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<document id="CBE26EE41AD2E199341F99F8FDBEE136" ID-DOI="10.1111/zoj.12300" ID-ISSN="0024-4082" ID-Zenodo-Dep="5458789" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630211112177" checkinUser="marcus" docAuthor="Scanferla, Agustín, Smith, Krister T. &amp; Schaal, Stephan F. K." docDate="2016" docId="03B8878EFFCDFFFC9242211BFD074F38" docLanguage="en" docName="zoj12300.pdf" docOrigin="Zoological Journal of the Linnean Society 176 (1)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/zoj.12300" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Messelophis variatus Baszio 2004" docType="treatment" docVersion="3" lastPageNumber="203" masterDocId="FF81FFF6FFD8FFEA913D243AFFD84D50" masterDocTitle="Revision of the cranial anatomy and phylogenetic relationships of the Eocene minute boas Messelophis variatus and Messelophis ermannorum (Serpentes, Booidea)" masterLastPageNumber="206" masterPageNumber="182" pageNumber="203" updateTime="1699187420293" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="17F1D5ACF7429619369E955D5118BC70">Revision of the cranial anatomy and phylogenetic relationships of the Eocene minute boas Messelophis variatus and Messelophis ermannorum (Serpentes, Booidea)</mods:title>
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<mods:namePart id="0FBACF8FC259AB58253D77327C36EDEB">Scanferla, Agustín</mods:namePart>
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<mods:namePart id="34B92B0808EAA276575675A699C71594">Smith, Krister T.</mods:namePart>
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<mods:namePart id="E885F18D277EC3B74D53B09D4209B46B">Schaal, Stephan F. K.</mods:namePart>
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<heading id="D0E681F4FFCDFFFF9242211BFAF74865" allCaps="true" box="[895,1327,1312,1334]" centered="true" fontSize="9" level="2" pageId="21" pageNumber="203" reason="9">
<taxonomicName id="4C114D1BFFCDFFFF9242211BFAF74865" authority="BASZIO, 2004" authorityName="Baszio" authorityYear="2004" box="[895,1327,1312,1334]" class="Reptilia" family="Boidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="183" phylum="Chordata" rank="species" species="variatus">
<emphasis id="B965EA8AFFCDFFFF9242211BFB574864" box="[895,1167,1313,1334]" italics="true" pageId="21" pageNumber="203">MESSELOPHIS VARIATUS</emphasis>
<bibRefCitation id="EF804B69FFCDFFFF95AA211AFAF74865" author="Baszio S" box="[1175,1327,1312,1333]" pageId="21" pageNumber="203" pagination="47 - 66" refId="ref13452" refString="Baszio S. 2004. Messelophis variatus n. gen. nov. sp. from the Eocene of Messel: a tropidopheine snake with affinities to Erycinae (Boidae). Courier Forschungsinstitut Senckenberg 252: 47 - 66." type="journal article" year="2004">BASZIO, 2004</bibRefCitation>
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<emphasis id="B965EA8AFFCDFFFF921E217DFC5E480C" box="[803,902,1351,1372]" italics="true" pageId="21" pageNumber="203">
<typeStatus id="54AA883AFFCDFFFF921E217DFC5E480C" box="[803,902,1351,1372]" pageId="21" pageNumber="203" type="holotype">Holotype</typeStatus>
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</paragraph>
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<materialsCitation id="3B793CC5FFCDFFFF921E215FFB7B48E8" collectionCode="SMF" pageId="21" pageNumber="203" specimenCode="ME 1828" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED00AE5DFFCDFFFF921E215FFC81482A" box="[803,857,1381,1402]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="21" pageNumber="203">SMF</collectionCode>
<specimenCode id="DBB79EE3FFCDFFFF9262215CFC19482A" box="[863,961,1381,1403]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="21" pageNumber="203">ME 1828</specimenCode>
a+b, an articulated, almost complete skeleton, and
<collectionCode id="ED00AE5DFFCDFFFF92AA21BEFC1648C9" box="[919,974,1412,1433]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="21" pageNumber="203">SMF</collectionCode>
<specimenCode id="DBB79EE3FFCDFFFF92E821BEFBE448C9" box="[981,1084,1412,1433]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="21" pageNumber="203">ME 1828</specimenCode>
c, four isolated vertebrae extracted from the same specimen.
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<emphasis id="B965EA8AFFCDFFFF921E21DEFBC848A9" box="[803,1040,1508,1529]" italics="true" pageId="21" pageNumber="203">Locality and horizon</emphasis>
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Middle Messel Formation, Messel Pit near Darmstadt (
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), middle Eocene (
<collectionCode id="ED00AE5DFFCDFFFF9590221BFB0D4B66" box="[1197,1237,1569,1590]" country="0" name="Mohonk Preserve, Inc." pageId="21" pageNumber="203">MP</collectionCode>
11).
</paragraph>
<paragraph id="8BAE3698FFCDFFFF921E2259FBDA4B28" blockId="21.[803,1419,1635,1902]" box="[803,1026,1635,1656]" pageId="21" pageNumber="203">
<emphasis id="B965EA8AFFCDFFFF921E2259FBDA4B28" box="[803,1026,1635,1656]" italics="true" pageId="21" pageNumber="203">Emended diagnosis</emphasis>
</paragraph>
<paragraph id="8BAE3698FFCDFFFC921E22B8FD074F38" blockId="21.[803,1419,1635,1902]" lastBlockId="22.[166,782,197,617]" lastPageId="22" lastPageNumber="204" pageId="21" pageNumber="203">
A minute booid snake that can be distinguished from all other members of
<taxonomicName id="4C114D1BFFCDFFFF9526229AFB544BE5" authorityName="LINNAEUS" authorityYear="1758" box="[1051,1164,1696,1717]" pageId="21" pageNumber="204" rank="subOrder" subOrder="Serpentes">Serpentes</taxonomicName>
by the following combination of characters: dentate premaxilla with ascending process; lacrimal duct opens ventrally; large finger-like medial foot process; postorbital is applied to parietal and frontal; parietal with parietal table and a pointed posterior process in sagittal crest; maxilla with one large maxillary foramen, and teeth dimin- ishing in size posteriorly; ectopterygoid with forked maxillary process; parietal table present; supraorbital process of parietal well developed; supratemporal short, with a conspicuous lip in its contact region with the quadrate; anterior dorsal and ventral processes of prootic weakly developed (i.e. anterior trigeminal foramen anteriorly delimited by parietal); laterosphenoid present; 300 precloacal and 6880 cloacalpostcloacal vertebrae; neural spine of precloacal vertebrae pointlike, shifted to the posteriormost part of the neural arch; prezygapophyseal process weakly developed; paracotylar foramina absent; haemal keel absent throughout the vertebral column; and paired laminar haemapophyses in postcloacal vertebrae present.
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