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<document id="02B31A8A9ED5231D2DB089063A54037A" ID-DOI="10.1206/0003-0090.469.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="14658466" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736967130833" checkinUser="felipe" docAuthor="Engel, Michael S. &amp; Xie, Jiaying" docDate="2024" docId="03B887F1CD24FF87FF36FB9B0AFFF914" docLanguage="en" docName="BulAmeMusNatHis.2024.B469.1-80.pdf" docOrigin="Bulletin of the American Museum of Natural History 2024 (469)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-2024/issue-469/0003-0090.469.1.1/The-Bee-Fauna-of-Eocene-Fushun-Amber-Hymenoptera-Apoidea/10.1206/0003-0090.469.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.10:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="10" docTitle="Glyptosmia hemiaspis Engel &amp; Xie 2024, new species" docType="treatment" docVersion="2" lastPageNumber="28" masterDocId="FF81FF89CD3CFF9BFFB9FF800E43FFDC" masterDocTitle="The Bee Fauna Of Eocene Fushun Amber (Hymenoptera: Apoidea)" masterLastPageNumber="81" masterPageNumber="1" pageNumber="24" updateTime="1736969989509" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="FA88EEB049BC64936D55AE5F243BF68C">
<mods:title id="AF9D9F9D171E6F5ABA0142D83259CC6A">The Bee Fauna Of Eocene Fushun Amber (Hymenoptera: Apoidea)</mods:title>
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<mods:name id="05B4F4B599E3A3ABC4BDFAE6419CA3F7" type="personal">
<mods:role id="5EF4CB6649AC9FADE9B47EE2CF3A0195">
<mods:roleTerm id="5000CB818AEB04DC83F3528F2118D372">Author</mods:roleTerm>
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<mods:namePart id="047D50AABCA993BC743EE853B8208C58">Engel, Michael S.</mods:namePart>
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<mods:namePart id="BDD43C4F82C6189D5C85B35F095F6527">Xie, Jiaying</mods:namePart>
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<mods:typeOfResource id="F931163F43ABD6EA959DE4859F4E429A">text</mods:typeOfResource>
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<mods:title id="3DF64BD61C13FCFA0AB21FE2E56D902C">Bulletin of the American Museum of Natural History</mods:title>
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<mods:date id="487226F8E3EC26A4452B6D895EC53581">2024</mods:date>
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<mods:number id="7F5C41E8A18CC6EB146AC1E16335D927">2024-10-23</mods:number>
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<mods:number id="8524C249C34E5CA1A49B8ACADDEFEBB7">2024</mods:number>
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<mods:detail id="BDC5ED015DBF24A90A9A1916EB85A22F" type="issue">
<mods:number id="19190B1DD946F042A6A9D4A6A218FA7F">469</mods:number>
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<mods:url id="7C5CDD9B8FE67339258065DCDBAEF1DA">https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-2024/issue-469/0003-0090.469.1.1/The-Bee-Fauna-of-Eocene-Fushun-Amber-Hymenoptera-Apoidea/10.1206/0003-0090.469.1.1.full</mods:url>
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<mods:classification id="A28BF600DA0BE6B65E9F18DF116A7FD5">journal article</mods:classification>
<mods:identifier id="9F65400F77BCFF102C7266C9014D23C6" type="DOI">10.1206/0003-0090.469.1.1</mods:identifier>
<mods:identifier id="42835931C0EE699A7DD9641C5F4C8573" type="ISSN">0003-0090</mods:identifier>
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<treatment id="03B887F1CD24FF87FF36FB9B0AFFF914" LSID="urn:lsid:plazi:treatment:03B887F1CD24FF87FF36FB9B0AFFF914" httpUri="http://treatment.plazi.org/id/03B887F1CD24FF87FF36FB9B0AFFF914" lastPageId="28" lastPageNumber="28" pageId="24" pageNumber="24">
<subSubSection id="C30B656CCD24FF83FF36FB9B0C1BFBEF" box="[143,600,1051,1075]" pageId="24" pageNumber="24" type="nomenclature">
<paragraph id="8BAE36E7CD24FF83FF36FB9B0C1BFBEF" blockId="24.[143,600,1051,1075]" box="[143,600,1051,1075]" pageId="24" pageNumber="24">
<heading id="D0E6818BCD24FF83FF36FB9B0C1BFBEF" box="[143,600,1051,1075]" centered="true" fontSize="9" level="2" pageId="24" pageNumber="24" reason="2">
<taxonomicName id="4C114D64CD24FF83FF36FB9B0F8FFBEF" authority="Engel" authorityName="Engel &amp; Xie" authorityYear="2024" box="[143,460,1051,1075]" class="Insecta" family="Megachilidae" genus="Glyptosmia" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="hemiaspis" status="sp. nov.">
<emphasis id="B965EAF5CD24FF83FF36FB9B0FC5FBEF" bold="true" box="[143,390,1051,1075]" italics="true" pageId="24" pageNumber="24">Glyptosmia hemiaspis</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="A256578ECD24FF83FE61FB9C0C1BFBEF" box="[472,600,1051,1075]" pageId="24" pageNumber="24" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30B656CCD24FF83FE9EFBCD0F82FBB9" box="[295,449,1101,1125]" pageId="24" pageNumber="24" type="description">
<paragraph id="8BAE36E7CD24FF83FE9EFBCD0F82FBB9" blockId="24.[295,449,1101,1125]" box="[295,449,1101,1125]" pageId="24" pageNumber="24">
<heading id="D0E6818BCD24FF83FE9EFBCD0F82FBB9" box="[295,449,1101,1125]" centered="true" fontSize="9" level="2" pageId="24" pageNumber="24" reason="2">
<figureCitation id="132A2A62CD24FF83FE9EFBCD0F82FBB9" box="[295,449,1101,1125]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStart-3="FIGURE 16" captionStart-4="FIGURE 17" captionStart-5="FIGURE 18" captionStart-6="FIGURE 19" captionStartId-0="25.[108,194,1596,1617]" captionStartId-1="26.[107,189,1031,1052]" captionStartId-2="27.[108,194,1318,1339]" captionStartId-3="28.[107,193,1324,1345]" captionStartId-4="30.[107,193,819,840]" captionStartId-5="31.[109,195,1344,1365]" captionStartId-6="32.[107,190,953,974]" captionTargetBox-0="[108,1212,224,1568]" captionTargetBox-1="[107,1211,224,1005]" captionTargetBox-2="[108,1212,224,1292]" captionTargetBox-3="[107,1211,224,1296]" captionTargetBox-4="[107,1211,224,792]" captionTargetBox-5="[108,1212,224,1318]" captionTargetBox-6="[107,1211,224,927]" captionTargetId-0="figure-36@25.[108,1212,224,1568]" captionTargetId-1="figure-286@26.[107,1211,224,1005]" captionTargetId-2="figure-159@27.[108,1212,224,1292]" captionTargetId-3="figure-139@28.[107,1211,224,1296]" captionTargetId-4="figure-434@30.[107,1211,224,792]" captionTargetId-5="figure-92@31.[108,1212,224,1318]" captionTargetId-6="figure-324@32.[107,1211,224,927]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionTargetPageId-3="28" captionTargetPageId-4="30" captionTargetPageId-5="31" captionTargetPageId-6="32" captionText-0="FIGURE 13. Photographs of holotype (CNU-HYM-LF-2023-002) female of Glyptosmia hemiaspis, n. gen. and sp., in Fushun amber. A. Left dorsal oblique view. B. Right lateral oblique view." captionText-1="FIGURE 14. Photographs of head of Glyptosmia hemiaspis. A. Oblique facial view from upper right. B. Facial view." captionText-2="FIGURE 15. Oblique dorsal view of mesoscutum and mesoscutellum of Glyptosmia hemiaspis." captionText-3="FIGURE 16. Profiles of Glyptosmia hemiaspis. A. Mesoscutellum, metanotum, and propodeum. B. Metasomal terga I and II and base of tergum III." captionText-4="FIGURE 17. Forewing of Glyptosmia hemiaspis." captionText-5="FIGURE 18. Photographs of leg podites of Glyptosmia hemiaspis. A. Dorsal view of metatibia, metatarsus, and metapretarsus. B. Apical view of metatibial spurs (note also hooked scopal setae in background). C. Detail of metapretarsus, dorsal view. D. Detail of metapretarsal claws, lateral view, arrow points to inner (subapical) ramus of left claw." captionText-6="FIGURE 19. Photographs of metasomal apex of Glyptosmia hemiaspis. A. Ventral view. B. Left oblique lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.14658496" figureDoi-1="http://doi.org/10.5281/zenodo.14658498" figureDoi-2="http://doi.org/10.5281/zenodo.14658502" figureDoi-3="http://doi.org/10.5281/zenodo.14658504" figureDoi-4="http://doi.org/10.5281/zenodo.14658508" figureDoi-5="http://doi.org/10.5281/zenodo.14658510" figureDoi-6="http://doi.org/10.5281/zenodo.14658512" httpUri-0="https://zenodo.org/record/14658496/files/figure.png" httpUri-1="https://zenodo.org/record/14658498/files/figure.png" httpUri-2="https://zenodo.org/record/14658502/files/figure.png" httpUri-3="https://zenodo.org/record/14658504/files/figure.png" httpUri-4="https://zenodo.org/record/14658508/files/figure.png" httpUri-5="https://zenodo.org/record/14658510/files/figure.png" httpUri-6="https://zenodo.org/record/14658512/files/figure.png" pageId="24" pageNumber="24">Figures 13–19</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30B656CCD24FF83FF35FBFF0C66FB4A" box="[140,549,1150,1175]" pageId="24" pageNumber="24" type="diagnosis">
<paragraph id="8BAE36E7CD24FF83FF35FBFF0C66FB4A" blockId="24.[108,637,1150,1736]" box="[140,549,1150,1175]" pageId="24" pageNumber="24">DIAGNOSIS: As for the genus (above).</paragraph>
</subSubSection>
<subSubSection id="C30B656CCD24FF87FF35FB200F14F9B9" lastPageId="28" lastPageNumber="28" pageId="24" pageNumber="24" type="description">
<paragraph id="8BAE36E7CD24FF83FF35FB200DA9FBCF" blockId="24.[108,637,1150,1736]" lastBlockId="24.[684,1214,226,1738]" pageId="24" pageNumber="24">
DESCRIPTION: FEMALE. Total body length (as preserved)
<quantity id="4CE99B02CD24FF83FEB6FB400F30FB04" box="[271,371,1216,1240]" metricMagnitude="-3" metricUnit="m" metricValue="4.36" pageId="24" pageNumber="24" unit="mm" value="4.36">4.36 mm</quantity>
; forewing length (based on the left forewing as the right wing is taphonomically stretched)
<quantity id="4CE99B02CD24FF83FEF7FA830FF3FAC6" box="[334,432,1283,1307]" metricMagnitude="-3" metricUnit="m" metricValue="2.27" pageId="24" pageNumber="24" unit="mm" value="2.27">2.27 mm</quantity>
. Integument black, shining (fig. 13); wing veins dark brown, membranes hyaline and clear (fig. 17). Head slightly longer than wide, length
<quantity id="4CE99B02CD24FF83FE37FAE60FB6FAA1" box="[398,501,1382,1406]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="24" pageNumber="24" unit="mm" value="1.03">1.03 mm</quantity>
, width
<quantity id="4CE99B02CD24FF83FDF7FAE60ED9FA42" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="24" pageNumber="24" unit="mm" value="0.81">0.81 mm</quantity>
; labrum about as long as wide, with truncate apical margin; mandible elongate, with distinct apical tooth and larger preapical edentate margin (apex similar to
<taxonomicName id="4C114D64CD24FF83FEA6FA6A0FC7F9DE" authorityName="Cockerell" authorityYear="1909" box="[287,388,1514,1538]" class="Insecta" family="Megachilidae" genus="Glyptapis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAF5CD24FF83FEA6FA6A0FC7F9DE" box="[287,388,1514,1538]" italics="true" pageId="24" pageNumber="24">Glyptapis</emphasis>
</taxonomicName>
except mandible more elongate); malar space linear; maxillary palpus tetramerous; labial palpomeres I and II long and flattened, albeit each shorter than prementum; clypeus flat, not extending below lower tangent of compound eyes; single subantennal sulcus straight to lower outer angle of torulus (visible faintly from right side; base of sulcus faint on left side), sulcus slightly longer than torular diameter; compound eye bare, slightly broader than gena in profile, inner margins straight and slightly converging below; upper interorbital distance greater than lower interorbital distance; scape short, length
<quantity id="4CE99B02CD24FF83FC3AFE280DABFE1C" box="[899,1000,424,448]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="24" pageNumber="24" unit="mm" value="0.18">0.18 mm</quantity>
, shorter than torulocellar distance; flagellomere I shorter than combined lengths of flagellomeres II and III (fig. 14B), flagellomeres II and III of equal lengths; preoccipital ridge sharply angled dorsally, but not carinate, rounded laterally. Mesoscutum length
<quantity id="4CE99B02CD24FF83FD4EFDEF0D1BFD5A" box="[759,856,623,647]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="24" pageNumber="24" unit="mm" value="0.88">0.88 mm</quantity>
, mesoscutellum length
<quantity id="4CE99B02CD24FF83FBEFFDEF0AFBFD5A" box="[1110,1208,623,647]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="24" unit="mm" value="0.3">0.30 mm</quantity>
; intertegular distance
<quantity id="4CE99B02CD24FF83FC23FD0F0A43FD7B" box="[922,1024,655,679]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="24" pageNumber="24" unit="mm" value="0.85">0.85 mm</quantity>
; parapsidal lines short, linear; transverse mesoscutal-mesoscutellar sulcus deeply impressed and wide, seemingly weakly foveolate within depression (challenging to observe as it is difficult to illuminate properly); tegula oval, brown and semitranslucent, with weak punctures. Forewing (fig. 17) with basal vein (1M) noticeably arched, orthogonal to 1Cu; hind wing with six distal hamuli arranged in a single series; jugal lobe more than 0.5× but less than 0.65× length of anal lobe (challenging to see precise measurement).
</paragraph>
<paragraph id="8BAE36E7CD24FF81FD74FB9C0EE5F9DF" blockId="24.[684,1214,226,1738]" lastBlockId="26.[108,638,1085,1737]" lastPageId="26" lastPageNumber="26" pageId="24" pageNumber="24">Labrum, clypeus, and supraclypeal area with dense, small punctures, punctures separated by much less than a puncture width, nearly contiguous in some places, integument between punctures apparently smooth and shining. Lower face punctured as on clypeus, frons with similar punctures but more spaced (fig. 14), separated by 0.5–1.0× a puncture width, although those on central frons denser, integument between punctures smooth and shining; ocellocular area (fig. 14A) and vertex with punctures separated by 0.5–1.2× a puncture width; gena apparently with punctures as on vertex. Mesosoma with dense punctures (figs. 13A, 15), punctures coarser and slightly larger than those of head, punctures nearly contiguous on mesoscutum, mesoscutellum, and pleura, those of pleura seemingly slightly larger than those of mesoscutum (challenging to see clearly given poor state of preservation), integument between punctures smooth and shining. Metanotum irregularly roughened with two rounded tubercles on either side of midline. Basal area of propodeum areolate, with broad seemingly impunctate triangular area on upper posterior surface. Metasomal terga II–V with pregradular surfaces depressed and polished relative to disc and apical marginal zones, pregradular areas smooth to finely imbricate with some scattered, shallow, small punctures apically; tergal discs imbricate with small punctures separated by a puncture width (fig. 16B), although laterally punctures denser, anterior-facing surface of tergum I with punctures much sparser and integument between smooth (fig. 16B).</paragraph>
<caption id="DF6E666FCD25FF82FFD5F9BC0DD6F9B3" ID-DOI="http://doi.org/10.5281/zenodo.14658496" ID-Zenodo-Dep="14658496" httpUri="https://zenodo.org/record/14658496/files/figure.png" pageId="25" pageNumber="25" startId="25.[108,194,1596,1617]" targetBox="[108,1212,224,1568]" targetPageId="25" targetType="figure">
<paragraph id="8BAE36E7CD25FF82FFD5F9BC0DD6F9B3" blockId="25.[108,1212,1594,1647]" pageId="25" pageNumber="25">
FIGURE 13. Photographs of holotype (CNU-HYM-LF-2023-002) female of
<taxonomicName id="4C114D64CD25FF82FCD0F9BA0A01F98E" authorityName="Engel &amp; Xie" authorityYear="2024" box="[873,1090,1594,1618]" class="Insecta" family="Megachilidae" genus="Glyptosmia" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="hemiaspis" status="n. gen. and sp.">
<emphasis id="B965EAF5CD25FF82FCD0F9BA0A01F98E" box="[873,1090,1594,1618]" italics="true" pageId="25" pageNumber="25">Glyptosmia hemiaspis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A256578ECD25FF82FBF4F9BA0ECAF9B3" pageId="25" pageNumber="25" rank="species">n. gen. and sp.</taxonomicNameLabel>
, in Fushun amber.
<emphasis id="B965EAF5CD25FF82FEF6F9D70F2BF9B3" bold="true" box="[335,360,1623,1647]" pageId="25" pageNumber="25">A.</emphasis>
Left dorsal oblique view.
<emphasis id="B965EAF5CD25FF82FDD7F9D70CC6F9B3" bold="true" box="[622,645,1623,1647]" pageId="25" pageNumber="25">B.</emphasis>
Right lateral oblique view.
</paragraph>
</caption>
<caption id="DF6E666FCD26FF81FFD2FB870AFAFBC1" ID-DOI="http://doi.org/10.5281/zenodo.14658498" ID-Zenodo-Dep="14658498" box="[107,1209,1029,1053]" httpUri="https://zenodo.org/record/14658498/files/figure.png" pageId="26" pageNumber="26" startId="26.[107,189,1031,1052]" targetBox="[107,1211,224,1005]" targetPageId="26" targetType="figure">
<paragraph id="8BAE36E7CD26FF81FFD2FB870AFAFBC1" blockId="26.[107,1209,1029,1053]" box="[107,1209,1029,1053]" pageId="26" pageNumber="26">
FIGURE 14. Photographs of head of
<taxonomicName id="4C114D64CD26FF81FE75FB850CDBFBC1" authorityName="Engel &amp; Xie" authorityYear="2024" box="[460,664,1029,1053]" class="Insecta" family="Megachilidae" genus="Glyptosmia" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="26" phylum="Arthropoda" rank="species" species="hemiaspis">
<emphasis id="B965EAF5CD26FF81FE75FB850CDBFBC1" box="[460,664,1029,1053]" italics="true" pageId="26" pageNumber="26">Glyptosmia hemiaspis</emphasis>
</taxonomicName>
.
<emphasis id="B965EAF5CD26FF81FD1AFB850CF8FBC1" bold="true" box="[675,699,1029,1053]" pageId="26" pageNumber="26">A.</emphasis>
Oblique facial view from upper right.
<emphasis id="B965EAF5CD26FF81FB97FB850A07FBC1" bold="true" box="[1070,1092,1029,1053]" pageId="26" pageNumber="26">B.</emphasis>
Facial view.
</paragraph>
</caption>
<paragraph id="8BAE36E7CD26FF87FF35F98C0FB0F998" blockId="26.[108,638,1085,1737]" lastBlockId="28.[108,636,1415,1736]" lastPageId="28" lastPageNumber="28" pageId="26" pageNumber="26">Head (fig. 14) with numerous, short, fine, white, erect to suberect, minutely branched setae, such setae not obscuring integument, setae denser in antennal basin and supraantennal area; such setae present on labrum, labrum without distinct patches of setae; clypeus and supraclypeal area with sparsely scattered short, simple setae; frons with scattered, short, simple to minutely branched setae, such setae denser in antennal basins and becoming sparser in ocellocular area and vertex. Mesoscutum with scattered, short, simple, erect, pale setae, such setae slightly longer on mesoscutellar surface except setae along margin of axilla and mesoscutellum denser, long to elongate, lightly fuscous, and minutely branched; pleura with scattered, short, erect, largely simple setae, such setae slightly longer posteriorly and on lateral surface of propodeum; basal area of propodeum without setae. Legs with short, simple or minutely branched, pale, suberect setae except mesotibia with such setae more numerous, slightly longer, and intermixed with some thicker setae, setae on inner surface distinctly longer; mesobasitarsus with similar setae except noticeably longer than those of mesotibial outer surface; metatibia with scopa composed of abundant elongate erect setae, those of outer surface minutely branched apically or some simple, those of inner surface as on outer surface except intermixed with numerous elongate, simple, characteristically hooked setae; metabasitarsus with abundant erect to suberect, elongate, minutely branched, pale setae; meso- and metafemora with sparse, short, largely simple to minutely branched setae, such setae especially minute and simple on ventral and posterior apical surfaces; meso- and metabasitarsi with setae similar to those of corresponding tibiae. Metasomal terga with scattered, short, fine, decumbent, pale setae, such setae sparse on anterior-facing surface of tergum I, intermixed with slightly longer, erect to suberect setae, some with minute branches, laterally; tergum II onward with short discal setae intermixed with somewhat more erect setae and slightly denser apically (but not forming setal bands); narrow apical marginal zones without setae, imbricate; tergum VI with setae denser medioapically; metasomal scopa on sterna II–V composed of bands of long simple erect setae; sternum VI without scopal setae, with dense brush of minute plumose fuscous setae apically, apical margin narrowly truncate; sting simple, sting sheaths thick, short, with some apical setae.</paragraph>
<caption id="DF6E666FCD27FF80FFD5FAA60A6FFAE0" ID-DOI="http://doi.org/10.5281/zenodo.14658502" ID-Zenodo-Dep="14658502" box="[108,1068,1316,1340]" httpUri="https://zenodo.org/record/14658502/files/figure.png" pageId="27" pageNumber="27" startId="27.[108,194,1318,1339]" targetBox="[108,1212,224,1292]" targetPageId="27" targetType="figure">
<paragraph id="8BAE36E7CD27FF80FFD5FAA60A6FFAE0" blockId="27.[108,1068,1316,1340]" box="[108,1068,1316,1340]" pageId="27" pageNumber="27">
FIGURE 15. Oblique dorsal view of mesoscutum and mesoscutellum of
<taxonomicName id="4C114D64CD27FF80FCF7FAA40A65FAE0" authorityName="Engel &amp; Xie" authorityYear="2024" box="[846,1062,1316,1340]" class="Insecta" family="Megachilidae" genus="Glyptosmia" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="27" phylum="Arthropoda" rank="species" species="hemiaspis">
<emphasis id="B965EAF5CD27FF80FCF7FAA40A65FAE0" box="[846,1062,1316,1340]" italics="true" pageId="27" pageNumber="27">Glyptosmia hemiaspis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF6E666FCD20FF87FFD2FAAC0FA6FA83" ID-DOI="http://doi.org/10.5281/zenodo.14658504" ID-Zenodo-Dep="14658504" httpUri="https://zenodo.org/record/14658504/files/figure.png" pageId="28" pageNumber="28" startId="28.[107,193,1324,1345]" targetBox="[107,1211,224,1296]" targetPageId="28" targetType="figure">
<paragraph id="8BAE36E7CD20FF87FFD2FAAC0FA6FA83" blockId="28.[107,1211,1322,1376]" pageId="28" pageNumber="28">
FIGURE 16. Profiles of
<taxonomicName id="4C114D64CD20FF87FEEEFAAA0C6CFA9E" authorityName="Engel &amp; Xie" authorityYear="2024" box="[343,559,1322,1346]" class="Insecta" family="Megachilidae" genus="Glyptosmia" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="28" phylum="Arthropoda" rank="species" species="hemiaspis">
<emphasis id="B965EAF5CD20FF87FEEEFAAA0C6CFA9E" box="[343,559,1322,1346]" italics="true" pageId="28" pageNumber="28">Glyptosmia hemiaspis</emphasis>
</taxonomicName>
.
<emphasis id="B965EAF5CD20FF87FD83FAAA0C10FA9E" bold="true" box="[570,595,1322,1346]" pageId="28" pageNumber="28">A.</emphasis>
Mesoscutellum, metanotum, and propodeum.
<emphasis id="B965EAF5CD20FF87FB96FAAA0A06FA9E" bold="true" box="[1071,1093,1322,1346]" pageId="28" pageNumber="28">B.</emphasis>
Metasomal terga I and II and base of tergum III.
</paragraph>
</caption>
<paragraph id="8BAE36E7CD20FF87FF35F9CE0F14F9B9" blockId="28.[108,636,1415,1736]" box="[140,343,1613,1638]" pageId="28" pageNumber="28">MALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30B656CCD20FF87FF35F9EF0AFFF914" pageId="28" pageNumber="28" type="materials_examined">
<paragraph id="8BAE36E7CD20FF87FF35F9EF0CB5F998" blockId="28.[108,636,1415,1736]" lastBlockId="28.[683,1214,1415,1736]" pageId="28" pageNumber="28">
<materialsCitation id="3B793CBACD20FF87FF35F9EF0CB5F998" country="China" county="Capital Normal University" location="Female" municipality="College of Life Sciences" pageId="28" pageNumber="28" specimenCount="1" stateProvince="Liaoning" typeStatus="holotype">
<typeStatus id="54AA8845CD20FF87FF35F9EF0F41F95B" box="[140,258,1647,1671]" pageId="28" pageNumber="28" type="holotype">HOLOTYPE</typeStatus>
:
<location id="8ECE603CCD20FF87FEB7F9EE0F1DF95A" LSID="urn:lsid:plazi:treatment:03B887F1CD24FF87FF36FB9B0AFFF914:8ECE603CCD20FF87FEB7F9EE0F1DF95A" box="[270,350,1646,1670]" country="China" county="Capital Normal University" municipality="College of Life Sciences" name="Female" pageId="28" pageNumber="28" stateProvince="Liaoning">Female</location>
, CNU-HYM-LF-2023-002 (fig. 13), in a single piece of amber without syninclusions,
<location id="8ECE603CCD20FF87FF51F9300F7AF914" LSID="urn:lsid:plazi:treatment:03B887F1CD24FF87FF36FB9B0AFFF914:8ECE603CCD20FF87FF51F9300F7AF914" box="[232,313,1712,1736]" country="China" county="Capital Normal University" municipality="College of Life Sciences" name="Fushun" pageId="28" pageNumber="28" stateProvince="Liaoning">Fushun</location>
coalfield,
<collectingRegion id="49D5F805CD20FF87FE15F9300C3BF914" box="[428,632,1712,1736]" country="China" name="Liaoning" pageId="28" pageNumber="28">Liaoning Province</collectingRegion>
, northeastern
<collectingCountry id="F3067677CD20FF87FCEDFA070DDFFA43" box="[852,924,1415,1439]" name="China" pageId="28" pageNumber="28">China</collectingCountry>
,
<location id="8ECE603CCD20FF87FC0BFA070AF9FA43" LSID="urn:lsid:plazi:treatment:03B887F1CD24FF87FF36FB9B0AFFF914:8ECE603CCD20FF87FC0BFA070AF9FA43" box="[946,1210,1415,1439]" country="China" county="Capital Normal University" municipality="College of Life Sciences" name="Guchengzi Formation" pageId="28" pageNumber="28" stateProvince="Liaoning">Guchengzi Formation</location>
,
<location id="8ECE603CCD20FF87FD15FA280D49FA1C" LSID="urn:lsid:plazi:treatment:03B887F1CD24FF87FF36FB9B0AFFF914:8ECE603CCD20FF87FD15FA280D49FA1C" box="[684,778,1448,1472]" country="China" county="Capital Normal University" municipality="College of Life Sciences" name="Ypresian" pageId="28" pageNumber="28" stateProvince="Liaoning">Ypresian</location>
(Eocene); deposited in the fossil insect collection of the
<location id="8ECE603CCD20FF87FCDAFA4A0C9BF9DE" LSID="urn:lsid:plazi:treatment:03B887F1CD24FF87FF36FB9B0AFFF914:8ECE603CCD20FF87FCDAFA4A0C9BF9DE" country="China" county="Capital Normal University" municipality="College of Life Sciences" name="Key Laboratory of Insect Evolution" pageId="28" pageNumber="28" stateProvince="Liaoning">Key Laboratory of Insect Evolution</location>
and
<location id="8ECE603CCD20FF87FCB4FA6A0A56F9DE" LSID="urn:lsid:plazi:treatment:03B887F1CD24FF87FF36FB9B0AFFF914:8ECE603CCD20FF87FCB4FA6A0A56F9DE" box="[781,1045,1514,1538]" country="China" county="Capital Normal University" municipality="College of Life Sciences" name="Environmental Changes" pageId="28" pageNumber="28" stateProvince="Liaoning">Environmental Changes</location>
,
<collectingMunicipality id="6BCAAC9DCD20FF87FBA6FA6A0D4EF9FF" pageId="28" pageNumber="28">College of Life Sciences</collectingMunicipality>
,
<collectingCounty id="62CF4E6BCD20FF87FCA7F98B0A16F9FF" box="[798,1109,1547,1571]" pageId="28" pageNumber="28">Capital Normal University</collectingCounty>
,
<collectingRegion id="49D5F805CD20FF87FBDEF98B0AFBF9FF" box="[1127,1208,1547,1571]" country="China" name="Beijing" pageId="28" pageNumber="28">Beijing</collectingRegion>
,
<collectingCountry id="F3067677CD20FF87FD15F9AC0CB1F998" box="[684,754,1580,1604]" name="China" pageId="28" pageNumber="28">China</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="8BAE36E7CD20FF87FD75F9CD0AFFF914" blockId="28.[683,1214,1415,1736]" pageId="28" pageNumber="28">
The
<typeStatus id="54AA8845CD20FF87FD44F9CD0D19F9B9" box="[765,858,1613,1637]" pageId="28" pageNumber="28" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C114D64CD20FF87FC38F9CD0A26F9B9" authorityName="Engel &amp; Xie" authorityYear="2024" box="[897,1125,1613,1637]" class="Insecta" family="Megachilidae" genus="Glyptosmia" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="28" phylum="Arthropoda" rank="species" species="hemiaspis">
<emphasis id="B965EAF5CD20FF87FC38F9CD0A26F9B9" box="[897,1125,1613,1637]" italics="true" pageId="28" pageNumber="28">Glyptosmia hemiaspis</emphasis>
</taxonomicName>
is challenging to interpret given its state of preservation (fig. 13). The bee is preserved with the head slightly pulled forward and slightly detached from meso-
</paragraph>
</subSubSection>
</treatment>
</document>
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