307 lines
37 KiB
XML
307 lines
37 KiB
XML
<document id="C2B6BED128394496F94A81800DFEB71C" ID-DOI="10.1206/0003-0090.469.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="14658466" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736967130833" checkinUser="felipe" docAuthor="Engel, Michael S. & Xie, Jiaying" docDate="2024" docId="03B887F1CD14FFB4FD52FA6A0F07FB06" docLanguage="en" docName="BulAmeMusNatHis.2024.B469.1-80.pdf" docOrigin="Bulletin of the American Museum of Natural History 2024 (469)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-2024/issue-469/0003-0090.469.1.1/The-Bee-Fauna-of-Eocene-Fushun-Amber-Hymenoptera-Apoidea/10.1206/0003-0090.469.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.10:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="10" docTitle="Thyreomelikertes kongi Engel & Xie, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="47" masterDocId="FF81FF89CD3CFF9BFFB9FF800E43FFDC" masterDocTitle="The Bee Fauna Of Eocene Fushun Amber (Hymenoptera: Apoidea)" masterLastPageNumber="81" masterPageNumber="1" pageNumber="40" updateTime="1736969989509" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="7B1CAF0950198FF411B826F646C3D4D5">
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<mods:title id="006275D3749D8738D03AAC927B5788DB">The Bee Fauna Of Eocene Fushun Amber (Hymenoptera: Apoidea)</mods:title>
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</mods:titleInfo>
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<mods:name id="04BD3D64E1AA259DDA69756ACB3FCFB1" type="personal">
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<mods:roleTerm id="7C79D84AC599D9EEA64F926368134E2D">Author</mods:roleTerm>
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<mods:namePart id="40D7D6F5CAEB7AD02D43BD11CA9D956D">Engel, Michael S.</mods:namePart>
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</mods:name>
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<mods:name id="1C8BBC355D30B0F87B97BAB31732B413" type="personal">
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<mods:role id="DFB5E68FE5570527D83B25FBF0F65706">
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<mods:roleTerm id="AD618EF8F41F0B32C743AAC995343205">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2E47D1EE68BBFD4AC8F0DAA5447C0560">Xie, Jiaying</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E5AFCE97D74DA42A02CCBEE55758D208">text</mods:typeOfResource>
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<mods:title id="60BB3F00FC672B7989CC9024855CEDA4">Bulletin of the American Museum of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="CCDB4205221ABF36CA90D27F0D2DD281">
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<mods:date id="E063B74D4785A2467585F2FD4F6064CD">2024</mods:date>
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<mods:number id="84C544C0780604F2243083F83E8916FF">2024-10-23</mods:number>
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<mods:detail id="2A1B95C5BD943B241ADA7239E3423DEB" type="volume">
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<mods:number id="AA4CD59EECBD151252A44515C8EED098">2024</mods:number>
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<mods:detail id="9B66DE25F406CEE7FE43D17D4C856F03" type="issue">
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<mods:number id="C3306ADF8D7404E80B1BAD9030560A89">469</mods:number>
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<mods:start id="F141373FD311AE91262D0B65F427F580">1</mods:start>
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<mods:location id="E9B6D9F853503004320087C2DD3AE326">
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<mods:url id="A88D6D9AF2A54863DF6417096B188417">https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-2024/issue-469/0003-0090.469.1.1/The-Bee-Fauna-of-Eocene-Fushun-Amber-Hymenoptera-Apoidea/10.1206/0003-0090.469.1.1.full</mods:url>
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</mods:location>
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<mods:classification id="AC040A3A0132C8E9D8CD7372E491BA6E">journal article</mods:classification>
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<mods:identifier id="72604EC3EA4278840A8A538A9F2693AB" type="DOI">10.1206/0003-0090.469.1.1</mods:identifier>
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<mods:identifier id="D4AF4854930017401F99C03A75F46400" type="ISSN">0003-0090</mods:identifier>
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<treatment id="03B887F1CD14FFB4FD52FA6A0F07FB06" LSID="urn:lsid:plazi:treatment:03B887F1CD14FFB4FD52FA6A0F07FB06" httpUri="http://treatment.plazi.org/id/03B887F1CD14FFB4FD52FA6A0F07FB06" lastPageId="47" lastPageNumber="47" pageId="40" pageNumber="40">
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<subSubSection id="C30B656CCD14FFB3FD52FA6A0A3EF9DE" box="[747,1149,1514,1538]" pageId="40" pageNumber="40" type="nomenclature">
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<paragraph id="8BAE36E7CD14FFB3FD52FA6A0A3EF9DE" blockId="40.[747,1149,1514,1538]" box="[747,1149,1514,1538]" pageId="40" pageNumber="40">
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<heading id="D0E6818BCD14FFB3FD52FA6A0A3EF9DE" box="[747,1149,1514,1538]" centered="true" fontSize="9" level="2" pageId="40" pageNumber="40" reason="2">
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<taxonomicName id="4C114D64CD14FFB3FD52FA6A0DACF9DE" authorityName="Engel & Xie" authorityYear="2024" box="[747,1007,1514,1538]" class="Insecta" family="Apidae" genus="Thyreomelikertes" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="40" phylum="Arthropoda" rank="species" species="kongi" status="sp. nov.">
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<emphasis id="B965EAF5CD14FFB3FD52FA6A0DACF9DE" bold="true" box="[747,1007,1514,1538]" italics="true" pageId="40" pageNumber="40">Thyreomelikertes kongi</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A256578ECD14FFB3FC45FA6B0A3EF9DE" box="[1020,1149,1514,1538]" pageId="40" pageNumber="40" rank="species">new species</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30B656CCD14FFB3FCDEF99C0A42F9E8" box="[871,1025,1564,1588]" pageId="40" pageNumber="40" type="description">
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<paragraph id="8BAE36E7CD14FFB3FCDEF99C0A42F9E8" blockId="40.[871,1025,1564,1588]" box="[871,1025,1564,1588]" pageId="40" pageNumber="40">
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<heading id="D0E6818BCD14FFB3FCDEF99C0A42F9E8" box="[871,1025,1564,1588]" centered="true" fontSize="9" level="2" pageId="40" pageNumber="40" reason="2">
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<figureCitation id="132A2A62CD14FFB3FCDEF99C0A42F9E8" box="[871,1025,1564,1588]" captionStart-0="FIGURE 26" captionStart-1="FIGURE 27" captionStart-2="FIGURE 28" captionStart-3="FIGURE 29" captionStart-4="FIGURE 30" captionStart-5="FIGURE 31" captionStartId-0="44.[107,194,1594,1615]" captionStartId-1="45.[108,195,1537,1558]" captionStartId-2="46.[109,195,726,747]" captionStartId-3="48.[108,194,1599,1620]" captionStartId-4="49.[108,194,1594,1615]" captionStartId-5="50.[108,194,1594,1615]" captionTargetBox-0="[120,1201,228,1549]" captionTargetBox-1="[108,1212,224,1505]" captionTargetBox-2="[107,1211,224,696]" captionTargetBox-3="[107,1211,224,1568]" captionTargetBox-4="[108,1212,224,1568]" captionTargetBox-5="[108,1212,224,1568]" captionTargetId-0="figure-9@44.[107,1211,224,1568]" captionTargetId-1="figure-8@45.[108,1212,224,1505]" captionTargetId-2="figure-405@46.[107,1211,224,696]" captionTargetId-3="figure-9@48.[107,1211,224,1568]" captionTargetId-4="figure-8@49.[108,1212,224,1568]" captionTargetId-5="figure-9@50.[108,1212,224,1568]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionTargetPageId-2="46" captionTargetPageId-3="48" captionTargetPageId-4="49" captionTargetPageId-5="50" captionText-0="FIGURE 26. Photographs of entire amber piece with holotype (CNU-HYM-LF-2023-003) and paratypes (CNU-HYM-LF-2023-004 and CNU-HYM-LF-2023-005) of Thyreomelikertes kongi, n. sp. A. Upper view, with holotype at right. B. Lower view, with holotype at left." captionText-1="FIGURE 27. Photographs of holotype (CNU-HYM-LF-2023-003) worker of Thyreomelikertes kongi, with microhymenopteran syninclusion. A. Left lateral-posterior oblique view. B. Dorsal view." captionText-2="FIGURE 28. Photograph of forewing venation of holotype (CNU-HYM-LF-2023-003) worker of Thyreomelikertes kongi." captionText-3="FIGURE 29. Photographs of holotype (CNU-HYM-LF-2023-003) worker of Thyreomelikertes kongi. A. Mesotibia, dorsal view. B. Metatibia, prolateral view." captionText-4="FIGURE 30. Photographs of holotype (CNU-HYM-LF-2023-003) worker of Thyreomelikertes kongi. A. Metatibia, metatarsus, and metapretarsus, retrolateral view. B. Detail of retromarginal fringe of metatibia, retrolateral view." captionText-5="FIGURE 31. Photographs of paratype workers of Thyreomelikertes kongi. A. Two workers together, left worker (CNU-HYM-LF-2023-005) in posterior view, right worker (CNU-HYM-LF-2023-004) in facial view. B. Facial view of CNU-HYM-LF-2023-004. C. Retromarginal fringe of CNU-HYM-LF-2023-005." figureDoi-0="http://doi.org/10.5281/zenodo.14658527" figureDoi-1="http://doi.org/10.5281/zenodo.14658529" figureDoi-2="http://doi.org/10.5281/zenodo.14658531" figureDoi-3="http://doi.org/10.5281/zenodo.14658533" figureDoi-4="http://doi.org/10.5281/zenodo.14658535" figureDoi-5="http://doi.org/10.5281/zenodo.14658537" httpUri-0="https://zenodo.org/record/14658527/files/figure.png" httpUri-1="https://zenodo.org/record/14658529/files/figure.png" httpUri-2="https://zenodo.org/record/14658531/files/figure.png" httpUri-3="https://zenodo.org/record/14658533/files/figure.png" httpUri-4="https://zenodo.org/record/14658535/files/figure.png" httpUri-5="https://zenodo.org/record/14658537/files/figure.png" pageId="40" pageNumber="40">Figures 26–31</figureCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30B656CCD14FFB2FD75F9CD0D55F9B9" lastPageId="41" lastPageNumber="41" pageId="40" pageNumber="40" type="diagnosis">
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<paragraph id="8BAE36E7CD14FFB2FD75F9CD0D55F9B9" blockId="40.[684,1213,1613,1736]" lastBlockId="41.[684,1212,1514,1736]" lastPageId="41" lastPageNumber="41" pageId="40" pageNumber="40">
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DIAGNOSIS: This species, which is known from three individuals (figs. 26, 27, 31A), can be distinguished by the distinctly more elongate fringe, fundal, and corbicular setae on the metatibia (figs. 27, 29, 30, 31A, B), the setae much longer than the maximum metatibal width. Likewise, the setae of the mesotibia are much longer than the maximum mesotibial width (up to about 1.5× mesotibial width) and none of the setae are capitate. The species is overall smaller than that of the
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<typeStatus id="54AA8845CD15FFB2FE44F90F0C68F97B" box="[509,555,1679,1703]" pageId="41" pageNumber="41">type</typeStatus>
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species (refer to metrics) and the apical margin of the metabasitarsus is concave (fig. 30A) (rather than straight). The facial setae, particularly on the clypeus, are seemingly denser than in
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<emphasis id="B965EAF5CD15FFB2FBA0F9AC0AFFF998" box="[1049,1212,1580,1604]" italics="true" pageId="41" pageNumber="41">
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T.
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<taxonomicName id="4C114D64CD15FFB2FB8DF9AC0AFFF998" authorityName="Engel & Xie" authorityYear="2024" box="[1076,1212,1580,1604]" class="Insecta" family="Apidae" genus="Thyreomelikertes" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="41" phylum="Arthropoda" rank="species" species="electrosinicus">electrosinicus</taxonomicName>
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</emphasis>
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(fig. 31B).
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</paragraph>
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</subSubSection>
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<caption id="DF6E666FCD15FFB2FFD5FA320A04FA14" ID-DOI="http://doi.org/10.5281/zenodo.14658521" ID-Zenodo-Dep="14658521" box="[108,1095,1456,1480]" httpUri="https://zenodo.org/record/14658521/files/figure.png" pageId="41" pageNumber="41" startId="41.[108,194,1458,1479]" targetBox="[109,1213,224,1431]" targetPageId="41" targetType="figure">
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<paragraph id="8BAE36E7CD15FFB2FFD5FA320A04FA14" blockId="41.[108,1095,1456,1480]" box="[108,1095,1456,1480]" pageId="41" pageNumber="41">
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FIGURE 23. Photographs of
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<taxonomicName id="4C114D64CD15FFB2FE2BFA300C82FA14" authorityName="Engel & Xie" authorityYear="2024" box="[402,705,1456,1480]" class="Insecta" family="Apidae" genus="Thyreomelikertes" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="41" phylum="Arthropoda" rank="species" species="electrosinicus">
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<emphasis id="B965EAF5CD15FFB2FE2BFA300C82FA14" box="[402,705,1456,1480]" italics="true" pageId="41" pageNumber="41">Thyreomelikertes electrosinicus</emphasis>
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</taxonomicName>
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.
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<emphasis id="B965EAF5CD15FFB2FD77FA300CA4FA14" bold="true" box="[718,743,1456,1480]" pageId="41" pageNumber="41">A.</emphasis>
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Mesosomal dorsum.
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<emphasis id="B965EAF5CD15FFB2FC7DFA300D98FA14" bold="true" box="[964,987,1456,1480]" pageId="41" pageNumber="41">B.</emphasis>
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Forewing.
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</paragraph>
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</caption>
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<subSubSection id="C30B656CCD15FFB5FD75F9EE0A7FFA81" lastPageId="46" lastPageNumber="46" pageId="41" pageNumber="41" type="description">
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<paragraph id="8BAE36E7CD15FFB5FD75F9EE0F25F95B" blockId="41.[684,1212,1514,1736]" lastBlockId="46.[108,638,820,1737]" lastPageId="46" lastPageNumber="46" pageId="41" pageNumber="41">
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DESCRIPTION: As described for
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<taxonomicName id="4C114D64CD15FFB2FB88F9EE0D34F97B" authorityName="Engel & Xie" authorityYear="2024" class="Insecta" family="Apidae" genus="Thyreomelikertes" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="41" phylum="Arthropoda" rank="species" species="electrosinicus">
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<emphasis id="B965EAF5CD15FFB2FB88F9EE0D34F97B" italics="true" pageId="41" pageNumber="41">Thyreomelikertes electrosinicus</emphasis>
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</taxonomicName>
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(above), with the following exceptions: WORKER. Total body length (as preserved)
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<quantity id="4CE99B02CD12FFB5FF7AFCB40F65FC90" box="[195,294,820,844]" metricMagnitude="-3" metricUnit="m" metricValue="3.51" pageId="46" pageNumber="46" unit="mm" value="3.51">3.51 mm</quantity>
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; forewing length (as preserved)
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<quantity id="4CE99B02CD12FFB5FFD7FCD50E8CFCB1" box="[110,207,853,877]" metricMagnitude="-3" metricUnit="m" metricValue="3.78" pageId="46" pageNumber="46" unit="mm" value="3.78">3.78 mm</quantity>
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. Head slightly wider than long (fig. 31B), length (summit of vertex to clypeal apical margin)
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<quantity id="4CE99B02CD12FFB5FFD5FC170E8EFC73" box="[108,205,919,943]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="46" pageNumber="46" unit="mm" value="1.16">1.16 mm</quantity>
|
||
(
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||
<typeStatus id="54AA8845CD12FFB5FF64FC170F7FFC73" box="[221,316,919,943]" pageId="46" pageNumber="46" type="paratype">paratype</typeStatus>
|
||
<quantity id="4CE99B02CD12FFB5FEFDFC170FEBFC73" box="[324,424,919,943]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="46" pageNumber="46" unit="mm" value="1.21">1.21 mm</quantity>
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), width (maximum width across compound eyes)
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<quantity id="4CE99B02CD12FFB5FE13FC380C4BFC0C" box="[426,520,952,976]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="46" pageNumber="46" unit="mm" value="1.66">1.66 mm</quantity>
|
||
(
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||
<typeStatus id="54AA8845CD12FFB5FDACFC380C3EFC0C" box="[533,637,952,976]" pageId="46" pageNumber="46" type="paratype">paratypes</typeStatus>
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||
<quantity id="4CE99B02CD12FFB5FFD5FC590F4FFC2E" box="[108,268,985,1010]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" metricValueMax="1.46" metricValueMin="1.32" pageId="46" pageNumber="46" unit="mm" value="1.39" valueMax="1.46" valueMin="1.32">1.32–1.46 mm</quantity>
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||
). Scape elongate, length
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||
<quantity id="4CE99B02CD12FFB5FDA3FC590C3FFC2D" box="[538,636,985,1009]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="46" pageNumber="46" unit="mm" value="0.47">0.47 mm</quantity>
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||
(artificially stretched) (
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||
<typeStatus id="54AA8845CD12FFB5FEDFFC7B0F8CFBCF" box="[358,463,1019,1043]" pageId="46" pageNumber="46" type="paratype">paratypes</typeStatus>
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||
<quantity id="4CE99B02CD12FFB5FE60FC7B0C3BFBCE" box="[473,632,1019,1043]" metricMagnitude="-4" metricUnit="m" metricValue="3.05" metricValueMax="3.8" metricValueMin="2.3" pageId="46" pageNumber="46" unit="mm" value="0.305" valueMax="0.38" valueMin="0.23">0.23–0.38 mm</quantity>
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||
, both damaged), longer than torulocellar distance; flagellomere I longer than flagellomere II, shorter than combined lengths of flagellomeres II and III, flagellomere II shorter than flagellomere III. Mesoscutum medial length
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||
<quantity id="4CE99B02CD12FFB5FE2CFB200FB6FB6B" box="[405,501,1184,1208]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="46" pageNumber="46" unit="mm" value="0.57">0.57 mm</quantity>
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||
; intertegular distance
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||
<quantity id="4CE99B02CD12FFB5FF72FB410F68FB04" box="[203,299,1217,1241]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="46" pageNumber="46" unit="mm" value="0.67">0.67 mm</quantity>
|
||
(
|
||
<typeStatus id="54AA8845CD12FFB5FE83FB410FDAFB05" box="[314,409,1217,1241]" pageId="46" pageNumber="46" type="paratype">paratype</typeStatus>
|
||
<quantity id="4CE99B02CD12FFB5FE19FB410C40FB05" box="[416,515,1217,1241]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="46" pageNumber="46" unit="mm" value="0.55">0.55 mm</quantity>
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||
); mesoscutellum medial length
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||
<quantity id="4CE99B02CD12FFB5FEEBFB620FF7FB25" box="[338,436,1250,1274]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="46" pageNumber="46" unit="mm" value="0.36">0.36 mm</quantity>
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||
. Metatibia slender, length
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||
<quantity id="4CE99B02CD12FFB5FF0FFA830F56FAC6" box="[182,277,1283,1307]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="46" pageNumber="46" unit="mm" value="1.5">1.50 mm</quantity>
|
||
(
|
||
<typeStatus id="54AA8845CD12FFB5FE9BFA830FC9FAC7" box="[290,394,1283,1307]" pageId="46" pageNumber="46" type="paratype">paratypes</typeStatus>
|
||
<quantity id="4CE99B02CD12FFB5FE29FA830C6FFAC7" box="[400,556,1283,1307]" metricMagnitude="-3" metricUnit="m" metricValue="1.36" metricValueMax="1.61" metricValueMin="1.11" pageId="46" pageNumber="46" unit="mm" value="1.36" valueMax="1.61" valueMin="1.11">1.11–1.61 mm</quantity>
|
||
), maximum width
|
||
<quantity id="4CE99B02CD12FFB5FF4AFAA40F10FAE0" box="[243,339,1316,1340]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="46" pageNumber="46" unit="mm" value="0.45">0.45 mm</quantity>
|
||
(
|
||
<typeStatus id="54AA8845CD12FFB5FEDBFAA40F89FAE0" box="[354,458,1316,1340]" pageId="46" pageNumber="46" type="paratype">paratypes</typeStatus>
|
||
<quantity id="4CE99B02CD12FFB5FE6BFAA40C33FAE0" box="[466,624,1316,1340]" metricMagnitude="-4" metricUnit="m" metricValue="2.9499999999999997" metricValueMax="3.7" metricValueMin="2.2" pageId="46" pageNumber="46" unit="mm" value="0.295" valueMax="0.37" valueMin="0.22">0.22–0.37 mm</quantity>
|
||
); metabasitarsus longer than wide, length
|
||
<quantity id="4CE99B02CD12FFB5FDA7FAC50C3DFA81" box="[542,638,1349,1373]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="46" unit="mm" value="0.5">0.50 mm</quantity>
|
||
(
|
||
<typeStatus id="54AA8845CD12FFB5FFCDFAE60E97FAA2" box="[116,212,1382,1406]" pageId="46" pageNumber="46" type="paratype">paratype</typeStatus>
|
||
<quantity id="4CE99B02CD12FFB5FF64FAE60F07FAA2" box="[221,324,1382,1406]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="46" pageNumber="46" unit="mm" value="0.34">0.34 mm</quantity>
|
||
), maximum width
|
||
<quantity id="4CE99B02CD12FFB5FDA3FAE60C3DFAA2" box="[538,638,1382,1406]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="46" pageNumber="46" unit="mm" value="0.37">0.37 mm</quantity>
|
||
(
|
||
<typeStatus id="54AA8845CD12FFB5FFCAFA070E91FA43" box="[115,210,1415,1439]" pageId="46" pageNumber="46" type="paratype">paratype</typeStatus>
|
||
<quantity id="4CE99B02CD12FFB5FF61FA070F7AFA43" box="[216,313,1415,1439]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="46" pageNumber="46" unit="mm" value="0.22">0.22 mm</quantity>
|
||
), apical margin concave. Forewing (fig. 28) with basal vein (1M) slightly proximad 1cu-a, pterostigma maximum width at midlength; marginal cell apex well separated from anterior wing margin, with nebulous appendiculate vein; 3Rs much shorter than both r-rs and 4Rs; 1rs-m arched; 2m-cu proximal 2rs-m by about 2–3× vein width.
|
||
</paragraph>
|
||
<caption id="DF6E666FCD16FFB1FFD2FA3C0F26FA33" ID-DOI="http://doi.org/10.5281/zenodo.14658523" ID-Zenodo-Dep="14658523" httpUri="https://zenodo.org/record/14658523/files/figure.png" pageId="42" pageNumber="42" startId="42.[107,193,1468,1489]" targetBox="[108,1212,224,1442]" targetPageId="42" targetType="figure">
|
||
<paragraph id="8BAE36E7CD16FFB1FFD2FA3C0F26FA33" blockId="42.[107,1211,1466,1519]" pageId="42" pageNumber="42">
|
||
FIGURE 24. Photographs of
|
||
<taxonomicName id="4C114D64CD16FFB1FE29FA3A0CFCFA0E" authorityName="Engel & Xie" authorityYear="2024" box="[400,703,1466,1490]" class="Insecta" family="Apidae" genus="Thyreomelikertes" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="42" phylum="Arthropoda" rank="species" species="electrosinicus">
|
||
<emphasis id="B965EAF5CD16FFB1FE29FA3A0CFCFA0E" box="[400,703,1466,1490]" italics="true" pageId="42" pageNumber="42">Thyreomelikertes electrosinicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B965EAF5CD16FFB1FD75FA3A0CA7FA0E" bold="true" box="[716,740,1466,1490]" pageId="42" pageNumber="42">A.</emphasis>
|
||
Mesotibia and mesotarsus.
|
||
<emphasis id="B965EAF5CD16FFB1FBB8FA3A0A54FA0E" bold="true" box="[1025,1047,1466,1490]" pageId="42" pageNumber="42">B.</emphasis>
|
||
Detail of mesotibia, note capitate setae.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6E666FCD17FFB0FFD5F9BC0C8CF9B3" ID-DOI="http://doi.org/10.5281/zenodo.14658525" ID-Zenodo-Dep="14658525" httpUri="https://zenodo.org/record/14658525/files/figure.png" pageId="43" pageNumber="43" startId="43.[108,194,1596,1617]" targetBox="[108,1212,224,1568]" targetPageId="43" targetType="figure">
|
||
<paragraph id="8BAE36E7CD17FFB0FFD5F9BC0C8CF9B3" blockId="43.[108,1214,1594,1647]" pageId="43" pageNumber="43">
|
||
FIGURE 25. Photographs of
|
||
<taxonomicName id="4C114D64CD17FFB0FE2FF9BA0C86F98E" authorityName="Engel & Xie" authorityYear="2024" box="[406,709,1594,1618]" class="Insecta" family="Apidae" genus="Thyreomelikertes" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="43" phylum="Arthropoda" rank="species" species="electrosinicus">
|
||
<emphasis id="B965EAF5CD17FFB0FE2FF9BA0C86F98E" box="[406,709,1594,1618]" italics="true" pageId="43" pageNumber="43">Thyreomelikertes electrosinicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B965EAF5CD17FFB0FD6AF9BA0CA8F98E" bold="true" box="[723,747,1594,1618]" pageId="43" pageNumber="43">A.</emphasis>
|
||
Metatibia, metatarsus, and metapretarsus.
|
||
<emphasis id="B965EAF5CD17FFB0FB11F9BA0AFDF98E" bold="true" box="[1192,1214,1594,1618]" pageId="43" pageNumber="43">B.</emphasis>
|
||
Detail of metatibial setae and bristles on proventral margin.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6E666FCD10FFB7FFD2F9BA0C84F957" ID-DOI="http://doi.org/10.5281/zenodo.14658527" ID-Zenodo-Dep="14658527" httpUri="https://zenodo.org/record/14658527/files/figure.png" pageId="44" pageNumber="44" startId="44.[107,194,1594,1615]" targetBox="[120,1201,228,1549]" targetPageId="44" targetType="figure">
|
||
<paragraph id="8BAE36E7CD10FFB7FFD2F9BA0C84F957" blockId="44.[107,1211,1592,1675]" pageId="44" pageNumber="44">
|
||
FIGURE 26. Photographs of entire amber piece with holotype (CNU-HYM-LF-2023-003) and paratypes (CNU-HYM-LF-2023-004 and CNU-HYM-LF-2023-005) of
|
||
<taxonomicName id="4C114D64CD10FFB7FD5CF9D60D8EF9B2" authorityName="Engel & Xie" authorityYear="2024" box="[741,973,1622,1646]" class="Insecta" family="Apidae" genus="Thyreomelikertes" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="44" phylum="Arthropoda" rank="species" species="kongi" status="sp. nov.">
|
||
<emphasis id="B965EAF5CD10FFB7FD5CF9D60D8EF9B2" box="[741,973,1622,1646]" italics="true" pageId="44" pageNumber="44">Thyreomelikertes kongi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A256578ECD10FFB7FC65F9D60A56F9B1" box="[988,1045,1622,1645]" pageId="44" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
|
||
<emphasis id="B965EAF5CD10FFB7FBA4F9D50A76F9B1" bold="true" box="[1053,1077,1621,1645]" pageId="44" pageNumber="44">A.</emphasis>
|
||
Upper view, with holotype at right.
|
||
<emphasis id="B965EAF5CD10FFB7FEEEF9F30F2EF957" bold="true" box="[343,365,1651,1675]" pageId="44" pageNumber="44">B.</emphasis>
|
||
Lower view, with holotype at left.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6E666FCD11FFB6FFD5F9810DB3F9E8" ID-DOI="http://doi.org/10.5281/zenodo.14658529" ID-Zenodo-Dep="14658529" httpUri="https://zenodo.org/record/14658529/files/figure.png" pageId="45" pageNumber="45" startId="45.[108,195,1537,1558]" targetBox="[108,1212,224,1505]" targetPageId="45" targetType="figure">
|
||
<paragraph id="8BAE36E7CD11FFB6FFD5F9810DB3F9E8" blockId="45.[108,1212,1535,1589]" pageId="45" pageNumber="45">
|
||
FIGURE 27. Photographs of holotype (CNU-HYM-LF-2023-003) worker of
|
||
<taxonomicName id="4C114D64CD11FFB6FC2DFA7F0A3CF9CB" authorityName="Engel & Xie" authorityYear="2024" box="[916,1151,1535,1559]" class="Insecta" family="Apidae" genus="Thyreomelikertes" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="45" phylum="Arthropoda" rank="species" species="kongi">
|
||
<emphasis id="B965EAF5CD11FFB6FC2DFA7F0A3CF9CB" box="[916,1151,1535,1559]" italics="true" pageId="45" pageNumber="45">Thyreomelikertes kongi</emphasis>
|
||
</taxonomicName>
|
||
, with microhymenopteran syninclusion.
|
||
<emphasis id="B965EAF5CD11FFB6FE68F99C0FAAF9E8" bold="true" box="[465,489,1564,1588]" pageId="45" pageNumber="45">A.</emphasis>
|
||
Left lateral-posterior oblique view.
|
||
<emphasis id="B965EAF5CD11FFB6FCECF99D0D2FF9E9" bold="true" box="[853,876,1565,1589]" pageId="45" pageNumber="45">B.</emphasis>
|
||
Dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6E666FCD12FFB5FFD4FD560EB5FCD5" ID-DOI="http://doi.org/10.5281/zenodo.14658531" ID-Zenodo-Dep="14658531" httpUri="https://zenodo.org/record/14658531/files/figure.png" pageId="46" pageNumber="46" startId="46.[109,195,726,747]" targetBox="[107,1211,224,696]" targetPageId="46" targetType="figure">
|
||
<paragraph id="8BAE36E7CD12FFB5FFD4FD560EB5FCD5" blockId="46.[108,1212,724,778]" pageId="46" pageNumber="46">
|
||
FIGURE 28. Photograph of forewing venation of holotype (CNU-HYM-LF-2023-003) worker of
|
||
<taxonomicName id="4C114D64CD12FFB5FBEBFD540EACFCD6" authorityName="Engel & Xie" authorityYear="2024" class="Insecta" family="Apidae" genus="Thyreomelikertes" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="46" phylum="Arthropoda" rank="species" species="kongi">
|
||
<emphasis id="B965EAF5CD12FFB5FBEBFD540EACFCD6" italics="true" pageId="46" pageNumber="46">Thyreomelikertes kongi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAE36E7CD12FFB5FF35F9100A0BFAE0" blockId="46.[108,638,820,1737]" lastBlockId="46.[684,1214,820,1737]" pageId="46" pageNumber="46">Mesotibia with dense, erect setae and bristles (fig. 29A), on dorsal surface, such setae and bristles progressively longer from base to apex, setae with some minute branches, retrolateral surface with bristles sparse and setae longer, longer than mesotibial width, such setae with abundant minute branches, no capitate setae; metatibia with dense elongate (distinctly longer than metatibial width), minutely branched setae forming retromarginal fringe (figs. 29B, 30, 31A), similar but shorter setae along proventral margin, retromarginal fringe sparsely intermixed with long, erect bristles and such bristles sparse along anterior edge of proventral surface and inferiorly on prolateral surface, proventral surface largely asetose, prolateral surface with sparsely scattered long, erect, simple setae and bristles, such setae similar on profundal and corbicular surfaces.</paragraph>
|
||
<paragraph id="8BAE36E7CD12FFB5FD75FAC50A7FFA81" blockId="46.[684,1214,820,1737]" box="[716,1084,1349,1373]" pageId="46" pageNumber="46">QUEEN AND MALE. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B656CCD12FFB4FD75FAE60FEBFBAA" lastPageId="47" lastPageNumber="47" pageId="46" pageNumber="46" type="materials_examined">
|
||
<paragraph id="8BAE36E7CD12FFB5FD75FAE60DB6F95B" blockId="46.[684,1214,820,1737]" pageId="46" pageNumber="46">
|
||
<materialsCitation id="3B793CBACD12FFB5FD75FAE60DB6F95B" country="China" county="Capital Normal University" location="Female" municipality="College of Life Sciences" pageId="46" pageNumber="46" specimenCount="1" stateProvince="Liaoning" typeStatus="holotype">
|
||
<typeStatus id="54AA8845CD12FFB5FD75FAE60D05FAA2" box="[716,838,1382,1406]" pageId="46" pageNumber="46" type="holotype">HOLOTYPE</typeStatus>
|
||
:
|
||
<location id="8ECE603CCD12FFB5FCD9FAE60DF2FAA2" LSID="urn:lsid:plazi:treatment:03B887F1CD14FFB4FD52FA6A0F07FB06:8ECE603CCD12FFB5FCD9FAE60DF2FAA2" box="[864,945,1382,1406]" country="China" county="Capital Normal University" municipality="College of Life Sciences" name="Female" pageId="46" pageNumber="46" stateProvince="Liaoning">Female</location>
|
||
worker, CNU-HYM- LF-2023-003 (figs. 26, 27), in a single piece of amber with two further bees as syninclusions,
|
||
<location id="8ECE603CCD12FFB5FD15FA490CBFFA3D" LSID="urn:lsid:plazi:treatment:03B887F1CD14FFB4FD52FA6A0F07FB06:8ECE603CCD12FFB5FD15FA490CBFFA3D" box="[684,764,1481,1505]" country="China" county="Capital Normal University" municipality="College of Life Sciences" name="Fushun" pageId="46" pageNumber="46" stateProvince="Liaoning">Fushun</location>
|
||
coalfield,
|
||
<collectingRegion id="49D5F805CD12FFB5FCDFFA490A68FA3D" box="[870,1067,1481,1505]" country="China" name="Liaoning" pageId="46" pageNumber="46">Liaoning Province</collectingRegion>
|
||
, northeastern
|
||
<collectingCountry id="F3067677CD12FFB5FD15FA6A0CB3F9DE" box="[684,752,1514,1538]" name="China" pageId="46" pageNumber="46">China</collectingCountry>
|
||
,
|
||
<location id="8ECE603CCD12FFB5FD43FA6A0DA4F9DE" LSID="urn:lsid:plazi:treatment:03B887F1CD14FFB4FD52FA6A0F07FB06:8ECE603CCD12FFB5FD43FA6A0DA4F9DE" box="[762,999,1514,1538]" country="China" county="Capital Normal University" municipality="College of Life Sciences" name="Guchengzi Formation" pageId="46" pageNumber="46" stateProvince="Liaoning">Guchengzi Formation</location>
|
||
,
|
||
<location id="8ECE603CCD12FFB5FC4AFA6A0A0CF9DE" LSID="urn:lsid:plazi:treatment:03B887F1CD14FFB4FD52FA6A0F07FB06:8ECE603CCD12FFB5FC4AFA6A0A0CF9DE" box="[1011,1103,1514,1538]" country="China" county="Capital Normal University" municipality="College of Life Sciences" name="Ypresian" pageId="46" pageNumber="46" stateProvince="Liaoning">Ypresian</location>
|
||
(Eocene); deposited in the fossil insect collection of the
|
||
<location id="8ECE603CCD12FFB5FB2DF98C0DB5F998" LSID="urn:lsid:plazi:treatment:03B887F1CD14FFB4FD52FA6A0F07FB06:8ECE603CCD12FFB5FB2DF98C0DB5F998" country="China" county="Capital Normal University" municipality="College of Life Sciences" name="Key Laboratory of Insect Evolution" pageId="46" pageNumber="46" stateProvince="Liaoning">Key Laboratory of Insect Evolution</location>
|
||
and
|
||
<location id="8ECE603CCD12FFB5FB94F9AC0D6AF9BA" LSID="urn:lsid:plazi:treatment:03B887F1CD14FFB4FD52FA6A0F07FB06:8ECE603CCD12FFB5FB94F9AC0D6AF9BA" country="China" county="Capital Normal University" municipality="College of Life Sciences" name="Environmental Changes" pageId="46" pageNumber="46" stateProvince="Liaoning">Environmental Changes</location>
|
||
,
|
||
<collectingMunicipality id="6BCAAC9DCD12FFB5FC8AF9CE0A6EF9BA" box="[819,1069,1614,1638]" pageId="46" pageNumber="46">College of Life Sciences</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62CF4E6BCD12FFB5FB8EF9CE0D0AF95B" pageId="46" pageNumber="46">Capital Normal University</collectingCounty>
|
||
,
|
||
<collectingRegion id="49D5F805CD12FFB5FCEDF9EF0DE2F95B" box="[852,929,1647,1671]" country="China" name="Beijing" pageId="46" pageNumber="46">Beijing</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3067677CD12FFB5FC15F9EF0DB2F95B" box="[940,1009,1647,1671]" name="China" pageId="46" pageNumber="46">China</collectingCountry>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BAE36E7CD12FFB4FD74F9100FA6FE3D" blockId="46.[684,1214,820,1737]" lastBlockId="47.[108,637,226,1242]" lastPageId="47" lastPageNumber="47" pageId="46" pageNumber="46">
|
||
The
|
||
<typeStatus id="54AA8845CD12FFB5FD44F9100D1FF974" box="[765,860,1680,1704]" pageId="46" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
is preserved in a relatively clear piece of amber and has two syninclusions of the same species as well as a tiny chalcidoid wasp near the wing apex. The
|
||
<typeStatus id="54AA8845CD13FFB4FE30FE830FA9FEC7" box="[393,490,259,283]" pageId="47" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
is located at one end of the piece, while the two other bees are clustered together and overlapping each other. The
|
||
<typeStatus id="54AA8845CD13FFB4FF1CFEE60F4BFEA2" box="[165,264,358,382]" pageId="47" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
is well preserved and can be observed in all orientations except ventral. The wings are obliquely folded over the body and the legs tucked up alongside the body.
|
||
</paragraph>
|
||
<paragraph id="8BAE36E7CD13FFB4FF34FE6A0C3EFCD7" blockId="47.[108,637,226,1242]" pageId="47" pageNumber="47">
|
||
<materialsCitation id="3B793CBACD13FFB4FF34FE6A0E93FD74" country="China" county="Ypresian" location="Fushun" municipality="Guchengzi Formation" pageId="47" pageNumber="47" specimenCount="2" stateProvince="Liaoning" typeStatus="paratype">
|
||
<typeStatus id="54AA8845CD13FFB4FF34FE6A0F4AFDDE" box="[141,265,490,514]" pageId="47" pageNumber="47" type="paratype">PARATYPES</typeStatus>
|
||
: Female workers, CNU-HYM- LF-2023-004 and CNU-HYM-LF-2023-005, in the same piece of amber with the
|
||
<typeStatus id="54AA8845CD13FFB4FE50FDAC0C0BFD98" box="[489,584,556,580]" pageId="47" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
(fig. 26),
|
||
<location id="8ECE603CCD13FFB4FF24FDCD0EADFDB9" LSID="urn:lsid:plazi:treatment:03B887F1CD14FFB4FD52FA6A0F07FB06:8ECE603CCD13FFB4FF24FDCD0EADFDB9" box="[157,238,589,613]" country="China" county="Ypresian" municipality="Guchengzi Formation" name="Fushun" pageId="47" pageNumber="47" stateProvince="Liaoning">Fushun</location>
|
||
coalfield,
|
||
<collectingRegion id="49D5F805CD13FFB4FEE6FDCD0C6AFDB9" box="[351,553,589,613]" country="China" name="Liaoning" pageId="47" pageNumber="47">Liaoning Province</collectingRegion>
|
||
, northeastern
|
||
<collectingCountry id="F3067677CD13FFB4FF7CFDEE0F48FD5A" box="[197,267,622,646]" name="China" pageId="47" pageNumber="47">China</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="6BCAAC9DCD13FFB4FEA0FDEE0C4CFD5A" box="[281,527,622,646]" pageId="47" pageNumber="47">Guchengzi Formation</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62CF4E6BCD13FFB4FDA4FDEE0C3FFD5A" box="[541,636,622,646]" pageId="47" pageNumber="47">Ypresian</collectingCounty>
|
||
(Eocene)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B793CBACD13FFB4FF63FD0F0C3BFCD7" country="China" location="Key Laboratory of Insect Evolution" municipality="College of Life Sciences" pageId="47" pageNumber="47" specimenCount="1" stateProvince="Beijing" typeStatus="paratype">
|
||
deposited in the fossil insect collection of the
|
||
<location id="8ECE603CCD13FFB4FF00FD310C0AFD14" LSID="urn:lsid:plazi:treatment:03B887F1CD14FFB4FD52FA6A0F07FB06:8ECE603CCD13FFB4FF00FD310C0AFD14" box="[185,585,688,713]" country="China" municipality="College of Life Sciences" name="Key Laboratory of Insect Evolution" pageId="47" pageNumber="47" stateProvince="Beijing">Key Laboratory of Insect Evolution</location>
|
||
and Environmental Changes,
|
||
<collectingMunicipality id="6BCAAC9DCD13FFB4FE2DFD520EEAFCD6" pageId="47" pageNumber="47">College of Life Sciences</collectingMunicipality>
|
||
, Capital Normal University,
|
||
<collectingRegion id="49D5F805CD13FFB4FE62FD730C69FCD7" box="[475,554,755,779]" country="China" name="Beijing" pageId="47" pageNumber="47">Beijing</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3067677CD13FFB4FD8AFD730C3BFCD7" box="[563,632,755,779]" name="China" pageId="47" pageNumber="47">China</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAE36E7CD13FFB4FF35FC940FEBFBAA" blockId="47.[108,637,226,1242]" pageId="47" pageNumber="47">
|
||
The
|
||
<typeStatus id="54AA8845CD13FFB4FF7BFC940F6EFCF0" box="[194,301,788,812]" pageId="47" pageNumber="47" type="paratype">paratypes</typeStatus>
|
||
are distinctly less well preserved relative to the
|
||
<typeStatus id="54AA8845CD13FFB4FEDFFCB50F84FC91" box="[358,455,821,845]" pageId="47" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
and, given that they are partially entwined as well as damaged through compression or missing sclerites, it was not possible to see each structure as well as in the
|
||
<typeStatus id="54AA8845CD13FFB4FFD5FC390E8DFC0D" box="[108,206,953,977]" pageId="47" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
. It was also not possible to get all of the same measurements from the
|
||
<typeStatus id="54AA8845CD13FFB4FE7BFC5B0C68FC2F" box="[450,555,987,1011]" pageId="47" pageNumber="47" type="paratype">paratypes</typeStatus>
|
||
as was possible from the
|
||
<typeStatus id="54AA8845CD13FFB4FEFFFC7B0FE8FBCF" box="[326,427,1019,1043]" pageId="47" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
. From observable characters, the
|
||
<typeStatus id="54AA8845CD13FFB4FEB6FB9D0F34FBE9" box="[271,375,1053,1077]" pageId="47" pageNumber="47" type="paratype">paratypes</typeStatus>
|
||
agree perfectly with the
|
||
<typeStatus id="54AA8845CD13FFB4FFD5FBBD0E8FFB89" box="[108,204,1085,1109]" pageId="47" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
and so we are confident in their conspecificity with the
|
||
<typeStatus id="54AA8845CD13FFB4FEFBFBDE0FE7FBAA" box="[322,420,1118,1142]" pageId="47" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B656CCD13FFB4FF34FB000F07FB06" pageId="47" pageNumber="47" type="etymology">
|
||
<paragraph id="8BAE36E7CD13FFB4FF34FB000F07FB06" blockId="47.[108,637,226,1242]" pageId="47" pageNumber="47">ETYMOLOGY: The specific epithet honors Chuijin Kong, who graciously donated the specimens for this study.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |