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<document id="2639080F20AE3B43E69ABCE6A628F53C" ID-DOI="10.11646/phytotaxa.602.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8141984" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1689235114003" checkinUser="plazi" docAuthor="Fernandez-Hilario, Robin, Goldenberg, Renato &amp; Michelangeli, Fabián A." docDate="2023" docId="03B887DAFFB2FFBAFF62C71AFA6DF7A2" docLanguage="en" docName="phytotaxa.602.1.1.pdf" docOrigin="Phytotaxa 602 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.602.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Meriania" docType="key" docVersion="3" lastPageNumber="13" masterDocId="FF81FFA2FFB9FFB6FFEAC236FFC7FFAB" masterDocTitle="A synopsis of Meriania (Melastomataceae: Merianieae) in Peru" masterLastPageNumber="101" masterPageNumber="1" pageNumber="12" updateTime="1689341265902" updateUser="juliana">
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<mods:title id="70F5D39ECFBBD6501CB6CFEB26E4DC26">A synopsis of Meriania (Melastomataceae: Merianieae) in Peru</mods:title>
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<mods:namePart id="EFBD39BB4664B516FA5593BBF3875A6A">Fernandez-Hilario, Robin</mods:namePart>
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<mods:affiliation id="C235DD9755EDFAE078743E7C011AAFC1">Programa de Pós-Graduação em Botânica, Universidade Federal do Paraná, Caixa Postal 19031, CEP 81531 - 970, Curitiba, PR, Brazil. &amp; División de Ecología Vegetal - CORBIDI, Calle Santa Rita 105 Of. 2, Urb. Huertos de San Antonio Monterrico, Surco, Lima, Perú. &amp; rfernandez @ corbidi. org; https: // orcid. org / 0000 - 0001 - 8234 - 9502</mods:affiliation>
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<mods:namePart id="B08187E2DA22190384F2B8E5C4554D3B">Goldenberg, Renato</mods:namePart>
<mods:nameIdentifier id="578E59333EEABC848EE8B6652F63D8B7" type="ORCID">0000-0002-7047-6720</mods:nameIdentifier>
<mods:affiliation id="7D53030D5FAD9B2C47B805D33F4EF633">Programa de Pós-Graduação em Botânica, Universidade Federal do Paraná, Caixa Postal 19031, CEP 81531 - 970, Curitiba, PR, Brazil. &amp; renato. goldenberg @ gmail. com; https: // orcid. org / 0000 - 0002 - 7047 - 6720</mods:affiliation>
<mods:nameIdentifier id="C212DFA73AB99D1A53330A6977627FD0" type="email">renato.goldenberg@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B49819696B50E6DAB934E05B189DC48A">Michelangeli, Fabián A.</mods:namePart>
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<mods:affiliation id="BBEEEC42AF66CB96C87239A59CA77C41">Programa de Pós-Graduação em Botânica, Universidade Federal do Paraná, Caixa Postal 19031, CEP 81531 - 970, Curitiba, PR, Brazil. &amp; The New York Botanical Garden, Bronx, NY 104458 USA. &amp; Programa de Pós-Graduação em Botânica, Universidade Federal do Paraná, Caixa Postal 19031, CEP 81531 - 970, Curitiba, PR, Brazil. &amp; fabian @ nybg. org; https: // orcid. org / 0000 - 0001 - 7348 - 143 X Corresponding author: rfernandez @ corbidi. org &amp; Programa de Pós-Graduação em Botânica, Universidade Federal do Paraná, Caixa Postal 19031, CEP 81531 - 970, Curitiba, PR, Brazil.</mods:affiliation>
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<mods:title id="7E7EE60629E39B7CEB10E99322B095DA">Phytotaxa</mods:title>
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<mods:date id="29AD556624F9F47182D8284BD7050524">2023</mods:date>
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<mods:number id="6DF941CFC4A53A87B0374C7C82B912B4">602</mods:number>
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<subSubSection id="C30B6547FFB2FFBDFF62C71AFDA6FAED" box="[136,609,1324,1350]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BAE36CCFFB2FFBDFF62C71AFDA6FAED" blockId="11.[136,609,1324,1350]" box="[136,609,1324,1350]" pageId="11" pageNumber="12">
<heading id="D0E681A0FFB2FFBDFF62C71AFDA6FAED" bold="true" box="[136,609,1324,1350]" fontSize="36" level="2" pageId="11" pageNumber="12" reason="6">
<emphasis id="B965EADEFFB2FFBDFF62C71AFDA6FAED" bold="true" box="[136,609,1324,1350]" pageId="11" pageNumber="12">
Key to the species of
<taxonomicName id="4C114D4FFFB2FFBDFE64C71AFE39FAED" ID-CoL="5P8S" ID-ENA="119892" box="[398,510,1324,1350]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="genus">
<emphasis id="B965EADEFFB2FFBDFE64C71AFE39FAED" bold="true" box="[398,510,1324,1350]" italics="true" pageId="11" pageNumber="12">Meriania</emphasis>
</taxonomicName>
in
<collectingCountry id="F306765CFFB2FFBDFDCFC71AFDA6FAED" box="[549,609,1324,1350]" name="Peru" pageId="11" pageNumber="12">Peru</collectingCountry>
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<keyStep id="30E52E69FFB2FFBDFF62C747FA6CFA14" pageId="11" pageNumber="12">
<paragraph id="8BAE36CCFFB2FFBDFF62C747FA6DFA2D" blockId="11.[136,1452,1392,2077]" box="[136,1450,1392,1414]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C747FA6DFA2D" box="[136,1450,1392,1414]" pageId="11" pageNumber="12">
1. Leaf blades with dentate-undulate, revolute auricles at the base (
<figureCitation id="132A2A49FFB2FFBDFCD6C746FC40FA2D" box="[828,903,1392,1414]" captionStart="FIGURE 4" captionStartId="8.[136,229,954,976]" captionTargetBox="[151,1433,194,929]" captionTargetId="figure-20@8.[150,1435,193,931]" captionTargetPageId="8" captionText="FIGURE 4. Morphological characteristics of Peruvian Meriania. A. Bullate adaxial leaf surface (M. sanguinea; R. Fernandez-Hilario et al. 1896). B. Punctiform abaxial leaf surface (M. cuzcoana; W. Farfán et al. 976). C. Glabrous abaxial leaf surface (M. callosa; R. Fernandez-Hilario et al. 2055). D. Furfuraceous abaxial leaf surface (M. vilcabambensis; L. Valenzuela 7494). E. Puberulent abaxial leaf surface (M. drakei; R. Fernandez-Hilario et al. 1775). F. Pubescent abaxial leaf surface (M. bicentenaria; A. Monteagudo et al. 6960). G. Tomentose abaxial leaf surface (M. tomentosa; R. Fernandez-Hilario et al. 1905). H. Setulose abaxial leaf surface (M. radula; D. Paredes et al. 604). I. Villose abaxial leaf surface (M. sumatika; I. Huamantupa et al. 2060). J. Hirsute adaxial leaf surface (M. hirsuta; S. Baldeón &amp; J. Campos 5373). K. Peltate leaf base (M. peltata; R. Fernandez-Hilario et al. 2093). L. Leaf base with dentate-undulate revolute auricles (M. cuzcoana; L. Valenzuela et al. 6904)." figureDoi="http://doi.org/10.5281/zenodo.8141996" httpUri="https://zenodo.org/record/8141996/files/figure.png" pageId="11" pageNumber="12">Fig. 4L</figureCitation>
), and punctiform abaxial surfaces (
<figureCitation id="132A2A49FFB2FFBDFB32C746FAE5FA2D" box="[1240,1314,1392,1414]" captionStart="FIGURE 4" captionStartId="8.[136,229,954,976]" captionTargetBox="[151,1433,194,929]" captionTargetId="figure-20@8.[150,1435,193,931]" captionTargetPageId="8" captionText="FIGURE 4. Morphological characteristics of Peruvian Meriania. A. Bullate adaxial leaf surface (M. sanguinea; R. Fernandez-Hilario et al. 1896). B. Punctiform abaxial leaf surface (M. cuzcoana; W. Farfán et al. 976). C. Glabrous abaxial leaf surface (M. callosa; R. Fernandez-Hilario et al. 2055). D. Furfuraceous abaxial leaf surface (M. vilcabambensis; L. Valenzuela 7494). E. Puberulent abaxial leaf surface (M. drakei; R. Fernandez-Hilario et al. 1775). F. Pubescent abaxial leaf surface (M. bicentenaria; A. Monteagudo et al. 6960). G. Tomentose abaxial leaf surface (M. tomentosa; R. Fernandez-Hilario et al. 1905). H. Setulose abaxial leaf surface (M. radula; D. Paredes et al. 604). I. Villose abaxial leaf surface (M. sumatika; I. Huamantupa et al. 2060). J. Hirsute adaxial leaf surface (M. hirsuta; S. Baldeón &amp; J. Campos 5373). K. Peltate leaf base (M. peltata; R. Fernandez-Hilario et al. 2093). L. Leaf base with dentate-undulate revolute auricles (M. cuzcoana; L. Valenzuela et al. 6904)." figureDoi="http://doi.org/10.5281/zenodo.8141996" httpUri="https://zenodo.org/record/8141996/files/figure.png" pageId="11" pageNumber="12">Fig. 4B</figureCitation>
)
<taxonomicName id="4C114D4FFFB2FFBDFAC6C747FA6DFA2D" box="[1324,1450,1393,1414]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="cuzcoana">
<emphasis id="B965EADEFFB2FFBDFAC6C747FA6DFA2D" bold="true" box="[1324,1450,1393,1414]" italics="true" pageId="11" pageNumber="12">M. cuzcoana</emphasis>
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</paragraph>
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- Leaf blades without auricles (but
<taxonomicName id="4C114D4FFFB2FFBDFDF9C7B8FDA9FA09" box="[531,622,1421,1443]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis id="B965EADEFFB2FFBDFDF9C7B8FDA9FA09" box="[531,622,1421,1443]" italics="true" pageId="11" pageNumber="12">M. rigida</emphasis>
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and
<taxonomicName id="4C114D4FFFB2FFBDFD76C7B8FCE3FA08" box="[668,804,1422,1443]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="zunacensis">
<emphasis id="B965EADEFFB2FFBDFD76C7B8FCE3FA08" box="[668,804,1422,1443]" italics="true" pageId="11" pageNumber="12">M. zunacensis</emphasis>
</taxonomicName>
with revolute bases, and
<taxonomicName id="4C114D4FFFB2FFBDFBFCC7B8FB60FA09" box="[1046,1191,1421,1443]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="franciscana">
<emphasis id="B965EADEFFB2FFBDFBFCC7B8FB60FA09" box="[1046,1191,1421,1443]" italics="true" pageId="11" pageNumber="12">M. franciscana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D4FFFB2FFBDFB3FC7B8FAEAFA09" box="[1237,1325,1421,1443]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="neillii">
<emphasis id="B965EADEFFB2FFBDFB3FC7B8FAEAFA09" box="[1237,1325,1421,1443]" italics="true" pageId="11" pageNumber="12">M. neillii</emphasis>
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with slightly revolute bases), and abaxial surface not punctiform ..........................................................................................................................2
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<paragraph id="8BAE36CCFFB2FFBDFF62C7F1FA6CFA77" blockId="11.[136,1452,1392,2077]" box="[136,1451,1479,1501]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C7F1FA6CFA77" box="[136,1451,1479,1501]" pageId="11" pageNumber="12">2. Inflorescences solitary, dichasia 3 or 9-flowered (often reduced to 5-flowered) ...............................................................................3</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB2FFBDFF62C7D2FA6CFA51" blockId="11.[136,1452,1392,2077]" box="[136,1451,1508,1530]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C7D2FA6CFA51" box="[136,1451,1508,1530]" pageId="11" pageNumber="12">
- Inflorescences in panicles with more than 10 flowers (but see comments below
<taxonomicName id="4C114D4FFFB2FFBDFC24C7D2FBF5FA52" box="[974,1074,1508,1529]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="hirsuta">
<emphasis id="B965EADEFFB2FFBDFC24C7D2FBF5FA52" box="[974,1074,1508,1529]" italics="true" pageId="11" pageNumber="12">M. hirsuta</emphasis>
</taxonomicName>
)...........................................................5
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<keyStep id="30E52E69FFB2FFBDFF62C437FA6CF999" pageId="11" pageNumber="12">
<paragraph id="8BAE36CCFFB2FFBDFF62C437FA6CF9BD" blockId="11.[136,1452,1392,2077]" box="[136,1451,1536,1558]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C437FA6CF9BD" box="[136,1451,1536,1558]" pageId="11" pageNumber="12">
3. Calyx calyptrate with irregular dehiscence ...................................................................................................................
<taxonomicName id="4C114D4FFFB2FFBDFACBC437FA6CF9BD" authorityName="Rob. Fern., R. Goldenb. &amp; Michelang. Ademas" authorityYear="2023" baseAuthorityName="Rob. Fern., R. Goldenb. &amp; Michelang. Ademas" baseAuthorityYear="2023" box="[1313,1451,1536,1558]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="sessilifolia">
<emphasis id="B965EADEFFB2FFBDFACBC437FA6CF9BD" bold="true" box="[1313,1451,1536,1558]" italics="true" pageId="11" pageNumber="12">M. sessilifolia</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB2FFBDFF62C428FA6CF999" blockId="11.[136,1452,1392,2077]" box="[136,1451,1565,1587]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C428FA6CF999" box="[136,1451,1565,1587]" pageId="11" pageNumber="12">- Calyx lobed with regular dehiscence..................................................................................................................................................4</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8BAE36CCFFB2FFBDFF62C40CFA6CF9C7" blockId="11.[136,1452,1392,2077]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C40CFA6CF9C7" pageId="11" pageNumber="12">
4. Terminal branches and leaf blades glabrous; leaf blades
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long; hypanthium terete; calyx with acute dorsal projections .. ..........................................................................................................................................................................................
<taxonomicName id="4C114D4FFFB2FFBDFADDC461FA6CF9C7" box="[1335,1451,1623,1644]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="speciosa">
<emphasis id="B965EADEFFB2FFBDFADDC461FA6CF9C7" bold="true" box="[1335,1451,1623,1644]" italics="true" pageId="11" pageNumber="12">M. speciosa</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BAE36CCFFB2FFBDFF62C442FA6CF90D" blockId="11.[136,1452,1392,2077]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C442FA6CF90D" pageId="11" pageNumber="12">
- Terminal branches and abaxial leaf blades moderately to densely furfuraceous; leaf blades
<quantity id="4CE99B29FFB2FFBDFBAAC442FB6CF922" box="[1088,1195,1652,1673]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="8.2" metricValueMin="2.3" pageId="11" pageNumber="12" unit="cm" value="5.25" valueMax="8.2" valueMin="2.3">2.38.2 cm</quantity>
long; hypanthium slightly costate; calyx with acicular dorsal projections (
<figureCitation id="132A2A49FFB2FFBDFD7DC4A6FD25F90D" box="[663,738,1680,1702]" captionStart="FIGURE 5" captionStartId="9.[136,229,1613,1635]" captionTargetBox="[155,1429,217,1583]" captionTargetId="figure-18@9.[150,1435,209,1589]" captionTargetPageId="9" captionText="FIGURE 5. Morphological characteristics of Peruvian Meriania. A. Calyx lacking dorsal projections (M. hexamera; I. Revilla 3484). B. Calyx with callose dorsal projections (left), and detail of a dorsal projection (right) (M. callosa; R. Fernandez-Hilario et al. 2055). C. Calyx with whitish callose dorsal projections (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Calyx with claw-shaped dorsal projections (M. dazae; J. L. Marcelo-Peña et al. 6568). E. Calyx with blunt dorsal projections (M. rugosa; H. van der Werff 17006). F. Subcalyptrate calyx with small claw-shaped projections and irregular dehiscence (left), and detail of the subcalyptrate calyx apex (right) (M. vasquezii; R. Vásquez et al. 45480). G. Calyptrate calyx with circumscissile dehiscence (M. escalerensis; M. Ríos et al. 3316). H. Calyx with aciculate dorsal projections and slightly costate hypanthium (M. prunifolia; J. Schunke-Vigo 11712). I. Costate hypanthium with 10 evident ribs (left) and apical view (right) (M. sumatika; P. Núñez &amp; J. Arque 8369)." figureDoi="http://doi.org/10.5281/zenodo.8142002" httpUri="https://zenodo.org/record/8142002/files/figure.png" pageId="11" pageNumber="12">Fig. 5H</figureCitation>
) ..............................................................................................
<taxonomicName id="4C114D4FFFB2FFBDFACFC4A7FA6CF90D" box="[1317,1451,1680,1702]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="prunifolia">
<emphasis id="B965EADEFFB2FFBDFACFC4A7FA6CF90D" bold="true" box="[1317,1451,1680,1702]" italics="true" pageId="11" pageNumber="12">M. prunifolia</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB2FFBDFF62C49BFA6CF89D" pageId="11" pageNumber="12">
<paragraph id="8BAE36CCFFB2FFBDFF62C49BFA6CF94B" blockId="11.[136,1452,1392,2077]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C49BFA6CF94B" pageId="11" pageNumber="12">
5. Leaf blades bullate (
<figureCitation id="132A2A49FFB2FFBDFE4EC49BFE30F969" box="[420,503,1709,1731]" captionStart="FIGURE 4" captionStartId="8.[136,229,954,976]" captionTargetBox="[151,1433,194,929]" captionTargetId="figure-20@8.[150,1435,193,931]" captionTargetPageId="8" captionText="FIGURE 4. Morphological characteristics of Peruvian Meriania. A. Bullate adaxial leaf surface (M. sanguinea; R. Fernandez-Hilario et al. 1896). B. Punctiform abaxial leaf surface (M. cuzcoana; W. Farfán et al. 976). C. Glabrous abaxial leaf surface (M. callosa; R. Fernandez-Hilario et al. 2055). D. Furfuraceous abaxial leaf surface (M. vilcabambensis; L. Valenzuela 7494). E. Puberulent abaxial leaf surface (M. drakei; R. Fernandez-Hilario et al. 1775). F. Pubescent abaxial leaf surface (M. bicentenaria; A. Monteagudo et al. 6960). G. Tomentose abaxial leaf surface (M. tomentosa; R. Fernandez-Hilario et al. 1905). H. Setulose abaxial leaf surface (M. radula; D. Paredes et al. 604). I. Villose abaxial leaf surface (M. sumatika; I. Huamantupa et al. 2060). J. Hirsute adaxial leaf surface (M. hirsuta; S. Baldeón &amp; J. Campos 5373). K. Peltate leaf base (M. peltata; R. Fernandez-Hilario et al. 2093). L. Leaf base with dentate-undulate revolute auricles (M. cuzcoana; L. Valenzuela et al. 6904)." figureDoi="http://doi.org/10.5281/zenodo.8141996" httpUri="https://zenodo.org/record/8141996/files/figure.png" pageId="11" pageNumber="12">Fig. 4A</figureCitation>
); corolla campanulate and deep red; stamen connectives with dorsal-basal appendages almost perpendicular to thecae (
<figureCitation id="132A2A49FFB2FFBDFE2AC4FCFDCCF94B" box="[448,523,1738,1760]" captionStart="FIGURE 6" captionStartId="10.[136,229,1633,1655]" captionTargetBox="[156,1429,267,1603]" captionTargetId="figure-20@10.[150,1435,193,1609]" captionTargetPageId="10" captionText="FIGURE 6. Morphological characteristics of Peruvian Meriania. A. Stamen with a triangular descending dorso-basal appendage (black arrow) but no dorsal appendage (M. tetraquetra; R. Fernandez-Hilario &amp; A. Vásquez 251). B. Stamen with a triangular descending dorso-basal appendage and a blunt ascending dorsal appendage (blue arrow) (M. megaphylla; A. Weberbauer 7048). C. Stamen with an acuminate descending dorso-basal appendage (black arrow) and a dorsal appendage as a mere hump (blue arrow) (M. franciscana; C. Díaz &amp; S. Fernández 10150). D. Stamen with a triangular descending dorso-basal appendage and a dorsal appendage as a mere hump, and detail of the dorsal appendage (M. rugosa; F. A. Michelangeli et al. 1725). E. Stamen with a triangular descending dorso-basal appendage and a dentiform dorsal appendage, and detail of the dorsal appendage (M. hexamera; I. Revilla 3484). F. Stamen with a multilobed descending dorso-basal appendage and an apically trilobed ascending dorsal appendage, and detail of the dorsal appendage apex (M. microflora; I. Huamantupa et al. 15552). G. Stamen with an almost perpendicular dorso-basal appendage to the theca, and detail of the prolonged connective below the theca (M. tetragona; R. Fernandez-Hilario et al. 2092). H. Stamen with inflated (bulbous) connective (blue arrow) (M. bicentenaria; R. Villanueva-Espinoza 675). I. Antesepalous stamen with a laterally expanded dorso-basal appendage (left) and antepetalous stamen with a dorso-basal appendage not laterally expanded (right) (M. vilcabambensis; L. Valenzuela 7494)." figureDoi="http://doi.org/10.5281/zenodo.8142006" httpUri="https://zenodo.org/record/8142006/files/figure.png" pageId="11" pageNumber="12">Fig. 6G</figureCitation>
).......................................................................................................................................................6
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB2FFBDFF62C4D1FA6CF89D" blockId="11.[136,1452,1392,2077]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C4D1FA6CF89D" pageId="11" pageNumber="12">
- Leaf blades flat to bullate; corolla spreading or campanulate, white, pink, pink-orange to reddish-orange, or fuchsia to reddish-purple but never deep red (except red in
<taxonomicName id="4C114D4FFFB2FFBDFD8FC532FD20F8B2" box="[613,743,1796,1817]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="rubriflora">
<emphasis id="B965EADEFFB2FFBDFD8FC532FD20F8B2" box="[613,743,1796,1817]" italics="true" pageId="11" pageNumber="12">M. rubriflora</emphasis>
</taxonomicName>
); stamen connectives with dorsal-basal appendages not perpendicular to thecae..................................................................................................................................................................................................9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB2FFBDFF62C50BFA6CF8C4" pageId="11" pageNumber="12">
<paragraph id="8BAE36CCFFB2FFBDFF62C50BFA6CF8F8" blockId="11.[136,1452,1392,2077]" box="[136,1451,1853,1875]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C50BFA6CF8F8" box="[136,1451,1853,1875]" pageId="11" pageNumber="12">6. Nodes with developed interpetiolar flaps...........................................................................................................................................7</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB2FFBDFF62C56DFA6CF8C4" blockId="11.[136,1452,1392,2077]" box="[136,1451,1882,1904]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C56DFA6CF8C4" box="[136,1451,1882,1904]" pageId="11" pageNumber="12">- Nodes with only interpetiolar lines, not flaps.....................................................................................................................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB2FFBDFF62C541FA6CF849" pageId="11" pageNumber="12">
<paragraph id="8BAE36CCFFB2FFBDFF62C541FA6CF802" blockId="11.[136,1452,1392,2077]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C541FA6CF802" pageId="11" pageNumber="12">
7. Internodes quadrangular and 4-winged (
<figureCitation id="132A2A49FFB2FFBDFDA6C541FD50F827" box="[588,663,1911,1933]" captionStart="FIGURE 3" captionStartId="6.[136,229,1742,1764]" captionTargetBox="[217,1370,1054,1715]" captionTargetId="figure-317@6.[216,1372,1053,1717]" captionTargetPageId="6" captionText="FIGURE 3. Morphological characteristics of Peruvian Meriania. A. Interpetiolar line (M. tomentosa; R. W. Bussmann et al. 17068). B. Interpetiolar flap (M. zunacensis; R. Fernandez-Hilario et al. 1920). C. Winged internode (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Winged internode (M. tetragona; R. Fernandez-Hilario et al. 2092). E. Adaxial projection (scutum) in the transition zone from the petiole to the midvein (M. zunacensis; R. Fernandez-Hilario et al. 1920). F. Swollen adaxial projection on the petiole apex (M. drakei; R. Fernandez-Hilario et al. 1775). G. Liguliform abaxial projections in the transition zone from the petiole to the midvein (M. sanguinea; R. Fernandez-Hilario et al. 1896). H. Tuberculate abaxial projections in the transition zone from the petiole to the midvein (M. tetragona; F. A. Michelangeli et al. 1739). I. Spreading corolla (M. rigida; R. Fernandez-Hilario et al. 1931). J. Campanulate corollas (M. dazae; J. L. Marcelo-Peña et al. 6568). K. Fruits with mature ovaries exceeding the hypanthia length (M. sanguinea; F.A. Michelangeli et al. 2743). L. Fruits with mature ovaries completely concealed by the hypanthia (M. tomentosa; F. A. Michelangeli et al. 1799). Photos by Rainer W. Bussmann (A); Robin Fernandez-Hilario (BG and IJ); and Fabián A. Michelangeli (H, K and L)." figureDoi="http://doi.org/10.5281/zenodo.8141994" httpUri="https://zenodo.org/record/8141994/files/figure.png" pageId="11" pageNumber="12">Fig. 3D</figureCitation>
); petioles with abaxial tuberculate projections (rarely horn-shaped) on the transition zone from the petiole to the midvein (
<figureCitation id="132A2A49FFB2FFBDFDD7C5A2FD4FF802" box="[573,648,1940,1962]" captionStart="FIGURE 3" captionStartId="6.[136,229,1742,1764]" captionTargetBox="[217,1370,1054,1715]" captionTargetId="figure-317@6.[216,1372,1053,1717]" captionTargetPageId="6" captionText="FIGURE 3. Morphological characteristics of Peruvian Meriania. A. Interpetiolar line (M. tomentosa; R. W. Bussmann et al. 17068). B. Interpetiolar flap (M. zunacensis; R. Fernandez-Hilario et al. 1920). C. Winged internode (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Winged internode (M. tetragona; R. Fernandez-Hilario et al. 2092). E. Adaxial projection (scutum) in the transition zone from the petiole to the midvein (M. zunacensis; R. Fernandez-Hilario et al. 1920). F. Swollen adaxial projection on the petiole apex (M. drakei; R. Fernandez-Hilario et al. 1775). G. Liguliform abaxial projections in the transition zone from the petiole to the midvein (M. sanguinea; R. Fernandez-Hilario et al. 1896). H. Tuberculate abaxial projections in the transition zone from the petiole to the midvein (M. tetragona; F. A. Michelangeli et al. 1739). I. Spreading corolla (M. rigida; R. Fernandez-Hilario et al. 1931). J. Campanulate corollas (M. dazae; J. L. Marcelo-Peña et al. 6568). K. Fruits with mature ovaries exceeding the hypanthia length (M. sanguinea; F.A. Michelangeli et al. 2743). L. Fruits with mature ovaries completely concealed by the hypanthia (M. tomentosa; F. A. Michelangeli et al. 1799). Photos by Rainer W. Bussmann (A); Robin Fernandez-Hilario (BG and IJ); and Fabián A. Michelangeli (H, K and L)." figureDoi="http://doi.org/10.5281/zenodo.8141994" httpUri="https://zenodo.org/record/8141994/files/figure.png" pageId="11" pageNumber="12">Fig. 3H</figureCitation>
) ..............................................................................................................
<taxonomicName id="4C114D4FFFB2FFBDFAC1C5A2FA6CF802" box="[1323,1451,1940,1961]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="tetragona">
<emphasis id="B965EADEFFB2FFBDFAC1C5A2FA6CF802" bold="true" box="[1323,1451,1940,1961]" italics="true" pageId="11" pageNumber="12">M. tetragona</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB2FFBDFF62C587FA6CF849" blockId="11.[136,1452,1392,2077]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C587FA6CF849" pageId="11" pageNumber="12">
- Internodes terete-quadrangular; petioles with abaxial liguliform projections on the transition zone from the petiole to the midvein (
<figureCitation id="132A2A49FFB2FFBDFF30C5FBFEE2F849" box="[218,293,1997,2019]" captionStart="FIGURE 3" captionStartId="6.[136,229,1742,1764]" captionTargetBox="[217,1370,1054,1715]" captionTargetId="figure-317@6.[216,1372,1053,1717]" captionTargetPageId="6" captionText="FIGURE 3. Morphological characteristics of Peruvian Meriania. A. Interpetiolar line (M. tomentosa; R. W. Bussmann et al. 17068). B. Interpetiolar flap (M. zunacensis; R. Fernandez-Hilario et al. 1920). C. Winged internode (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Winged internode (M. tetragona; R. Fernandez-Hilario et al. 2092). E. Adaxial projection (scutum) in the transition zone from the petiole to the midvein (M. zunacensis; R. Fernandez-Hilario et al. 1920). F. Swollen adaxial projection on the petiole apex (M. drakei; R. Fernandez-Hilario et al. 1775). G. Liguliform abaxial projections in the transition zone from the petiole to the midvein (M. sanguinea; R. Fernandez-Hilario et al. 1896). H. Tuberculate abaxial projections in the transition zone from the petiole to the midvein (M. tetragona; F. A. Michelangeli et al. 1739). I. Spreading corolla (M. rigida; R. Fernandez-Hilario et al. 1931). J. Campanulate corollas (M. dazae; J. L. Marcelo-Peña et al. 6568). K. Fruits with mature ovaries exceeding the hypanthia length (M. sanguinea; F.A. Michelangeli et al. 2743). L. Fruits with mature ovaries completely concealed by the hypanthia (M. tomentosa; F. A. Michelangeli et al. 1799). Photos by Rainer W. Bussmann (A); Robin Fernandez-Hilario (BG and IJ); and Fabián A. Michelangeli (H, K and L)." figureDoi="http://doi.org/10.5281/zenodo.8141994" httpUri="https://zenodo.org/record/8141994/files/figure.png" pageId="11" pageNumber="12">Fig. 3G</figureCitation>
) ........................................................................................................................................................................
<taxonomicName id="4C114D4FFFB2FFBDFAC9C5F8FA6CF849" box="[1315,1451,1997,2019]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="sanguinea">
<emphasis id="B965EADEFFB2FFBDFAC9C5F8FA6CF849" bold="true" box="[1315,1451,1997,2019]" italics="true" pageId="11" pageNumber="12">M. sanguinea</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB2FFBAFF62C5DCFA6DFF7A" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<paragraph id="8BAE36CCFFB2FFBDFF62C5DCFA6CF7B7" blockId="11.[136,1452,1392,2077]" pageId="11" pageNumber="12">
<keyLead id="30E095F9FFB2FFBDFF62C5DCFA6CF7B7" pageId="11" pageNumber="12">
8. Branches, petioles, and both leaf surfaces hirsute (
<figureCitation id="132A2A49FFB2FFBDFD45C5DCFD33F854" box="[687,756,2026,2048]" captionStart="FIGURE 4" captionStartId="8.[136,229,954,976]" captionTargetBox="[151,1433,194,929]" captionTargetId="figure-20@8.[150,1435,193,931]" captionTargetPageId="8" captionText="FIGURE 4. Morphological characteristics of Peruvian Meriania. A. Bullate adaxial leaf surface (M. sanguinea; R. Fernandez-Hilario et al. 1896). B. Punctiform abaxial leaf surface (M. cuzcoana; W. Farfán et al. 976). C. Glabrous abaxial leaf surface (M. callosa; R. Fernandez-Hilario et al. 2055). D. Furfuraceous abaxial leaf surface (M. vilcabambensis; L. Valenzuela 7494). E. Puberulent abaxial leaf surface (M. drakei; R. Fernandez-Hilario et al. 1775). F. Pubescent abaxial leaf surface (M. bicentenaria; A. Monteagudo et al. 6960). G. Tomentose abaxial leaf surface (M. tomentosa; R. Fernandez-Hilario et al. 1905). H. Setulose abaxial leaf surface (M. radula; D. Paredes et al. 604). I. Villose abaxial leaf surface (M. sumatika; I. Huamantupa et al. 2060). J. Hirsute adaxial leaf surface (M. hirsuta; S. Baldeón &amp; J. Campos 5373). K. Peltate leaf base (M. peltata; R. Fernandez-Hilario et al. 2093). L. Leaf base with dentate-undulate revolute auricles (M. cuzcoana; L. Valenzuela et al. 6904)." figureDoi="http://doi.org/10.5281/zenodo.8141996" httpUri="https://zenodo.org/record/8141996/files/figure.png" pageId="11" pageNumber="12">Fig. 4J</figureCitation>
); antesepalous stamen connectives with dorso-basal appendages laterally expanded.............................................................................................................................................................................
<taxonomicName id="4C114D4FFFB2FFBDFAA9CA31FA6CF7B7" box="[1347,1451,2054,2076]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="hirsuta">
<emphasis id="B965EADEFFB2FFBDFAA9CA31FA6CF7B7" bold="true" box="[1347,1451,2054,2076]" italics="true" pageId="11" pageNumber="12">M. hirsuta</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C296FA6DFF7A" blockId="12.[136,1452,159,2058]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C296FA6DFF7A" pageId="12" pageNumber="13">
- Branches, petioles, and abaxial leaf surfaces moderately to densely setulose (
<figureCitation id="132A2A49FFB5FFBAFC52C2A9FBC1FF1F" box="[952,1030,159,181]" captionStart="FIGURE 4" captionStartId="8.[136,229,954,976]" captionTargetBox="[151,1433,194,929]" captionTargetId="figure-20@8.[150,1435,193,931]" captionTargetPageId="8" captionText="FIGURE 4. Morphological characteristics of Peruvian Meriania. A. Bullate adaxial leaf surface (M. sanguinea; R. Fernandez-Hilario et al. 1896). B. Punctiform abaxial leaf surface (M. cuzcoana; W. Farfán et al. 976). C. Glabrous abaxial leaf surface (M. callosa; R. Fernandez-Hilario et al. 2055). D. Furfuraceous abaxial leaf surface (M. vilcabambensis; L. Valenzuela 7494). E. Puberulent abaxial leaf surface (M. drakei; R. Fernandez-Hilario et al. 1775). F. Pubescent abaxial leaf surface (M. bicentenaria; A. Monteagudo et al. 6960). G. Tomentose abaxial leaf surface (M. tomentosa; R. Fernandez-Hilario et al. 1905). H. Setulose abaxial leaf surface (M. radula; D. Paredes et al. 604). I. Villose abaxial leaf surface (M. sumatika; I. Huamantupa et al. 2060). J. Hirsute adaxial leaf surface (M. hirsuta; S. Baldeón &amp; J. Campos 5373). K. Peltate leaf base (M. peltata; R. Fernandez-Hilario et al. 2093). L. Leaf base with dentate-undulate revolute auricles (M. cuzcoana; L. Valenzuela et al. 6904)." figureDoi="http://doi.org/10.5281/zenodo.8141996" httpUri="https://zenodo.org/record/8141996/files/figure.png" pageId="12" pageNumber="13">Fig. 4H</figureCitation>
); stamen connectives from both cycles with dorso-basal appendages not laterally expanded ..................................................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFAA2C28DFA6DFF7A" box="[1352,1450,187,209]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="radula">
<emphasis id="B965EADEFFB5FFBAFAA2C28DFA6DFF7A" bold="true" box="[1352,1450,187,209]" italics="true" pageId="12" pageNumber="13">M. radula</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C2E1FA6CFEA3" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C2E1FA6CFF47" blockId="12.[136,1452,159,2058]" box="[136,1451,215,237]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C2E1FA6CFF47" box="[136,1451,215,237]" pageId="12" pageNumber="13">
9. Antepetalous stamens with inflated (bulbous) connectives (
<figureCitation id="132A2A49FFB5FFBAFCE1C2E1FC90FF46" box="[779,855,215,237]" captionStart="FIGURE 6" captionStartId="10.[136,229,1633,1655]" captionTargetBox="[156,1429,267,1603]" captionTargetId="figure-20@10.[150,1435,193,1609]" captionTargetPageId="10" captionText="FIGURE 6. Morphological characteristics of Peruvian Meriania. A. Stamen with a triangular descending dorso-basal appendage (black arrow) but no dorsal appendage (M. tetraquetra; R. Fernandez-Hilario &amp; A. Vásquez 251). B. Stamen with a triangular descending dorso-basal appendage and a blunt ascending dorsal appendage (blue arrow) (M. megaphylla; A. Weberbauer 7048). C. Stamen with an acuminate descending dorso-basal appendage (black arrow) and a dorsal appendage as a mere hump (blue arrow) (M. franciscana; C. Díaz &amp; S. Fernández 10150). D. Stamen with a triangular descending dorso-basal appendage and a dorsal appendage as a mere hump, and detail of the dorsal appendage (M. rugosa; F. A. Michelangeli et al. 1725). E. Stamen with a triangular descending dorso-basal appendage and a dentiform dorsal appendage, and detail of the dorsal appendage (M. hexamera; I. Revilla 3484). F. Stamen with a multilobed descending dorso-basal appendage and an apically trilobed ascending dorsal appendage, and detail of the dorsal appendage apex (M. microflora; I. Huamantupa et al. 15552). G. Stamen with an almost perpendicular dorso-basal appendage to the theca, and detail of the prolonged connective below the theca (M. tetragona; R. Fernandez-Hilario et al. 2092). H. Stamen with inflated (bulbous) connective (blue arrow) (M. bicentenaria; R. Villanueva-Espinoza 675). I. Antesepalous stamen with a laterally expanded dorso-basal appendage (left) and antepetalous stamen with a dorso-basal appendage not laterally expanded (right) (M. vilcabambensis; L. Valenzuela 7494)." figureDoi="http://doi.org/10.5281/zenodo.8142006" httpUri="https://zenodo.org/record/8142006/files/figure.png" pageId="12" pageNumber="13">Fig. 6H</figureCitation>
)..............................................................................................10
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C2C2FA6CFEA3" blockId="12.[136,1452,159,2058]" box="[136,1451,243,265]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C2C2FA6CFEA3" box="[136,1451,243,265]" pageId="12" pageNumber="13">- Stamens from both cycles without inflated (bulbous) connectives ..................................................................................................13</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8BAE36CCFFB5FFBAFF62C339FA6CFE8F" blockId="12.[136,1452,159,2058]" box="[136,1451,271,293]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C339FA6CFE8F" box="[136,1451,271,293]" pageId="12" pageNumber="13">
10. Leaf blades subpeltate to peltate (
<figureCitation id="132A2A49FFB5FFBAFDFFC339FDA5FE8F" box="[533,610,271,293]" captionStart="FIGURE 4" captionStartId="8.[136,229,954,976]" captionTargetBox="[151,1433,194,929]" captionTargetId="figure-20@8.[150,1435,193,931]" captionTargetPageId="8" captionText="FIGURE 4. Morphological characteristics of Peruvian Meriania. A. Bullate adaxial leaf surface (M. sanguinea; R. Fernandez-Hilario et al. 1896). B. Punctiform abaxial leaf surface (M. cuzcoana; W. Farfán et al. 976). C. Glabrous abaxial leaf surface (M. callosa; R. Fernandez-Hilario et al. 2055). D. Furfuraceous abaxial leaf surface (M. vilcabambensis; L. Valenzuela 7494). E. Puberulent abaxial leaf surface (M. drakei; R. Fernandez-Hilario et al. 1775). F. Pubescent abaxial leaf surface (M. bicentenaria; A. Monteagudo et al. 6960). G. Tomentose abaxial leaf surface (M. tomentosa; R. Fernandez-Hilario et al. 1905). H. Setulose abaxial leaf surface (M. radula; D. Paredes et al. 604). I. Villose abaxial leaf surface (M. sumatika; I. Huamantupa et al. 2060). J. Hirsute adaxial leaf surface (M. hirsuta; S. Baldeón &amp; J. Campos 5373). K. Peltate leaf base (M. peltata; R. Fernandez-Hilario et al. 2093). L. Leaf base with dentate-undulate revolute auricles (M. cuzcoana; L. Valenzuela et al. 6904)." figureDoi="http://doi.org/10.5281/zenodo.8141996" httpUri="https://zenodo.org/record/8141996/files/figure.png" pageId="12" pageNumber="13">Fig. 4K</figureCitation>
), and more than
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wide .........................................................................................11
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</paragraph>
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- Leaf blades neither subpeltate nor peltate, less than
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wide....................................................................................................12
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11. Leaf blades 22.828.7 ×
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, peltate, abaxial surfaces pubescent to setulose ....................................................
<taxonomicName id="4C114D4FFFB5FFBAFAADC371FA6CFEF6" box="[1351,1451,327,349]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="peltata">
<emphasis id="B965EADEFFB5FFBAFAADC371FA6CFEF6" bold="true" box="[1351,1451,327,349]" italics="true" pageId="12" pageNumber="13">M. peltata</emphasis>
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</paragraph>
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- Leaf blades 17.227.7 ×
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, subpeltate, abaxial surfaces puberulent ......................................................
<taxonomicName id="4C114D4FFFB5FFBAFAEEC355FA6DFED2" box="[1284,1450,355,377]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="ninakurorum">
<emphasis id="B965EADEFFB5FFBAFAEEC355FA6DFED2" bold="true" box="[1284,1450,355,377]" italics="true" pageId="12" pageNumber="13">M. ninakurorum</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C349FA6DFE42" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C349FA6CFE1A" blockId="12.[136,1452,159,2058]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C349FA6CFE1A" pageId="12" pageNumber="13">
12. Leaf abaxial surface densely pubescent, the trichomes evenly covering the entire surface (
<figureCitation id="132A2A49FFB5FFBAFBDBC349FBBDFE3F" box="[1073,1146,383,405]" captionStart="FIGURE 4" captionStartId="8.[136,229,954,976]" captionTargetBox="[151,1433,194,929]" captionTargetId="figure-20@8.[150,1435,193,931]" captionTargetPageId="8" captionText="FIGURE 4. Morphological characteristics of Peruvian Meriania. A. Bullate adaxial leaf surface (M. sanguinea; R. Fernandez-Hilario et al. 1896). B. Punctiform abaxial leaf surface (M. cuzcoana; W. Farfán et al. 976). C. Glabrous abaxial leaf surface (M. callosa; R. Fernandez-Hilario et al. 2055). D. Furfuraceous abaxial leaf surface (M. vilcabambensis; L. Valenzuela 7494). E. Puberulent abaxial leaf surface (M. drakei; R. Fernandez-Hilario et al. 1775). F. Pubescent abaxial leaf surface (M. bicentenaria; A. Monteagudo et al. 6960). G. Tomentose abaxial leaf surface (M. tomentosa; R. Fernandez-Hilario et al. 1905). H. Setulose abaxial leaf surface (M. radula; D. Paredes et al. 604). I. Villose abaxial leaf surface (M. sumatika; I. Huamantupa et al. 2060). J. Hirsute adaxial leaf surface (M. hirsuta; S. Baldeón &amp; J. Campos 5373). K. Peltate leaf base (M. peltata; R. Fernandez-Hilario et al. 2093). L. Leaf base with dentate-undulate revolute auricles (M. cuzcoana; L. Valenzuela et al. 6904)." figureDoi="http://doi.org/10.5281/zenodo.8141996" httpUri="https://zenodo.org/record/8141996/files/figure.png" pageId="12" pageNumber="13">Fig. 4F</figureCitation>
); petals fuchsia to light fuchsia; antepetalous stamen connectives without dorsal appendages ...................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFAE7C3ADFA6CFE1A" box="[1293,1451,411,433]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="bicentenaria">
<emphasis id="B965EADEFFB5FFBAFAE7C3ADFA6CFE1A" bold="true" box="[1293,1451,411,433]" italics="true" pageId="12" pageNumber="13">M. bicentenaria</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C38EFA6DFE42" blockId="12.[136,1452,159,2058]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C38EFA6DFE42" pageId="12" pageNumber="13">
- Leaf abaxial surface sparsely to densely puberulent, the trichomes not covering the entire surface; petals reddish-purple; antepetalous stamen connectives with blunt ascending dorsal appendages .....................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFAFEC3E5FA6DFE42" box="[1300,1450,467,489]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="franciscana">
<emphasis id="B965EADEFFB5FFBAFAFEC3E5FA6DFE42" bold="true" box="[1300,1450,467,489]" italics="true" pageId="12" pageNumber="13">M. franciscana</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C3D9FA6CFDDE" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C3D9FA6CFD8B" blockId="12.[136,1452,159,2058]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C3D9FA6CFD8B" pageId="12" pageNumber="13">
13. Stamens strongly dimorphic (connectives with different shapes, sizes and colors), antepetalous stamen connectives with apically trilobed (
<figureCitation id="132A2A49FFB5FFBAFEC1C03DFEB5FD8A" box="[299,370,523,545]" captionStart="FIGURE 6" captionStartId="10.[136,229,1633,1655]" captionTargetBox="[156,1429,267,1603]" captionTargetId="figure-20@10.[150,1435,193,1609]" captionTargetPageId="10" captionText="FIGURE 6. Morphological characteristics of Peruvian Meriania. A. Stamen with a triangular descending dorso-basal appendage (black arrow) but no dorsal appendage (M. tetraquetra; R. Fernandez-Hilario &amp; A. Vásquez 251). B. Stamen with a triangular descending dorso-basal appendage and a blunt ascending dorsal appendage (blue arrow) (M. megaphylla; A. Weberbauer 7048). C. Stamen with an acuminate descending dorso-basal appendage (black arrow) and a dorsal appendage as a mere hump (blue arrow) (M. franciscana; C. Díaz &amp; S. Fernández 10150). D. Stamen with a triangular descending dorso-basal appendage and a dorsal appendage as a mere hump, and detail of the dorsal appendage (M. rugosa; F. A. Michelangeli et al. 1725). E. Stamen with a triangular descending dorso-basal appendage and a dentiform dorsal appendage, and detail of the dorsal appendage (M. hexamera; I. Revilla 3484). F. Stamen with a multilobed descending dorso-basal appendage and an apically trilobed ascending dorsal appendage, and detail of the dorsal appendage apex (M. microflora; I. Huamantupa et al. 15552). G. Stamen with an almost perpendicular dorso-basal appendage to the theca, and detail of the prolonged connective below the theca (M. tetragona; R. Fernandez-Hilario et al. 2092). H. Stamen with inflated (bulbous) connective (blue arrow) (M. bicentenaria; R. Villanueva-Espinoza 675). I. Antesepalous stamen with a laterally expanded dorso-basal appendage (left) and antepetalous stamen with a dorso-basal appendage not laterally expanded (right) (M. vilcabambensis; L. Valenzuela 7494)." figureDoi="http://doi.org/10.5281/zenodo.8142006" httpUri="https://zenodo.org/record/8142006/files/figure.png" pageId="12" pageNumber="13">Fig. 6F</figureCitation>
) or sagittate ascending dorsal appendages ............................................................................................................14
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C01EFA6CFDDE" blockId="12.[136,1452,159,2058]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C01EFA6CFDDE" pageId="12" pageNumber="13">
- Stamens dimorphic (connectives with different shapes and sizes but with similar colors) or isomorphic, stamen connectives from both cycles without dorsal appendages (
<figureCitation id="132A2A49FFB5FFBAFD66C075FD1DFDF2" box="[652,730,579,601]" captionStart="FIGURE 6" captionStartId="10.[136,229,1633,1655]" captionTargetBox="[156,1429,267,1603]" captionTargetId="figure-20@10.[150,1435,193,1609]" captionTargetPageId="10" captionText="FIGURE 6. Morphological characteristics of Peruvian Meriania. A. Stamen with a triangular descending dorso-basal appendage (black arrow) but no dorsal appendage (M. tetraquetra; R. Fernandez-Hilario &amp; A. Vásquez 251). B. Stamen with a triangular descending dorso-basal appendage and a blunt ascending dorsal appendage (blue arrow) (M. megaphylla; A. Weberbauer 7048). C. Stamen with an acuminate descending dorso-basal appendage (black arrow) and a dorsal appendage as a mere hump (blue arrow) (M. franciscana; C. Díaz &amp; S. Fernández 10150). D. Stamen with a triangular descending dorso-basal appendage and a dorsal appendage as a mere hump, and detail of the dorsal appendage (M. rugosa; F. A. Michelangeli et al. 1725). E. Stamen with a triangular descending dorso-basal appendage and a dentiform dorsal appendage, and detail of the dorsal appendage (M. hexamera; I. Revilla 3484). F. Stamen with a multilobed descending dorso-basal appendage and an apically trilobed ascending dorsal appendage, and detail of the dorsal appendage apex (M. microflora; I. Huamantupa et al. 15552). G. Stamen with an almost perpendicular dorso-basal appendage to the theca, and detail of the prolonged connective below the theca (M. tetragona; R. Fernandez-Hilario et al. 2092). H. Stamen with inflated (bulbous) connective (blue arrow) (M. bicentenaria; R. Villanueva-Espinoza 675). I. Antesepalous stamen with a laterally expanded dorso-basal appendage (left) and antepetalous stamen with a dorso-basal appendage not laterally expanded (right) (M. vilcabambensis; L. Valenzuela 7494)." figureDoi="http://doi.org/10.5281/zenodo.8142006" httpUri="https://zenodo.org/record/8142006/files/figure.png" pageId="12" pageNumber="13">Fig. 6A</figureCitation>
), with dentiform dorsal appendages (
<figureCitation id="132A2A49FFB5FFBAFBA9C075FB49FDF2" box="[1091,1166,579,601]" captionStart="FIGURE 6" captionStartId="10.[136,229,1633,1655]" captionTargetBox="[156,1429,267,1603]" captionTargetId="figure-20@10.[150,1435,193,1609]" captionTargetPageId="10" captionText="FIGURE 6. Morphological characteristics of Peruvian Meriania. A. Stamen with a triangular descending dorso-basal appendage (black arrow) but no dorsal appendage (M. tetraquetra; R. Fernandez-Hilario &amp; A. Vásquez 251). B. Stamen with a triangular descending dorso-basal appendage and a blunt ascending dorsal appendage (blue arrow) (M. megaphylla; A. Weberbauer 7048). C. Stamen with an acuminate descending dorso-basal appendage (black arrow) and a dorsal appendage as a mere hump (blue arrow) (M. franciscana; C. Díaz &amp; S. Fernández 10150). D. Stamen with a triangular descending dorso-basal appendage and a dorsal appendage as a mere hump, and detail of the dorsal appendage (M. rugosa; F. A. Michelangeli et al. 1725). E. Stamen with a triangular descending dorso-basal appendage and a dentiform dorsal appendage, and detail of the dorsal appendage (M. hexamera; I. Revilla 3484). F. Stamen with a multilobed descending dorso-basal appendage and an apically trilobed ascending dorsal appendage, and detail of the dorsal appendage apex (M. microflora; I. Huamantupa et al. 15552). G. Stamen with an almost perpendicular dorso-basal appendage to the theca, and detail of the prolonged connective below the theca (M. tetragona; R. Fernandez-Hilario et al. 2092). H. Stamen with inflated (bulbous) connective (blue arrow) (M. bicentenaria; R. Villanueva-Espinoza 675). I. Antesepalous stamen with a laterally expanded dorso-basal appendage (left) and antepetalous stamen with a dorso-basal appendage not laterally expanded (right) (M. vilcabambensis; L. Valenzuela 7494)." figureDoi="http://doi.org/10.5281/zenodo.8142006" httpUri="https://zenodo.org/record/8142006/files/figure.png" pageId="12" pageNumber="13">Fig. 6E</figureCitation>
), with blunt ascending dorsal appendages (
<figureCitation id="132A2A49FFB5FFBAFEBAC069FE5EFDDE" box="[336,409,607,629]" captionStart="FIGURE 6" captionStartId="10.[136,229,1633,1655]" captionTargetBox="[156,1429,267,1603]" captionTargetId="figure-20@10.[150,1435,193,1609]" captionTargetPageId="10" captionText="FIGURE 6. Morphological characteristics of Peruvian Meriania. A. Stamen with a triangular descending dorso-basal appendage (black arrow) but no dorsal appendage (M. tetraquetra; R. Fernandez-Hilario &amp; A. Vásquez 251). B. Stamen with a triangular descending dorso-basal appendage and a blunt ascending dorsal appendage (blue arrow) (M. megaphylla; A. Weberbauer 7048). C. Stamen with an acuminate descending dorso-basal appendage (black arrow) and a dorsal appendage as a mere hump (blue arrow) (M. franciscana; C. Díaz &amp; S. Fernández 10150). D. Stamen with a triangular descending dorso-basal appendage and a dorsal appendage as a mere hump, and detail of the dorsal appendage (M. rugosa; F. A. Michelangeli et al. 1725). E. Stamen with a triangular descending dorso-basal appendage and a dentiform dorsal appendage, and detail of the dorsal appendage (M. hexamera; I. Revilla 3484). F. Stamen with a multilobed descending dorso-basal appendage and an apically trilobed ascending dorsal appendage, and detail of the dorsal appendage apex (M. microflora; I. Huamantupa et al. 15552). G. Stamen with an almost perpendicular dorso-basal appendage to the theca, and detail of the prolonged connective below the theca (M. tetragona; R. Fernandez-Hilario et al. 2092). H. Stamen with inflated (bulbous) connective (blue arrow) (M. bicentenaria; R. Villanueva-Espinoza 675). I. Antesepalous stamen with a laterally expanded dorso-basal appendage (left) and antepetalous stamen with a dorso-basal appendage not laterally expanded (right) (M. vilcabambensis; L. Valenzuela 7494)." figureDoi="http://doi.org/10.5281/zenodo.8142006" httpUri="https://zenodo.org/record/8142006/files/figure.png" pageId="12" pageNumber="13">Fig. 6B</figureCitation>
) or dorsal appendages as mere humps (
<figureCitation id="132A2A49FFB5FFBAFCFEC069FC6DFDDE" box="[788,938,607,629]" captionStart="FIGURE 6" captionStartId="10.[136,229,1633,1655]" captionTargetBox="[156,1429,267,1603]" captionTargetId="figure-20@10.[150,1435,193,1609]" captionTargetPageId="10" captionText="FIGURE 6. Morphological characteristics of Peruvian Meriania. A. Stamen with a triangular descending dorso-basal appendage (black arrow) but no dorsal appendage (M. tetraquetra; R. Fernandez-Hilario &amp; A. Vásquez 251). B. Stamen with a triangular descending dorso-basal appendage and a blunt ascending dorsal appendage (blue arrow) (M. megaphylla; A. Weberbauer 7048). C. Stamen with an acuminate descending dorso-basal appendage (black arrow) and a dorsal appendage as a mere hump (blue arrow) (M. franciscana; C. Díaz &amp; S. Fernández 10150). D. Stamen with a triangular descending dorso-basal appendage and a dorsal appendage as a mere hump, and detail of the dorsal appendage (M. rugosa; F. A. Michelangeli et al. 1725). E. Stamen with a triangular descending dorso-basal appendage and a dentiform dorsal appendage, and detail of the dorsal appendage (M. hexamera; I. Revilla 3484). F. Stamen with a multilobed descending dorso-basal appendage and an apically trilobed ascending dorsal appendage, and detail of the dorsal appendage apex (M. microflora; I. Huamantupa et al. 15552). G. Stamen with an almost perpendicular dorso-basal appendage to the theca, and detail of the prolonged connective below the theca (M. tetragona; R. Fernandez-Hilario et al. 2092). H. Stamen with inflated (bulbous) connective (blue arrow) (M. bicentenaria; R. Villanueva-Espinoza 675). I. Antesepalous stamen with a laterally expanded dorso-basal appendage (left) and antepetalous stamen with a dorso-basal appendage not laterally expanded (right) (M. vilcabambensis; L. Valenzuela 7494)." figureDoi="http://doi.org/10.5281/zenodo.8142006" httpUri="https://zenodo.org/record/8142006/files/figure.png" pageId="12" pageNumber="13">Fig. 6C and 6D</figureCitation>
)................................................................................15
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<keyStep id="30E52E69FFB5FFBAFF62C04DFA6DFD4E" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C04DFA6DFD06" blockId="12.[136,1452,159,2058]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C04DFA6DFD06" pageId="12" pageNumber="13">
14. Petioles with adaxial and lateral liguliform projections on the apex; corolla white; antepetalous stamen connectives with apically trilobed ascending dorsal appendages (
<figureCitation id="132A2A49FFB5FFBAFDADC0A1FD48FD06" box="[583,655,663,685]" captionStart="FIGURE 6" captionStartId="10.[136,229,1633,1655]" captionTargetBox="[156,1429,267,1603]" captionTargetId="figure-20@10.[150,1435,193,1609]" captionTargetPageId="10" captionText="FIGURE 6. Morphological characteristics of Peruvian Meriania. A. Stamen with a triangular descending dorso-basal appendage (black arrow) but no dorsal appendage (M. tetraquetra; R. Fernandez-Hilario &amp; A. Vásquez 251). B. Stamen with a triangular descending dorso-basal appendage and a blunt ascending dorsal appendage (blue arrow) (M. megaphylla; A. Weberbauer 7048). C. Stamen with an acuminate descending dorso-basal appendage (black arrow) and a dorsal appendage as a mere hump (blue arrow) (M. franciscana; C. Díaz &amp; S. Fernández 10150). D. Stamen with a triangular descending dorso-basal appendage and a dorsal appendage as a mere hump, and detail of the dorsal appendage (M. rugosa; F. A. Michelangeli et al. 1725). E. Stamen with a triangular descending dorso-basal appendage and a dentiform dorsal appendage, and detail of the dorsal appendage (M. hexamera; I. Revilla 3484). F. Stamen with a multilobed descending dorso-basal appendage and an apically trilobed ascending dorsal appendage, and detail of the dorsal appendage apex (M. microflora; I. Huamantupa et al. 15552). G. Stamen with an almost perpendicular dorso-basal appendage to the theca, and detail of the prolonged connective below the theca (M. tetragona; R. Fernandez-Hilario et al. 2092). H. Stamen with inflated (bulbous) connective (blue arrow) (M. bicentenaria; R. Villanueva-Espinoza 675). I. Antesepalous stamen with a laterally expanded dorso-basal appendage (left) and antepetalous stamen with a dorso-basal appendage not laterally expanded (right) (M. vilcabambensis; L. Valenzuela 7494)." figureDoi="http://doi.org/10.5281/zenodo.8142006" httpUri="https://zenodo.org/record/8142006/files/figure.png" pageId="12" pageNumber="13">Fig. 6F</figureCitation>
) ............................................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFACAC0A1FA6DFD06" box="[1312,1450,663,685]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="microflora">
<emphasis id="B965EADEFFB5FFBAFACAC0A1FA6DFD06" bold="true" box="[1312,1450,663,685]" italics="true" pageId="12" pageNumber="13">M. microflora</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C082FA6DFD4E" blockId="12.[136,1452,159,2058]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C082FA6DFD4E" pageId="12" pageNumber="13">
- Petioles without projections; corolla fuchsia; antepetalous stamen connectives with apically sagittate ascending dorsal appendages ......................................................................................................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFAC6C0F9FA6DFD4E" box="[1324,1450,719,741]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="urceolata">
<emphasis id="B965EADEFFB5FFBAFAC6C0F9FA6DFD4E" bold="true" box="[1324,1450,719,741]" italics="true" pageId="12" pageNumber="13">M. urceolata</emphasis>
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<keyStep id="30E52E69FFB5FFBAFF62C0DDFA6CFCB7" pageId="12" pageNumber="13">
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<keyLead id="30E095F9FFB5FFBAFF62C0DDFA6CFCAA" box="[136,1451,747,769]" pageId="12" pageNumber="13">
15. Corolla campanulate, pink-orange to reddish-orange (except fuchsia in
<taxonomicName id="4C114D4FFFB5FFBAFC95C0DAFC3FFCAA" box="[895,1016,748,769]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="vasquezii">
<emphasis id="B965EADEFFB5FFBAFC95C0DAFC3FFCAA" box="[895,1016,748,769]" italics="true" pageId="12" pageNumber="13">M. vasquezii</emphasis>
</taxonomicName>
and red in
<taxonomicName id="4C114D4FFFB5FFBAFB8EC0DAFB23FCAB" box="[1124,1252,747,769]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="rubriflora">
<emphasis id="B965EADEFFB5FFBAFB8EC0DAFB23FCAB" box="[1124,1252,747,769]" italics="true" pageId="12" pageNumber="13">M. rubriflora</emphasis>
</taxonomicName>
) (
<figureCitation id="132A2A49FFB5FFBAFB10C0DDFAFAFCAB" box="[1274,1341,747,769]" captionStart="FIGURE 3" captionStartId="6.[136,229,1742,1764]" captionTargetBox="[217,1370,1054,1715]" captionTargetId="figure-317@6.[216,1372,1053,1717]" captionTargetPageId="6" captionText="FIGURE 3. Morphological characteristics of Peruvian Meriania. A. Interpetiolar line (M. tomentosa; R. W. Bussmann et al. 17068). B. Interpetiolar flap (M. zunacensis; R. Fernandez-Hilario et al. 1920). C. Winged internode (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Winged internode (M. tetragona; R. Fernandez-Hilario et al. 2092). E. Adaxial projection (scutum) in the transition zone from the petiole to the midvein (M. zunacensis; R. Fernandez-Hilario et al. 1920). F. Swollen adaxial projection on the petiole apex (M. drakei; R. Fernandez-Hilario et al. 1775). G. Liguliform abaxial projections in the transition zone from the petiole to the midvein (M. sanguinea; R. Fernandez-Hilario et al. 1896). H. Tuberculate abaxial projections in the transition zone from the petiole to the midvein (M. tetragona; F. A. Michelangeli et al. 1739). I. Spreading corolla (M. rigida; R. Fernandez-Hilario et al. 1931). J. Campanulate corollas (M. dazae; J. L. Marcelo-Peña et al. 6568). K. Fruits with mature ovaries exceeding the hypanthia length (M. sanguinea; F.A. Michelangeli et al. 2743). L. Fruits with mature ovaries completely concealed by the hypanthia (M. tomentosa; F. A. Michelangeli et al. 1799). Photos by Rainer W. Bussmann (A); Robin Fernandez-Hilario (BG and IJ); and Fabián A. Michelangeli (H, K and L)." figureDoi="http://doi.org/10.5281/zenodo.8141994" httpUri="https://zenodo.org/record/8141994/files/figure.png" pageId="12" pageNumber="13">Fig. 3J</figureCitation>
).............16
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</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C13EFA6CFCB7" blockId="12.[136,1452,159,2058]" box="[136,1451,775,797]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C13EFA6CFCB7" box="[136,1451,775,797]" pageId="12" pageNumber="13">
- Corolla spreading, fuchsia to reddish-purple (
<figureCitation id="132A2A49FFB5FFBAFD6DC131FD0EFCB7" box="[647,713,775,797]" captionStart="FIGURE 3" captionStartId="6.[136,229,1742,1764]" captionTargetBox="[217,1370,1054,1715]" captionTargetId="figure-317@6.[216,1372,1053,1717]" captionTargetPageId="6" captionText="FIGURE 3. Morphological characteristics of Peruvian Meriania. A. Interpetiolar line (M. tomentosa; R. W. Bussmann et al. 17068). B. Interpetiolar flap (M. zunacensis; R. Fernandez-Hilario et al. 1920). C. Winged internode (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Winged internode (M. tetragona; R. Fernandez-Hilario et al. 2092). E. Adaxial projection (scutum) in the transition zone from the petiole to the midvein (M. zunacensis; R. Fernandez-Hilario et al. 1920). F. Swollen adaxial projection on the petiole apex (M. drakei; R. Fernandez-Hilario et al. 1775). G. Liguliform abaxial projections in the transition zone from the petiole to the midvein (M. sanguinea; R. Fernandez-Hilario et al. 1896). H. Tuberculate abaxial projections in the transition zone from the petiole to the midvein (M. tetragona; F. A. Michelangeli et al. 1739). I. Spreading corolla (M. rigida; R. Fernandez-Hilario et al. 1931). J. Campanulate corollas (M. dazae; J. L. Marcelo-Peña et al. 6568). K. Fruits with mature ovaries exceeding the hypanthia length (M. sanguinea; F.A. Michelangeli et al. 2743). L. Fruits with mature ovaries completely concealed by the hypanthia (M. tomentosa; F. A. Michelangeli et al. 1799). Photos by Rainer W. Bussmann (A); Robin Fernandez-Hilario (BG and IJ); and Fabián A. Michelangeli (H, K and L)." figureDoi="http://doi.org/10.5281/zenodo.8141994" httpUri="https://zenodo.org/record/8141994/files/figure.png" pageId="12" pageNumber="13">Fig. 3I</figureCitation>
) .....................................................................................................................22
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<keyStep id="30E52E69FFB5FFBAFF62C115FA6CFCFF" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C115FA6CFC93" blockId="12.[136,1452,159,2058]" box="[136,1451,803,825]" pageId="12" pageNumber="13">
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16. Calyx calyptrate or subcalyptrate, with irregular dehiscence (
<figureCitation id="132A2A49FFB5FFBAFCF6C115FCA3FC92" box="[796,868,803,825]" captionStart="FIGURE 5" captionStartId="9.[136,229,1613,1635]" captionTargetBox="[155,1429,217,1583]" captionTargetId="figure-18@9.[150,1435,209,1589]" captionTargetPageId="9" captionText="FIGURE 5. Morphological characteristics of Peruvian Meriania. A. Calyx lacking dorsal projections (M. hexamera; I. Revilla 3484). B. Calyx with callose dorsal projections (left), and detail of a dorsal projection (right) (M. callosa; R. Fernandez-Hilario et al. 2055). C. Calyx with whitish callose dorsal projections (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Calyx with claw-shaped dorsal projections (M. dazae; J. L. Marcelo-Peña et al. 6568). E. Calyx with blunt dorsal projections (M. rugosa; H. van der Werff 17006). F. Subcalyptrate calyx with small claw-shaped projections and irregular dehiscence (left), and detail of the subcalyptrate calyx apex (right) (M. vasquezii; R. Vásquez et al. 45480). G. Calyptrate calyx with circumscissile dehiscence (M. escalerensis; M. Ríos et al. 3316). H. Calyx with aciculate dorsal projections and slightly costate hypanthium (M. prunifolia; J. Schunke-Vigo 11712). I. Costate hypanthium with 10 evident ribs (left) and apical view (right) (M. sumatika; P. Núñez &amp; J. Arque 8369)." figureDoi="http://doi.org/10.5281/zenodo.8142002" httpUri="https://zenodo.org/record/8142002/files/figure.png" pageId="12" pageNumber="13">Fig. 5F</figureCitation>
) ...........................................................................................17
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</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C176FA6CFCFF" blockId="12.[136,1452,159,2058]" box="[136,1451,831,853]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C176FA6CFCFF" box="[136,1451,831,853]" pageId="12" pageNumber="13">- Calyx neither calyptrate nor subcalyptrate, with regular dehiscence ...............................................................................................20</keyLead>
</paragraph>
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<keyStep id="30E52E69FFB5FFBAFF62C16DFA6CFC27" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C16DFA6CFCDB" blockId="12.[136,1452,159,2058]" box="[136,1451,859,881]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C16DFA6CFCDB" box="[136,1451,859,881]" pageId="12" pageNumber="13">17. Calyx calyptrate, without dorsal projections....................................................................................................................................18</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C14EFA6CFC27" blockId="12.[136,1452,159,2058]" box="[136,1451,887,909]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C14EFA6CFC27" box="[136,1451,887,909]" pageId="12" pageNumber="13">
- Calyx subcalyptrate, with dorsal projections
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long (
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) ...........................................................................................19
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<keyLead id="30E095F9FFB5FFBAFF62C1A5FA6DFC03" box="[136,1450,915,937]" pageId="12" pageNumber="13">
18. Inflorescences with flowers in regular dichasia (3-flowered) in the branchlet ends; petals
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long ...............
<taxonomicName id="4C114D4FFFB5FFBAFACEC1A5FA6DFC03" box="[1316,1450,915,936]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="B965EADEFFB5FFBAFACEC1A5FA6DFC03" bold="true" box="[1316,1450,915,936]" italics="true" pageId="12" pageNumber="13">M. tomentosa</emphasis>
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<keyLead id="30E095F9FFB5FFBAFF62C186FA6DFC6E" box="[136,1450,943,965]" pageId="12" pageNumber="13">
- Inflorescences with flowers in 56-flowered umbels in the branchlet ends; petals
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long......................................
<taxonomicName id="4C114D4FFFB5FFBAFABEC199FA6DFC6E" box="[1364,1450,943,965]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="acida">
<emphasis id="B965EADEFFB5FFBAFABEC199FA6DFC6E" bold="true" box="[1364,1450,943,965]" italics="true" pageId="12" pageNumber="13">M. acida</emphasis>
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<keyStep id="30E52E69FFB5FFBAFF62C1FDFA6CFBB3" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C1FDFA6DFC56" blockId="12.[136,1452,159,2058]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C1FDFA6DFC56" pageId="12" pageNumber="13">
19. Leaf blades 10.812.7 ×
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, abaxial surface sparsely to moderately puberulent, trichomes not covering the entire surface... .............................................................................................................................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFAA2C1D1FA6DFC56" box="[1352,1450,999,1021]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="juanjil">
<emphasis id="B965EADEFFB5FFBAFAA2C1D1FA6DFC56" bold="true" box="[1352,1450,999,1021]" italics="true" pageId="12" pageNumber="13">M. juanjil</emphasis>
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</paragraph>
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- Leaf blades 16.523.5 ×
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, abaxial surface densely villose, evenly covering the entire surface ..................
<taxonomicName id="4C114D4FFFB5FFBAFAC5C635FA6CFBB3" box="[1327,1451,1027,1048]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="vasquezii">
<emphasis id="B965EADEFFB5FFBAFAC5C635FA6CFBB3" bold="true" box="[1327,1451,1027,1048]" italics="true" pageId="12" pageNumber="13">M. vasquezii</emphasis>
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<keyStep id="30E52E69FFB5FFBAFF62C629FA6CFBFB" pageId="12" pageNumber="13">
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<keyLead id="30E095F9FFB5FFBAFF62C629FA6CFB9E" box="[136,1451,1055,1077]" pageId="12" pageNumber="13">
20. Nodes with robust interpetiolar flaps; inflorescences pendulous; calyces with obsolete dorsal projections .................
<taxonomicName id="4C114D4FFFB5FFBAFACDC629FA6CFB9E" box="[1319,1451,1055,1077]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="rubriflora">
<emphasis id="B965EADEFFB5FFBAFACDC629FA6CFB9E" bold="true" box="[1319,1451,1055,1077]" italics="true" pageId="12" pageNumber="13">M. rubriflora</emphasis>
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</paragraph>
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<keyLead id="30E095F9FFB5FFBAFF62C60AFA6CFBFB" box="[136,1451,1083,1105]" pageId="12" pageNumber="13">
- Nodes without interpetiolar flaps; inflorescences erect; calyces with claw-shaped dorsal projections (
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) ...........................21
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<keyLead id="30E095F9FFB5FFBAFF62C661FA6CFBC6" box="[136,1451,1111,1133]" pageId="12" pageNumber="13">
21. Abaxial leaf surface tomentose with whitish to cream trichomes when dry; petals
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long ................................
<taxonomicName id="4C114D4FFFB5FFBAFABBC661FA6CFBC6" box="[1361,1451,1111,1133]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="dazae">
<emphasis id="B965EADEFFB5FFBAFABBC661FA6CFBC6" bold="true" box="[1361,1451,1111,1133]" italics="true" pageId="12" pageNumber="13">M. dazae</emphasis>
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<keyLead id="30E095F9FFB5FFBAFF62C642FA6CFB22" box="[136,1451,1139,1161]" pageId="12" pageNumber="13">
- Abaxial leaf surface pubescent with ferrugineous trichomes when dry; petals
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long..............................
<taxonomicName id="4C114D4FFFB5FFBAFAF7C645FA6CFB22" box="[1309,1451,1139,1161]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="bongarana">
<emphasis id="B965EADEFFB5FFBAFAF7C645FA6CFB22" bold="true" box="[1309,1451,1139,1161]" italics="true" pageId="12" pageNumber="13">M. bongarana</emphasis>
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<keyLead id="30E095F9FFB5FFBAFF62C6B9FA6CFB0E" box="[136,1451,1167,1189]" pageId="12" pageNumber="13">
22. Calyx calyptrate, with circumscissile dehiscence (
<figureCitation id="132A2A49FFB5FFBAFD58C6B9FD3AFB0E" box="[690,765,1167,1189]" captionStart="FIGURE 5" captionStartId="9.[136,229,1613,1635]" captionTargetBox="[155,1429,217,1583]" captionTargetId="figure-18@9.[150,1435,209,1589]" captionTargetPageId="9" captionText="FIGURE 5. Morphological characteristics of Peruvian Meriania. A. Calyx lacking dorsal projections (M. hexamera; I. Revilla 3484). B. Calyx with callose dorsal projections (left), and detail of a dorsal projection (right) (M. callosa; R. Fernandez-Hilario et al. 2055). C. Calyx with whitish callose dorsal projections (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Calyx with claw-shaped dorsal projections (M. dazae; J. L. Marcelo-Peña et al. 6568). E. Calyx with blunt dorsal projections (M. rugosa; H. van der Werff 17006). F. Subcalyptrate calyx with small claw-shaped projections and irregular dehiscence (left), and detail of the subcalyptrate calyx apex (right) (M. vasquezii; R. Vásquez et al. 45480). G. Calyptrate calyx with circumscissile dehiscence (M. escalerensis; M. Ríos et al. 3316). H. Calyx with aciculate dorsal projections and slightly costate hypanthium (M. prunifolia; J. Schunke-Vigo 11712). I. Costate hypanthium with 10 evident ribs (left) and apical view (right) (M. sumatika; P. Núñez &amp; J. Arque 8369)." figureDoi="http://doi.org/10.5281/zenodo.8142002" httpUri="https://zenodo.org/record/8142002/files/figure.png" pageId="12" pageNumber="13">Fig. 5G</figureCitation>
).......................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFAF9C6B9FA6CFB0E" box="[1299,1451,1167,1189]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="escalerensis">
<emphasis id="B965EADEFFB5FFBAFAF9C6B9FA6CFB0E" bold="true" box="[1299,1451,1167,1189]" italics="true" pageId="12" pageNumber="13">M. escalerensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C69AFA6CFB6B" blockId="12.[136,1452,159,2058]" box="[136,1451,1195,1217]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C69AFA6CFB6B" box="[136,1451,1195,1217]" pageId="12" pageNumber="13">- Calyx not calyptrate, with regular dehiscence..................................................................................................................................23</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C6F1FA6CFB53" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C6F1FA6CFB77" blockId="12.[136,1452,159,2058]" box="[136,1451,1223,1245]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C6F1FA6CFB77" box="[136,1451,1223,1245]" pageId="12" pageNumber="13">
23. Stamens dimorphic; antesepalous stamen connectives with descending dorso-basal appendages laterally expanded (
<figureCitation id="132A2A49FFB5FFBAFACCC6F1FAAFFB76" box="[1318,1384,1223,1245]" captionStart="FIGURE 6" captionStartId="10.[136,229,1633,1655]" captionTargetBox="[156,1429,267,1603]" captionTargetId="figure-20@10.[150,1435,193,1609]" captionTargetPageId="10" captionText="FIGURE 6. Morphological characteristics of Peruvian Meriania. A. Stamen with a triangular descending dorso-basal appendage (black arrow) but no dorsal appendage (M. tetraquetra; R. Fernandez-Hilario &amp; A. Vásquez 251). B. Stamen with a triangular descending dorso-basal appendage and a blunt ascending dorsal appendage (blue arrow) (M. megaphylla; A. Weberbauer 7048). C. Stamen with an acuminate descending dorso-basal appendage (black arrow) and a dorsal appendage as a mere hump (blue arrow) (M. franciscana; C. Díaz &amp; S. Fernández 10150). D. Stamen with a triangular descending dorso-basal appendage and a dorsal appendage as a mere hump, and detail of the dorsal appendage (M. rugosa; F. A. Michelangeli et al. 1725). E. Stamen with a triangular descending dorso-basal appendage and a dentiform dorsal appendage, and detail of the dorsal appendage (M. hexamera; I. Revilla 3484). F. Stamen with a multilobed descending dorso-basal appendage and an apically trilobed ascending dorsal appendage, and detail of the dorsal appendage apex (M. microflora; I. Huamantupa et al. 15552). G. Stamen with an almost perpendicular dorso-basal appendage to the theca, and detail of the prolonged connective below the theca (M. tetragona; R. Fernandez-Hilario et al. 2092). H. Stamen with inflated (bulbous) connective (blue arrow) (M. bicentenaria; R. Villanueva-Espinoza 675). I. Antesepalous stamen with a laterally expanded dorso-basal appendage (left) and antepetalous stamen with a dorso-basal appendage not laterally expanded (right) (M. vilcabambensis; L. Valenzuela 7494)." figureDoi="http://doi.org/10.5281/zenodo.8142006" httpUri="https://zenodo.org/record/8142006/files/figure.png" pageId="12" pageNumber="13">Fig. 6I</figureCitation>
) .....24
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C6D2FA6CFB53" blockId="12.[136,1452,159,2058]" box="[136,1451,1251,1273]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C6D2FA6CFB53" box="[136,1451,1251,1273]" pageId="12" pageNumber="13">- Stamens isomorphic; stamen connectives from both cycles with descending dorso-basal appendages not laterally expanded......28</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C6C9FA6CFA9A" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C6C9FA6CFABE" blockId="12.[136,1452,159,2058]" box="[136,1451,1279,1301]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C6C9FA6CFABE" box="[136,1451,1279,1301]" pageId="12" pageNumber="13">
24. Leaves subsessile (petioles up to
<quantity id="4CE99B29FFB5FFBAFDF8C6C9FD8EFABE" box="[530,585,1279,1301]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="13" unit="mm" value="3.0">3 mm</quantity>
long), base auriculate ..............................................................................
<taxonomicName id="4C114D4FFFB5FFBAFB13C6C9FA6CFABE" box="[1273,1451,1279,1301]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="amischophylla">
<emphasis id="B965EADEFFB5FFBAFB13C6C9FA6CFABE" bold="true" box="[1273,1451,1279,1301]" italics="true" pageId="12" pageNumber="13">M. amischophylla</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C72AFA6CFA9A" blockId="12.[136,1452,159,2058]" box="[136,1451,1307,1329]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C72AFA6CFA9A" box="[136,1451,1307,1329]" pageId="12" pageNumber="13">
- Leaves with petioles longer than
<quantity id="4CE99B29FFB5FFBAFDFBC72DFD8CFA9A" box="[529,587,1307,1329]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="13" unit="mm" value="8.0">8 mm</quantity>
, base acute, attenuate, or obtuse (rarely broadly obtuse in
<taxonomicName id="4C114D4FFFB5FFBAFB9DC72AFAD7FA9B" box="[1143,1296,1307,1329]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B965EADEFFB5FFBAFB9DC72AFAD7FA9B" box="[1143,1296,1307,1329]" italics="true" pageId="12" pageNumber="13">M. weberbaueri</emphasis>
</taxonomicName>
)....................25
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C701FA6CFAC2" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C701FA6CFAE6" blockId="12.[136,1452,159,2058]" box="[136,1451,1335,1357]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C701FA6CFAE6" box="[136,1451,1335,1357]" pageId="12" pageNumber="13">
25. Stamen connectives without dorsal appendages.........................................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFAE5C701FA6CFAE6" box="[1295,1451,1335,1357]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B965EADEFFB5FFBAFAE5C701FA6CFAE6" bold="true" box="[1295,1451,1335,1357]" italics="true" pageId="12" pageNumber="13">M. weberbaueri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C762FA6CFAC2" blockId="12.[136,1452,159,2058]" box="[136,1451,1363,1385]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C762FA6CFAC2" box="[136,1451,1363,1385]" pageId="12" pageNumber="13">
- Stamen connectives with blunt ascending dorsal appendages (
<figureCitation id="132A2A49FFB5FFBAFCCEC765FCAAFAC2" box="[804,877,1363,1385]" captionStart="FIGURE 6" captionStartId="10.[136,229,1633,1655]" captionTargetBox="[156,1429,267,1603]" captionTargetId="figure-20@10.[150,1435,193,1609]" captionTargetPageId="10" captionText="FIGURE 6. Morphological characteristics of Peruvian Meriania. A. Stamen with a triangular descending dorso-basal appendage (black arrow) but no dorsal appendage (M. tetraquetra; R. Fernandez-Hilario &amp; A. Vásquez 251). B. Stamen with a triangular descending dorso-basal appendage and a blunt ascending dorsal appendage (blue arrow) (M. megaphylla; A. Weberbauer 7048). C. Stamen with an acuminate descending dorso-basal appendage (black arrow) and a dorsal appendage as a mere hump (blue arrow) (M. franciscana; C. Díaz &amp; S. Fernández 10150). D. Stamen with a triangular descending dorso-basal appendage and a dorsal appendage as a mere hump, and detail of the dorsal appendage (M. rugosa; F. A. Michelangeli et al. 1725). E. Stamen with a triangular descending dorso-basal appendage and a dentiform dorsal appendage, and detail of the dorsal appendage (M. hexamera; I. Revilla 3484). F. Stamen with a multilobed descending dorso-basal appendage and an apically trilobed ascending dorsal appendage, and detail of the dorsal appendage apex (M. microflora; I. Huamantupa et al. 15552). G. Stamen with an almost perpendicular dorso-basal appendage to the theca, and detail of the prolonged connective below the theca (M. tetragona; R. Fernandez-Hilario et al. 2092). H. Stamen with inflated (bulbous) connective (blue arrow) (M. bicentenaria; R. Villanueva-Espinoza 675). I. Antesepalous stamen with a laterally expanded dorso-basal appendage (left) and antepetalous stamen with a dorso-basal appendage not laterally expanded (right) (M. vilcabambensis; L. Valenzuela 7494)." figureDoi="http://doi.org/10.5281/zenodo.8142006" httpUri="https://zenodo.org/record/8142006/files/figure.png" pageId="12" pageNumber="13">Fig. 6B</figureCitation>
)..........................................................................................26
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C759FA6CFA0B" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C759FA6CFA2E" blockId="12.[136,1452,159,2058]" box="[136,1451,1391,1413]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C759FA6CFA2E" box="[136,1451,1391,1413]" pageId="12" pageNumber="13">
26. Hypanthium 10-costate, with longitudinal ridges up to
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high (
<figureCitation id="132A2A49FFB5FFBAFC83C759FC6DFA2E" box="[873,938,1391,1413]" captionStart="FIGURE 5" captionStartId="9.[136,229,1613,1635]" captionTargetBox="[155,1429,217,1583]" captionTargetId="figure-18@9.[150,1435,209,1589]" captionTargetPageId="9" captionText="FIGURE 5. Morphological characteristics of Peruvian Meriania. A. Calyx lacking dorsal projections (M. hexamera; I. Revilla 3484). B. Calyx with callose dorsal projections (left), and detail of a dorsal projection (right) (M. callosa; R. Fernandez-Hilario et al. 2055). C. Calyx with whitish callose dorsal projections (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Calyx with claw-shaped dorsal projections (M. dazae; J. L. Marcelo-Peña et al. 6568). E. Calyx with blunt dorsal projections (M. rugosa; H. van der Werff 17006). F. Subcalyptrate calyx with small claw-shaped projections and irregular dehiscence (left), and detail of the subcalyptrate calyx apex (right) (M. vasquezii; R. Vásquez et al. 45480). G. Calyptrate calyx with circumscissile dehiscence (M. escalerensis; M. Ríos et al. 3316). H. Calyx with aciculate dorsal projections and slightly costate hypanthium (M. prunifolia; J. Schunke-Vigo 11712). I. Costate hypanthium with 10 evident ribs (left) and apical view (right) (M. sumatika; P. Núñez &amp; J. Arque 8369)." figureDoi="http://doi.org/10.5281/zenodo.8142002" httpUri="https://zenodo.org/record/8142002/files/figure.png" pageId="12" pageNumber="13">Fig. 5I</figureCitation>
)...............................................................
<taxonomicName id="4C114D4FFFB5FFBAFAC5C759FA6CFA2E" box="[1327,1451,1391,1413]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="sumatika">
<emphasis id="B965EADEFFB5FFBAFAC5C759FA6CFA2E" bold="true" box="[1327,1451,1391,1413]" italics="true" pageId="12" pageNumber="13">M. sumatika</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C7BAFA6CFA0B" blockId="12.[136,1452,159,2058]" box="[136,1451,1419,1441]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C7BAFA6CFA0B" box="[136,1451,1419,1441]" pageId="12" pageNumber="13">- Hypanthium terete, without ridges ...................................................................................................................................................27</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C791FA6CFA5E" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C791FA6CFA17" blockId="12.[136,1452,159,2058]" box="[136,1451,1447,1469]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C791FA6CFA17" box="[136,1451,1447,1469]" pageId="12" pageNumber="13">
27. Inflorescences, hypanthia, and calyces densely tomentose, the trichomes up to
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long.........................................
<taxonomicName id="4C114D4FFFB5FFBAFAD1C791FA6CFA17" box="[1339,1451,1447,1468]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="vargasii">
<emphasis id="B965EADEFFB5FFBAFAD1C791FA6CFA17" bold="true" box="[1339,1451,1447,1468]" italics="true" pageId="12" pageNumber="13">M. vargasii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C7F2FA6CFA5E" blockId="12.[136,1452,159,2058]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C7F2FA6CFA5E" pageId="12" pageNumber="13">
- Inflorescences, hypanthia, and calyces sparsely to densely furfuraceous, the trichomes up to
<quantity id="4CE99B29FFB5FFBAFBA0C7F5FB58FA72" box="[1098,1183,1475,1497]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="13" unit="mm" value="0.25">0.25 mm</quantity>
long (
<figureCitation id="132A2A49FFB5FFBAFB37C7F5FAEFFA73" box="[1245,1320,1475,1497]" captionStart="FIGURE 4" captionStartId="8.[136,229,954,976]" captionTargetBox="[151,1433,194,929]" captionTargetId="figure-20@8.[150,1435,193,931]" captionTargetPageId="8" captionText="FIGURE 4. Morphological characteristics of Peruvian Meriania. A. Bullate adaxial leaf surface (M. sanguinea; R. Fernandez-Hilario et al. 1896). B. Punctiform abaxial leaf surface (M. cuzcoana; W. Farfán et al. 976). C. Glabrous abaxial leaf surface (M. callosa; R. Fernandez-Hilario et al. 2055). D. Furfuraceous abaxial leaf surface (M. vilcabambensis; L. Valenzuela 7494). E. Puberulent abaxial leaf surface (M. drakei; R. Fernandez-Hilario et al. 1775). F. Pubescent abaxial leaf surface (M. bicentenaria; A. Monteagudo et al. 6960). G. Tomentose abaxial leaf surface (M. tomentosa; R. Fernandez-Hilario et al. 1905). H. Setulose abaxial leaf surface (M. radula; D. Paredes et al. 604). I. Villose abaxial leaf surface (M. sumatika; I. Huamantupa et al. 2060). J. Hirsute adaxial leaf surface (M. hirsuta; S. Baldeón &amp; J. Campos 5373). K. Peltate leaf base (M. peltata; R. Fernandez-Hilario et al. 2093). L. Leaf base with dentate-undulate revolute auricles (M. cuzcoana; L. Valenzuela et al. 6904)." figureDoi="http://doi.org/10.5281/zenodo.8141996" httpUri="https://zenodo.org/record/8141996/files/figure.png" pageId="12" pageNumber="13">Fig. 4D</figureCitation>
) .................... ...............................................................................................................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFB19C7E9FA6CFA5E" box="[1267,1451,1503,1525]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="vilcabambensis">
<emphasis id="B965EADEFFB5FFBAFB19C7E9FA6CFA5E" bold="true" box="[1267,1451,1503,1525]" italics="true" pageId="12" pageNumber="13">M. vilcabambensis</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C7CDFA6CF987" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C7CDFA6CF9BB" blockId="12.[136,1452,159,2058]" box="[136,1451,1531,1553]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C7CDFA6CF9BB" box="[136,1451,1531,1553]" pageId="12" pageNumber="13">
28. Nodes with robust interpetiolar flaps (sometimes small in
<taxonomicName id="4C114D4FFFB5FFBAFCEAC7CAFCA2F9BB" box="[768,869,1531,1553]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="callosa">
<emphasis id="B965EADEFFB5FFBAFCEAC7CAFCA2F9BB" box="[768,869,1531,1553]" italics="true" pageId="12" pageNumber="13">M. callosa</emphasis>
</taxonomicName>
) (
<figureCitation id="132A2A49FFB5FFBAFC90C7CDFC03F9BB" box="[890,964,1531,1553]" captionStart="FIGURE 3" captionStartId="6.[136,229,1742,1764]" captionTargetBox="[217,1370,1054,1715]" captionTargetId="figure-317@6.[216,1372,1053,1717]" captionTargetPageId="6" captionText="FIGURE 3. Morphological characteristics of Peruvian Meriania. A. Interpetiolar line (M. tomentosa; R. W. Bussmann et al. 17068). B. Interpetiolar flap (M. zunacensis; R. Fernandez-Hilario et al. 1920). C. Winged internode (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Winged internode (M. tetragona; R. Fernandez-Hilario et al. 2092). E. Adaxial projection (scutum) in the transition zone from the petiole to the midvein (M. zunacensis; R. Fernandez-Hilario et al. 1920). F. Swollen adaxial projection on the petiole apex (M. drakei; R. Fernandez-Hilario et al. 1775). G. Liguliform abaxial projections in the transition zone from the petiole to the midvein (M. sanguinea; R. Fernandez-Hilario et al. 1896). H. Tuberculate abaxial projections in the transition zone from the petiole to the midvein (M. tetragona; F. A. Michelangeli et al. 1739). I. Spreading corolla (M. rigida; R. Fernandez-Hilario et al. 1931). J. Campanulate corollas (M. dazae; J. L. Marcelo-Peña et al. 6568). K. Fruits with mature ovaries exceeding the hypanthia length (M. sanguinea; F.A. Michelangeli et al. 2743). L. Fruits with mature ovaries completely concealed by the hypanthia (M. tomentosa; F. A. Michelangeli et al. 1799). Photos by Rainer W. Bussmann (A); Robin Fernandez-Hilario (BG and IJ); and Fabián A. Michelangeli (H, K and L)." figureDoi="http://doi.org/10.5281/zenodo.8141994" httpUri="https://zenodo.org/record/8141994/files/figure.png" pageId="12" pageNumber="13">Fig. 3B</figureCitation>
) ...........................................................................29
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C42EFA6CF987" blockId="12.[136,1452,159,2058]" box="[136,1451,1559,1581]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C42EFA6CF987" box="[136,1451,1559,1581]" pageId="12" pageNumber="13">- Nodes without interpetiolar flaps .....................................................................................................................................................31</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C405FA6CF9CF" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C405FA6CF9E2" blockId="12.[136,1452,159,2058]" box="[136,1451,1587,1609]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C405FA6CF9E2" box="[136,1451,1587,1609]" pageId="12" pageNumber="13">
29. Internodes quadrangular and 4-winged; stamen connectives with blunt ascending dorsal appendages (
<figureCitation id="132A2A49FFB5FFBAFB4AC405FB2EF9E2" box="[1184,1257,1587,1609]" captionStart="FIGURE 6" captionStartId="10.[136,229,1633,1655]" captionTargetBox="[156,1429,267,1603]" captionTargetId="figure-20@10.[150,1435,193,1609]" captionTargetPageId="10" captionText="FIGURE 6. Morphological characteristics of Peruvian Meriania. A. Stamen with a triangular descending dorso-basal appendage (black arrow) but no dorsal appendage (M. tetraquetra; R. Fernandez-Hilario &amp; A. Vásquez 251). B. Stamen with a triangular descending dorso-basal appendage and a blunt ascending dorsal appendage (blue arrow) (M. megaphylla; A. Weberbauer 7048). C. Stamen with an acuminate descending dorso-basal appendage (black arrow) and a dorsal appendage as a mere hump (blue arrow) (M. franciscana; C. Díaz &amp; S. Fernández 10150). D. Stamen with a triangular descending dorso-basal appendage and a dorsal appendage as a mere hump, and detail of the dorsal appendage (M. rugosa; F. A. Michelangeli et al. 1725). E. Stamen with a triangular descending dorso-basal appendage and a dentiform dorsal appendage, and detail of the dorsal appendage (M. hexamera; I. Revilla 3484). F. Stamen with a multilobed descending dorso-basal appendage and an apically trilobed ascending dorsal appendage, and detail of the dorsal appendage apex (M. microflora; I. Huamantupa et al. 15552). G. Stamen with an almost perpendicular dorso-basal appendage to the theca, and detail of the prolonged connective below the theca (M. tetragona; R. Fernandez-Hilario et al. 2092). H. Stamen with inflated (bulbous) connective (blue arrow) (M. bicentenaria; R. Villanueva-Espinoza 675). I. Antesepalous stamen with a laterally expanded dorso-basal appendage (left) and antepetalous stamen with a dorso-basal appendage not laterally expanded (right) (M. vilcabambensis; L. Valenzuela 7494)." figureDoi="http://doi.org/10.5281/zenodo.8142006" httpUri="https://zenodo.org/record/8142006/files/figure.png" pageId="12" pageNumber="13">Fig. 6B</figureCitation>
) .....
<taxonomicName id="4C114D4FFFB5FFBAFAFDC405FA6CF9E2" box="[1303,1451,1587,1609]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="megaphylla">
<emphasis id="B965EADEFFB5FFBAFAFDC405FA6CF9E2" bold="true" box="[1303,1451,1587,1609]" italics="true" pageId="12" pageNumber="13">M. megaphylla</emphasis>
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</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C466FA6CF9CF" blockId="12.[136,1452,159,2058]" box="[136,1451,1615,1637]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C466FA6CF9CF" box="[136,1451,1615,1637]" pageId="12" pageNumber="13">- Internodes quadrangular, but not winged; stamen connectives with dorsal appendages as mere humps or obsolete......................30</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C45DFA6DF912" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C45DFA6CF937" blockId="12.[136,1452,159,2058]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C45DFA6CF937" pageId="12" pageNumber="13">
30. Petioles
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long, with an adaxial projection (scutum) on the transition zone from the petiole to the midvein (
<figureCitation id="132A2A49FFB5FFBAFADAC45DFABFF92B" box="[1328,1400,1643,1665]" captionStart="FIGURE 3" captionStartId="6.[136,229,1742,1764]" captionTargetBox="[217,1370,1054,1715]" captionTargetId="figure-317@6.[216,1372,1053,1717]" captionTargetPageId="6" captionText="FIGURE 3. Morphological characteristics of Peruvian Meriania. A. Interpetiolar line (M. tomentosa; R. W. Bussmann et al. 17068). B. Interpetiolar flap (M. zunacensis; R. Fernandez-Hilario et al. 1920). C. Winged internode (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Winged internode (M. tetragona; R. Fernandez-Hilario et al. 2092). E. Adaxial projection (scutum) in the transition zone from the petiole to the midvein (M. zunacensis; R. Fernandez-Hilario et al. 1920). F. Swollen adaxial projection on the petiole apex (M. drakei; R. Fernandez-Hilario et al. 1775). G. Liguliform abaxial projections in the transition zone from the petiole to the midvein (M. sanguinea; R. Fernandez-Hilario et al. 1896). H. Tuberculate abaxial projections in the transition zone from the petiole to the midvein (M. tetragona; F. A. Michelangeli et al. 1739). I. Spreading corolla (M. rigida; R. Fernandez-Hilario et al. 1931). J. Campanulate corollas (M. dazae; J. L. Marcelo-Peña et al. 6568). K. Fruits with mature ovaries exceeding the hypanthia length (M. sanguinea; F.A. Michelangeli et al. 2743). L. Fruits with mature ovaries completely concealed by the hypanthia (M. tomentosa; F. A. Michelangeli et al. 1799). Photos by Rainer W. Bussmann (A); Robin Fernandez-Hilario (BG and IJ); and Fabián A. Michelangeli (H, K and L)." figureDoi="http://doi.org/10.5281/zenodo.8141994" httpUri="https://zenodo.org/record/8141994/files/figure.png" pageId="12" pageNumber="13">Fig. 3E</figureCitation>
)....... ......................................................................................................................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFAF7C4B1FA6CF937" box="[1309,1451,1671,1692]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="zunacensis">
<emphasis id="B965EADEFFB5FFBAFAF7C4B1FA6CF937" bold="true" box="[1309,1451,1671,1692]" italics="true" pageId="12" pageNumber="13">M. zunacensis</emphasis>
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</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C492FA6DF912" blockId="12.[136,1452,159,2058]" box="[136,1450,1699,1721]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C492FA6DF912" box="[136,1450,1699,1721]" pageId="12" pageNumber="13">
- Petioles
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long, without projections .....................................................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFAAEC495FA6DF912" box="[1348,1450,1699,1721]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="callosa">
<emphasis id="B965EADEFFB5FFBAFAAEC495FA6DF912" bold="true" box="[1348,1450,1699,1721]" italics="true" pageId="12" pageNumber="13">M. callosa</emphasis>
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</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C489FA6CF95B" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C489FA6CF97E" blockId="12.[136,1452,159,2058]" box="[136,1451,1727,1749]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C489FA6CF97E" box="[136,1451,1727,1749]" pageId="12" pageNumber="13">
31. Petioles with an apical adaxial projection (
<figureCitation id="132A2A49FFB5FFBAFD81C489FD73F97F" box="[619,692,1727,1749]" captionStart="FIGURE 3" captionStartId="6.[136,229,1742,1764]" captionTargetBox="[217,1370,1054,1715]" captionTargetId="figure-317@6.[216,1372,1053,1717]" captionTargetPageId="6" captionText="FIGURE 3. Morphological characteristics of Peruvian Meriania. A. Interpetiolar line (M. tomentosa; R. W. Bussmann et al. 17068). B. Interpetiolar flap (M. zunacensis; R. Fernandez-Hilario et al. 1920). C. Winged internode (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Winged internode (M. tetragona; R. Fernandez-Hilario et al. 2092). E. Adaxial projection (scutum) in the transition zone from the petiole to the midvein (M. zunacensis; R. Fernandez-Hilario et al. 1920). F. Swollen adaxial projection on the petiole apex (M. drakei; R. Fernandez-Hilario et al. 1775). G. Liguliform abaxial projections in the transition zone from the petiole to the midvein (M. sanguinea; R. Fernandez-Hilario et al. 1896). H. Tuberculate abaxial projections in the transition zone from the petiole to the midvein (M. tetragona; F. A. Michelangeli et al. 1739). I. Spreading corolla (M. rigida; R. Fernandez-Hilario et al. 1931). J. Campanulate corollas (M. dazae; J. L. Marcelo-Peña et al. 6568). K. Fruits with mature ovaries exceeding the hypanthia length (M. sanguinea; F.A. Michelangeli et al. 2743). L. Fruits with mature ovaries completely concealed by the hypanthia (M. tomentosa; F. A. Michelangeli et al. 1799). Photos by Rainer W. Bussmann (A); Robin Fernandez-Hilario (BG and IJ); and Fabián A. Michelangeli (H, K and L)." figureDoi="http://doi.org/10.5281/zenodo.8141994" httpUri="https://zenodo.org/record/8141994/files/figure.png" pageId="12" pageNumber="13">Fig. 3F</figureCitation>
); petals abaxially moderately to densely puberulent ...........................
<taxonomicName id="4C114D4FFFB5FFBAFAA1C489FA6CF97E" box="[1355,1451,1727,1749]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="drakei">
<emphasis id="B965EADEFFB5FFBAFAA1C489FA6CF97E" bold="true" box="[1355,1451,1727,1749]" italics="true" pageId="12" pageNumber="13">M. drakei</emphasis>
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</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C4EAFA6CF95B" blockId="12.[136,1452,159,2058]" box="[136,1451,1755,1777]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C4EAFA6CF95B" box="[136,1451,1755,1777]" pageId="12" pageNumber="13">- Petioles without projections; petals glabrous ...................................................................................................................................32</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C4C1FA6CF883" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C4C1FA6CF8A7" blockId="12.[136,1452,159,2058]" box="[136,1451,1783,1805]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C4C1FA6CF8A7" box="[136,1451,1783,1805]" pageId="12" pageNumber="13">32. Leaf blades with entire margins, and the adaxial surfaces flat.........................................................................................................33</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C522FA6CF883" blockId="12.[136,1452,159,2058]" box="[136,1451,1811,1833]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C522FA6CF883" box="[136,1451,1811,1833]" pageId="12" pageNumber="13">- Leaf blades with denticulate margins, and the adaxial surfaces flat to bullate ................................................................................34</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C519FA6CF8CA" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C519FA6DF8EE" blockId="12.[136,1452,159,2058]" box="[136,1450,1839,1861]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C519FA6DF8EE" box="[136,1450,1839,1861]" pageId="12" pageNumber="13">
33. Flowers 6-merous; calyx without dorsal projections (
<figureCitation id="132A2A49FFB5FFBAFD25C519FCDDF8EE" box="[719,794,1839,1861]" captionStart="FIGURE 5" captionStartId="9.[136,229,1613,1635]" captionTargetBox="[155,1429,217,1583]" captionTargetId="figure-18@9.[150,1435,209,1589]" captionTargetPageId="9" captionText="FIGURE 5. Morphological characteristics of Peruvian Meriania. A. Calyx lacking dorsal projections (M. hexamera; I. Revilla 3484). B. Calyx with callose dorsal projections (left), and detail of a dorsal projection (right) (M. callosa; R. Fernandez-Hilario et al. 2055). C. Calyx with whitish callose dorsal projections (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Calyx with claw-shaped dorsal projections (M. dazae; J. L. Marcelo-Peña et al. 6568). E. Calyx with blunt dorsal projections (M. rugosa; H. van der Werff 17006). F. Subcalyptrate calyx with small claw-shaped projections and irregular dehiscence (left), and detail of the subcalyptrate calyx apex (right) (M. vasquezii; R. Vásquez et al. 45480). G. Calyptrate calyx with circumscissile dehiscence (M. escalerensis; M. Ríos et al. 3316). H. Calyx with aciculate dorsal projections and slightly costate hypanthium (M. prunifolia; J. Schunke-Vigo 11712). I. Costate hypanthium with 10 evident ribs (left) and apical view (right) (M. sumatika; P. Núñez &amp; J. Arque 8369)." figureDoi="http://doi.org/10.5281/zenodo.8142002" httpUri="https://zenodo.org/record/8142002/files/figure.png" pageId="12" pageNumber="13">Fig. 5A</figureCitation>
).............................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFABAC519FA6DF8EE" box="[1360,1450,1839,1861]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="neillii">
<emphasis id="B965EADEFFB5FFBAFABAC519FA6DF8EE" bold="true" box="[1360,1450,1839,1861]" italics="true" pageId="12" pageNumber="13">M. neillii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C57AFA6CF8CA" blockId="12.[136,1452,159,2058]" box="[136,1451,1867,1889]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C57AFA6CF8CA" box="[136,1451,1867,1889]" pageId="12" pageNumber="13">
- Flowers 5-merous; calyx with thick, callose dorsal projections...........................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFAA5C57DFA6CF8CA" box="[1359,1451,1867,1889]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis id="B965EADEFFB5FFBAFAA5C57DFA6CF8CA" bold="true" box="[1359,1451,1867,1889]" italics="true" pageId="12" pageNumber="13">M. rigida</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C551FA6CF832" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C551FA6CF8D7" blockId="12.[136,1452,159,2058]" box="[136,1451,1895,1917]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C551FA6CF8D7" box="[136,1451,1895,1917]" pageId="12" pageNumber="13">
34. Leaf venation with 1 pair of secondary veins (lateral nerves) and an additional pair of faint submarginal veins..............
<taxonomicName id="4C114D4FFFB5FFBAFAAFC551FA6CF8D7" box="[1349,1451,1895,1916]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="rugosa">
<emphasis id="B965EADEFFB5FFBAFAAFC551FA6CF8D7" bold="true" box="[1349,1451,1895,1916]" italics="true" pageId="12" pageNumber="13">M. rugosa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C5B2FA6CF832" blockId="12.[136,1452,159,2058]" box="[136,1451,1923,1945]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C5B2FA6CF832" box="[136,1451,1923,1945]" pageId="12" pageNumber="13">- Leaf venation with 2 pairs of secondary veins (lateral nerves) and an additional pair of faint submarginal veins .........................35</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E69FFB5FFBAFF62C5A9FA6DF7A2" pageId="12" pageNumber="13">
<paragraph id="8BAE36CCFFB5FFBAFF62C5A9FA6CF87B" blockId="12.[136,1452,159,2058]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C5A9FA6CF87B" pageId="12" pageNumber="13">
35. Internodes quadrangular and 4-winged, wings
<quantity id="4CE99B29FFB5FFBAFD7EC5A9FD32F81E" box="[660,757,1951,1973]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.5" metricValueMin="1.0" pageId="12" pageNumber="13" unit="mm" value="2.25" valueMax="3.5" valueMin="1.0">13.5 mm</quantity>
high (
<figureCitation id="132A2A49FFB5FFBAFCDEC5A9FC46F81F" box="[820,897,1951,1973]" captionStart="FIGURE 3" captionStartId="6.[136,229,1742,1764]" captionTargetBox="[217,1370,1054,1715]" captionTargetId="figure-317@6.[216,1372,1053,1717]" captionTargetPageId="6" captionText="FIGURE 3. Morphological characteristics of Peruvian Meriania. A. Interpetiolar line (M. tomentosa; R. W. Bussmann et al. 17068). B. Interpetiolar flap (M. zunacensis; R. Fernandez-Hilario et al. 1920). C. Winged internode (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Winged internode (M. tetragona; R. Fernandez-Hilario et al. 2092). E. Adaxial projection (scutum) in the transition zone from the petiole to the midvein (M. zunacensis; R. Fernandez-Hilario et al. 1920). F. Swollen adaxial projection on the petiole apex (M. drakei; R. Fernandez-Hilario et al. 1775). G. Liguliform abaxial projections in the transition zone from the petiole to the midvein (M. sanguinea; R. Fernandez-Hilario et al. 1896). H. Tuberculate abaxial projections in the transition zone from the petiole to the midvein (M. tetragona; F. A. Michelangeli et al. 1739). I. Spreading corolla (M. rigida; R. Fernandez-Hilario et al. 1931). J. Campanulate corollas (M. dazae; J. L. Marcelo-Peña et al. 6568). K. Fruits with mature ovaries exceeding the hypanthia length (M. sanguinea; F.A. Michelangeli et al. 2743). L. Fruits with mature ovaries completely concealed by the hypanthia (M. tomentosa; F. A. Michelangeli et al. 1799). Photos by Rainer W. Bussmann (A); Robin Fernandez-Hilario (BG and IJ); and Fabián A. Michelangeli (H, K and L)." figureDoi="http://doi.org/10.5281/zenodo.8141994" httpUri="https://zenodo.org/record/8141994/files/figure.png" pageId="12" pageNumber="13">Fig. 3C</figureCitation>
); calyx with callose dorsal projections, whitish and much lighter than the rest of the calyx and hypanthium when dry (
<figureCitation id="132A2A49FFB5FFBAFCF8C58DFC9BF87A" box="[786,860,1979,2001]" captionStart="FIGURE 5" captionStartId="9.[136,229,1613,1635]" captionTargetBox="[155,1429,217,1583]" captionTargetId="figure-18@9.[150,1435,209,1589]" captionTargetPageId="9" captionText="FIGURE 5. Morphological characteristics of Peruvian Meriania. A. Calyx lacking dorsal projections (M. hexamera; I. Revilla 3484). B. Calyx with callose dorsal projections (left), and detail of a dorsal projection (right) (M. callosa; R. Fernandez-Hilario et al. 2055). C. Calyx with whitish callose dorsal projections (M. penningtonii; R. Fernandez-Hilario et al. 2072). D. Calyx with claw-shaped dorsal projections (M. dazae; J. L. Marcelo-Peña et al. 6568). E. Calyx with blunt dorsal projections (M. rugosa; H. van der Werff 17006). F. Subcalyptrate calyx with small claw-shaped projections and irregular dehiscence (left), and detail of the subcalyptrate calyx apex (right) (M. vasquezii; R. Vásquez et al. 45480). G. Calyptrate calyx with circumscissile dehiscence (M. escalerensis; M. Ríos et al. 3316). H. Calyx with aciculate dorsal projections and slightly costate hypanthium (M. prunifolia; J. Schunke-Vigo 11712). I. Costate hypanthium with 10 evident ribs (left) and apical view (right) (M. sumatika; P. Núñez &amp; J. Arque 8369)." figureDoi="http://doi.org/10.5281/zenodo.8142002" httpUri="https://zenodo.org/record/8142002/files/figure.png" pageId="12" pageNumber="13">Fig. 5C</figureCitation>
)......................................................................
<taxonomicName id="4C114D4FFFB5FFBAFAE1C58DFA6CF87B" box="[1291,1451,1979,2000]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="penningtonii">
<emphasis id="B965EADEFFB5FFBAFAE1C58DFA6CF87B" bold="true" box="[1291,1451,1979,2000]" italics="true" pageId="12" pageNumber="13">M. penningtonii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAE36CCFFB5FFBAFF62C5EEFA6DF7A2" blockId="12.[136,1452,159,2058]" pageId="12" pageNumber="13">
<keyLead id="30E095F9FFB5FFBAFF62C5EEFA6DF7A2" pageId="12" pageNumber="13">
- Internodes quadrangular, without wings; calyx with small conic dorsal projections, of the same color as the rest of the calyx and hypanthium when dry ...................................................................................................................................................
<taxonomicName id="4C114D4FFFB5FFBAFAF6C5C2FA6DF7A2" box="[1308,1450,2036,2057]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="tetraquetra">
<emphasis id="B965EADEFFB5FFBAFAF6C5C2FA6DF7A2" bold="true" box="[1308,1450,2036,2057]" italics="true" pageId="12" pageNumber="13">M. tetraquetra</emphasis>
</taxonomicName>
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</paragraph>
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</subSubSection>
</treatment>
</document>