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<mods:title id="C66F4CF813E1D08E8DF31848B3C21426">A review of the genus Colias Fabricius, 1807 (Lepidoptera: Pieridae) from Qinghai with descriptions of a new subspecies of Colias thrasibulus Fruhstorfer, 1910</mods:title>
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<taxonomicName id="4C114D7DFFBCA60AFF14AAA860DCFD68" authority="wanda Grum-Grshimailo, 1907" authorityName="wanda Grum-Grshimailo" authorityYear="1907" box="[151,711,654,681]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="wanda" subSpecies="wanda">
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<emphasis id="B965EAECFFBCA60AFF14AAA86389FD69" bold="true" box="[151,402,654,681]" italics="true" pageId="4" pageNumber="455">Colias wanda wanda</emphasis>
<bibRefCitation id="EF804B0FFFBCA60AFE19AAA860DCFD68" author="Grum-Grshimailo, G. E." box="[410,711,654,680]" pageId="4" pageNumber="455" refId="ref15257" refString="Grum-Grshimailo, G. E. (1907) [A description of a journey to West-China. Round Kuku-noor, via Nan-Shan, Bej-Shan and along the Eastern Tien-Schan back to the Motherland]. Vol. 3. Izdanie imperatorskogo russkogo geographicheskogo obshestva, St. Petersburg, 537 pp., 1 map, 54 figs. [in Russian]" type="book" year="1907">Grum-Grshimailo, 1907</bibRefCitation>
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<paragraph id="8BAE36FEFFBCA60AFF14AA9560C9FD0E" blockId="4.[151,722,654,718]" box="[151,722,690,718]" pageId="4" pageNumber="455">
(
<figureCitation id="132A2A7BFFBCA60AFF1CAA9562F5FD0D" box="[159,238,691,717]" captionStart="FIGURES 13" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[217,1370,181,1920]" captionTargetId="figure-19@13.[217,1370,181,1920]" captionTargetPageId="13" captionText="FIGURES 13. Habitus of Colias taxa ♂♂ &amp; ♀♀ at same scale (Upper half- upper side; lower half- underside). 1—Colias adelaidae adelaidae; 2—Colias arida cakana; 3—Colias wanda wanda. Serial number unique for each specimen. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.6333775" httpUri="https://zenodo.org/record/6333775/files/figure.png" pageId="4" pageNumber="455">Figs. 3</figureCitation>
,
<figureCitation id="132A2A7BFFBCA60AFF78AA946310FD0D" box="[251,267,690,717]" captionStart="FIGURES 45" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[210,1376,181,1919]" captionTargetId="figure-21@14.[210,1376,181,1919]" captionTargetPageId="14" captionText="FIGURES 45. Habitus of Colias taxa ♂♂ &amp; ♀♀ at same scale (Upper half- upper side; lower half- underside). 4—Colias arida cakana; 5—Colias wanda wanda. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.6333777" httpUri="https://zenodo.org/record/6333777/files/figure.png" pageId="4" pageNumber="455">5</figureCitation>
,
<figureCitation id="132A2A7BFFBCA60AFE94AA95633CFD0D" box="[279,295,691,717]" captionStart="FIGURES 67" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[204,1383,181,1920]" captionTargetId="figure-19@15.[204,1383,181,1920]" captionTargetPageId="15" captionText="FIGURES 67. Habitus of Colias taxa ♂♂ at same scale (Upper half- upper side; lower half- underside). 6—Colias arida cakana; 7—Colias wanda wanda. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.6333779" httpUri="https://zenodo.org/record/6333779/files/figure.png" pageId="4" pageNumber="455">7</figureCitation>
,
<figureCitation id="132A2A7BFFBCA60AFEB0AA95634AFD0D" box="[307,337,691,717]" captionStart="FIGURE 41" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[216,1372,182,1918]" captionTargetId="figure-19@23.[216,1372,181,1920]" captionTargetPageId="23" captionText="FIGURE 41. ♂ genitalia and 8th tergum (with superuncus) in lateral view at same scale. Serial numbers corresponding to those of specimens. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6333795" httpUri="https://zenodo.org/record/6333795/files/figure.png" pageId="4" pageNumber="455">41</figureCitation>
,
<figureCitation id="132A2A7BFFBCA60AFEDEAA9563BEFD0D" box="[349,421,691,717]" captionStart-0="FIGURE 43" captionStart-1="FIGURE 44" captionStartId-0="25.[151,250,1943,1967]" captionStartId-1="26.[151,250,1943,1967]" captionTargetBox-0="[218,1370,181,1920]" captionTargetBox-1="[225,1362,188,1913]" captionTargetId-0="figure-19@25.[218,1370,181,1920]" captionTargetId-1="figure-21@26.[225,1362,181,1920]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURE 43. 8th tergum in dorsal view at same scale to show superuncus. Scale bar = 1 mm." captionText-1="FIGURE 44. ♂ genitalia in dorsal view at same scale. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6333799" figureDoi-1="http://doi.org/10.5281/zenodo.6333803" httpUri-0="https://zenodo.org/record/6333799/files/figure.png" httpUri-1="https://zenodo.org/record/6333803/files/figure.png" pageId="4" pageNumber="455">4344</figureCitation>
,
<figureCitation id="132A2A7BFFBCA60AFE32AA9563E2FD0D" box="[433,505,691,717]" captionStart-0="FIGURE 46" captionStart-1="FIGURE 47" captionStartId-0="28.[151,250,1943,1967]" captionStartId-1="29.[151,250,1943,1967]" captionTargetBox-0="[216,1372,181,1909]" captionTargetBox-1="[220,1368,193,1919]" captionTargetId-0="figure-21@28.[216,1372,181,1920]" captionTargetId-1="figure-19@29.[220,1368,181,1920]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionText-0="FIGURE 46. Ring in lateral view and scaphium in ventral view at same scale. Scale bar = 1 mm." captionText-1="FIGURE 47. Phallus in lateral view and coecum-process in full-face view at same scale. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6333809" figureDoi-1="http://doi.org/10.5281/zenodo.6333811" httpUri-0="https://zenodo.org/record/6333809/files/figure.png" httpUri-1="https://zenodo.org/record/6333811/files/figure.png" pageId="4" pageNumber="455">4647</figureCitation>
,
<figureCitation id="132A2A7BFFBCA60AFD86AA946039FD0D" box="[517,546,690,717]" captionStart="FIGURE 49" captionStartId="31.[151,250,1943,1967]" captionTargetBox="[234,1353,181,1920]" captionTargetId="figure-19@31.[234,1353,181,1920]" captionTargetPageId="31" captionText="FIGURE 49. Juxta in full-face view (nearly posterior view) at same scale. Scale bar = 1 mm. FIGURE 50. Valva in inner lateral and anterior views at same scale. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6333815" httpUri="https://zenodo.org/record/6333815/files/figure.png" pageId="4" pageNumber="455">49</figureCitation>
50, 5253, 68)
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<treatmentCitationGroup id="AB0111D0FFBCA60AFF14AAD166FFFCCF" box="[151,1252,759,783]" pageId="4" pageNumber="455">
<treatmentCitation id="0AB010EFFFBCA60AFF14AAD16088FCCF" author="Grum-Grshimailo, G. E." box="[151,659,759,783]" page="13" pageId="4" pageNumber="455" year="1907">
<taxonomicName id="4C114D7DFFBCA60AFF14AAD16088FCCF" authority="Grum-Grshimailo, 1907: 13" authorityName="Grum-Grshimailo" authorityPageNumber="13" authorityYear="1907" box="[151,659,759,783]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="eogene" subSpecies="wanda">
<emphasis id="B965EAECFFBCA60AFF14AAD16369FCCF" box="[151,370,759,783]" italics="true" pageId="4" pageNumber="455">Colias eogene wanda</emphasis>
<bibRefCitation id="EF804B0FFFBCA60AFEFBAAD16088FCCF" author="Grum-Grshimailo, G. E." box="[376,659,759,783]" pageId="4" pageNumber="455" refId="ref15257" refString="Grum-Grshimailo, G. E. (1907) [A description of a journey to West-China. Round Kuku-noor, via Nan-Shan, Bej-Shan and along the Eastern Tien-Schan back to the Motherland]. Vol. 3. Izdanie imperatorskogo russkogo geographicheskogo obshestva, St. Petersburg, 537 pp., 1 map, 54 figs. [in Russian]" type="book" year="1907">Grum-Grshimailo, 1907: 13</bibRefCitation>
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(TL: Hachen, Lajishan,
<collectingRegion id="49D5F81CFFBCA60AFC0FAAD161F9FCCF" box="[908,994,759,783]" country="China" name="Qinghai" pageId="4" pageNumber="455">Qinghai</collectingRegion>
:
<bibRefCitation id="EF804B0FFFBCA60AFC68AAD166C3FCCF" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[1003,1240,759,783]" pageId="4" pageNumber="455" refId="ref15180" refString="Grieshuber, J., Worthy, R. &amp; Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias. Tshikolovets Publications, Pardubice, 438 pp." type="book" year="2012">
Grieshuber
<emphasis id="B965EAECFFBCA60AFBE1AADE6680FCCF" box="[1122,1179,759,783]" italics="true" pageId="4" pageNumber="455">et al.,</emphasis>
2012
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).
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</paragraph>
<paragraph id="8BAE36FEFFBCA60AFF14AB30662FFC68" blockId="4.[151,1437,759,1428]" pageId="4" pageNumber="455">
<treatmentCitationGroup id="AB0111D0FFBCA60AFF14AB30662FFC68" pageId="4" pageNumber="455">
<treatmentCitation id="0AB010EFFFBCA60AFF14AB30601EFCEE" author="Kotzsch, H." box="[151,517,790,814]" page="236" pageId="4" pageNumber="455" year="1930">
<taxonomicName id="4C114D7DFFBCA60AFF14AB30601EFCEE" ID-CoL="5ZLPW" authority="Kotzsch, 1930: 236" authorityName="Kotzsch" authorityPageNumber="236" authorityYear="1930" box="[151,517,790,814]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="baeckeri">
<emphasis id="B965EAECFFBCA60AFF14AB30632CFCEE" box="[151,311,790,814]" italics="true" pageId="4" pageNumber="455">Colias baeckeri</emphasis>
<bibRefCitation id="EF804B0FFFBCA60AFEBDAB30601EFCEE" author="Kotzsch, H." box="[318,517,790,814]" pageId="4" pageNumber="455" pagination="236 - 237" refId="ref15698" refString="Kotzsch, H. (1930) Ein neuer Colias aus dem Richthofengebirge. Entomologische Zeitschrift, Frankfurt, 43 (19), 236 - 237." type="journal article" year="1930">Kotzsch, 1930: 236</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(TL: Dingtsiang-miau = Dingqiangmiao,
<collectingRegion id="49D5F81CFFBCA60AFC32AB3061EDFCEE" box="[945,1014,790,814]" country="China" name="Gansu" pageId="4" pageNumber="455">Gansu</collectingRegion>
);
<treatmentCitation id="0AB010EFFFBCA60AFB8BAB3066C4FCEE" author="Verhulst, J." box="[1032,1247,790,814]" page="587" pageId="4" pageNumber="455" year="1994">
<bibRefCitation id="EF804B0FFFBCA60AFB8BAB3066C4FCEE" author="Verhulst, J." box="[1032,1247,790,814]" pageId="4" pageNumber="455" pagination="587 - 590" refId="ref16347" refString="Verhulst, J. (1994 b) Fiches specifiques des Colias F. (Colias data sheets). COLIAS BAECKERI. Lambillionea, 94 (4), 587 - 590. [in French]" type="journal article" year="1994">Verhulst, 1994b: 587</bibRefCitation>
</treatmentCitation>
, 589- figs. for
<specimenCount id="9D17FD77FFBCA60AFAF5AB306787FCED" box="[1398,1436,790,813]" pageId="4" pageNumber="455" type="male">♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFBCA60AFF61AB136312FC8C" box="[226,265,821,844]" pageId="4" pageNumber="455" type="female">♀♀</specimenCount>
;
<treatmentCitation id="0AB010EFFFBCA60AFE94AB1263EAFC8D" author="Grieshuber, J." box="[279,497,820,845]" page="47" pageId="4" pageNumber="455" year="1999">
<bibRefCitation id="EF804B0FFFBCA60AFE94AB1263EAFC8D" author="Grieshuber, J." box="[279,497,820,845]" pageId="4" pageNumber="455" pagination="41 - 48" refId="ref14631" refString="Grieshuber, J. (1999) Zur Artengruppe Colias arida Alpheraky, 1889, adelaidae Verhulst, 1991, baeckeri Kotzsch, 1930 und stoliczkana Moore, 1878. Beschreibung einer neuen Unterart von Colias adelaidae (Lepidoptera: Pieridae). Lambillionea, 99 (1), 41 - 48. [in Germany]" type="journal article" year="1999">Grieshuber, 1999: 47</bibRefCitation>
</treatmentCitation>
, synonymic list;
<bibRefCitation id="EF804B0FFFBCA60AFD1CAB12612CFC8D" author="Verhulst, J." box="[671,823,820,845]" pageId="4" pageNumber="455" refId="ref16453" refString="Verhulst, J. (2001) Les Colias du Globe. Monograph of the genus Colias. Plates. Goecke &amp; Evers, Kelterns, 45 pp., 183 pls. [in French]" type="book" year="2001">Verhulst, 2001</bibRefCitation>
: pl. 55, figs. 124 for
<specimenCount id="9D17FD77FFBCA60AFB9EAB136658FC8C" box="[1053,1091,821,844]" pageId="4" pageNumber="455" type="male">♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFBCA60AFBE5AB136696FC8C" box="[1126,1165,821,844]" pageId="4" pageNumber="455" type="female">♀♀</specimenCount>
;
<treatmentCitation id="0AB010EFFFBCA60AFB19AB126720FC8D" author="Dietz, M." box="[1178,1339,820,845]" page="93" pageId="4" pageNumber="455" year="2002">
<bibRefCitation id="EF804B0FFFBCA60AFB19AB126720FC8D" author="Dietz, M." box="[1178,1339,820,845]" pageId="4" pageNumber="455" refId="ref14565" refString="Dietz, M. (2002) Pieridae. Colias Fabricius 1807. Species and subspecies of the Palaearctic Region. Part 1. Selbstverlag, Solms, 154 pp. [in Germany]" type="book" year="2002">Dietz, 2002: 93</bibRefCitation>
</treatmentCitation>
, 95- figs. 3, 612 for
<specimenCount id="9D17FD77FFBCA60AFEC3AB72637DFCAB" box="[320,358,852,875]" pageId="4" pageNumber="455" type="male">♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFBCA60AFE04AB7263B5FCAB" box="[391,430,852,875]" pageId="4" pageNumber="455" type="female">♀♀</specimenCount>
;
<treatmentCitation id="0AB010EFFFBCA60AFE38AB75611CFCAB" author="Grieshuber, J. &amp; Lamas, G." box="[443,775,851,875]" page="164" pageId="4" pageNumber="455" year="2007">
<bibRefCitation id="EF804B0FFFBCA60AFE38AB75611CFCAB" author="Grieshuber, J. &amp; Lamas, G." box="[443,775,851,875]" pageId="4" pageNumber="455" pagination="131 - 171" refId="ref15140" refString="Grieshuber, J. &amp; Lamas, G. (2007) A Synonymic List of the Genus Colias Fabricius, 1807 (Lepidoptera: Pieridae). Mitteilungen der Munchner Entomologischen Gesellschaft, 97, 131 - 171." type="journal article" year="2007">Grieshuber &amp; Lamas, 2007: 164</bibRefCitation>
</treatmentCitation>
, synonymic list;
<treatmentCitation id="0AB010EFFFBCA60AFC35AB7566D1FCAB" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[950,1226,851,875]" page="57" pageId="4" pageNumber="455" year="2012">
<bibRefCitation id="EF804B0FFFBCA60AFC35AB7566D1FCAB" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[950,1226,851,875]" pageId="4" pageNumber="455" refId="ref15180" refString="Grieshuber, J., Worthy, R. &amp; Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias. Tshikolovets Publications, Pardubice, 438 pp." type="book" year="2012">
Grieshuber
<emphasis id="B965EAECFFBCA60AFBADAB72667CFCAB" box="[1070,1127,851,875]" italics="true" pageId="4" pageNumber="455">et al.,</emphasis>
2012: 57
</bibRefCitation>
</treatmentCitation>
, discussions on
<typeStatus id="54AA885CFFBCA60AFAF2AB726786FCAC" box="[1393,1437,852,876]" pageId="4" pageNumber="455">type</typeStatus>
series,
<typeStatus id="54AA885CFFBCA60AFE8DAB546321FC4A" box="[270,314,882,906]" pageId="4" pageNumber="455">type</typeStatus>
locality and historical overview on taxonomic status (these various historical taxonomic changes in literature are not repeatedly cited herein);
<bibRefCitation id="EF804B0FFFBCA60AFD93ABB660D0FC68" author="Grieshuber, J." box="[528,715,912,936]" pageId="4" pageNumber="455" refId="ref14869" refString="Grieshuber, J. (2016 b) Specimen of Colias in Museum fur Naturkunde Berlin. Available from: https: // commons. wikimedia. org / wiki / Category: Specimen _ of _ Colias _ in _ Museum _ f % C 3 % BCr _ Naturkunde _ Berlin (accessed 23 October 2021)" type="url" year="2016">Grieshuber, 2016b</bibRefCitation>
&amp;
<bibRefCitation id="EF804B0FFFBCA60AFD6FABB76138FC69" author="Grieshuber, J." box="[748,803,913,937]" pageId="4" pageNumber="455" refId="ref14928" refString="Grieshuber, J. (2020) Lepidoptera in the Zoologische Staatssammlung Munchen. Available from: https: // commons. wikimedia. org / wiki / Category: Lepidoptera _ in _ the _ Zoologische _ Staatssammlung _ M % C 3 % BCnchen (accessed 23 October 2021)" type="url" year="2020">2020</bibRefCitation>
: photos of
<typeStatus id="54AA885CFFBCA60AFC12ABB761A6FC69" box="[913,957,913,937]" pageId="4" pageNumber="455">type</typeStatus>
specimens.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAE36FEFFBCA60AFF14AB8960AEFC07" blockId="4.[151,1437,759,1428]" box="[151,693,943,967]" pageId="4" pageNumber="455">
<treatmentCitationGroup id="AB0111D0FFBCA60AFF14AB8960AEFC07" box="[151,693,943,967]" pageId="4" pageNumber="455">
<treatmentCitation id="0AB010EFFFBCA60AFF14AB8960AAFC07" author="Rose, K. &amp; Schulte, A." box="[151,689,943,967]" page="96" pageId="4" pageNumber="455" year="1992">
<taxonomicName id="4C114D7DFFBCA60AFF14AB8960AAFC07" authority=": Rose &amp; Schulte, 1992: 96" authorityName="Rose &amp; Schulte" authorityPageNumber="96" authorityYear="1992" box="[151,689,943,967]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="stoliczkana" subSpecies="wanda">
<emphasis id="B965EAECFFBCA60AFF14AB896387FC07" box="[151,412,943,967]" italics="true" pageId="4" pageNumber="455">Colias stoliczkana wanda</emphasis>
:
<bibRefCitation id="EF804B0FFFBCA60AFE2AAB9660AAFC07" author="Rose, K. &amp; Schulte, A." box="[425,689,943,967]" pageId="4" pageNumber="455" pagination="93 - 104" refId="ref16020" refString="Rose, K. &amp; Schulte, A. (1992) Ein Beitrag zur systematischen Stellung und subspezifischen Gliederung von Colias arida Alpheraky 1889. Nachrichten des entomologischen Vereins Apollo, Neue Folge, 13 (2), 93 - 104, pls. 1 - 2. [in Germany]" type="journal article" year="1992">Rose &amp; Schulte, 1992: 96</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAE36FEFFBCA60AFF14ABE86096FC26" blockId="4.[151,1437,759,1428]" box="[151,653,974,998]" pageId="4" pageNumber="455">
<treatmentCitationGroup id="AB0111D0FFBCA60AFF14ABE86096FC26" box="[151,653,974,998]" pageId="4" pageNumber="455">
<treatmentCitation id="0AB010EFFFBCA60AFF14ABE86092FC26" author="Rose, K. &amp; Schulte, A." box="[151,649,974,998]" page="98" pageId="4" pageNumber="455" year="1992">
<taxonomicName id="4C114D7DFFBCA60AFF14ABE86092FC26" ID-CoL="5ZLPW" authority=": Rose &amp; Schulte, 1992: 98" authorityName="Rose &amp; Schulte" authorityPageNumber="98" authorityYear="1992" box="[151,649,974,998]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="arida" subSpecies="baeckeri">
<emphasis id="B965EAECFFBCA60AFF14ABE8636FFC26" box="[151,372,974,998]" italics="true" pageId="4" pageNumber="455">Colias arida baeckeri</emphasis>
:
<bibRefCitation id="EF804B0FFFBCA60AFE02ABE86092FC26" author="Rose, K. &amp; Schulte, A." box="[385,649,974,998]" pageId="4" pageNumber="455" pagination="93 - 104" refId="ref16020" refString="Rose, K. &amp; Schulte, A. (1992) Ein Beitrag zur systematischen Stellung und subspezifischen Gliederung von Colias arida Alpheraky 1889. Nachrichten des entomologischen Vereins Apollo, Neue Folge, 13 (2), 93 - 104, pls. 1 - 2. [in Germany]" type="journal article" year="1992">Rose &amp; Schulte, 1992: 98</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAE36FEFFBCA60AFF14ABCA617CFBC4" blockId="4.[151,1437,759,1428]" box="[151,871,1004,1029]" pageId="4" pageNumber="455">
<treatmentCitationGroup id="AB0111D0FFBCA60AFF14ABCA617CFBC4" box="[151,871,1004,1029]" pageId="4" pageNumber="455">
<treatmentCitation id="0AB010EFFFBCA60AFF14ABCA6068FBC5" author="Hoshiai, A." box="[151,627,1004,1029]" page="10" pageId="4" pageNumber="455" year="1996">
<taxonomicName id="4C114D7DFFBCA60AFF14ABCA6068FBC5" ID-CoL="5ZLPW" authority=": Hoshiai, 1996: 10" authorityName="Hoshiai" authorityPageNumber="10" authorityYear="1996" box="[151,627,1004,1029]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="stoliczkana" subSpecies="baeckeri">
<emphasis id="B965EAECFFBCA60AFF14ABCA63B4FBC4" box="[151,431,1004,1028]" italics="true" pageId="4" pageNumber="455">Colias stoliczkana baeckeri</emphasis>
:
<bibRefCitation id="EF804B0FFFBCA60AFE3EABCA6068FBC5" author="Hoshiai, A." box="[445,627,1004,1029]" pageId="4" pageNumber="455" pagination="8 - 10" refId="ref15351" refString="Hoshiai, A. (1996) Notes of Colias from China and surrounding area (1). In: Koiwaya, S. (Ed.), Studies of Chinese Butterflies. Vol. 3. S. Koiwaya, Tokyo, pp. 211 - 217, pls. 8 - 10. [in Japanese]" type="book chapter" year="1996">Hoshiai, 1996: 10</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 6265 for
<specimenCount id="9D17FD77FFBCA60AFC99ABCB6136FBC4" box="[794,813,1005,1028]" pageId="4" pageNumber="455" type="male"></specimenCount>
&amp;
<specimenCount id="9D17FD77FFBCA60AFCCDABCB6179FBC4" box="[846,866,1005,1028]" pageId="4" pageNumber="455" type="female"></specimenCount>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAE36FEFFBCA60AFF14AC2D612DFBE3" blockId="4.[151,1437,759,1428]" box="[151,822,1035,1059]" pageId="4" pageNumber="455">
<treatmentCitationGroup id="AB0111D0FFBCA60AFF14AC2D612DFBE3" box="[151,822,1035,1059]" pageId="4" pageNumber="455">
<taxonomicName id="4C114D7DFFBCA60AFF14AC2D6349FBE3" ID-CoL="X2GP" authorityName="Moore" authorityYear="1878" box="[151,338,1035,1059]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="stoliczkana">
<emphasis id="B965EAECFFBCA60AFF14AC2D6349FBE3" box="[151,338,1035,1059]" italics="true" pageId="4" pageNumber="455">Colias stoliczkana</emphasis>
</taxonomicName>
ssp.:
<treatmentCitation id="0AB010EFFFBCA60AFE0FAC2D6058FBE3" author="Hoshiai, A." box="[396,579,1035,1059]" page="10" pageId="4" pageNumber="455" year="1996">
<bibRefCitation id="EF804B0FFFBCA60AFE0FAC2D6058FBE3" author="Hoshiai, A." box="[396,579,1035,1059]" pageId="4" pageNumber="455" pagination="8 - 10" refId="ref15351" refString="Hoshiai, A. (1996) Notes of Colias from China and surrounding area (1). In: Koiwaya, S. (Ed.), Studies of Chinese Butterflies. Vol. 3. S. Koiwaya, Tokyo, pp. 211 - 217, pls. 8 - 10. [in Japanese]" type="book chapter" year="1996">Hoshiai, 1996: 10</bibRefCitation>
</treatmentCitation>
, figs. 5457 for
<specimenCount id="9D17FD77FFBCA60AFD69AC2A60E6FBE3" box="[746,765,1036,1059]" pageId="4" pageNumber="455" type="male"></specimenCount>
&amp;
<specimenCount id="9D17FD77FFBCA60AFC9EAC2A612AFBE3" box="[797,817,1036,1059]" pageId="4" pageNumber="455" type="female"></specimenCount>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAE36FEFFBCA60AFF14AC0C63ECFBBF" blockId="4.[151,1437,759,1428]" pageId="4" pageNumber="455">
<treatmentCitationGroup id="AB0111D0FFBCA60AFF14AC0C63ECFBBF" pageId="4" pageNumber="455">
<treatmentCitation id="0AB010EFFFBCA60AFF14AC0C605CFB82" author="Kocman, S." box="[151,583,1066,1090]" page="56" pageId="4" pageNumber="455" year="1999">
<taxonomicName id="4C114D7DFFBCA60AFF14AC0C605CFB82" ID-CoL="5GJXX" authority="Kocman, 1999: 56" authorityName="Kocman" authorityPageNumber="56" authorityYear="1999" box="[151,583,1066,1090]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="baeckeri" subSpecies="paskoi">
<emphasis id="B965EAECFFBCA60AFF14AC0C639BFB82" box="[151,384,1066,1090]" italics="true" pageId="4" pageNumber="455">Colias baeckeri paskoi</emphasis>
<bibRefCitation id="EF804B0FFFBCA60AFE04AC0C605CFB82" author="Kocman, S." box="[391,583,1066,1090]" pageId="4" pageNumber="455" pagination="47 - 64" refId="ref15579" refString="Kocman, S. (1999) Descriptions of some new subspecies of Lepidoptera from China. Wallace, 5, 47 - 64, pls. 6 - 7." type="journal article" year="1999">Kocman, 1999: 56</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 8, figs. 34 (TL: Chiehku = Yushu, S.
<collectingRegion id="49D5F81CFFBCA60AFC78AC0C664AFB82" box="[1019,1105,1066,1090]" country="China" name="Qinghai" pageId="4" pageNumber="455">Qinghai</collectingRegion>
);
<treatmentCitation id="0AB010EFFFBCA60AFBE0AC0C6718FB82" author="Dietz, M." box="[1123,1283,1066,1090]" page="93" pageId="4" pageNumber="455" year="2002">
<bibRefCitation id="EF804B0FFFBCA60AFBE0AC0C6718FB82" author="Dietz, M." box="[1123,1283,1066,1090]" pageId="4" pageNumber="455" refId="ref14565" refString="Dietz, M. (2002) Pieridae. Colias Fabricius 1807. Species and subspecies of the Palaearctic Region. Part 1. Selbstverlag, Solms, 154 pp. [in Germany]" type="book" year="2002">Dietz, 2002: 93</bibRefCitation>
</treatmentCitation>
, synonymy for
<taxonomicName id="4C114D7DFFBCA60AFF44AC6F6320FBA0" ID-CoL="5ZLPW" authorityName="Kotzsch" authorityYear="1930" box="[199,315,1096,1120]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="baeckeri">
<emphasis id="B965EAECFFBCA60AFF44AC6F6320FBA0" box="[199,315,1096,1120]" italics="true" pageId="4" pageNumber="455">C. baeckeri</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AB010EFFFBCA60AFECBAC6E6094FBA0" author="Grieshuber, J. &amp; Lamas, G." box="[328,655,1096,1121]" page="164" pageId="4" pageNumber="455" year="2007">
<bibRefCitation id="EF804B0FFFBCA60AFECBAC6E6094FBA0" author="Grieshuber, J. &amp; Lamas, G." box="[328,655,1096,1121]" pageId="4" pageNumber="455" pagination="131 - 171" refId="ref15140" refString="Grieshuber, J. &amp; Lamas, G. (2007) A Synonymic List of the Genus Colias Fabricius, 1807 (Lepidoptera: Pieridae). Mitteilungen der Munchner Entomologischen Gesellschaft, 97, 131 - 171." type="journal article" year="2007">Grieshuber &amp; Lamas, 2007: 164</bibRefCitation>
</treatmentCitation>
, synonymic list;
<treatmentCitation id="0AB010EFFFBCA60AFCB9AC6E664CFBA1" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[826,1111,1096,1121]" page="177" pageId="4" pageNumber="455" year="2012">
<bibRefCitation id="EF804B0FFFBCA60AFCB9AC6E664CFBA1" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[826,1111,1096,1121]" pageId="4" pageNumber="455" refId="ref15180" refString="Grieshuber, J., Worthy, R. &amp; Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias. Tshikolovets Publications, Pardubice, 438 pp." type="book" year="2012">
Grieshuber
<emphasis id="B965EAECFFBCA60AFC32AC6F61F2FBA0" box="[945,1001,1096,1120]" italics="true" pageId="4" pageNumber="455">et al.,</emphasis>
2012: 177
</bibRefCitation>
</treatmentCitation>
, discussions on
<typeStatus id="54AA885CFFBCA60AFB7AAC6F673EFBA1" box="[1273,1317,1097,1121]" pageId="4" pageNumber="455">type</typeStatus>
series,
<typeStatus id="54AA885CFFBCA60AFAF2AC6F6786FBA1" box="[1393,1437,1097,1121]" pageId="4" pageNumber="455">type</typeStatus>
locality and taxonomic status.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAE36FEFFBCA60AFF14ACA0616DFB3A" blockId="4.[151,1437,759,1428]" pageId="4" pageNumber="455">
<treatmentCitationGroup id="AB0111D0FFBCA60AFF14ACA0616DFB3A" pageId="4" pageNumber="455">
<treatmentCitation id="0AB010EFFFBCA60AFF14ACA060D0FB5E" author="Grieshuber, J. &amp; Churkin, S." box="[151,715,1158,1182]" page="251" pageId="4" pageNumber="455" year="2003">
<taxonomicName id="4C114D7DFFBCA60AFF14ACA060D0FB5E" authority=": Grieshuber &amp; Churkin, 2003: 249 - 251" authorityName=": Grieshuber &amp; Churkin" authorityPageNumber="249 - 251" authorityYear="2003" box="[151,715,1158,1182]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="wanda">
<emphasis id="B965EAECFFBCA60AFF14ACA0633DFB5E" box="[151,294,1158,1182]" italics="true" pageId="4" pageNumber="455">Colias wanda</emphasis>
:
<bibRefCitation id="EF804B0FFFBCA60AFEB6ACA060D0FB5E" author="Grieshuber, J. &amp; Churkin, S." box="[309,715,1158,1182]" pageId="4" pageNumber="455" pagination="244 - 271" refId="ref14980" refString="Grieshuber, J. &amp; Churkin, S. (2003) The lectotypes of Colias diva Grum-Grshimailo, 1891, Colias wanda Grum-Grshimailo, 1907, Colias grumi Alpheraky, 1897, Colias cocandica maja Grum-Grshimailo, 1891, Colias cocandica tatarica Bang- Haas, 1915, and Colias tamerlana mongola Alpheraky, 1897 (Lepidoptera: Pieridae). Helios, Moscow, 4, 244 - 271, pls. 15 - 16." type="journal article" year="2003">Grieshuber &amp; Churkin, 2003: 249251</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<typeStatus id="54AA885CFFBCA60AFD54ACA0612FFB5E" box="[727,820,1158,1182]" pageId="4" pageNumber="455" type="lectotype">lectotype</typeStatus>
designation;
<treatmentCitation id="0AB010EFFFBCA60AFC42ACA06707FB5E" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[961,1308,1158,1182]" page="235" pageId="4" pageNumber="455" year="2012">
<bibRefCitation id="EF804B0FFFBCA60AFC42ACA06707FB5E" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[961,1308,1158,1182]" pageId="4" pageNumber="455" refId="ref15180" refString="Grieshuber, J., Worthy, R. &amp; Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias. Tshikolovets Publications, Pardubice, 438 pp." type="book" year="2012">
Grieshuber
<emphasis id="B965EAECFFBCA60AFBB8ACA1666EFB5E" box="[1083,1141,1158,1182]" italics="true" pageId="4" pageNumber="455">et al.,</emphasis>
2012: 235237
</bibRefCitation>
</treatmentCitation>
, discussions on type series, type locality and taxonomic status, records from Tsona, S.
<collectingRegion id="49D5F81CFFBCA60AFC3FAC8261E8FB7C" box="[956,1011,1188,1212]" country="China" name="Xizang" pageId="4" pageNumber="455">Tibet</collectingRegion>
, 239, record from W.
<collectingRegion id="49D5F81CFFBCA60AFB56AC82673CFB7C" box="[1237,1319,1188,1212]" country="China" name="Sichuan" pageId="4" pageNumber="455">Sichuan</collectingRegion>
(no further information), pl. 23, figs. 112 for
<specimenCount id="9D17FD77FFBCA60AFDA5ACE26057FB1B" box="[550,588,1220,1243]" pageId="4" pageNumber="455" type="male">♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFBCA60AFDE9ACE2608BFB1B" box="[618,656,1220,1243]" pageId="4" pageNumber="455" type="female">♀♀</specimenCount>
(type specimens);
<treatmentCitation id="0AB010EFFFBCA60AFCCDACE5663EFB1B" author="Grieshuber, J" box="[846,1061,1219,1243]" page="60" pageId="4" pageNumber="455" year="2014">
<bibRefCitation id="EF804B0FFFBCA60AFCCDACE5663EFB1B" author="Grieshuber, J" box="[846,1061,1219,1243]" pageId="4" pageNumber="455" refId="ref14730" refString="Grieshuber, J (2014) Guide to the butterflies of the Palearctic region. Pieridae Part II. Omnes Artes, Milano, 86 pp." type="book" year="2014">Grieshuber, 2014: 60</bibRefCitation>
</treatmentCitation>
, figs. for
<specimenCount id="9D17FD77FFBCA60AFB07ACE266B1FB1B" box="[1156,1194,1220,1243]" pageId="4" pageNumber="455" type="male">♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFBCA60AFB4AACE266EBFB1B" box="[1225,1264,1220,1243]" pageId="4" pageNumber="455" type="female">♀♀</specimenCount>
, record from Qia la, Chamdo, E.
<collectingRegion id="49D5F81CFFBCA60AFEE7ACC46387FB3A" box="[356,412,1250,1274]" country="China" name="Xizang" pageId="4" pageNumber="455">Tibet</collectingRegion>
;
<bibRefCitation id="EF804B0FFFBCA60AFE25ACC4607FFB3A" author="Grieshuber, J." box="[422,612,1250,1274]" pageId="4" pageNumber="455" refId="ref14798" refString="Grieshuber, J. (2016 a) Specimen of Colias in Museum of Zoology, the Zoological Institute of the Russian Academy of Sciences (Saint Petersburg). Available from: https: // commons. wikimedia. org / wiki / Category: Specimen _ of _ Colias _ in _ Museum _ of _ Zoology _ (Saint _ Petersburg) (accessed 23 October 2021)" type="url" year="2016">Grieshuber, 2016a</bibRefCitation>
: photos of type specimens.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAE36FEFFBCA60AFF14AD2661F7FAD8" blockId="4.[151,1437,759,1428]" box="[151,1004,1280,1305]" pageId="4" pageNumber="455">
<treatmentCitationGroup id="AB0111D0FFBCA60AFF14AD2661F7FAD8" box="[151,1004,1280,1305]" pageId="4" pageNumber="455">
<treatmentCitation id="0AB010EFFFBCA60AFF14AD2660FCFAD8" author="Grieshuber, J. &amp; Churkin, S." box="[151,743,1280,1305]" page="253" pageId="4" pageNumber="455" year="2003">
<taxonomicName id="4C114D7DFFBCA60AFF14AD2660FCFAD8" ID-CoL="5ZLPW" authority=": Grieshuber &amp; Churkin, 2003: 253" authorityName="Grieshuber &amp; Churkin" authorityPageNumber="253" authorityYear="2003" box="[151,743,1280,1305]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="wanda" subSpecies="baeckeri">
<emphasis id="B965EAECFFBCA60AFF14AD26639AFAD8" box="[151,385,1280,1304]" italics="true" pageId="4" pageNumber="455">Colias wanda baeckeri</emphasis>
:
<bibRefCitation id="EF804B0FFFBCA60AFE0DAD2660FCFAD8" author="Grieshuber, J. &amp; Churkin, S." box="[398,743,1280,1305]" pageId="4" pageNumber="455" pagination="244 - 271" refId="ref14980" refString="Grieshuber, J. &amp; Churkin, S. (2003) The lectotypes of Colias diva Grum-Grshimailo, 1891, Colias wanda Grum-Grshimailo, 1907, Colias grumi Alpheraky, 1897, Colias cocandica maja Grum-Grshimailo, 1891, Colias cocandica tatarica Bang- Haas, 1915, and Colias tamerlana mongola Alpheraky, 1897 (Lepidoptera: Pieridae). Helios, Moscow, 4, 244 - 271, pls. 15 - 16." type="journal article" year="2003">Grieshuber &amp; Churkin, 2003: 253</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, synonymy for
<taxonomicName id="4C114D7DFFBCA60AFC06AD2761FCFAD8" authorityName=": Grieshuber &amp; Churkin" authorityYear="2003" box="[901,999,1280,1304]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="wanda">
<emphasis id="B965EAECFFBCA60AFC06AD2761FCFAD8" box="[901,999,1280,1304]" italics="true" pageId="4" pageNumber="455">C. wanda</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAE36FEFFBCA60AFF14AD3961CCFAF7" blockId="4.[151,1437,759,1428]" box="[151,983,1311,1336]" pageId="4" pageNumber="455">
<treatmentCitationGroup id="AB0111D0FFBCA60AFF14AD3961CCFAF7" box="[151,983,1311,1336]" pageId="4" pageNumber="455">
<treatmentCitation id="0AB010EFFFBCA60AFF14AD3960C9FAF7" author="Grieshuber, J. &amp; Churkin, S." box="[151,722,1311,1335]" page="253" pageId="4" pageNumber="455" year="2003">
<taxonomicName id="4C114D7DFFBCA60AFF14AD3960C9FAF7" ID-CoL="5GJXX" authority=": Grieshuber &amp; Churkin, 2003: 253" authorityName="Grieshuber &amp; Churkin" authorityPageNumber="253" authorityYear="2003" box="[151,722,1311,1335]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="wanda" subSpecies="paskoi">
<emphasis id="B965EAECFFBCA60AFF14AD396377FAF7" box="[151,364,1311,1335]" italics="true" pageId="4" pageNumber="455">Colias wanda paskoi</emphasis>
:
<bibRefCitation id="EF804B0FFFBCA60AFEFAAD3960C9FAF7" author="Grieshuber, J. &amp; Churkin, S." box="[377,722,1311,1335]" pageId="4" pageNumber="455" pagination="244 - 271" refId="ref14980" refString="Grieshuber, J. &amp; Churkin, S. (2003) The lectotypes of Colias diva Grum-Grshimailo, 1891, Colias wanda Grum-Grshimailo, 1907, Colias grumi Alpheraky, 1897, Colias cocandica maja Grum-Grshimailo, 1891, Colias cocandica tatarica Bang- Haas, 1915, and Colias tamerlana mongola Alpheraky, 1897 (Lepidoptera: Pieridae). Helios, Moscow, 4, 244 - 271, pls. 15 - 16." type="journal article" year="2003">Grieshuber &amp; Churkin, 2003: 253</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, synonymy for
<taxonomicName id="4C114D7DFFBCA60AFCF3AD0661C9FAF7" authorityName=": Grieshuber &amp; Churkin" authorityYear="2003" box="[880,978,1311,1335]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="wanda">
<emphasis id="B965EAECFFBCA60AFCF3AD0661C9FAF7" box="[880,978,1311,1335]" italics="true" pageId="4" pageNumber="455">C. wanda</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAE36FEFFBCA60AFF14AD186017FA53" blockId="4.[151,1437,759,1428]" pageId="4" pageNumber="455">
<treatmentCitationGroup id="AB0111D0FFBCA60AFF14AD186017FA53" pageId="4" pageNumber="455">
<treatmentCitation id="0AB010EFFFBCA60AFF14AD18600FFA96" author="Miller, M. A." box="[151,532,1342,1366]" page="482" pageId="4" pageNumber="455" year="2009">
<taxonomicName id="4C114D7DFFBCA60AFF14AD18600FFA96" authority="Miller, 2009: 482" authorityName="Miller" authorityPageNumber="482" authorityYear="2009" box="[151,532,1342,1366]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="wernerbacki">
<emphasis id="B965EAECFFBCA60AFF14AD186345FA96" box="[151,350,1342,1366]" italics="true" pageId="4" pageNumber="455">Colias wernerbacki</emphasis>
<bibRefCitation id="EF804B0FFFBCA60AFEE0AD18600FFA96" author="Miller, M. A." box="[355,532,1342,1366]" pageId="4" pageNumber="455" pagination="478 - 486" refId="ref15817" refString="Miller, M. A. (2009) Colilas jomooseri spec. nov. und Colilas wernerbacki spec. nov., zwei neue Gelblingsarten vom Himalaya- Gebirge, Bhutan. Atalanta, 40 (3 / 4), 478 - 486." type="journal article" year="2009">Miller, 2009: 482</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 5, figs. 2-2b (TL:
<collectingCountry id="F306766EFFBCA60AFD73AD186122FA96" box="[752,825,1342,1366]" name="Bhutan" pageId="4" pageNumber="455">Bhutan</collectingCountry>
(West), Himalaya-Gebirge, Paro-Gebiet, nordl. Chomalhari-Hange, Umg. Jangothank);
<treatmentCitation id="0AB010EFFFBCA60AFE66AD7A6117FAB5" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[485,780,1372,1397]" page="239" pageId="4" pageNumber="455" year="2012">
<bibRefCitation id="EF804B0FFFBCA60AFE66AD7A6117FAB5" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[485,780,1372,1397]" pageId="4" pageNumber="455" refId="ref15180" refString="Grieshuber, J., Worthy, R. &amp; Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias. Tshikolovets Publications, Pardubice, 438 pp." type="book" year="2012">
Grieshuber
<emphasis id="B965EAECFFBCA60AFDDDAD7B6088FAB4" box="[606,659,1372,1396]" italics="true" pageId="4" pageNumber="455">et al.</emphasis>
, 2012: 239
</bibRefCitation>
</treatmentCitation>
, discussions on
<typeStatus id="54AA885CFFBCA60AFC35AD7B61F9FAB5" box="[950,994,1373,1397]" pageId="4" pageNumber="455">type</typeStatus>
series,
<typeStatus id="54AA885CFFBCA60AFBB0AD7B6644FAB5" box="[1075,1119,1373,1397]" pageId="4" pageNumber="455">type</typeStatus>
locality and taxonomic status, synonymy for
<taxonomicName id="4C114D7DFFBCA60AFED8AD5A601DFA53" authority="wanda" authorityName="wanda Grum-Grshimailo" authorityYear="1907" box="[347,518,1403,1427]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="wanda" subSpecies="wanda">
<emphasis id="B965EAECFFBCA60AFED8AD5A601DFA53" box="[347,518,1403,1427]" italics="true" pageId="4" pageNumber="455">C. wanda wanda</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30B6575FFBCA60AFF14ADE766FAF94F" pageId="4" pageNumber="455" type="materials_examined">
<paragraph id="8BAE36FEFFBCA60AFF14ADE766FAF94F" blockId="4.[151,1437,1473,2040]" pageId="4" pageNumber="455">
<emphasis id="B965EAECFFBCA60AFF14ADE76366FA1B" bold="true" box="[151,381,1473,1499]" pageId="4" pageNumber="455">Material examined</emphasis>
.
<materialsCitation id="3B793CA3FFBCA60AFE0DADE763A1FA3F" ID-GBIF-Occurrence="3698883347" collectingDate="2008-07-27" collectingDateMax="2008-07-29" collectingDateMin="2008-07-27" collectionCode="CSK, CHH" collectorName="K. Song" country="China" county="Qilian County" location="Babao" municipality="Niuxinshan" pageId="4" pageNumber="455" specimenCount="15" specimenCount-female="2" specimenCount-male="13" stateProvince="Qinghai">
<collectingCountry id="F306766EFFBCA60AFE0DADE763F0FA1B" box="[398,491,1473,1499]" name="China" pageId="4" pageNumber="455">CHINA</collectingCountry>
:
<collectingRegion id="49D5F81CFFBCA60AFE7AADE76043FA1B" box="[505,600,1473,1499]" country="China" name="Qinghai" pageId="4" pageNumber="455">Qinghai</collectingRegion>
:
<specimenCount id="9D17FD77FFBCA60AFDE5ADE760ADFA1C" box="[614,694,1473,1500]" pageId="4" pageNumber="455" type="male">13 ♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFBCA60AFD62ADE76138FA1C" box="[737,803,1473,1500]" pageId="4" pageNumber="455" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="ED00AE3BFFBCA60AFCB6ADE76174FA1B" box="[821,879,1473,1499]" pageId="4" pageNumber="455">CSK</collectionCode>
,
<collectionCode id="ED00AE3BFFBCA60AFCFCADE761A5FA1B" box="[895,958,1473,1499]" pageId="4" pageNumber="455">CHH</collectionCode>
),
<collectingMunicipality id="6BCAAC84FFBCA60AFC57ADE76643FA1B" box="[980,1112,1473,1499]" pageId="4" pageNumber="455">Niuxinshan</collectingMunicipality>
,
<location id="8ECE6025FFBCA60AFBE4ADE766AAFA1B" LSID="urn:lsid:plazi:treatment:03B887E8FFBCA60BFF14AAA860CDFE6D:8ECE6025FFBCA60AFBE4ADE766AAFA1B" box="[1127,1201,1473,1499]" country="China" county="Qilian County" municipality="Niuxinshan" name="Babao" pageId="4" pageNumber="455" stateProvince="Qinghai">Babao</location>
,
<collectingCounty id="62CF4E72FFBCA60AFB43ADE7677FFA1B" box="[1216,1380,1473,1499]" pageId="4" pageNumber="455">Qilian County</collectingCounty>
,
<date id="FFAF103EFFBCA60AFAF1ADE76338FA3F" pageId="4" pageNumber="455" value="2008-07-27" valueMax="2008-07-29" valueMin="2008-07-27">
<collectingDate id="EFEBE9D6FFBCA60AFAF1ADE76338FA3F" pageId="4" pageNumber="455" value="2008-07-27" valueMax="2008-07-29" valueMin="2008-07-27">27 29.VII.2008</collectingDate>
</date>
,
<collectorName id="26E45328FFBCA60AFEAEADC06393FA3F" box="[301,392,1509,1536]" pageId="4" pageNumber="455">K. Song</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793CA3FFBCA60AFE46ADC366C3FA3F" ID-GBIF-Occurrence="3698883312" box="[453,1240,1509,1536]" collectingDate="2008-07-24" collectingDateMax="2008-07-26" collectingDateMin="2008-07-24" collectionCode="CSK" collectorName="K. Song" country="China" county="Qilian County" location="Erlonggou" pageId="4" pageNumber="455" specimenCount="6" specimenCount-male="6" stateProvince="Qinghai">
<specimenCount id="9D17FD77FFBCA60AFE46ADC36018F9C0" box="[453,515,1509,1536]" pageId="4" pageNumber="455" type="male">6 ♂♂</specimenCount>
(
<collectionCode id="ED00AE3BFFBCA60AFD92ADC36057FA3F" box="[529,588,1509,1535]" pageId="4" pageNumber="455">CSK</collectionCode>
),
<location id="8ECE6025FFBCA60AFDDCADC360C3FA3F" LSID="urn:lsid:plazi:treatment:03B887E8FFBCA60BFF14AAA860CDFE6D:8ECE6025FFBCA60AFDDCADC360C3FA3F" box="[607,728,1509,1535]" country="China" county="Qilian County" name="Erlonggou" pageId="4" pageNumber="455" stateProvince="Qinghai">Erlonggou</location>
,
<collectingCounty id="62CF4E72FFBCA60AFD61ADC36199FA3F" box="[738,898,1509,1535]" pageId="4" pageNumber="455">Qilian County</collectingCounty>
,
<date id="FFAF103EFFBCA60AFC08ADC3665BFA3F" box="[907,1088,1509,1535]" pageId="4" pageNumber="455" value="2008-07-24" valueMax="2008-07-26" valueMin="2008-07-24">
<collectingDate id="EFEBE9D6FFBCA60AFC08ADC3665BFA3F" box="[907,1088,1509,1535]" pageId="4" pageNumber="455" value="2008-07-24" valueMax="2008-07-26" valueMin="2008-07-24">2426.VII.2008</collectingDate>
</date>
,
<collectorName id="26E45328FFBCA60AFBC8ADC066BDFA3F" box="[1099,1190,1509,1536]" pageId="4" pageNumber="455">K. Song</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793CA3FFBCA60AFB60ADC3662FF9E3" ID-GBIF-Occurrence="3698883314" collectingDate="24 - 26. VII. 2010 &amp; 26. VII. 2015" collectionCode="CSK, CHH" collectorName="K. Song" country="China" county="Dulan County" location="Dulan County" pageId="4" pageNumber="455" specimenCount="47" specimenCount-female="14" specimenCount-male="33" stateProvince="Qinghai">
<specimenCount id="9D17FD77FFBCA60AFB60ADC36734F9C0" box="[1251,1327,1509,1536]" pageId="4" pageNumber="455" type="male">33 ♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFBCA60AFAD3ADC36787F9C0" box="[1360,1436,1509,1536]" pageId="4" pageNumber="455" type="female">14 ♀♀</specimenCount>
(
<collectionCode id="ED00AE3BFFBCA60AFF1CAE2F62C2F9E3" box="[159,217,1545,1571]" pageId="4" pageNumber="455">CSK</collectionCode>
,
<collectionCode id="ED00AE3BFFBCA60AFF66AE2F633FF9E3" box="[229,292,1545,1571]" pageId="4" pageNumber="455">CHH</collectionCode>
), NE. of
<collectingCounty id="62CF4E72FFBCA60AFE08AE2F6031F9E3" box="[395,554,1545,1571]" pageId="4" pageNumber="455">Dulan County</collectingCounty>
,
<collectingDate id="EFEBE9D6FFBCA60AFDB6AE2F6183F9E4" box="[565,920,1545,1572]" isEnumeration="true" pageId="4" pageNumber="455">
<date id="FFAF103EFFBCA60AFDB6AE2F60F3F9E3" box="[565,744,1545,1571]" pageId="4" pageNumber="455" value="2010-07-24" valueMax="2010-07-26" valueMin="2010-07-24">2426.VII.2010</date>
&amp;
<date id="FFAF103EFFBCA60AFC8EAE2F6183F9E4" box="[781,920,1545,1572]" pageId="4" pageNumber="455" value="2015-07-26">26.VII.2015</date>
</collectingDate>
,
<collectorName id="26E45328FFBCA60AFC27AE2C661AF9E3" box="[932,1025,1545,1572]" pageId="4" pageNumber="455">K. Song</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793CA3FFBCA60AFBC3AE2F6078F987" ID-GBIF-Occurrence="3698883306" collectingDate="2021-07-15" collectionCode="CSK" collectorName="K. Song" country="China" county="Dulan County" location="Dulan County" pageId="4" pageNumber="455" specimenCount="15" specimenCount-female="4" specimenCount-male="11" stateProvince="Qinghai">
<specimenCount id="9D17FD77FFBCA60AFBC3AE2F6697F9E4" box="[1088,1164,1545,1572]" pageId="4" pageNumber="455" type="male">11 ♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFBCA60AFB32AE2F66EBF9E4" box="[1201,1264,1545,1572]" pageId="4" pageNumber="455" type="female">4 ♀♀</specimenCount>
(
<collectionCode id="ED00AE3BFFBCA60AFA83AE2F6720F9E3" box="[1280,1339,1545,1571]" pageId="4" pageNumber="455">CSK</collectionCode>
), SW. of
<collectingCounty id="62CF4E72FFBCA60AFF14AE0B632DF987" box="[151,310,1581,1607]" pageId="4" pageNumber="455">Dulan County</collectingCounty>
,
<date id="FFAF103EFFBCA60AFEC3AE0B63D0F988" box="[320,459,1581,1608]" pageId="4" pageNumber="455" value="2021-07-15">
<collectingDate id="EFEBE9D6FFBCA60AFEC3AE0B63D0F988" box="[320,459,1581,1608]" pageId="4" pageNumber="455" value="2021-07-15">15.VII.2021</collectingDate>
</date>
,
<collectorName id="26E45328FFBCA60AFE55AE08602AF987" box="[470,561,1581,1608]" pageId="4" pageNumber="455">K. Song</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793CA3FFBCA60AFDEDAE0B6783F987" ID-GBIF-Occurrence="3698883348" box="[622,1432,1581,1608]" collectingDate="2020-07-04" collectingDateMax="2020-07-10" collectingDateMin="2020-07-04" collectionCode="CSK" collectorName="Y. Yang" country="China" location="Tianjun" municipality="Guanjiaoshan" pageId="4" pageNumber="455" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Qinghai">
<specimenCount id="9D17FD77FFBCA60AFDEDAE0B608CF988" box="[622,663,1581,1608]" pageId="4" pageNumber="455" type="male">1 ♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFBCA60AFD3AAE0B60ECF988" box="[697,759,1581,1608]" pageId="4" pageNumber="455" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="ED00AE3BFFBCA60AFC85AE0B615AF987" box="[774,833,1581,1607]" pageId="4" pageNumber="455">CSK</collectionCode>
),
<collectingMunicipality id="6BCAAC84FFBCA60AFCD7AE0B61EAF987" box="[852,1009,1581,1607]" pageId="4" pageNumber="455">Guanjiaoshan</collectingMunicipality>
,
<location id="8ECE6025FFBCA60AFC79AE0B6648F987" LSID="urn:lsid:plazi:treatment:03B887E8FFBCA60BFF14AAA860CDFE6D:8ECE6025FFBCA60AFC79AE0B6648F987" box="[1018,1107,1581,1607]" country="China" municipality="Guanjiaoshan" name="Tianjun" pageId="4" pageNumber="455" stateProvince="Qinghai">Tianjun</location>
,
<date id="FFAF103EFFBCA60AFBDDAE0B671EF987" box="[1118,1285,1581,1607]" pageId="4" pageNumber="455" value="2020-07-04" valueMax="2020-07-10" valueMin="2020-07-04">
<collectingDate id="EFEBE9D6FFBCA60AFBDDAE0B671EF987" box="[1118,1285,1581,1607]" pageId="4" pageNumber="455" value="2020-07-04" valueMax="2020-07-10" valueMin="2020-07-04">410.VII.2020</collectingDate>
</date>
,
<collectorName id="26E45328FFBCA60AFA8CAE08677EF988" box="[1295,1381,1582,1608]" pageId="4" pageNumber="455">Y. Yang</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793CA3FFBCA60AFF14AE77663AF9AB" ID-GBIF-Occurrence="3698883334" box="[151,1057,1617,1644]" collectingDate="2020-07-30" collectionCode="CHH" collectorName="S. - Y. Lang" country="China" location="Tianjun" municipality="Guanjiaoshan" pageId="4" pageNumber="455" specimenCount="1" specimenCount-female="1" stateProvince="Qinghai">
<specimenCount id="9D17FD77FFBCA60AFF14AE7762D9F9AC" box="[151,194,1617,1644]" pageId="4" pageNumber="455" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED00AE3BFFBCA60AFF51AE776309F9AB" box="[210,274,1617,1643]" pageId="4" pageNumber="455">CHH</collectionCode>
),
<collectingMunicipality id="6BCAAC84FFBCA60AFEA4AE7763DFF9AB" box="[295,452,1617,1643]" pageId="4" pageNumber="455">Guanjiaoshan</collectingMunicipality>
,
<quantity id="4CE99B1BFFBCA60AFE53AE776003F9AB" box="[464,536,1617,1643]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="455" unit="km" value="20.0">20 km</quantity>
SW. of
<location id="8ECE6025FFBCA60AFDF5AE7760D4F9AB" LSID="urn:lsid:plazi:treatment:03B887E8FFBCA60BFF14AAA860CDFE6D:8ECE6025FFBCA60AFDF5AE7760D4F9AB" box="[630,719,1617,1643]" country="China" municipality="Guanjiaoshan" name="Tianjun" pageId="4" pageNumber="455" stateProvince="Qinghai">Tianjun</location>
,
<date id="FFAF103EFFBCA60AFD58AE77617DF9AB" box="[731,870,1617,1643]" pageId="4" pageNumber="455" value="2020-07-30">
<collectingDate id="EFEBE9D6FFBCA60AFD58AE77617DF9AB" box="[731,870,1617,1643]" pageId="4" pageNumber="455" value="2020-07-30">30.VII.2020</collectingDate>
</date>
,
<collectorName id="26E45328FFBCA60AFCF0AE7761F6F9AC" box="[883,1005,1617,1644]" pageId="4" pageNumber="455">S.-Y. Lang</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793CA3FFBCA60AFBAEAE776386F94F" ID-GBIF-Occurrence="3698883309" collectingDate="2020-07-05" collectionCode="CHH" collectorName="Z. Yao" country="China" location="Huashixia" pageId="4" pageNumber="455" specimenCount="7" specimenCount-female="1" specimenCount-male="6" stateProvince="Qinghai">
<specimenCount id="9D17FD77FFBCA60AFBAEAE776675F9AC" box="[1069,1134,1617,1644]" pageId="4" pageNumber="455" type="male">6 ♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFBCA60AFB17AE7766A4F9AC" box="[1172,1215,1617,1644]" pageId="4" pageNumber="455" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED00AE3BFFBCA60AFB4CAE776714F9AB" box="[1231,1295,1617,1643]" pageId="4" pageNumber="455">CHH</collectionCode>
),
<location id="8ECE6025FFBCA60AFAA7AE776783F9AB" LSID="urn:lsid:plazi:treatment:03B887E8FFBCA60BFF14AAA860CDFE6D:8ECE6025FFBCA60AFAA7AE776783F9AB" box="[1316,1432,1617,1643]" country="China" name="Huashixia" pageId="4" pageNumber="455" stateProvince="Qinghai">Huashixia</location>
,
<date id="FFAF103EFFBCA60AFF14AE53630EF950" box="[151,277,1653,1680]" pageId="4" pageNumber="455" value="2020-07-05">
<collectingDate id="EFEBE9D6FFBCA60AFF14AE53630EF950" box="[151,277,1653,1680]" pageId="4" pageNumber="455" value="2020-07-05">5.VII.2020</collectingDate>
</date>
,
<collectorName id="26E45328FFBCA60AFEA3AE506371F950" box="[288,362,1654,1680]" pageId="4" pageNumber="455">Z. Yao</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793CA3FFBCA60AFE2AAE5366CEF94F" ID-GBIF-Occurrence="3698883331" box="[425,1237,1653,1680]" collectingDate="2018-07-10" collectionCode="CSK" collectorName="K. Song" country="China" county="On" location="Xiahe" municipality="Tongren" pageId="4" pageNumber="455" specimenCount="3" specimenCount-male="3" stateProvince="Qinghai">
<specimenCount id="9D17FD77FFBCA60AFE2AAE5363F3F950" box="[425,488,1653,1680]" pageId="4" pageNumber="455" type="male">3 ♂♂</specimenCount>
(
<collectionCode id="ED00AE3BFFBCA60AFE74AE536029F94F" box="[503,562,1653,1679]" pageId="4" pageNumber="455">CSK</collectionCode>
),
<collectingCounty id="62CF4E72FFBCA60AFDC5AE536073F94F" box="[582,616,1653,1679]" pageId="4" pageNumber="455">On</collectingCounty>
road from
<collectingMunicipality id="6BCAAC84FFBCA60AFD66AE506159F950" box="[741,834,1654,1680]" pageId="4" pageNumber="455">Tongren</collectingMunicipality>
to
<location id="8ECE6025FFBCA60AFCE5AE5361B7F94F" LSID="urn:lsid:plazi:treatment:03B887E8FFBCA60BFF14AAA860CDFE6D:8ECE6025FFBCA60AFCE5AE5361B7F94F" box="[870,940,1653,1679]" country="China" county="On" municipality="Tongren" name="Xiahe" pageId="4" pageNumber="455" stateProvince="Qinghai">Xiahe</location>
,
<date id="FFAF103EFFBCA60AFC35AE536659F94F" box="[950,1090,1653,1679]" pageId="4" pageNumber="455" value="2018-07-10">
<collectingDate id="EFEBE9D6FFBCA60AFC35AE536659F94F" box="[950,1090,1653,1679]" pageId="4" pageNumber="455" value="2018-07-10">10.VII.2018</collectingDate>
</date>
,
<collectorName id="26E45328FFBCA60AFBCEAE5066B1F94F" box="[1101,1194,1653,1680]" pageId="4" pageNumber="455">K. Song</collectorName>
leg
</materialsCitation>
..
</paragraph>
</subSubSection>
<subSubSection id="C30B6575FFBCA60BFF44AEBF60CDFE6D" lastPageId="5" lastPageNumber="456" pageId="4" pageNumber="455" type="diagnosis">
<paragraph id="8BAE36FEFFBCA60AFF44AEBF66F8F918" blockId="4.[151,1437,1473,2040]" pageId="4" pageNumber="455">
<emphasis id="B965EAECFFBCA60AFF44AEBF6359F973" bold="true" box="[199,322,1689,1715]" pageId="4" pageNumber="455">Diagnosis.</emphasis>
For differences between
<taxonomicName id="4C114D7DFFBCA60AFDDEAEBC6101F973" authority="wanda" authorityName="wanda Grum-Grshimailo" authorityYear="1907" box="[605,794,1689,1716]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="wanda" subSpecies="wanda">
<emphasis id="B965EAECFFBCA60AFDDEAEBC6101F973" box="[605,794,1689,1716]" italics="true" pageId="4" pageNumber="455">C. wanda wanda</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D7DFFBCA60AFCCCAEBC661FF973" authorityName="Rose &amp; Schulte" authorityYear="1992" box="[847,1028,1689,1716]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="arida" subSpecies="cakana">
<emphasis id="B965EAECFFBCA60AFCCCAEBC661FF973" box="[847,1028,1689,1716]" italics="true" pageId="4" pageNumber="455">C. arida cakana</emphasis>
</taxonomicName>
, see under
<taxonomicName id="4C114D7DFFBCA60AFB00AEBC6723F973" authorityName="Rose &amp; Schulte" authorityYear="1992" box="[1155,1336,1689,1716]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="arida" subSpecies="cakana">
<emphasis id="B965EAECFFBCA60AFB00AEBC6723F973" box="[1155,1336,1689,1716]" italics="true" pageId="4" pageNumber="455">C. arida cakana</emphasis>
</taxonomicName>
. For differences between
<taxonomicName id="4C114D7DFFBCA60AFEE3AE986006F917" authority="wanda" authorityName="wanda Grum-Grshimailo" authorityYear="1907" box="[352,541,1725,1752]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="wanda" subSpecies="wanda">
<emphasis id="B965EAECFFBCA60AFEE3AE986006F917" box="[352,541,1725,1752]" italics="true" pageId="4" pageNumber="455">C. wanda wanda</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D7DFFBCA60AFDD0AE98614CF917" authority="adelaidae" authorityName="adelaidae Verhulst" authorityYear="1991" box="[595,855,1725,1752]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="adelaidae" subSpecies="adelaidae">
<emphasis id="B965EAECFFBCA60AFDD0AE98614CF917" box="[595,855,1725,1752]" italics="true" pageId="4" pageNumber="455">C. adelaidae adelaidae</emphasis>
</taxonomicName>
, see under
<taxonomicName id="4C114D7DFFBCA60AFC54AE9866C0F917" authority="adelaidae" authorityName="adelaidae Verhulst" authorityYear="1991" box="[983,1243,1725,1752]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="adelaidae" subSpecies="adelaidae">
<emphasis id="B965EAECFFBCA60AFC54AE9866C0F917" box="[983,1243,1725,1752]" italics="true" pageId="4" pageNumber="455">C. adelaidae adelaidae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAE36FEFFBCA60BFF44AEC76089FF39" blockId="4.[151,1437,1473,2040]" lastBlockId="5.[151,1436,151,429]" lastPageId="5" lastPageNumber="456" pageId="4" pageNumber="455">
<emphasis id="B965EAECFFBCA60AFF44AEC7636BF93B" bold="true" box="[199,368,1761,1787]" pageId="4" pageNumber="455">Identification.</emphasis>
The photos of the type specimens of both
<taxonomicName id="4C114D7DFFBCA60AFCCFAEC761FCF93B" authorityName=": Grieshuber &amp; Churkin" authorityYear="2003" box="[844,999,1761,1787]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="wanda">
<emphasis id="B965EAECFFBCA60AFCCFAEC761FCF93B" box="[844,999,1761,1787]" italics="true" pageId="4" pageNumber="455">Colias wanda</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D7DFFBCA60AFB9EAEC46685F93B" authorityName="Kotzsch" authorityYear="1930" box="[1053,1182,1761,1788]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="baeckeri">
<emphasis id="B965EAECFFBCA60AFB9EAEC46685F93B" box="[1053,1182,1761,1788]" italics="true" pageId="4" pageNumber="455">C. baeckeri</emphasis>
</taxonomicName>
presented by
<bibRefCitation id="EF804B0FFFBCA60AFAB9AEC7631EF8E0" author="Grieshuber, J." pageId="4" pageNumber="455" refId="ref14757" refString="Grieshuber, J. (2015) Images by Josef Grieshuber. Available from: https: // commons. wikimedia. org / wiki / Category: Images _ by _ Josef _ Grieshuber (accessed 23 October 2021)" type="url" year="2015">Grieshuber (2015</bibRefCitation>
,
<bibRefCitation id="EF804B0FFFBCA60AFE93AF23634DF8DF" author="Grieshuber, J." box="[272,342,1797,1823]" pageId="4" pageNumber="455" refId="ref14798" refString="Grieshuber, J. (2016 a) Specimen of Colias in Museum of Zoology, the Zoological Institute of the Russian Academy of Sciences (Saint Petersburg). Available from: https: // commons. wikimedia. org / wiki / Category: Specimen _ of _ Colias _ in _ Museum _ of _ Zoology _ (Saint _ Petersburg) (accessed 23 October 2021)" type="url" year="2016">2016a</bibRefCitation>
,
<bibRefCitation id="EF804B0FFFBCA60AFEE2AF236380F8DF" author="Grieshuber, J." box="[353,411,1797,1823]" pageId="4" pageNumber="455" refId="ref14928" refString="Grieshuber, J. (2020) Lepidoptera in the Zoologische Staatssammlung Munchen. Available from: https: // commons. wikimedia. org / wiki / Category: Lepidoptera _ in _ the _ Zoologische _ Staatssammlung _ M % C 3 % BCnchen (accessed 23 October 2021)" type="url" year="2020">2020</bibRefCitation>
) on the internet are consulted. One of the male specimens from Tongren area (
<figureCitation id="132A2A7BFFBCA60AFB7DAF236724F8E0" box="[1278,1343,1797,1824]" captionStart="FIGURES 67" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[204,1383,181,1920]" captionTargetId="figure-19@15.[204,1383,181,1920]" captionTargetPageId="15" captionText="FIGURES 67. Habitus of Colias taxa ♂♂ at same scale (Upper half- upper side; lower half- underside). 6—Colias arida cakana; 7—Colias wanda wanda. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.6333779" httpUri="https://zenodo.org/record/6333779/files/figure.png" pageId="4" pageNumber="455">Fig. 7</figureCitation>
- Specimen C27), neighbouring to the type locality of
<taxonomicName id="4C114D7DFFBCA60AFD28AF0C610CF883" authorityName=": Grieshuber &amp; Churkin" authorityYear="2003" box="[683,791,1833,1860]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="wanda">
<emphasis id="B965EAECFFBCA60AFD28AF0C610CF883" box="[683,791,1833,1860]" italics="true" pageId="4" pageNumber="455">C. wanda</emphasis>
</taxonomicName>
, is almost identical with the
<typeStatus id="54AA885CFFBCA60AFBDEAF0F66DFF883" box="[1117,1220,1833,1859]" pageId="4" pageNumber="455" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C114D7DFFBCA60AFB6AAF0C674DF883" authorityName=": Grieshuber &amp; Churkin" authorityYear="2003" box="[1257,1366,1833,1860]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="wanda">
<emphasis id="B965EAECFFBCA60AFB6AAF0C674DF883" box="[1257,1366,1833,1860]" italics="true" pageId="4" pageNumber="455">C. wanda</emphasis>
</taxonomicName>
in the wing-characters. Male and female specimens from Qilian County are in common with some of the type specimens of
<taxonomicName id="4C114D7DFFBCA60AFF35AF54632CF84B" authorityName="Kotzsch" authorityYear="1930" box="[182,311,1905,1932]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="baeckeri">
<emphasis id="B965EAECFFBCA60AFF35AF54632CF84B" box="[182,311,1905,1932]" italics="true" pageId="4" pageNumber="455">C. baeckeri</emphasis>
</taxonomicName>
. The authors agree with
<bibRefCitation id="EF804B0FFFBCA60AFDCCAF576186F84B" author="Grieshuber, J. &amp; Churkin, S." box="[591,925,1905,1931]" pageId="4" pageNumber="455" pagination="244 - 271" refId="ref14980" refString="Grieshuber, J. &amp; Churkin, S. (2003) The lectotypes of Colias diva Grum-Grshimailo, 1891, Colias wanda Grum-Grshimailo, 1907, Colias grumi Alpheraky, 1897, Colias cocandica maja Grum-Grshimailo, 1891, Colias cocandica tatarica Bang- Haas, 1915, and Colias tamerlana mongola Alpheraky, 1897 (Lepidoptera: Pieridae). Helios, Moscow, 4, 244 - 271, pls. 15 - 16." type="journal article" year="2003">Grieshuber &amp; Churkin (2003)</bibRefCitation>
in synonymizing
<taxonomicName id="4C114D7DFFBCA60AFBEBAF5466F2F84B" authorityName="Kotzsch" authorityYear="1930" box="[1128,1257,1905,1932]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="baeckeri">
<emphasis id="B965EAECFFBCA60AFBEBAF5466F2F84B" box="[1128,1257,1905,1932]" italics="true" pageId="4" pageNumber="455">C. baeckeri</emphasis>
</taxonomicName>
with
<taxonomicName id="4C114D7DFFBCA60AFAA9AF54678DF84B" authorityName=": Grieshuber &amp; Churkin" authorityYear="2003" box="[1322,1430,1905,1932]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="wanda">
<emphasis id="B965EAECFFBCA60AFAA9AF54678DF84B" box="[1322,1430,1905,1932]" italics="true" pageId="4" pageNumber="455">C. wanda</emphasis>
</taxonomicName>
, considering the great individual variations in male and the consistence of the wing characters in female. The male specimens from Huashixia area are a little different in the more frequently greenish ground color on hindwing underside, but such difference is not out of the individual variations of
<taxonomicName id="4C114D7DFFBCA60AFC1AAFF8664DF837" authority="wanda" authorityName="wanda Grum-Grshimailo" authorityYear="1907" box="[921,1110,2013,2040]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="wanda" subSpecies="wanda">
<emphasis id="B965EAECFFBCA60AFC1AAFF8664DF837" box="[921,1110,2013,2040]" italics="true" pageId="4" pageNumber="455">C. wanda wanda</emphasis>
</taxonomicName>
. The female specimens from Huashixia are in common with the females from the other localities in
<collectingRegion id="49D5F81CFFBDA60BFC4BA8B1663DFF71" box="[968,1062,151,177]" country="China" name="Qinghai" pageId="5" pageNumber="456">Qinghai</collectingRegion>
. Specimens from Dulan area are larger than in other populations of
<taxonomicName id="4C114D7DFFBDA60BFD9AA89D609EFF15" authorityName=": Grieshuber &amp; Churkin" authorityYear="2003" box="[537,645,187,213]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="wanda">
<emphasis id="B965EAECFFBDA60BFD9AA89D609EFF15" box="[537,645,187,213]" italics="true" pageId="5" pageNumber="456">C. wanda</emphasis>
</taxonomicName>
, often as large as the specimens of
<taxonomicName id="4C114D7DFFBDA60BFB8DA89D66D8FF15" authorityName="Rose &amp; Schulte" authorityYear="1992" box="[1038,1219,187,213]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="subSpecies" species="arida" subSpecies="cakana">
<emphasis id="B965EAECFFBDA60BFB8DA89D66D8FF15" box="[1038,1219,187,213]" italics="true" pageId="5" pageNumber="456">C. arida cakana</emphasis>
</taxonomicName>
; they are identified as
<taxonomicName id="4C114D7DFFBDA60BFF35A8F96368FF39" authority="wanda" authorityName="wanda Grum-Grshimailo" authorityYear="1907" box="[182,371,223,249]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="subSpecies" species="wanda" subSpecies="wanda">
<emphasis id="B965EAECFFBDA60BFF35A8F96368FF39" box="[182,371,223,249]" italics="true" pageId="5" pageNumber="456">C. wanda wanda</emphasis>
</taxonomicName>
on the female characters.
</paragraph>
<paragraph id="8BAE36FEFFBDA60BFF44A92463ECFEDD" blockId="5.[151,1436,151,429]" box="[199,503,258,285]" pageId="5" pageNumber="456">
<emphasis id="B965EAECFFBDA60BFF44A924637FFEDC" bold="true" box="[199,356,258,284]" pageId="5" pageNumber="456">Flight period</emphasis>
. Whole July.
</paragraph>
<paragraph id="8BAE36FEFFBDA60BFF44A90063C8FE81" blockId="5.[151,1436,151,429]" box="[199,467,294,321]" pageId="5" pageNumber="456">
<emphasis id="B965EAECFFBDA60BFF44A9006332FE80" bold="true" box="[199,297,294,320]" pageId="5" pageNumber="456">Altitude</emphasis>
.
<quantity id="4CE99B1BFFBDA60BFEB4A90163D5FE81" box="[311,462,295,321]" metricMagnitude="3" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" pageId="5" pageNumber="456" unit="m" value="4050.0" valueMax="4300.0" valueMin="3800.0">38004300m</quantity>
.
</paragraph>
<paragraph id="8BAE36FEFFBDA60BFF44A96C60CDFE6D" blockId="5.[151,1436,151,429]" pageId="5" pageNumber="456">
<emphasis id="B965EAECFFBDA60BFF44A96C63A0FEA5" bold="true" box="[199,443,330,357]" pageId="5" pageNumber="456">Distributional notes.</emphasis>
The junior author collected
<specimenCount id="9D17FD77FFBDA60BFD70A96D612AFEA5" box="[755,817,331,357]" pageId="5" pageNumber="456" type="male">3 ♂♂</specimenCount>
from a locality of Tongren (Fig. 68), close to Xiahe, the
<typeStatus id="54AA885CFFBDA60BFF14A94962DCFE49" box="[151,199,367,393]" pageId="5" pageNumber="456">type</typeStatus>
locality of
<taxonomicName id="4C114D7DFFBDA60BFEC9A94963C2FE49" authorityName="Verhulst" authorityYear="1991" box="[330,473,367,393]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="adelaidae">
<emphasis id="B965EAECFFBDA60BFEC9A94963C2FE49" box="[330,473,367,393]" italics="true" pageId="5" pageNumber="456">C. adelaidae</emphasis>
</taxonomicName>
. He also collected a series of specimens from a locality of Dulan (Fig. 68), ca
<quantity id="4CE99B1BFFBDA60BFAD6A9486787FE49" box="[1365,1436,366,393]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="456" unit="km" value="15.0">15 km</quantity>
away from a new locality of
<taxonomicName id="4C114D7DFFBDA60BFE5AA9B56073FE6D" authorityName="Verhulst" authorityYear="1991" box="[473,616,403,429]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="adelaidae">
<emphasis id="B965EAECFFBDA60BFE5AA9B56073FE6D" box="[473,616,403,429]" italics="true" pageId="5" pageNumber="456">C. adelaidae</emphasis>
</taxonomicName>
at Dulan.
</paragraph>
</subSubSection>
</treatment>
</document>