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<document id="F6A9B09BBBC29234B1CF4E747D05B763" ID-CLB-Dataset="307344" ID-DOI="10.5252/geodiversitas2025v47a1" ID-GBIF-Dataset="0b3bc518-cb63-4adb-8e55-a3cfa9a92aed" ID-ISSN="1638-9395" ID-Zenodo-Dep="14620093" ID-ZooBank="urn:lsid:zoobank.org:pub:4D31AE72-DD52-4E81-93D3-DD581D63396D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1736412153764" checkinUser="plazi" docAuthor="Martin, Jean-Paul Saint, Charbonnier, Sylvain, Martin, Simona Saint, Cazes, Lilian &amp; André, Jean-Pierre" docDate="2025" docId="03B86E7CFA276E323B97F961E5A9FD30" docLanguage="en" docName="geodiversitas.2025.47.1.pdf" docOrigin="Geodiversitas 47 (1)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/geodiversitas2025v47a1.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Palaeopascichnus linearis" docType="treatment" docVersion="2" lastPageNumber="8" masterDocId="FF811604FA2F6E3B3815FF95E006FFDF" masterDocTitle="New records of Palaeopaschichnus Palij, 1976 from the Ediacaran of Romania" masterLastPageNumber="16" masterPageNumber="1" pageNumber="7" updateTime="1736430649321" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="F18EE5500DC5D92A62499EF3C251B277">New records of Palaeopaschichnus Palij, 1976 from the Ediacaran of Romania</mods:title>
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<mods:namePart id="22B43DF42EF8282B07AFC67AA39E1283">Martin, Jean-Paul Saint</mods:namePart>
<mods:affiliation id="3AD7732D1E32C1E316D5E5761DA70993">Centre de Recherche en Paléontologie (UMR 7207 CR 2 P), CNRS, MNHN, Sorbonne Université, Muséum national dHistoire naturelle, Département Origines &amp; Évolution, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:namePart id="3CE63C4550519B595396B58FD6ADC9C3">Charbonnier, Sylvain</mods:namePart>
<mods:affiliation id="45986968C09304ED6A2A63825DDB73D8">Centre de Recherche en Paléontologie (UMR 7207 CR 2 P), CNRS, MNHN, Sorbonne Université, Muséum national dHistoire naturelle, Département Origines &amp; Évolution, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:namePart id="307EEEFBDD100BAFFA8101AF70421DF5">Martin, Simona Saint</mods:namePart>
<mods:affiliation id="73D4C61C10EA0AFC42DCE9CC2A933E9E">Centre de Recherche en Paléontologie (UMR 7207 CR 2 P), CNRS, MNHN, Sorbonne Université, Muséum national dHistoire naturelle, Département Origines &amp; Évolution, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:namePart id="339EE7E500340DB797C722F610B0D7F4">Cazes, Lilian</mods:namePart>
<mods:affiliation id="AEEDEAE0B7344628A9C04F274B022562">Centre de Recherche en Paléontologie (UMR 7207 CR 2 P), CNRS, MNHN, Sorbonne Université, Muséum national dHistoire naturelle, Département Origines &amp; Évolution, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:namePart id="0E1D1FEA268B4A2F5077495629BE8F1E">André, Jean-Pierre</mods:namePart>
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<mods:date id="326F713A09A281A4E44EDA24DC914B7E">2025</mods:date>
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<treatment id="03B86E7CFA276E323B97F961E5A9FD30" ID-DOI="http://doi.org/10.5281/zenodo.14622003" ID-Zenodo-Dep="14622003" LSID="urn:lsid:plazi:treatment:03B86E7CFA276E323B97F961E5A9FD30" httpUri="http://treatment.plazi.org/id/03B86E7CFA276E323B97F961E5A9FD30" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
<subSubSection id="C30B8CE1FA276E333B97F961E55CF8D0" box="[898,1370,1780,1807]" pageId="8" pageNumber="7" type="nomenclature">
<paragraph id="8BAEDF6AFA276E333B97F961E55CF8D0" blockId="8.[898,1370,1780,1839]" box="[898,1370,1780,1807]" pageId="8" pageNumber="7">
<taxonomicName id="4C11A4E9FA276E333B97F961E55CF8D0" authority="(Fedonkin, 1976)" baseAuthorityName="Fedonkin" baseAuthorityYear="1976" box="[898,1370,1780,1807]" genus="Palaeopascichnus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="7" rank="species" species="linearis">
<emphasis id="B9650378FA276E333B97F961E495F8D0" bold="true" box="[898,1171,1780,1807]" italics="true" pageId="8" pageNumber="7">Palaeopascichnus linearis</emphasis>
(
<bibRefCitation id="EF80A29BFA276E333CB7F960E554F8D0" author="FEDONKIN M. A." box="[1186,1362,1781,1807]" pageId="8" pageNumber="7" pagination="129 - 32" refId="ref8075" refString="FEDONKIN M. A. 1976. - Sledy mnogokletotchnykh iz valdaiskoi serii. Izvestiya Akademii Nauk SSSR, Seriya Geologicheskaya 4: 129 - 32 (in Russian)." type="journal article" year="1976">Fedonkin, 1976</bibRefCitation>
)
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</paragraph>
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<subSubSection id="C30B8CE1FA276E333C22F880E4A3F8F0" box="[1079,1189,1813,1839]" pageId="8" pageNumber="7" type="description">
<paragraph id="8BAEDF6AFA276E333C22F880E4A3F8F0" blockId="8.[898,1370,1780,1839]" box="[1079,1189,1813,1839]" pageId="8" pageNumber="7">
(
<figureCitation id="132AC3EFFA276E333C55F880E486F8F0" box="[1088,1152,1813,1839]" captionStart="FIG" captionStartId="10.[132,143,1340,1357]" captionTargetBox="[132,1456,215,1298]" captionTargetId="figure-312@10.[132,1456,215,1298]" captionTargetPageId="10" captionText="FIG. 6. — Specimens from Casimcea surface S1: A, View in situ of the Palaeopascichnus Palij, 1976 specimen; B, photogrammetric representation of the cast showing the poorly marked ridges at the surface (black arrow), the network of arcuate meandering elements and the “empty” meandering parallel imprints (white arrow); C, detailed view showing the well-marked outline of segments,the clear superposition of segments (a), the subdivision of segments with elements increasing in size just before the subdivision (b); D, detail of S1 specimen (close up of B, white arrow) with arcuate elements and imprint with parallel lines of constant width (c. 1 cm), without internal structures. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14620107" httpUri="https://zenodo.org/record/14620107/files/figure.png" pageId="8" pageNumber="7">Figs 6</figureCitation>
;
<figureCitation id="132AC3EFFA276E333C9BF880E49AF8F0" box="[1166,1180,1813,1839]" captionStart="FIG" captionStartId="11.[132,143,1205,1222]" captionTargetBox="[136,1437,205,1177]" captionTargetId="figure-538@11.[151,1437,219,1165]" captionText="FIG. 7. — A, Specimens from Casimcea surface S2 with a meandering succession of slightly arcuate element (black arrow) poorly expressed and imprint with parallel lines of constant width (c. 1 cm), without internal structures (white arrow) similar to Fig. 6D; B, same picture as A with specimens highlighted;C, D, speci- men from Rhaman; C, meandering succession of arcuate elements (arrow) on S2 surface; D, detail of C. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14622006" httpUri="https://zenodo.org/record/14622006/files/figure.png" pageId="8" pageNumber="7">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B8CE1FA276E333B38F8DBE504F834" pageId="8" pageNumber="7" type="reference_group">
<paragraph id="8BAEDF6AFA276E333B38F8DBE504F834" blockId="8.[811,1457,1870,2027]" pageId="8" pageNumber="7">
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. — See details in
<bibRefCitation id="EF80A29BFA276E333C45F8DBE540F8B9" author="KOLESNIKOV A. V. &amp; LIU A. G. &amp; DANELIAN T. &amp; GRAZHDANKIN D. V." box="[1104,1350,1870,1894]" pageId="8" pageNumber="7" pagination="197 - 205" refId="ref10664" refString="KOLESNIKOV A. V., LIU A. G., DANELIAN T. &amp; GRAZHDANKIN D. V. 2018 a. - A reassessment of the problematic Ediacaran genus Orbisiana Sokolov 1976. Precambrian Research 316: 197 - 205. https: // doi. org / 10.1016 / j. precamres. 2018.08.011" type="book chapter" year="2018">
Kolesnikov
<emphasis id="B9650378FA276E333CD6F8DAE4F5F8B9" box="[1219,1267,1870,1894]" italics="true" pageId="8" pageNumber="7">et al.</emphasis>
(2018a)
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and
<bibRefCitation id="EF80A29BFA276E333D6EF8DBE43DF85E" author="KOLESNIKOV A. V. &amp; DESIATKIN V." pageId="8" pageNumber="7" pagination="1175 - 1191" refId="ref10554" refString="KOLESNIKOV A. V. &amp; DESIATKIN V. 2022. - Taxonomy and palaeoenvironmental distribution of palaeopascichnids. Geological Magazine 159: 1175 - 1191. https: // doi. org / 10.1017 / S 0016756822000437" type="book chapter" year="2022">Kolesnikov &amp; Desiatkin (2022)</bibRefCitation>
. The original name
<taxonomicName id="4C11A4E9FA276E333D0AF8FCE410F844" authority="Fedonkin, 1976" authorityName="Fedonkin" authorityYear="1976" genus="Margaritichnus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="7" rank="species" species="linearis">
<emphasis id="B9650378FA276E333D0AF8FCE377F844" italics="true" pageId="8" pageNumber="7">Margaritichnus linearis</emphasis>
<bibRefCitation id="EF80A29BFA276E333B6DF816E410F844" author="FEDONKIN M. A." box="[888,1046,1923,1947]" pageId="8" pageNumber="7" pagination="129 - 32" refId="ref8075" refString="FEDONKIN M. A. 1976. - Sledy mnogokletotchnykh iz valdaiskoi serii. Izvestiya Akademii Nauk SSSR, Seriya Geologicheskaya 4: 129 - 32 (in Russian)." type="journal article" year="1976">Fedonkin, 1976</bibRefCitation>
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was first proposed for a supposed trace fossil (chains of faecal pellets). The species was later revised with the diagnosis emended to correspond to a body fossil (see
<bibRefCitation id="EF80A29BFA276E333D56F82CE398F834" author="KOLESNIKOV A. V. &amp; LIU A. G. &amp; DANELIAN T. &amp; GRAZHDANKIN D. V." pageId="8" pageNumber="7" pagination="197 - 205" refId="ref10664" refString="KOLESNIKOV A. V., LIU A. G., DANELIAN T. &amp; GRAZHDANKIN D. V. 2018 a. - A reassessment of the problematic Ediacaran genus Orbisiana Sokolov 1976. Precambrian Research 316: 197 - 205. https: // doi. org / 10.1016 / j. precamres. 2018.08.011" type="book chapter" year="2018">
Kolesnikov
<emphasis id="B9650378FA276E333B38F841E35AF834" box="[813,860,2003,2027]" italics="true" pageId="8" pageNumber="7">et al.</emphasis>
2018
<bibRefCitation id="EF80A29BFA276E333B86F846E398F834" author="KOLESNIKOV A. V. &amp; DESIATKIN V." box="[915,926,2003,2027]" pageId="8" pageNumber="7" pagination="1175 - 1191" refId="ref10554" refString="KOLESNIKOV A. V. &amp; DESIATKIN V. 2022. - Taxonomy and palaeoenvironmental distribution of palaeopascichnids. Geological Magazine 159: 1175 - 1191. https: // doi. org / 10.1017 / S 0016756822000437" type="book chapter" year="2022">a</bibRefCitation>
</bibRefCitation>
and Kolesnikov &amp; Desiatkin 2022).
</paragraph>
</subSubSection>
<subSubSection id="C30B8CE1FA266E323891FF4DE2A3FE76" pageId="9" pageNumber="8" type="diagnosis">
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<smallCapsWord id="8D4849B6FA266E3238FFFF48E148FF31" baselines="234" box="[234,334,221,238]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="diagnosis" pageId="9" pageNumber="8">DIAGNOSIS</smallCapsWord>
. — Test agglutinated, elongated, curved or rectilinear, occasionally branching, consists of a single series of globular or ellipsoidal chambers
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in width. The series occasionally diverge dichotomously. Chambers are relatively consistent in size within a series or gradually increase in width before diverging, but the length-to width ratio of the chambers is relatively constant along the series. The wall thickness does not exceed
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. Number of chambers in a series ranges between 3 and 5 and 30-40.
</paragraph>
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<collectionCode id="ED0047AFFA266E3238A9FE53E0E7FE08" box="[188,225,454,471]" pageId="9" pageNumber="8">AGE</collectionCode>
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. — Late Ediacaran.
</paragraph>
<paragraph id="8BAEDF6AFA266E323891FE65E111FDE2" blockId="9.[132,775,496,573]" pageId="9" pageNumber="8">
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EXAMINED SPECIMENS. —
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<collectingCountry id="F3069FFAFA266E32396BFE64E1DEFDD6" box="[382,472,497,521]" name="Romania" pageId="9" pageNumber="8">Romania</collectingCountry>
</emphasis>
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(
<emphasis id="B9650378FA266E323A73FE64E299FDD7" box="[614,671,497,520]" italics="true" pageId="9" pageNumber="8">in situ</emphasis>
specimens and cast);
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;
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.
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<materialsCitation id="3B79D537FA266E323A0EFD9FE111FDE2" country="Romania" location="Rahman" pageId="9" pageNumber="8" specimenCount="1">
<location id="8ECE89B1FA266E323A0EFD9FE277FDFD" LSID="urn:lsid:plazi:treatment:03B86E7CFA276E323B97F961E5A9FD30:8ECE89B1FA266E323A0EFD9FE277FDFD" box="[539,625,522,546]" country="Romania" name="Rahman" pageId="9" pageNumber="8">Rahman</location>
;
<specimenCount id="9D1714E3FA266E323A6CFD9EE2E0FDFD" box="[633,742,522,547]" count="1" pageId="9" pageNumber="8" type="generic">1 specimen</specimenCount>
(
<emphasis id="B9650378FA266E323AE6FD9EE0A3FDE2" italics="true" pageId="9" pageNumber="8">in situ</emphasis>
specimen).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30B8CE1FA266E323891FDE3E2F7F894" pageId="9" pageNumber="8" type="description">
<paragraph id="8BAEDF6AFA266E323891FDE3E113FD51" blockId="9.[130,777,630,1613]" box="[132,277,630,656]" pageId="9" pageNumber="8">
<smallCapsWord id="8D4849B6FA266E323891FDE3E113FD51" baselines="649,649" box="[132,277,630,656]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="9" pageNumber="8">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8BAEDF6AFA266E323891FD03E149FD70" blockId="9.[130,777,630,1613]" box="[132,335,662,688]" pageId="9" pageNumber="8">
<emphasis id="B9650378FA266E323891FD03E149FD70" box="[132,335,662,688]" italics="true" pageId="9" pageNumber="8">Casimcea specimens</emphasis>
</paragraph>
<paragraph id="8BAEDF6AFA266E323891FD20E1CCFA72" blockId="9.[130,777,630,1613]" pageId="9" pageNumber="8">
The specimen from the S1 surface is preserved as positive epirelief visible on a roughly rectangular area (
<emphasis id="B9650378FA266E323A7BFD43E279FD30" box="[622,639,726,751]" italics="true" pageId="9" pageNumber="8">c.</emphasis>
30 ×
<quantity id="4CE9728FFA266E323AAEFD43E306FD2F" box="[699,768,726,752]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="8" unit="cm" value="70.0">70 cm</quantity>
) exhibiting some slightly marked ripples (
<figureCitation id="132AC3EFFA266E323A33FD60E296FCD0" box="[550,656,757,784]" captionStart="FIG" captionStartId="10.[132,143,1340,1357]" captionTargetBox="[132,1456,215,1298]" captionTargetId="figure-312@10.[132,1456,215,1298]" captionTargetPageId="10" captionText="FIG. 6. — Specimens from Casimcea surface S1: A, View in situ of the Palaeopascichnus Palij, 1976 specimen; B, photogrammetric representation of the cast showing the poorly marked ridges at the surface (black arrow), the network of arcuate meandering elements and the “empty” meandering parallel imprints (white arrow); C, detailed view showing the well-marked outline of segments,the clear superposition of segments (a), the subdivision of segments with elements increasing in size just before the subdivision (b); D, detail of S1 specimen (close up of B, white arrow) with arcuate elements and imprint with parallel lines of constant width (c. 1 cm), without internal structures. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14620107" httpUri="https://zenodo.org/record/14620107/files/figure.png" pageId="9" pageNumber="8">Fig. 6A, B</figureCitation>
). It consists of meandering segments with parallel edges, of fairly constant width (
<quantity id="4CE9728FFA266E3238C6FCA0E113FC8F" box="[211,277,821,848]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="8" unit="mm" value="8.0">8 mm</quantity>
in average). The segments are composed of a series of generally consecutive apparently incurved-arcuate elements, about
<quantity id="4CE9728FFA266E323950FCE0E18CFC50" box="[325,394,885,911]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="8" unit="mm" value="4.0">4 mm</quantity>
thick, with low positive epirelief (
<figureCitation id="132AC3EFFA266E323898FC00E0E7FC70" box="[141,225,917,943]" captionStart="FIG" captionStartId="10.[132,143,1340,1357]" captionTargetBox="[132,1456,215,1298]" captionTargetId="figure-312@10.[132,1456,215,1298]" captionTargetPageId="10" captionText="FIG. 6. — Specimens from Casimcea surface S1: A, View in situ of the Palaeopascichnus Palij, 1976 specimen; B, photogrammetric representation of the cast showing the poorly marked ridges at the surface (black arrow), the network of arcuate meandering elements and the “empty” meandering parallel imprints (white arrow); C, detailed view showing the well-marked outline of segments,the clear superposition of segments (a), the subdivision of segments with elements increasing in size just before the subdivision (b); D, detail of S1 specimen (close up of B, white arrow) with arcuate elements and imprint with parallel lines of constant width (c. 1 cm), without internal structures. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14620107" httpUri="https://zenodo.org/record/14620107/files/figure.png" pageId="9" pageNumber="8">Fig. 6C</figureCitation>
). They were considered as chambers by
<bibRefCitation id="EF80A29BFA266E323A9AFC00E110FC10" author="KOLESNIKOV A. V. &amp; ROGOV V. I. &amp; BYKOVA N. V. &amp; DANELIAN T. &amp; CLAUSEN S. &amp; MASLOV A. V. &amp; GRAZHDANKIN D. V." pageId="9" pageNumber="8" pagination="24 - 37" refId="ref10722" refString="KOLESNIKOV A. V., ROGOV V. I., BYKOVA N. V., DANELIAN T., CLAUSEN S., MASLOV A. V. &amp; GRAZHDANKIN D. V. 2018 b. - The oldest skeletal macroscopic organism Palaeopascichnuslinearis. Precambrian Research 316: 24 - 37. https: // doi. org / 10.1016 / j. precamres. 2018.07. 017" type="book chapter" year="2018">
Kolesnikov
<emphasis id="B9650378FA266E323891FC20E0B1FC11" box="[132,183,949,975]" italics="true" pageId="9" pageNumber="8">et al.</emphasis>
(2018b)
</bibRefCitation>
. Elements are separated by a shallow incurved depression (
<emphasis id="B9650378FA266E323916FC40E112FC31" box="[259,276,981,1006]" italics="true" pageId="9" pageNumber="8">c.</emphasis>
<quantity id="4CE9728FFA266E32390DFC40E151FC30" box="[280,343,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="8" unit="mm" value="1.0">1 mm</quantity>
wide). The parallel edges of the segments are often visible and marked by a fine negative epirelief groove (
<quantity id="4CE9728FFA266E32389FFB81E0D9FBF0" box="[138,223,1044,1071]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="8" unit="mm" value="0.5">0.5 mm</quantity>
wide;
<figureCitation id="132AC3EFFA266E323934FB81E172FBF1" box="[289,372,1044,1070]" captionStart="FIG" captionStartId="10.[132,143,1340,1357]" captionTargetBox="[132,1456,215,1298]" captionTargetId="figure-312@10.[132,1456,215,1298]" captionTargetPageId="10" captionText="FIG. 6. — Specimens from Casimcea surface S1: A, View in situ of the Palaeopascichnus Palij, 1976 specimen; B, photogrammetric representation of the cast showing the poorly marked ridges at the surface (black arrow), the network of arcuate meandering elements and the “empty” meandering parallel imprints (white arrow); C, detailed view showing the well-marked outline of segments,the clear superposition of segments (a), the subdivision of segments with elements increasing in size just before the subdivision (b); D, detail of S1 specimen (close up of B, white arrow) with arcuate elements and imprint with parallel lines of constant width (c. 1 cm), without internal structures. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14620107" httpUri="https://zenodo.org/record/14620107/files/figure.png" pageId="9" pageNumber="8">Fig. 6C</figureCitation>
). One imprint differs by the lack of the successive elements between the two parallel lines (
<figureCitation id="132AC3EFFA266E323AB1FBA1E2FFFB91" box="[676,761,1076,1102]" captionStart="FIG" captionStartId="10.[132,143,1340,1357]" captionTargetBox="[132,1456,215,1298]" captionTargetId="figure-312@10.[132,1456,215,1298]" captionTargetPageId="10" captionText="FIG. 6. — Specimens from Casimcea surface S1: A, View in situ of the Palaeopascichnus Palij, 1976 specimen; B, photogrammetric representation of the cast showing the poorly marked ridges at the surface (black arrow), the network of arcuate meandering elements and the “empty” meandering parallel imprints (white arrow); C, detailed view showing the well-marked outline of segments,the clear superposition of segments (a), the subdivision of segments with elements increasing in size just before the subdivision (b); D, detail of S1 specimen (close up of B, white arrow) with arcuate elements and imprint with parallel lines of constant width (c. 1 cm), without internal structures. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14620107" httpUri="https://zenodo.org/record/14620107/files/figure.png" pageId="9" pageNumber="8">Fig. 6D</figureCitation>
). The photogrammetry of the mold reveals the bumpy appearance of the surface and the arrangement of the fossil (
<figureCitation id="132AC3EFFA266E323AA6FBE1E09CFB71" captionStart="FIG" captionStartId="10.[132,143,1340,1357]" captionTargetBox="[132,1456,215,1298]" captionTargetId="figure-312@10.[132,1456,215,1298]" captionTargetPageId="10" captionText="FIG. 6. — Specimens from Casimcea surface S1: A, View in situ of the Palaeopascichnus Palij, 1976 specimen; B, photogrammetric representation of the cast showing the poorly marked ridges at the surface (black arrow), the network of arcuate meandering elements and the “empty” meandering parallel imprints (white arrow); C, detailed view showing the well-marked outline of segments,the clear superposition of segments (a), the subdivision of segments with elements increasing in size just before the subdivision (b); D, detail of S1 specimen (close up of B, white arrow) with arcuate elements and imprint with parallel lines of constant width (c. 1 cm), without internal structures. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14620107" httpUri="https://zenodo.org/record/14620107/files/figure.png" pageId="9" pageNumber="8">Fig. 6B, C</figureCitation>
). The lateral delimitation of segments seems to correspond to a tubular structure enclosing the arcuate elements (
<figureCitation id="132AC3EFFA266E323AB9FB21E2FCFB11" box="[684,762,1204,1230]" captionStart="FIG" captionStartId="10.[132,143,1340,1357]" captionTargetBox="[132,1456,215,1298]" captionTargetId="figure-312@10.[132,1456,215,1298]" captionTargetPageId="10" captionText="FIG. 6. — Specimens from Casimcea surface S1: A, View in situ of the Palaeopascichnus Palij, 1976 specimen; B, photogrammetric representation of the cast showing the poorly marked ridges at the surface (black arrow), the network of arcuate meandering elements and the “empty” meandering parallel imprints (white arrow); C, detailed view showing the well-marked outline of segments,the clear superposition of segments (a), the subdivision of segments with elements increasing in size just before the subdivision (b); D, detail of S1 specimen (close up of B, white arrow) with arcuate elements and imprint with parallel lines of constant width (c. 1 cm), without internal structures. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14620107" httpUri="https://zenodo.org/record/14620107/files/figure.png" pageId="9" pageNumber="8">Fig. 6B</figureCitation>
). In several cases, a divergence of the segments occurs (
<figureCitation id="132AC3EFFA266E323ABCFB41E2FCFB31" box="[681,762,1236,1262]" captionStart="FIG" captionStartId="10.[132,143,1340,1357]" captionTargetBox="[132,1456,215,1298]" captionTargetId="figure-312@10.[132,1456,215,1298]" captionTargetPageId="10" captionText="FIG. 6. — Specimens from Casimcea surface S1: A, View in situ of the Palaeopascichnus Palij, 1976 specimen; B, photogrammetric representation of the cast showing the poorly marked ridges at the surface (black arrow), the network of arcuate meandering elements and the “empty” meandering parallel imprints (white arrow); C, detailed view showing the well-marked outline of segments,the clear superposition of segments (a), the subdivision of segments with elements increasing in size just before the subdivision (b); D, detail of S1 specimen (close up of B, white arrow) with arcuate elements and imprint with parallel lines of constant width (c. 1 cm), without internal structures. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14620107" httpUri="https://zenodo.org/record/14620107/files/figure.png" pageId="9" pageNumber="8">Fig. 6C</figureCitation>
). Just before the branching, the terminal element is wider than the preceding elements and those of the two divergent segments, up to
<quantity id="4CE9728FFA266E323904FAA6E14CFA91" box="[273,330,1331,1358]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="8" unit="cm" value="1.0">1 cm</quantity>
. In two cases, a segment can cross another but overlapping (
<figureCitation id="132AC3EFFA266E32395CFAC6E1A4FAB2" box="[329,418,1363,1389]" captionStart="FIG" captionStartId="10.[132,143,1340,1357]" captionTargetBox="[132,1456,215,1298]" captionTargetId="figure-312@10.[132,1456,215,1298]" captionTargetPageId="10" captionText="FIG. 6. — Specimens from Casimcea surface S1: A, View in situ of the Palaeopascichnus Palij, 1976 specimen; B, photogrammetric representation of the cast showing the poorly marked ridges at the surface (black arrow), the network of arcuate meandering elements and the “empty” meandering parallel imprints (white arrow); C, detailed view showing the well-marked outline of segments,the clear superposition of segments (a), the subdivision of segments with elements increasing in size just before the subdivision (b); D, detail of S1 specimen (close up of B, white arrow) with arcuate elements and imprint with parallel lines of constant width (c. 1 cm), without internal structures. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14620107" httpUri="https://zenodo.org/record/14620107/files/figure.png" pageId="9" pageNumber="8">Fig. 6C</figureCitation>
). This superposition may result from the crushing of two distinct parts of the initial body, not located in the same plane.
</paragraph>
<paragraph id="8BAEDF6AFA266E32388EFA26E2B9F992" blockId="9.[130,777,630,1613]" pageId="9" pageNumber="8">
The largest surface S2 exhibits discreet meandering imprint with parallel lines and segmented elements (
<figureCitation id="132AC3EFFA266E323A5BFA46E299FA32" box="[590,671,1491,1517]" captionStart="FIG" captionStartId="11.[132,143,1205,1222]" captionTargetBox="[136,1437,205,1177]" captionTargetId="figure-538@11.[151,1437,219,1165]" captionText="FIG. 7. — A, Specimens from Casimcea surface S2 with a meandering succession of slightly arcuate element (black arrow) poorly expressed and imprint with parallel lines of constant width (c. 1 cm), without internal structures (white arrow) similar to Fig. 6D; B, same picture as A with specimens highlighted;C, D, speci- men from Rhaman; C, meandering succession of arcuate elements (arrow) on S2 surface; D, detail of C. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14622006" httpUri="https://zenodo.org/record/14622006/files/figure.png" pageId="9" pageNumber="8">Fig. 7A</figureCitation>
). Another imprint, perpendicular to the previous one, with two parallel lines without distinguishable constitutive elements (
<figureCitation id="132AC3EFFA266E323AA5F987E2F9F9F2" box="[688,767,1554,1581]" captionStart="FIG" captionStartId="11.[132,143,1205,1222]" captionTargetBox="[136,1437,205,1177]" captionTargetId="figure-538@11.[151,1437,219,1165]" captionText="FIG. 7. — A, Specimens from Casimcea surface S2 with a meandering succession of slightly arcuate element (black arrow) poorly expressed and imprint with parallel lines of constant width (c. 1 cm), without internal structures (white arrow) similar to Fig. 6D; B, same picture as A with specimens highlighted;C, D, speci- men from Rhaman; C, meandering succession of arcuate elements (arrow) on S2 surface; D, detail of C. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14622006" httpUri="https://zenodo.org/record/14622006/files/figure.png" pageId="9" pageNumber="8">Fig. 7B</figureCitation>
) is here also present as for the S1 surface (
<figureCitation id="132AC3EFFA266E323A29F9A7E2B7F992" box="[572,689,1586,1613]" captionStart="FIG" captionStartId="10.[132,143,1340,1357]" captionTargetBox="[132,1456,215,1298]" captionTargetId="figure-312@10.[132,1456,215,1298]" captionTargetPageId="10" captionText="FIG. 6. — Specimens from Casimcea surface S1: A, View in situ of the Palaeopascichnus Palij, 1976 specimen; B, photogrammetric representation of the cast showing the poorly marked ridges at the surface (black arrow), the network of arcuate meandering elements and the “empty” meandering parallel imprints (white arrow); C, detailed view showing the well-marked outline of segments,the clear superposition of segments (a), the subdivision of segments with elements increasing in size just before the subdivision (b); D, detail of S1 specimen (close up of B, white arrow) with arcuate elements and imprint with parallel lines of constant width (c. 1 cm), without internal structures. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14620107" httpUri="https://zenodo.org/record/14620107/files/figure.png" pageId="9" pageNumber="8">Fig. 6B, D</figureCitation>
).
</paragraph>
<paragraph id="8BAEDF6AFA266E323891F9E7E139F953" blockId="9.[132,319,1650,1676]" box="[132,319,1650,1676]" pageId="9" pageNumber="8">
<heading id="D0E66806FA266E323891F9E7E139F953" box="[132,319,1650,1676]" fontSize="11" level="2" pageId="9" pageNumber="8" reason="8">
<emphasis id="B9650378FA266E323891F9E7E139F953" box="[132,319,1650,1676]" italics="true" pageId="9" pageNumber="8">Rahman specimen</emphasis>
</heading>
</paragraph>
<paragraph id="8BAEDF6AFA266E323891F907E2F7F894" blockId="9.[130,776,1682,1868]" pageId="9" pageNumber="8">
The specimen is clearly visible only over a length of about
<quantity id="4CE9728FFA266E323AD4F907E302F972" box="[705,772,1682,1709]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="8" unit="cm" value="15.0">15 cm</quantity>
. It is probably only part of a larger body fossil. It corresponds to a slightly sinuous imprint with parallel edges and constant width (
<emphasis id="B9650378FA266E3238C1F966E0E3F8D3" box="[212,229,1779,1804]" italics="true" pageId="9" pageNumber="8">c.</emphasis>
<quantity id="4CE9728FFA266E3238F8F967E12FF8D3" box="[237,297,1778,1804]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="8" unit="cm" value="1.0">1 cm</quantity>
). It is composed of millimeter thick arcuate elements in slightly marked positive relief (
<figureCitation id="132AC3EFFA266E323A51F887E2B1F8F3" box="[580,695,1810,1836]" captionStart="FIG" captionStartId="11.[132,143,1205,1222]" captionTargetBox="[136,1437,205,1177]" captionTargetId="figure-538@11.[151,1437,219,1165]" captionText="FIG. 7. — A, Specimens from Casimcea surface S2 with a meandering succession of slightly arcuate element (black arrow) poorly expressed and imprint with parallel lines of constant width (c. 1 cm), without internal structures (white arrow) similar to Fig. 6D; B, same picture as A with specimens highlighted;C, D, speci- men from Rhaman; C, meandering succession of arcuate elements (arrow) on S2 surface; D, detail of C. Photos: J. P. Saint Martin." figureDoi="http://doi.org/10.5281/zenodo.14622006" httpUri="https://zenodo.org/record/14622006/files/figure.png" pageId="9" pageNumber="8">Fig. 7C, D</figureCitation>
). These characteristics are similar to those on Casimcea S2 surface.
</paragraph>
</subSubSection>
<subSubSection id="C30B8CE1FA266E323891F8E7E5A9FD30" pageId="9" pageNumber="8" type="discussion">
<paragraph id="8BAEDF6AFA266E323891F8E7E100F855" blockId="9.[132,777,1906,2028]" box="[132,262,1906,1932]" pageId="9" pageNumber="8">
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</paragraph>
<paragraph id="8BAEDF6AFA266E323891F804E5A9FD30" blockId="9.[132,777,1906,2028]" lastBlockId="9.[811,1457,215,752]" pageId="9" pageNumber="8">
According to recent studies, the Romanian specimens have palaeopascichnid characteristics: 1) bedding plane-parallel modular fossils consisting of simple or more complex series of closely spaced millimetric circular, sausage- or kidney-shaped units (
<bibRefCitation id="EF80A29BFA266E323B64FF6DE42CFECE" author="JENSEN S. &amp; HOGSTROM A. E. S. &amp; HOYBERGET M. &amp; MEINHOLD G. &amp; MCILROY D. &amp; EBBESTAD J. O. R. &amp; TAYLOR W. L. &amp; AGIC H. &amp; PALACIOS T." box="[881,1066,247,274]" pageId="9" pageNumber="8" pagination="1253 - 1261" refId="ref10155" refString="JENSEN S., HOGSTROM A. E. S., HOYBERGET M., MEINHOLD G., MCILROY D., EBBESTAD J. O. R., TAYLOR W. L., AGIC H. &amp; PALACIOS T. 2018. - New occurrences of Palaeopascichnus from the Stahpogieddi Formation, Arctic Norway, and their bearing on the age of Varanger Ice Age. Canadian Journal of Earth Sciences 55: 1253 - 1261. https: // doi. org / 10.1139 / cjes- 2018 - 0035" type="journal article" year="2018">
Jensen
<emphasis id="B9650378FA266E323BAEFF6DE3EBFECE" box="[955,1005,247,273]" italics="true" pageId="9" pageNumber="8">et al.</emphasis>
2018
</bibRefCitation>
); 2) macroscopic modular chain-like fossils (
<bibRefCitation id="EF80A29BFA266E323B6EFE82E435FEEE" author="KOLESNIKOV A. V." box="[891,1075,279,305]" pageId="9" pageNumber="8" pagination="1 - 5" refId="ref10512" refString="KOLESNIKOV A. V. 2019. - Stratigraphic correlation potential of the Ediacaran palaeopascichnids. Estudios Geologicos 75 (2), e 102: 1 - 5. https: // doi. org / 10.3989 / egeol. 43588.557" type="book chapter" year="2019">Kolesnikov 2019</bibRefCitation>
), serially arranged, millimetre-scale allantoid chambers (
<bibRefCitation id="EF80A29BFA266E323C1BFEA2E4DAFE8E" author="HAWCO J. B. &amp; KENCHINGTON, C. G. &amp; MC &amp; ILROY D." box="[1038,1244,311,337]" pageId="9" pageNumber="8" pagination="657 - 673" refId="ref9385" refString="HAWCO J. B., KENCHINGTON, C. G. &amp; MC ILROY D. 2019. - A quantitative and statistical discrimination of morphotaxa within the Ediacaran genus Palaeopascichnus. Papers in Palaeontology 7 (2): 657 - 673. https: // doi. org / 10.1002 / spp 2.1290" type="journal article" year="2019">
Hawco
<emphasis id="B9650378FA266E323C71FEADE49FFE8E" box="[1124,1177,311,337]" italics="true" pageId="9" pageNumber="8">et al.</emphasis>
2019
</bibRefCitation>
); and 3) serially or cluster-like arranged, millimetre- to centimetre-scale globular or allantoid chambers (
<bibRefCitation id="EF80A29BFA266E323C37FEE2E564FE4E" author="KOLESNIKOV A. V. &amp; DESIATKIN V." box="[1058,1378,375,401]" pageId="9" pageNumber="8" pagination="1175 - 1191" refId="ref10554" refString="KOLESNIKOV A. V. &amp; DESIATKIN V. 2022. - Taxonomy and palaeoenvironmental distribution of palaeopascichnids. Geological Magazine 159: 1175 - 1191. https: // doi. org / 10.1017 / S 0016756822000437" type="book chapter" year="2022">Kolesnikov &amp; Desiatkin 2022</bibRefCitation>
). Morphometric and taxonomic analyses characterized and discriminated different species of
<taxonomicName id="4C11A4E9FA266E323C29FE23E4ECFE0F" authorityName="Palij" authorityYear="1976" box="[1084,1258,438,464]" genus="Palaeopascichnus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="8" rank="genus">
<emphasis id="B9650378FA266E323C29FE23E4ECFE0F" box="[1084,1258,438,464]" italics="true" pageId="9" pageNumber="8">Palaeopascichnus</emphasis>
</taxonomicName>
or morphospecies (
<bibRefCitation id="EF80A29BFA266E323B23FE42E3FBFE2F" author="HAWCO J. B. &amp; KENCHINGTON, C. G. &amp; MC &amp; ILROY D." box="[822,1021,470,497]" pageId="9" pageNumber="8" pagination="657 - 673" refId="ref9385" refString="HAWCO J. B., KENCHINGTON, C. G. &amp; MC ILROY D. 2019. - A quantitative and statistical discrimination of morphotaxa within the Ediacaran genus Palaeopascichnus. Papers in Palaeontology 7 (2): 657 - 673. https: // doi. org / 10.1002 / spp 2.1290" type="journal article" year="2019">
Hawco
<emphasis id="B9650378FA266E323B9FFE42E3B8FE2F" box="[906,958,470,496]" italics="true" pageId="9" pageNumber="8">et al.</emphasis>
2019
</bibRefCitation>
;
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<bibRefCitation id="EF80A29BFA266E323CB2FE43E5AEFE2F" author="KOLESNIKOV A. V. &amp; ROGOV V. I. &amp; BYKOVA N. V. &amp; DANELIAN T. &amp; CLAUSEN S. &amp; MASLOV A. V. &amp; GRAZHDANKIN D. V." box="[1191,1448,470,496]" pageId="9" pageNumber="8" pagination="24 - 37" refId="ref10722" refString="KOLESNIKOV A. V., ROGOV V. I., BYKOVA N. V., DANELIAN T., CLAUSEN S., MASLOV A. V. &amp; GRAZHDANKIN D. V. 2018 b. - The oldest skeletal macroscopic organism Palaeopascichnuslinearis. Precambrian Research 316: 24 - 37. https: // doi. org / 10.1016 / j. precamres. 2018.07. 017" type="book chapter" year="2018">
Kolesnikov
<emphasis id="B9650378FA266E323D32FE42E55DFE2F" box="[1319,1371,470,496]" italics="true" pageId="9" pageNumber="8">et al.</emphasis>
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<bibRefCitation id="EF80A29BFA266E323B38FE63E3E4FDCF" author="KOLESNIKOV A. V." box="[813,994,502,528]" pageId="9" pageNumber="8" pagination="1 - 5" refId="ref10512" refString="KOLESNIKOV A. V. 2019. - Stratigraphic correlation potential of the Ediacaran palaeopascichnids. Estudios Geologicos 75 (2), e 102: 1 - 5. https: // doi. org / 10.3989 / egeol. 43588.557" type="book chapter" year="2019">Kolesnikov 2019</bibRefCitation>
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Desiatkin
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). Based on these works, the Romanian specimens are here assigned to
<taxonomicName id="4C11A4E9FA266E323BE2FDA3E5B6FD8F" authority="(Fedonkin, 1976)" baseAuthorityName="Fedonkin" baseAuthorityYear="1976" box="[1015,1456,566,592]" genus="Palaeopascichnus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="8" rank="species" species="linearis">
<emphasis id="B9650378FA266E323BE2FDA3E4F6FD8F" box="[1015,1264,566,592]" italics="true" pageId="9" pageNumber="8">Palaeopascichnus linearis</emphasis>
(
<bibRefCitation id="EF80A29BFA266E323CE8FDA3E5A1FD8F" author="FEDONKIN M. A." box="[1277,1447,566,592]" pageId="9" pageNumber="8" pagination="129 - 32" refId="ref8075" refString="FEDONKIN M. A. 1976. - Sledy mnogokletotchnykh iz valdaiskoi serii. Izvestiya Akademii Nauk SSSR, Seriya Geologicheskaya 4: 129 - 32 (in Russian)." type="journal article" year="1976">Fedonkin, 1976</bibRefCitation>
)
</taxonomicName>
because of their relative consistency in the size of arcuate elements (considered as chambers) throughout the series, which is not seen in the
<typeStatus id="54AA61C8FA266E323BF5FD03E408FD6F" box="[992,1038,662,688]" pageId="9" pageNumber="8">type</typeStatus>
species.
<taxonomicName id="4C11A4E9FA266E323C7CFD03E56EFD6F" baseAuthorityName="Fedonkin" baseAuthorityYear="1976" box="[1129,1384,662,688]" genus="Palaeopascichnus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="8" rank="species" species="linearis">
<emphasis id="B9650378FA266E323C7CFD03E56EFD6F" box="[1129,1384,662,688]" italics="true" pageId="9" pageNumber="8">Palaeopascichnus linearis</emphasis>
</taxonomicName>
shares common characters with the
<typeStatus id="54AA61C8FA266E323C48FD23E48DFD0F" box="[1117,1163,694,720]" pageId="9" pageNumber="8">type</typeStatus>
species, such as the possible branching of the series, as observed in the Romanian specimens.
</paragraph>
</subSubSection>
</treatment>
</document>