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<mods:title id="7C0226839235094807DCE1A1CF272FE9">Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China</mods:title>
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<mods:affiliation id="5C6D9154D7A69885531611FE65A1D5EE">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659</mods:affiliation>
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<mods:affiliation id="486BCFCC374FA4E12566111CCAE9F059">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659</mods:affiliation>
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[3]
<taxonomicName id="4C114D05FFB7FFB59EB1C89F9D854B8B" authority="Ma &amp; Liu, 2021" authorityName="Ma &amp; Liu" authorityYear="2021" box="[193,675,1693,1720]" class="Insecta" family="Chrysopidae" genus="Ankylopteryx" kingdom="Animalia" order="Neuroptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="diffluens">
<emphasis id="B965EA94FFB7FFB59EB1C89F9EF24B8B" bold="true" box="[193,468,1693,1720]" italics="true" pageId="16" pageNumber="17">Ankylopteryx diffluens</emphasis>
<bibRefCitation id="EF804B77FFB7FFB59FACC89C9D854B8B" author="Ma, Y. L. &amp; Liu, X. Y." box="[476,675,1694,1720]" pageId="16" pageNumber="17" pagination="425 - 433" refId="ref89814" refString="Ma, Y. L. &amp; Liu, X. Y. (2021) Two new species of the green lacewing subgenus Ankylopteryx Brauer, 1864 (s. str.) (Neuroptera, Chrysopidae) from China. Zootaxa, 4941 (3), 425 - 433. https: // doi. org / 10.11646 / zootaxa. 4941.3.7" type="journal article" year="2021">Ma &amp; Liu, 2021</bibRefCitation>
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</paragraph>
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<paragraph id="8BAE3686FFB7FFB59EE7C8C09E3F4BEF" blockId="16.[151,675,1693,1756]" box="[151,281,1730,1756]" pageId="16" pageNumber="17">
(
<figureCitation id="132A2A03FFB7FFB59EEFC8C09FCE4BEF" box="[159,232,1730,1756]" captionStart="FIGURE 9" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[200,1388,191,1875]" captionTargetId="figure-19@17.[190,1398,181,1885]" captionTargetPageId="17" captionText="FIGURE 9. Ankylopteryx difluens Ma &amp; Liu, 2021.A. Dorsal habitus; B. Head, frontal view; C. Head, lateral view; D. Thorax, dorsal view; E. Male genitalia, lateral view; F. Gonarcus and entoprocessus, dorsal view; G. Gonarcus and entoprocessus, lateral view; H. Pseudopenis, dorsal view; I. Pseudopenis, lateral view; J. Female genitalia, lateral view; K. Subgenitale; L. Spermatheca. Scale bar: 5 mm (A), 1 mm (BD)." figureDoi="http://doi.org/10.5281/zenodo.14240526" httpUri="https://zenodo.org/record/14240526/files/figure.png" pageId="16" pageNumber="17">Figs 9</figureCitation>
,
<figureCitation id="132A2A03FFB7FFB59E84C8C09E374BEF" box="[244,273,1730,1756]" captionStart="FIGURE 79" captionStartId="142.[151,250,1894,1918]" captionTargetBox="[196,1392,1098,1870]" captionTargetId="figure-46@142.[196,1392,1098,1870]" captionTargetPageId="142" captionText="FIGURE 79. Distribution map of species of Ankylopteryx from China." figureDoi="http://doi.org/10.5281/zenodo.14240704" httpUri="https://zenodo.org/record/14240704/files/figure.png" pageId="16" pageNumber="17">79</figureCitation>
)
</paragraph>
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<paragraph id="8BAE3686FFB7FFB59EE7C9049BE14A2D" blockId="16.[151,1223,1798,1823]" box="[151,1223,1798,1823]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB0111A8FFB7FFB59EE7C9049BE14A2D" box="[151,1223,1798,1823]" pageId="16" pageNumber="17">
<treatmentCitation id="0AB01097FFB7FFB59EE7C9049D7A4A2D" author="Ma, Y. L. &amp; Liu, X. Y." box="[151,604,1798,1823]" page="426" pageId="16" pageNumber="17" year="2021">
<taxonomicName id="4C114D05FFB7FFB59EE7C9049D7A4A2D" authority="Ma &amp; Liu, 2021: 426" authorityName="Ma &amp; Liu" authorityPageNumber="426" authorityYear="2021" box="[151,604,1798,1823]" class="Insecta" family="Chrysopidae" genus="Ankylopteryx" kingdom="Animalia" order="Neuroptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="diffluens">
<emphasis id="B965EA94FFB7FFB59EE7C9049E5C4A2D" box="[151,378,1798,1822]" italics="true" pageId="16" pageNumber="17">Ankylopteryx diffluens</emphasis>
<materialsCitation id="3B793CDBFFB7FFB59FF1C9059D7A4A2D" box="[385,604,1798,1823]" country="China" location="Menglun" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<bibRefCitation id="EF804B77FFB7FFB59FF1C9059D7A4A2D" author="Ma, Y. L. &amp; Liu, X. Y." box="[385,604,1798,1823]" pageId="16" pageNumber="17" pagination="425 - 433" refId="ref89814" refString="Ma, Y. L. &amp; Liu, X. Y. (2021) Two new species of the green lacewing subgenus Ankylopteryx Brauer, 1864 (s. str.) (Neuroptera, Chrysopidae) from China. Zootaxa, 4941 (3), 425 - 433. https: // doi. org / 10.11646 / zootaxa. 4941.3.7" type="journal article" year="2021">Ma &amp; Liu, 2021: 426</bibRefCitation>
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. Type locality:
<collectingCountry id="F3067616FFB7FFB59C87C9049C124A2D" box="[759,820,1798,1822]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
(
<collectingRegion id="49D5F864FFB7FFB59D32C9059CB44A2D" box="[834,914,1799,1822]" country="China" name="Yunnan" pageId="16" pageNumber="17">Yunnan</collectingRegion>
,
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).
<typeStatus id="54AA8824FFB7FFB59A7DC9049B4D4A2D" box="[1037,1131,1798,1822]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
in CAU.
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</paragraph>
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<subSubSection id="C30B650DFFB7FFB59EE7C94E9DC44AC4" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BAE3686FFB7FFB59EE7C94E9DC44AC4" blockId="16.[151,1436,1868,2039]" pageId="16" pageNumber="17">
<emphasis id="B965EA94FFB7FFB59EE7C94E9E344A55" bold="true" box="[151,274,1868,1894]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
This species is characterized by the presence of the reddish-black spots below the antennae, the scape and pedicel each with reddish-brown stripe on both the outer and inner side, the pronotum, mesoprescutum and mesoscutum with brown spots on the anterolateral angle, the pro- and mesotibia with median black spot, the wings with scattered brown shadings, the quadrate cell
<emphasis id="B965EA94FFB7FFB59CC4C9BB9DF64AE1" box="[692,720,1977,2002]" italics="true" pageId="16" pageNumber="17">im</emphasis>
, the male entoprocessus attached at about basal 1/3 of gonarcus, and the slightly projected apex of the entoprocessus.
</paragraph>
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<caption id="DF6E660EFFB6FFB49EE7C9779DAD4ACA" ID-DOI="http://doi.org/10.5281/zenodo.14240526" ID-Zenodo-Dep="14240526" httpUri="https://zenodo.org/record/14240526/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1909,1933]" targetBox="[200,1388,191,1875]" targetPageId="17" targetType="figure">
<paragraph id="8BAE3686FFB6FFB49EE7C9779DAD4ACA" blockId="17.[151,1437,1909,2041]" pageId="17" pageNumber="18">
<emphasis id="B965EA94FFB6FFB49EE7C9779E344ABE" bold="true" box="[151,274,1909,1933]" pageId="17" pageNumber="18">FIGURE 9.</emphasis>
<taxonomicName id="4C114D05FFB6FFB49F68C9779D874ABE" authority="Ma &amp; Liu, 2021" authorityName="Ma &amp; Liu" authorityYear="2021" box="[280,673,1909,1933]" class="Insecta" family="Chrysopidae" genus="Ankylopteryx" kingdom="Animalia" order="Neuroptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="difluens">
<emphasis id="B965EA94FFB6FFB49F68C9779ED24ABE" box="[280,500,1909,1933]" italics="true" pageId="17" pageNumber="18">Ankylopteryx difluens</emphasis>
<bibRefCitation id="EF804B77FFB6FFB49F8AC9779D874ABE" author="Ma, Y. L. &amp; Liu, X. Y." box="[506,673,1909,1933]" pageId="17" pageNumber="18" pagination="425 - 433" refId="ref89814" refString="Ma, Y. L. &amp; Liu, X. Y. (2021) Two new species of the green lacewing subgenus Ankylopteryx Brauer, 1864 (s. str.) (Neuroptera, Chrysopidae) from China. Zootaxa, 4941 (3), 425 - 433. https: // doi. org / 10.11646 / zootaxa. 4941.3.7" type="journal article" year="2021">Ma &amp; Liu, 2021</bibRefCitation>
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. A. Dorsal habitus; B. Head, frontal view; C. Head, lateral view; D. Thorax, dorsal view; E. Male genitalia, lateral view; F. Gonarcus and entoprocessus, dorsal view; G. Gonarcus and entoprocessus, lateral view; H. Pseudopenis, dorsal view; I. Pseudopenis, lateral view; J. Female genitalia, lateral view; K. Subgenitale; L. Spermatheca. Scale bar: 5 mm (A), 1 mm (BD).
</paragraph>
</caption>
<subSubSection id="C30B650DFFB5FFB79EB7CE949C454EDE" pageId="18" pageNumber="19" type="description">
<paragraph id="8BAE3686FFB5FFB79EB7CE949A314D82" blockId="18.[151,1437,150,1185]" box="[199,1303,150,177]" pageId="18" pageNumber="19">
<emphasis id="B965EA94FFB5FFB79EB7CE949E7F4D83" bold="true" box="[199,345,150,176]" pageId="18" pageNumber="19">Description.</emphasis>
Male. Body length
<quantity id="4CE99B63FFB5FFB79C48CE959DB84D82" box="[568,670,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="18" pageNumber="19" unit="mm" value="7.8">7.80 mm</quantity>
, forewing length
<quantity id="4CE99B63FFB5FFB79D12CE959CF34D82" box="[866,981,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="18" pageNumber="19" unit="mm" value="11.9">11.90 mm</quantity>
, hindwing length 11.00 mm.
</paragraph>
<paragraph id="8BAE3686FFB5FFB79EB7CEB99C934C2E" blockId="18.[151,1437,150,1185]" pageId="18" pageNumber="19">Head yellow, vertex raised. Compound eyes blackish-brown. Scape and pedicel with reddish-brown stripe on outer and inner side, flagellum yellow, reddish-black spots below antennae. Clypeal margin with brown stripes. Gena marked brown spots. Maxillary and labial palpi brownish yellow.</paragraph>
<paragraph id="8BAE3686FFB5FFB79EB7CF259B964C56" blockId="18.[151,1437,150,1185]" pageId="18" pageNumber="19">Pronotum anterior margin short than posterior margin, with small brown spots on anterolateral angle; mesoprescutum and mesoscutum with brown spots on anterolateral angle, metanotum yellow.</paragraph>
<paragraph id="8BAE3686FFB5FFB79EB7CF6D9AAE4CBA" blockId="18.[151,1437,150,1185]" box="[199,1416,367,393]" pageId="18" pageNumber="19">Legs with dense setae. Pro- and mesotibia with median black spot. Pretarsal claw brown, with basal dilation.</paragraph>
<paragraph id="8BAE3686FFB5FFB79EB7CF919C124F0E" blockId="18.[151,1437,150,1185]" pageId="18" pageNumber="19">
Forewing broad, with brown shadings, especially on radial crossveins, base of Rs, inner gradate series, psm-psc crossveins and CuP. Pterostigma pale brown. Costal area broad basally, with 2325 crossveins; 11 radial crossveins present.
<emphasis id="B965EA94FFB5FFB79E88CFD99E3D4CC6" box="[248,283,475,501]" italics="true" pageId="18" pageNumber="19">m1</emphasis>
smaller than
<emphasis id="B965EA94FFB5FFB79FC8CFD99EFC4CC6" box="[440,474,475,501]" italics="true" pageId="18" pageNumber="19">m2</emphasis>
,
<emphasis id="B965EA94FFB5FFB79F9ACFD99D204CC7" box="[490,518,475,500]" italics="true" pageId="18" pageNumber="19">im</emphasis>
quadrate; 78 crossveins present between Psm and Psc.
<emphasis id="B965EA94FFB5FFB79AE3CFD99B884CC6" box="[1171,1198,475,501]" italics="true" pageId="18" pageNumber="19">c1</emphasis>
smaller than
<emphasis id="B965EA94FFB5FFB79B3BCFD99A434CC6" box="[1355,1381,475,501]" italics="true" pageId="18" pageNumber="19">c2</emphasis>
,
<emphasis id="B965EA94FFB5FFB79B05CFD99ABA4CC6" box="[1397,1436,475,501]" italics="true" pageId="18" pageNumber="19">dcc</emphasis>
opened. A1 forked, A2 and A3 simple. Inner gradate series with 67 crossveins, outer gradate series with 78 crossveins. Faint shadings on the forking of marginal veins.
</paragraph>
<paragraph id="8BAE3686FFB5FFB79EB7CC459E5B4FFE" blockId="18.[151,1437,150,1185]" pageId="18" pageNumber="19">Hindwing narrow, with shadings on inner gradate series, CuP and posterior marginal veins. Pterostigma pale brown. Costal area with 19 crossveins; 89 radial crossveins and 6 psm-psc crossveins present. A1 forked, A2 and A3 simple. Gradate series with pale brown shadings, inner gradate series with 46 crossveins, outer gradate series with 56 crossveins.</paragraph>
<paragraph id="8BAE3686FFB5FFB79EB7CCD59B0F4E6E" blockId="18.[151,1437,150,1185]" pageId="18" pageNumber="19">Abdomen with dense setae. Tergum VII and sternum VII subquadrate in lateral view; tergum VIII shorter than tergum VII; sternum VIII fused with sternum IX; tergum IX fused with ectoproct. Callus cerci ovoid. Gonarcus arcuate, lateral arms narrow at base and wide at tip; entoprocessus slightly curved in dorsal view, attached at about basal 1/3 of gonarcus, narrowed and projected at tip; pseudopenis widest near tip.</paragraph>
<paragraph id="8BAE3686FFB5FFB79EB7CD659BBF4EB2" blockId="18.[151,1437,150,1185]" box="[199,1177,871,897]" pageId="18" pageNumber="19">
Female. Body length
<quantity id="4CE99B63FFB5FFB79FC8CD659D384EB2" box="[440,542,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="18" pageNumber="19" unit="mm" value="8.2">8.20 mm</quantity>
, forewing length 13.00 mm, hindwing length
<quantity id="4CE99B63FFB5FFB79A50CD659BB34EB2" box="[1056,1173,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="18" pageNumber="19" unit="mm" value="12.4">12.40 mm</quantity>
.
</paragraph>
<paragraph id="8BAE3686FFB5FFB79EB7CD899C454EDE" blockId="18.[151,1437,150,1185]" pageId="18" pageNumber="19">Sternum VII obviously longer than tergum VII; tergum VII subquadrate in lateral view; tergum VIII shorter than tergum VII; tergum IX fused with ectoproct. Callus cerci ovoid. Subgenitale bilobed apically; spermatheca thick; vela and ventral impression absent; duct long and curved.</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFB5FFB79EB7CDF49C154906" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BAE3686FFB5FFB79EB7CDF49C154906" blockId="18.[151,1437,150,1185]" pageId="18" pageNumber="19">
<emphasis id="B965EA94FFB5FFB79EB7CDF49E104923" bold="true" box="[199,310,1014,1040]" pageId="18" pageNumber="19">Material.</emphasis>
<materialsCitation id="3B793CDBFFB5FFB79F4CCDF59E4F4906" collectingDate="2019-04-25" collectionCode="CAU" collectorName="Menglun, No." country="China" elevation="780" location="Hongyu Li" municipality="Xishuangbanna" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54AA8824FFB5FFB79F4CCDF59E824922" box="[316,420,1015,1041]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
<specimenCount id="9D17FD0FFFB5FFB79FDACDF59EE74922" box="[426,449,1015,1041]" count="1" pageId="18" pageNumber="19" type="male"></specimenCount>
,
<collectingCountry id="F3067616FFB5FFB79FBCCDF59D344922" box="[460,530,1015,1041]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
,
<collectingRegion id="49D5F864FFB5FFB79C6BCDF59D524922" box="[539,628,1015,1041]" country="China" name="Yunnan" pageId="18" pageNumber="19">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BCAACFCFFB5FFB79C0DCDF59C084922" box="[637,814,1015,1041]" pageId="18" pageNumber="19">Xishuangbanna</collectingMunicipality>
,
<collectorName id="26E45350FFB5FFB79D48CDF59CF24922" box="[824,980,1015,1041]" pageId="18" pageNumber="19">Menglun, No.</collectorName>
55 experimental site,
<quantity id="4CE99B63FFB5FFB79AB7CDF59A294922" box="[1223,1295,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="18" pageNumber="19" unit="m" value="780.0">
<elevation id="003CD1B5FFB5FFB79AB7CDF59A294922" box="[1223,1295,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="18" pageNumber="19" unit="m" value="780.0">780 m</elevation>
</quantity>
,
<date id="FFAF1046FFB5FFB79B69CDF49ABE4922" box="[1305,1432,1014,1041]" pageId="18" pageNumber="19" value="2019-04-25">
<collectingDate id="EFEBE9AEFFB5FFB79B69CDF49ABE4922" box="[1305,1432,1014,1041]" pageId="18" pageNumber="19" value="2019-04-25">2019.IV.25</collectingDate>
</date>
,
<location id="8ECE605DFFB5FFB79EE7CA199E374906" LSID="urn:lsid:plazi:treatment:03B88790FFB7FFB79EE7C89C9E484992:8ECE605DFFB5FFB79EE7CA199E374906" box="[151,273,1051,1077]" country="China" municipality="Xishuangbanna" name="Hongyu Li" pageId="18" pageNumber="19" stateProvince="Yunnan">Hongyu Li</location>
(
<collectionCode id="ED00AE43FFB5FFB79F50CA199E474906" box="[288,353,1051,1077]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="18" pageNumber="19">CAU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B793CDBFFB5FFB79F04CA199C094906" box="[372,815,1051,1077]" collectionCode="CAU" country="China" pageId="18" pageNumber="19" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54AA8824FFB5FFB79F04CA199EFC4906" box="[372,474,1051,1077]" pageId="18" pageNumber="19" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D17FD0FFFB5FFB79F95CA199D2E4906" box="[485,520,1051,1077]" count="1" pageId="18" pageNumber="19" type="female">1♀</specimenCount>
, same as
<typeStatus id="54AA8824FFB5FFB79C05CA199DF14906" box="[629,727,1051,1077]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED00AE43FFB5FFB79C96CA199C014906" box="[742,807,1051,1077]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="18" pageNumber="19">CAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFB5FFB79EB7CA3C9D07496A" box="[199,545,1086,1113]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BAE3686FFB5FFB79EB7CA3C9D07496A" blockId="18.[151,1437,150,1185]" box="[199,545,1086,1113]" pageId="18" pageNumber="19">
<emphasis id="B965EA94FFB5FFB79EB7CA3C9E47496B" bold="true" box="[199,353,1086,1112]" pageId="18" pageNumber="19">Distribution.</emphasis>
<collectingCountry id="F3067616FFB5FFB79F17CA3D9E8C496A" box="[359,426,1087,1113]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
(
<collectingRegion id="49D5F864FFB5FFB79FCBCA3D9D30496A" box="[443,534,1087,1113]" country="China" name="Yunnan" pageId="18" pageNumber="19">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFB5FFB79EB7CA619E484992" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BAE3686FFB5FFB79EB7CA619E484992" blockId="18.[151,1437,150,1185]" pageId="18" pageNumber="19">
<emphasis id="B965EA94FFB5FFB79EB7CA619E1D494E" bold="true" box="[199,315,1123,1149]" pageId="18" pageNumber="19">Remarks.</emphasis>
This species can be identified easily by the reddish stripes around the antennae and the scattered shadings on wings.
</paragraph>
</subSubSection>
</treatment>
</document>