1317,266,1874]" captionTargetId="figure-78@12.[270,1317,266,1875]" captionTargetPageId="12" captionText="Fig. 5. Scanning electron micrographs of the endoterga of midbody rings. A. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). B. Z. silens sp. nov., holotype (FMNH-INS 3119887). C. Z. ambatovaky sp. nov., holotype (CASENT 9068294/1). D. Z. beanka sp. nov., holotype (FMNH- INS 3119897). E. Z. voahangy sp. nov., holotype (FMNH-INS 3119921). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5075834" httpUri="https://zenodo.org/record/5075834/files/figure.png" pageId="24" pageNumber="25">5D</figureCitation>
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,
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<figureCitation id="132DE6F1FFA6E02C1D76FCAA811EFCDE" box="[880,895,849,875]" captionStart="Fig" captionStartId="25.[189,232,1751,1777]" captionTargetBox="[269,1327,269,1734]" captionTargetId="figure-143@25.[249,1339,261,1742]" captionTargetPageId="25" captionText="Fig. 8. Zoosphaerium beanka sp. nov., holotype (FMNH-INS 3119906). A. right leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, posterior view.E. Left anterior gonopod, lateral view.F. Left anterior gonopod, anterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075840" httpUri="https://zenodo.org/record/5075840/files/figure.png" pageId="24" pageNumber="25">8</figureCitation>
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</paragraph>
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<subSubSection id="C30CA9FFFFA6E02C1EBBFC69860BFB64" pageId="24" pageNumber="25" type="diagnosis">
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<paragraph id="8BA9FA74FFA6E02C1EBBFC698357FC19" blockId="24.[189,310,914,940]" box="[189,310,914,940]" pageId="24" pageNumber="25">
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<heading id="D0E14D18FFA6E02C1EBBFC698357FC19" bold="true" box="[189,310,914,940]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
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<emphasis id="B9622666FFA6E02C1EBBFC698357FC19" bold="true" box="[189,310,914,940]" pageId="24" pageNumber="25">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA9FA74FFA6E02C1EBBFC3B860BFB64" blockId="24.[189,1399,960,1233]" pageId="24" pageNumber="25">
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The shape and position of the locking carinae place
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<taxonomicName id="4C1681F7FFA6E02C1D31FC3A81CCFC6F" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[823,941,960,986]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="beanka" status="sp. nov.">
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<emphasis id="B9622666FFA6E02C1D31FC3A81CCFC6F" box="[823,941,960,986]" italics="true" pageId="24" pageNumber="25">Z. beanka</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2519B1DFFA6E02C1DB2FC3A8670FC6E" box="[948,1041,961,987]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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in the
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<taxonomicName id="4C1681F7FFA6E02C1A6DFC3A8770FC6F" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1902" box="[1131,1297,960,986]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="platylabum">
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<emphasis id="B9622666FFA6E02C1A6DFC3A8770FC6F" box="[1131,1297,960,986]" italics="true" pageId="24" pageNumber="25">Z. platylabum</emphasis>
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</taxonomicName>
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speciesgroup, but the telopods of
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<taxonomicName id="4C1681F7FFA6E02C1FF7FC1F8007FC4B" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[497,614,996,1022]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="24" pageNumber="44" phylum="Arthropoda" rank="species" species="beanka" status="sp. nov.">
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Z.
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<emphasis id="B9622666FFA6E02C1C17FC1F8007FC4B" box="[529,614,996,1022]" italics="true" pageId="24" pageNumber="25">beanka</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2519B1DFFA6E02C1C6BFC1F80A6FC4B" box="[621,711,996,1022]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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have almost no similarities to those of any other described giant pill-millipede species except
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<taxonomicName id="4C1681F7FFA6E02C1C64FBF380ABFB94" authorityName="Wesener" authorityYear="2009" box="[610,714,1032,1057]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tsingy">
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<emphasis id="B9622666FFA6E02C1C64FBF380ABFB94" box="[610,714,1032,1057]" italics="true" pageId="24" pageNumber="25">Z. tsingy</emphasis>
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</taxonomicName>
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. Both species share characters of the posterior telopods, namely the presence of spines and at least one large membranous lobe on the movable finger, and an uncurved, wide immovable finger (2.7 times as long as wide) that is parallel to the movable finger.
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<taxonomicName id="4C1681F7FFA6E02C1EBBFB8B83A4FB3F" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[189,453,1136,1162]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="beanka" status="sp. nov.">
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<emphasis id="B9622666FFA6E02C1EBBFB8B83A4FB3F" box="[189,453,1136,1162]" italics="true" pageId="24" pageNumber="25">Zoosphaerium beanka</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2519B1DFFA6E02C1FC8FB8A804AFB3E" box="[462,555,1137,1163]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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differs from the latter in the light brown color (olive in
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<taxonomicName id="4C1681F7FFA6E02C1AD7FB8A8759FB3F" authorityName="Wesener" authorityYear="2009" box="[1233,1336,1137,1162]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tsingy">
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<emphasis id="B9622666FFA6E02C1AD7FB8A8759FB3F" box="[1233,1336,1137,1162]" italics="true" pageId="24" pageNumber="25">Z. tsingy</emphasis>
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</taxonomicName>
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), the presence of only four apical cones on the antenna (>
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<quantity id="4CEE5791FFA6E02C1D2FFB688104FB1B" box="[809,869,1171,1198]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="24" pageNumber="25" unit="in" value="15.0">15 in</quantity>
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<taxonomicName id="4C1681F7FFA6E02C1D6AFB6F81B3FB18" authorityName="Wesener" authorityYear="2009" box="[876,978,1172,1197]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tsingy">
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<emphasis id="B9622666FFA6E02C1D6AFB6F81B3FB18" box="[876,978,1172,1197]" italics="true" pageId="24" pageNumber="25">Z. tsingy</emphasis>
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</taxonomicName>
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) and the differently shaped anterior telopods, in which telopoditomere 3 lacks crenulated teeth (present in
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<taxonomicName id="4C1681F7FFA6E02C1DF3FB438638FB64" authorityName="Wesener" authorityYear="2009" box="[1013,1113,1208,1233]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tsingy">
|
||
<emphasis id="B9622666FFA6E02C1DF3FB438638FB64" box="[1013,1113,1208,1233]" italics="true" pageId="24" pageNumber="25">Z. tsingy</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30CA9FFFFA6E02C1EBBFB03831CFAD1" pageId="24" pageNumber="25" type="etymology">
|
||
<paragraph id="8BA9FA74FFA6E02C1EBBFB038325FAA7" blockId="24.[189,324,1272,1298]" box="[189,324,1272,1298]" pageId="24" pageNumber="25">
|
||
<heading id="D0E14D18FFA6E02C1EBBFB038325FAA7" bold="true" box="[189,324,1272,1298]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
|
||
<emphasis id="B9622666FFA6E02C1EBBFB038325FAA7" bold="true" box="[189,324,1272,1298]" pageId="24" pageNumber="25">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA6E02C1EBBFADC831CFAD1" blockId="24.[189,1398,1319,1380]" pageId="24" pageNumber="25">
|
||
‘Beanka’, noun in apposition, after the
|
||
<typeStatus id="54AD44D6FFA6E02C1C96FADC80A2FAF4" box="[656,707,1319,1345]" pageId="24" pageNumber="25">type</typeStatus>
|
||
and only known locality for this species, the dry forest of Beanka (
|
||
<figureCitation id="132DE6F1FFA6E02C1F20FAB1830CFAD1" box="[294,365,1354,1380]" captionStart="Fig" captionStartId="10.[189,232,1891,1917]" captionTargetBox="[228,1372,266,1870]" captionTargetId="figure-87@10.[202,1385,239,1881]" captionTargetPageId="10" captionText="Fig. 3. Distribution map of the newly described species of Zoosphaerium. Vegetation color marks modified from Moat & Smith 2007. 1 = Z. nigrum sp. nov.; 2 = Z. silens sp. nov.; 3 = Z. ambatovaky sp. nov.; 4 = Z. beanka sp. nov.; 5 = Z. voahangy sp. nov.; 6 = Z. masoala sp. nov.; 7 = Z. spinopiligerum sp. nov. Numbers followed by ‘a’ mark additional, non-type localities." figureDoi="http://doi.org/10.5281/zenodo.5075830" httpUri="https://zenodo.org/record/5075830/files/figure.png" pageId="24" pageNumber="25">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30CA9FFFFA6E02C1EBBFA70834CF98B" pageId="24" pageNumber="25" type="materials_examined">
|
||
<paragraph id="8BA9FA74FFA6E02C1EBBFA7083CBFA10" blockId="24.[189,426,1419,1445]" box="[189,426,1419,1445]" pageId="24" pageNumber="25">
|
||
<heading id="D0E14D18FFA6E02C1EBBFA7083CBFA10" bold="true" box="[189,426,1419,1445]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
|
||
<emphasis id="B9622666FFA6E02C1EBBFA7083CBFA10" bold="true" box="[189,426,1419,1445]" pageId="24" pageNumber="25">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA6E02C1EECFA41834CF98B" blockId="24.[189,1399,1466,1598]" pageId="24" pageNumber="25">
|
||
<materialsCitation id="3B7EF029FFA6E02C1EECFA418349F98B" ID-GBIF-Occurrence="3314642324" collectedFrom="slightly disturbed dry deciduous forest" collectingDate="2010-01-14" collectionCode="FMNH-INS" collectorName="Z. H. Harimpitia" country="Madagascar" elevation="220" latitude="-18.023611" location="Foret de Beanka" longLatPrecision="21" longitude="44.502224" pageId="24" pageNumber="25" specimenCode="FMNH-INS 3119897" specimenCount="1" specimenCount-male="1" stateProvince="Mahajanga" typeStatus="holotype">
|
||
<emphasis id="B9622666FFA6E02C1EECFA41833DFA61" bold="true" box="[234,348,1466,1492]" pageId="24" pageNumber="25">
|
||
<typeStatus id="54AD44D6FFA6E02C1EECFA41833DFA61" box="[234,348,1466,1492]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F301BAE4FFA6E02C1EBBFA2683EFFA42" box="[189,398,1501,1527]" name="Madagascar" pageId="24" pageNumber="25">MADAGASCAR</collectingCountry>
|
||
–
|
||
<emphasis id="B9622666FFA6E02C1FA9FA26805FFA42" bold="true" box="[431,574,1501,1527]" pageId="24" pageNumber="25">
|
||
<collectingRegion id="49D23496FFA6E02C1FA9FA26805FFA42" box="[431,574,1501,1527]" country="Madagascar" name="Mahajanga" pageId="24" pageNumber="25">Mahajanga</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D1031FDFFA6E02C1C5FFA258013FA4D" box="[601,626,1502,1528]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
|
||
;
|
||
<location id="8EC9ACAFFFA6E02C1C86FA268126FA42" LSID="urn:lsid:plazi:treatment:03BF4B62FFA6E02E1C67FCF18669F852:8EC9ACAFFFA6E02C1C86FA268126FA42" box="[640,839,1501,1527]" country="Madagascar" latitude="-18.023611" longLatPrecision="21" longitude="44.502224" name="Foret de Beanka" pageId="24" pageNumber="25" stateProvince="Mahajanga">Forêt de Beanka</location>
|
||
;
|
||
<geoCoordinate id="EE229CB3FFA6E02C1D55FA278184FA42" box="[851,997,1500,1528]" degrees="18" direction="south" minutes="01" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="25" value="-18.023611">18°01′25″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE229CB3FFA6E02C1DF5FA2786E6FA4D" box="[1011,1159,1500,1528]" degrees="44" direction="east" minutes="30" orientation="longitude" pageId="24" pageNumber="25" precision="15" seconds="08" value="44.502224">44°30′08″ E</geoCoordinate>
|
||
; alt.
|
||
<quantity id="4CEE5791FFA6E02C1AC4FA268770FA4D" box="[1218,1297,1501,1528]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="24" pageNumber="25" unit="m" value="220.0">
|
||
<elevation id="003B1D47FFA6E02C1AC4FA268770FA4D" box="[1218,1297,1501,1528]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="24" pageNumber="25" unit="m" value="220.0">220 m</elevation>
|
||
</quantity>
|
||
;
|
||
<collectedFrom id="380AB989FFA6E02C1B18FA26804DF9AF" pageId="24" pageNumber="25">slightly disturbed dry deciduous forest</collectedFrom>
|
||
;
|
||
<date id="FFA8DCB4FFA6E02C1C3CF9FA80B6F9AE" box="[570,727,1537,1563]" pageId="24" pageNumber="25" value="2010-01-14">
|
||
<collectingDate id="EFEC255CFFA6E02C1C3CF9FA80B6F9AE" box="[570,727,1537,1563]" pageId="24" pageNumber="25" value="2010-01-14">14 Jan. 2010</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26E39FA2FFA6E02C1CE3F9FA81C5F9AF" box="[741,932,1536,1563]" pageId="24" pageNumber="25">Z.H. Harimpitia</collectorName>
|
||
leg.; coll. no. Z.H.H-031;
|
||
<specimenCode id="DBB0520FFFA6E02C1AEFF9FA8349F98B" pageId="24" pageNumber="25">FMNH-INS 3119897</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30CA9FFFFA6E02E1EBBF99E8358F89C" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="description">
|
||
<paragraph id="8BA9FA74FFA6E02C1EBBF99E832FF9CA" blockId="24.[189,334,1637,1663]" box="[189,334,1637,1663]" pageId="24" pageNumber="25">
|
||
<heading id="D0E14D18FFA6E02C1EBBF99E832FF9CA" bold="true" box="[189,334,1637,1663]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
|
||
<emphasis id="B9622666FFA6E02C1EBBF99E832FF9CA" bold="true" box="[189,334,1637,1663]" pageId="24" pageNumber="25">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA6E02C1EBBF96F802CF967" blockId="24.[187,1398,1683,1746]" pageId="24" pageNumber="25">
|
||
BODY LENGTH. Male (
|
||
<typeStatus id="54AD44D6FFA6E02C1FBCF9688049F91B" box="[442,552,1683,1710]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
): length
|
||
<quantity id="4CEE5791FFA6E02C1C8FF9688092F91B" box="[649,755,1683,1710]" metricMagnitude="-2" metricUnit="m" metricValue="3.67" pageId="24" pageNumber="25" unit="mm" value="36.7">36.7 mm</quantity>
|
||
, width of thoracic shield up
|
||
<quantity id="4CEE5791FFA6E02C1A30F96F86FCF91B" box="[1078,1181,1684,1710]" metricMagnitude="-2" metricUnit="m" metricValue="2.21" pageId="24" pageNumber="25" unit="mm" value="22.1">22.1 mm</quantity>
|
||
(widest), height of thoracic shield
|
||
<quantity id="4CEE5791FFA6E02C1F69F94C83B7F964" box="[367,470,1719,1745]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="24" pageNumber="25" unit="mm" value="11.8">11.8 mm</quantity>
|
||
(highest).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA6E02C1EBBF90383B2F880" blockId="24.[189,1398,1784,1846]" pageId="24" pageNumber="25">
|
||
COLORATION. Head, collum, thoracic shield, tergites and anal shield light brown (
|
||
<figureCitation id="132DE6F1FFA6E02C1A96F9038690F8A7" box="[1168,1265,1784,1810]" captionStart="Fig" captionStartId="9.[189,232,1764,1790]" captionTargetBox="[198,1390,265,1713]" captionTargetId="figure-92@9.[189,1398,265,1719]" captionTargetPageId="9" captionText="Fig. 2. Habitus photographs of the seven new species of Zoosphaerium, lateral view. A. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). B. Z. silens sp. nov., holotype (FMNH-INS 3119887). C. Z. ambatovaky sp. nov., holotype (CASENT 9068294/1).D. Z. beanka sp. nov., holotype (FMNH-INS 3119897). E. Z. voahangy sp. nov., holotype (FMNH-INS 3119921). F. Z. masoala sp. nov., holotype (FMNH-INS 3195651). G. Z. spinopiligerum sp. nov., holotype (FMNH-INS 3119885). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5075828" httpUri="https://zenodo.org/record/5075828/files/figure.png" pageId="24" pageNumber="25">Fig. 2D</figureCitation>
|
||
). Antennae green, legs light brown.
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA6E02C1EBBF8A68716F82E" blockId="24.[189,1399,1885,1947]" pageId="24" pageNumber="25">HEAD. Eyes with more than 75 ommatidia. Antennae short, protruding back to leg 3. Length of antennomeres 1>2=3 =4=5 <6. A shallow groove present on antennomere 1, disc with 4 apical cones.</paragraph>
|
||
<paragraph id="8BA9FA74FFA6E02C1EBBF8398169F86E" blockId="24.[189,776,1985,2012]" box="[189,776,1985,2012]" pageId="24" pageNumber="25">GNATHOCHILARIUM AND MANDIBLE. Not dissected.</paragraph>
|
||
<caption id="DF69AAFCFFA7E02D1EBBF92C83C0F85D" ID-DOI="http://doi.org/10.5281/zenodo.5075840" ID-Zenodo-Dep="5075840" httpUri="https://zenodo.org/record/5075840/files/figure.png" pageId="25" pageNumber="26" startId="25.[189,232,1751,1777]" targetBox="[269,1327,269,1734]" targetPageId="25">
|
||
<paragraph id="8BA9FA74FFA7E02D1EBBF92C83C0F85D" blockId="25.[187,1399,1751,2024]" pageId="25" pageNumber="26">
|
||
<emphasis id="B9622666FFA7E02D1EBBF92C836AF944" bold="true" box="[189,267,1751,1777]" pageId="25" pageNumber="26">Fig. 8.</emphasis>
|
||
<taxonomicName id="4C1681F7FFA7E02D1F15F92C807BF944" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[275,538,1751,1777]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="beanka" status="sp. nov.">
|
||
<emphasis id="B9622666FFA7E02D1F15F92C807BF944" box="[275,538,1751,1777]" italics="true" pageId="25" pageNumber="26">Zoosphaerium beanka</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFA7E02D1C24F92380E1F947" box="[546,640,1752,1778]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (FMNH-INS 3119906).
|
||
<emphasis id="B9622666FFA7E02D1A1CF92C864FF944" bold="true" box="[1050,1070,1751,1777]" pageId="25" pageNumber="26">A</emphasis>
|
||
. right leg 9, posterior view.
|
||
<emphasis id="B9622666FFA7E02D1EB8F90182B0F8A1" bold="true" box="[190,209,1786,1812]" pageId="25" pageNumber="26">B</emphasis>
|
||
. Left coxa 2 with gonopore, posterior view.
|
||
<emphasis id="B9622666FFA7E02D1CF8F9018172F8A1" bold="true" box="[766,787,1786,1812]" pageId="25" pageNumber="26">C</emphasis>
|
||
. Left coxa 1 with stigmatic plate, posterior view.
|
||
<emphasis id="B9622666FFA7E02D1EBBF8E682B3F882" bold="true" box="[189,210,1821,1847]" pageId="25" pageNumber="26">D</emphasis>
|
||
. Left anterior gonopod, posterior view.
|
||
<emphasis id="B9622666FFA7E02D1C90F8E680CBF882" bold="true" box="[662,682,1821,1847]" pageId="25" pageNumber="26">E</emphasis>
|
||
. Left anterior gonopod, lateral view.
|
||
<emphasis id="B9622666FFA7E02D1A54F8E68605F882" bold="true" box="[1106,1124,1821,1847]" pageId="25" pageNumber="26">F</emphasis>
|
||
. Left anterior gonopod, anterior view.
|
||
<emphasis id="B9622666FFA7E02D1F6BF8BB83E5F8EF" bold="true" box="[365,388,1856,1882]" pageId="25" pageNumber="26">G</emphasis>
|
||
. Right posterior telopod, posterior view.
|
||
<emphasis id="B9622666FFA7E02D1D78F8BA81F4F8EE" bold="true" box="[894,917,1857,1883]" pageId="25" pageNumber="26">H</emphasis>
|
||
. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBFEF2834EFEDC" blockId="26.[188,1398,265,361]" pageId="26" pageNumber="27">
|
||
STIGMATIC PLATES. First stigmatic plate triangular, apex well rounded, nearly as wide as at base, slightly turned towards coxa. Hairs most abundant on apical margin, very few hairs on basal and lateral margins (
|
||
<figureCitation id="132DE6F1FFA4E02E1EC3FEB48343FEDC" box="[197,290,335,361]" captionStart="Fig" captionStartId="25.[189,232,1751,1777]" captionTargetBox="[269,1327,269,1734]" captionTargetId="figure-143@25.[249,1339,261,1742]" captionTargetPageId="25" captionText="Fig. 8. Zoosphaerium beanka sp. nov., holotype (FMNH-INS 3119906). A. right leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, posterior view.E. Left anterior gonopod, lateral view.F. Left anterior gonopod, anterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075840" httpUri="https://zenodo.org/record/5075840/files/figure.png" pageId="26" pageNumber="27">Fig. 8C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBFE638618FE06" blockId="26.[189,1145,408,435]" box="[189,1145,408,435]" pageId="26" pageNumber="27">PLEURITES. First pleurite slightly extending posteriorly in a well-rounded process.</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBFE1A8660FE4E" blockId="26.[189,1025,481,508]" box="[189,1025,481,508]" pageId="26" pageNumber="27">COLLUM. Margins covered with a few short hairs, central part glabrous.</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBFDD18107FDF1" blockId="26.[189,870,554,580]" box="[189,870,554,580]" pageId="26" pageNumber="27">THORACIC SHIELD. Surface like those of tergites, glabrous.</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBFD888754FD38" blockId="26.[189,1333,627,654]" box="[189,1333,627,654]" pageId="26" pageNumber="27">TERGITES. Smooth and glabrous, no small pits present. Paratergites projecting slightly posteriorly.</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBFD4780EAFCF4" blockId="26.[188,1399,700,833]" pageId="26" pageNumber="27">
|
||
ENDOTERGUM. Inner section with numerous short spines and isolated bristles. Between marginal ridge and inner area, a single row of sparse large, circular cuticular impressions. Externally one or two rows of marginal bristles, standing irregularly to each other. Bristles scaly, of medium length, longest reaching slightly above tergite margin (
|
||
<figureCitation id="132DE6F1FFA4E02E1C19FCDD801FFCF4" box="[543,638,806,833]" captionStart="Fig" captionStartId="12.[189,232,1891,1917]" captionTargetBox="[270,1317,266,1874]" captionTargetId="figure-78@12.[270,1317,266,1875]" captionTargetPageId="12" captionText="Fig. 5. Scanning electron micrographs of the endoterga of midbody rings. A. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). B. Z. silens sp. nov., holotype (FMNH-INS 3119887). C. Z. ambatovaky sp. nov., holotype (CASENT 9068294/1). D. Z. beanka sp. nov., holotype (FMNH- INS 3119897). E. Z. voahangy sp. nov., holotype (FMNH-INS 3119921). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5075834" httpUri="https://zenodo.org/record/5075834/files/figure.png" pageId="26" pageNumber="27">Fig. 5D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBFC94838DFC7A" blockId="26.[189,1400,879,976]" pageId="26" pageNumber="27">ANAL SHIELD. Rounded, neither bell-shaped nor tapered, glabrous. Ventral side of anal shield with two black locking carinae, located close to laterotergites. Anterior carina small, posterior carina 2–3 times as long as anterior carina.</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBFC058618FBEA" blockId="26.[189,1398,1022,1119]" pageId="26" pageNumber="27">
|
||
LEGS. Leg 1 with 6 or 7 ventral spines, leg 2 with 8, leg 3 with 10. First two leg pairs without an apical spine, spine present on leg 3. Leg pairs 4–21 with 9 or 10 ventral spines and an apical spine. On leg 9, femur 2 and tarsus 4.9 times as long as wide. All podomeres with setae (
|
||
<figureCitation id="132DE6F1FFA4E02E1A0BFBBE860DFBEA" box="[1037,1132,1093,1119]" captionStart="Fig" captionStartId="25.[189,232,1751,1777]" captionTargetBox="[269,1327,269,1734]" captionTargetId="figure-143@25.[249,1339,261,1742]" captionTargetPageId="25" captionText="Fig. 8. Zoosphaerium beanka sp. nov., holotype (FMNH-INS 3119906). A. right leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, posterior view.E. Left anterior gonopod, lateral view.F. Left anterior gonopod, anterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075840" httpUri="https://zenodo.org/record/5075840/files/figure.png" pageId="26" pageNumber="27">Fig. 8A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EECFB75806AFB1D" blockId="26.[187,1399,1166,1263]" box="[234,523,1166,1192]" pageId="26" pageNumber="27">
|
||
<emphasis id="B9622666FFA4E02E1EECFB75806AFB1D" bold="true" box="[234,523,1166,1192]" pageId="26" pageNumber="27">Male sexual characters</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBFB4A80B7FB5B" blockId="26.[187,1399,1166,1263]" pageId="26" pageNumber="27">
|
||
GONOPORE. Covered with a single undivided, rounded membranous plate, covering ¼ of surface of coxa. Gonopore located centrally on joint (
|
||
<figureCitation id="132DE6F1FFA4E02E1C6BFB2F80A9FB5B" box="[621,712,1236,1262]" captionStart="Fig" captionStartId="25.[189,232,1751,1777]" captionTargetBox="[269,1327,269,1734]" captionTargetId="figure-143@25.[249,1339,261,1742]" captionTargetPageId="25" captionText="Fig. 8. Zoosphaerium beanka sp. nov., holotype (FMNH-INS 3119906). A. right leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, posterior view.E. Left anterior gonopod, lateral view.F. Left anterior gonopod, anterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075840" httpUri="https://zenodo.org/record/5075840/files/figure.png" pageId="26" pageNumber="27">Fig. 8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBFAE680B9FA52" blockId="26.[189,1399,1309,1512]" pageId="26" pageNumber="27">
|
||
ANTERIOR TELOPODS. Harp with two stridulation ribs, mesal rib smaller than lateral one, both ribs well developed (
|
||
<figureCitation id="132DE6F1FFA4E02E1F4CFABB83C6FAEF" box="[330,423,1344,1370]" captionStart="Fig" captionStartId="25.[189,232,1751,1777]" captionTargetBox="[269,1327,269,1734]" captionTargetId="figure-143@25.[249,1339,261,1742]" captionTargetPageId="25" captionText="Fig. 8. Zoosphaerium beanka sp. nov., holotype (FMNH-INS 3119906). A. right leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, posterior view.E. Left anterior gonopod, lateral view.F. Left anterior gonopod, anterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075840" httpUri="https://zenodo.org/record/5075840/files/figure.png" pageId="26" pageNumber="27">Fig. 8F</figureCitation>
|
||
). First podomere 1.2 times as wide as long. Second podomere process lobe-like and slightly curved, with rounded edges, protruding up to half of third podomere height, apically with sclerotized spots. Third podomere long, as long as first; cavity mesally with numerous sclerotized spots and three thin, sclerotized spines; one sclerotized spot located close to tip. Podomeres 1–3 covered on both sides with numerous long, isolated hairs.
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBF9ED861BF908" blockId="26.[189,1400,1558,1725]" pageId="26" pageNumber="27">
|
||
POSTERIOR TELOPODS. Movable finger with a well-rounded tip, not tapering towards tip; hollowed-out inner margin with one non-sclerotized lobe and 3 or 4 sclerotized spines, posterior aspect with ca 14 small sclerotized teeth. Movable finger slightly longer than fixed finger, the latter straight, as wide as third podomere. Fixed finger at margin with numerous sclerotized spots. All podomeres covered on both sides with numerous, long isolated hairs, only tips of chela glabrous (
|
||
<figureCitation id="132DE6F1FFA4E02E1DECF958860CF908" box="[1002,1133,1699,1725]" captionStart="Fig" captionStartId="25.[189,232,1751,1777]" captionTargetBox="[269,1327,269,1734]" captionTargetId="figure-143@25.[249,1339,261,1742]" captionTargetPageId="25" captionText="Fig. 8. Zoosphaerium beanka sp. nov., holotype (FMNH-INS 3119906). A. right leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, posterior view.E. Left anterior gonopod, lateral view.F. Left anterior gonopod, anterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075840" httpUri="https://zenodo.org/record/5075840/files/figure.png" pageId="26" pageNumber="27">Fig. 8G–H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EECF9178324F8B3" blockId="26.[189,325,1772,1833]" box="[234,325,1772,1798]" pageId="26" pageNumber="27">
|
||
<emphasis id="B9622666FFA4E02E1EECF9178324F8B3" bold="true" box="[234,325,1772,1798]" pageId="26" pageNumber="27">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBF8F48358F89C" blockId="26.[189,325,1772,1833]" box="[189,313,1807,1833]" pageId="26" pageNumber="27">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30CA9FFFFA4E02E1EBBF8A38669F852" pageId="26" pageNumber="27" type="distribution">
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBF8A38390F8C7" blockId="26.[189,497,1880,1906]" box="[189,497,1880,1906]" pageId="26" pageNumber="27">
|
||
<heading id="D0E14D18FFA4E02E1EBBF8A38390F8C7" bold="true" box="[189,497,1880,1906]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
|
||
<emphasis id="B9622666FFA4E02E1EBBF8A38390F8C7" bold="true" box="[189,497,1880,1906]" pageId="26" pageNumber="27">Distribution and ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA4E02E1EBBF87C8669F852" blockId="26.[188,1398,1927,2024]" pageId="26" pageNumber="27">
|
||
<taxonomicName id="4C1681F7FFA4E02E1EBBF87C83A4F814" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[189,453,1927,1953]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="beanka" status="sp. nov.">
|
||
<emphasis id="B9622666FFA4E02E1EBBF87C83A4F814" box="[189,453,1927,1953]" italics="true" pageId="26" pageNumber="27">Zoosphaerium beanka</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFA4E02E1FC9F87C804DF814" box="[463,556,1927,1953]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is currently known only from the dry forest of Beanka (
|
||
<figureCitation id="132DE6F1FFA4E02E1ADCF87C8745F814" box="[1242,1316,1927,1953]" captionStart="Fig" captionStartId="10.[189,232,1891,1917]" captionTargetBox="[228,1372,266,1870]" captionTargetId="figure-87@10.[202,1385,239,1881]" captionTargetPageId="10" captionText="Fig. 3. Distribution map of the newly described species of Zoosphaerium. Vegetation color marks modified from Moat & Smith 2007. 1 = Z. nigrum sp. nov.; 2 = Z. silens sp. nov.; 3 = Z. ambatovaky sp. nov.; 4 = Z. beanka sp. nov.; 5 = Z. voahangy sp. nov.; 6 = Z. masoala sp. nov.; 7 = Z. spinopiligerum sp. nov. Numbers followed by ‘a’ mark additional, non-type localities." figureDoi="http://doi.org/10.5281/zenodo.5075830" httpUri="https://zenodo.org/record/5075830/files/figure.png" pageId="26" pageNumber="27">Fig. 3</figureCitation>
|
||
). With most known dry forest giant pill-millipede species being widespread (
|
||
<bibRefCitation id="EF878785FFA4E02E1DE4F8518769F871" author="Wesener T. & Wagele J. W." box="[994,1288,1962,1988]" pageId="26" pageNumber="27" pagination="5 - 82" refId="ref28537" refString="Wesener T. & Wagele J. W. 2008. The giant pill-millipedes of Madagascar: revision of the genus Zoosphaerium (Myriapoda, Diplopoda, Sphaerotheriida). Zoosystema 30 (1): 5 - 82." type="journal article" year="2008">Wesener & Wägele 2008</bibRefCitation>
|
||
; Wesener 2009), this species can be expected in surrounding dry forest fragments.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |