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<mods:title id="EF6912BCD99A4203C38D95D62AD06E2C">A review of subtribe Cymindidina Laporte, 1834 (Coleoptera: Carabidae: Lebiini) in Southwestern Saudi Arabia, with descriptions of two new species</mods:title>
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<treatment id="03B84B653D30FFB604F9C8C5F5ECFD61" ID-DOI="http://doi.org/10.5281/zenodo.6000051" ID-GBIF-Taxon="127696736" ID-Zenodo-Dep="6000051" LSID="urn:lsid:plazi:treatment:03B84B653D30FFB604F9C8C5F5ECFD61" httpUri="http://treatment.plazi.org/id/03B84B653D30FFB604F9C8C5F5ECFD61" lastPageNumber="168" pageId="11" pageNumber="168">
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<heading id="D0E64D1F3D30FFB604F9C8C5F7DEFF15" bold="true" box="[151,453,151,178]" fontSize="11" level="1" pageId="11" pageNumber="168" reason="1">
<taxonomicName id="4C1181F03D30FFB604F9C8C5F7DEFF15" ID-CoL="8KVPV" authority="Latreille, 1806" authorityName="Latreille" authorityYear="1806" box="[151,453,151,178]" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B96526613D30FFB604F9C8C5F7DEFF15" bold="true" box="[151,453,151,178]" pageId="11" pageNumber="168">
<emphasis id="B96526613D30FFB604F9C8C5F712FF16" bold="true" box="[151,265,151,177]" italics="true" pageId="11" pageNumber="168">Cymindis</emphasis>
Latreille, 1806
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<paragraph id="8BAEFA733D30FFB604F9C88CF48FFF53" blockId="11.[151,660,222,245]" box="[151,660,222,245]" pageId="11" pageNumber="168">
<emphasis id="B96526613D30FFB604F9C88CF739FF53" bold="true" box="[151,290,222,244]" pageId="11" pageNumber="168">Type species.</emphasis>
<taxonomicName id="4C1181F03D30FFB60547C8B2F48FFF53" authority="Geoffroy, 1785" authorityName="Geoffroy" authorityYear="1785" box="[297,660,222,245]" class="Insecta" family="Buprestidae" genus="Buprestis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="B96526613D30FFB60547C8B2F7E8FF53" box="[297,499,222,245]" italics="true" pageId="11" pageNumber="168">Buprestis humeralis</emphasis>
Geoffřoy, 1785
</taxonomicName>
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<paragraph id="8BAEFA733D30FFB604F9C973F5ECFD61" blockId="11.[151,1437,289,710]" pageId="11" pageNumber="168">
<taxonomicName id="4C1181F03D30FFB604F9C973F7B0FE9D" ID-CoL="8KVPV" authority="Latreille, 1806" authorityName="Latreille" authorityYear="1806" box="[151,427,289,314]" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B96526613D30FFB604F9C973F71AFE9E" box="[151,257,289,313]" italics="true" pageId="11" pageNumber="168">Cymindis</emphasis>
Latreille, 1806
</taxonomicName>
is the largest genus in subtribe
<taxonomicName id="4C1181F03D30FFB60766C973F581FE9D" ID-CoL="8LTBS" box="[776,922,289,314]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="subTribe" subTribe="Cymindidina">Cymindidina</taxonomicName>
with more than 220 taxa in 15 subgenera. It is the most speciose and widespread genus that extends to Nearctic, Neotropical, and Palaearctic zoogeographical regions (
<bibRefCitation id="EF8087823D30FFB60494C93BF7B7FE26" author="Bousquet" box="[250,428,361,385]" pageId="11" pageNumber="168" refString="Bousquet, Y. (2012) Catalogue of Geadephaga (Coleoptera; Adephaga) of America, north of Mexico. ZooKeys, 245, 1 - 1772. https: // doi. org / 10.3897 / zookeys. 245.3416" type="journal article" year="2012">Bousquet 2012</bibRefCitation>
), with exception of
<collectingCountry id="F306BAE33D30FFB606FBC93BF4E7FE25" box="[661,764,361,386]" name="Australia" pageId="11" pageNumber="168">Australia</collectingCountry>
(
<bibRefCitation id="EF8087823D30FFB60763C93BF5E4FE26" author="Ball" box="[781,1023,361,386]" pageId="11" pageNumber="168" refString="Ball, G. E &amp; Hilchie, G. J. (1983) Cymindine Lebiini of Authors: Redefinition and Reclassification of Genera (Coleoptera: Carabidae). Quaestiones Entomologicae, 19, 93 - 216." type="journal article" year="1983">Ball &amp; Hilchie 1983</bibRefCitation>
). The Palaearctic region contributes most of the diversity with 167 species (
<bibRefCitation id="EF8087823D30FFB6060BC9DFF4EDFE01" author="Kabak" box="[613,758,397,422]" pageId="11" pageNumber="168" refString="Kabak, I. (2003) Tribe Lebiini Bonelli, 1810 (Coleoptera; Carabidae). In: Catalogue of Palaearctic Coleoptera, Archostemata, Myxophaga, Adephaga. Fol. 1. Apollo Books Stenstrup, pp. 408 - 443." type="journal article" year="2003">Kabak 2003</bibRefCitation>
;
<bibRefCitation id="EF8087823D30FFB6076AC9DCF5AEFE01" author="Bousquet" box="[772,949,398,422]" pageId="11" pageNumber="168" refString="Bousquet, Y. (2012) Catalogue of Geadephaga (Coleoptera; Adephaga) of America, north of Mexico. ZooKeys, 245, 1 - 1772. https: // doi. org / 10.3897 / zookeys. 245.3416" type="journal article" year="2012">Bousquet 2012</bibRefCitation>
). The members of the subgenus
<taxonomicName id="4C1181F03D30FFB6015CC9DFF387FE02" ID-CoL="8KVPV" box="[1330,1436,397,421]" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B96526613D30FFB6015CC9DFF387FE02" box="[1330,1436,397,421]" italics="true" pageId="11" pageNumber="168">Cymindis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2569B1A3D30FFB604F9C9E0F73FFE6D" box="[151,292,433,458]" pageId="11" pageNumber="168" sensu="stricto">sensu stricto</taxonomicNameLabel>
from the South of the Arabian Peninsula can be separated from these genera and subgenera by: last labial palpomeres dilated and truncate at apex; temples pubescent; pronotum cordiform; elytra with complete striae; wings usually reduced; males with 13 dilated protarsomeres and with serrate adhesive setae underneath; tarsal claws pectinate (
<bibRefCitation id="EF8087823D30FFB605C8CA4FF449FD91" author="Lindroth" box="[422,594,541,566]" pageId="11" pageNumber="168" refString="Lindroth, C. H. (1974) Handbook for the identification of British Insects. Royal Entomological Society 2012, 4 (2). London. Available from: http: // www. royensoc. co. uk / publications / index. htm (accessed 10 January 2017)" type="journal article" year="1974">Lindroth 1974</bibRefCitation>
;
<bibRefCitation id="EF8087823D30FFB6060ACA4FF546FD91" author="Ball" box="[612,861,541,566]" pageId="11" pageNumber="168" refString="Ball, G. E &amp; Hilchie, G. J. (1983) Cymindine Lebiini of Authors: Redefinition and Reclassification of Genera (Coleoptera: Carabidae). Quaestiones Entomologicae, 19, 93 - 216." type="journal article" year="1983">Ball &amp; Hilchie 1983</bibRefCitation>
: Bousquet 2010). Hitherto only three species of
<taxonomicName id="4C1181F03D30FFB604F9CA13F71AFDFE" ID-CoL="8KVPV" box="[151,257,577,601]" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B96526613D30FFB604F9CA13F71AFDFE" box="[151,257,577,601]" italics="true" pageId="11" pageNumber="168">Cymindis</emphasis>
</taxonomicName>
have been recorded from
<collectingCountry id="F306BAE33D30FFB60648CA13F4A7FDFD" box="[550,700,577,602]" name="Saudi Arabia" pageId="11" pageNumber="168">Saudi Arabia</collectingCountry>
,
<taxonomicName id="4C1181F03D30FFB606A9CA13F5E7FDFE" ID-CoL="8MGK5" authority="Menetries 1832" authorityName="Menetries" authorityYear="1832" box="[711,1020,577,602]" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="species" species="andreae">
<emphasis id="B96526613D30FFB606A9CA13F558FDFE" box="[711,835,577,601]" italics="true" pageId="11" pageNumber="168">C. andreae</emphasis>
Ménétriés 1832
</taxonomicName>
, this species was recently transferred to the subgenus
<taxonomicName id="4C1181F03D30FFB6053CCA37F56BFDD9" authority="Motschulsky 1864 (Sundukov, 2005)" authorityName="Motschulsky 1864 (Sundukov" authorityYear="2005" box="[338,880,613,638]" class="Insecta" family="Carabidae" genus="Arrhostus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B96526613D30FFB6053CCA37F7DBFDDA" box="[338,448,613,637]" italics="true" pageId="11" pageNumber="168">Arrhostus</emphasis>
Motschulsky 1864 (
<bibRefCitation id="EF8087823D30FFB606C4CA37F573FDD9" author="Sundukov" box="[682,872,613,638]" pageId="11" pageNumber="168" refString="Sundukov, Yu. N. (2005) Arrhostus Motschulsky is a new subgenus of the genus Cymindis (Coleoptera, Carabidae). Russian Journal of Zoology, 84, 395 - 396. [in Russian]" type="journal article" year="2005">Sundukov, 2005</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C1181F03D30FFB60713CA34F2E1FDD9" ID-CoL="33757" authority="Dejean, 1825" authorityName="Dejean" authorityYear="1825" box="[893,1274,613,638]" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B96526613D30FFB60713CA34F58CFDDA" box="[893,919,614,637]" italics="true" pageId="11" pageNumber="168">C.</emphasis>
(
<emphasis id="B96526613D30FFB607C7CA34F5FDFDDA" box="[937,998,614,637]" italics="true" pageId="11" pageNumber="168">s. str.</emphasis>
)
<emphasis id="B96526613D30FFB60796CA37F241FDDA" box="[1016,1114,613,637]" italics="true" pageId="11" pageNumber="168">suturalis</emphasis>
<bibRefCitation id="EF8087823D30FFB6000DCA37F2E1FDD9" author="Dejean" box="[1123,1274,613,638]" pageId="11" pageNumber="168" refString="Dejean, P. F. M. A. (1825) Species general des Coleopteres, de la collection de M. le Comte Dejean. Tome Premier. Crevot, Paris, sxx + 463." type="book chapter" year="1825">Dejean, 1825</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EF8087823D30FFB60164CA34F38CFDD9" author="Mateu" box="[1290,1431,613,638]" pageId="11" pageNumber="168" refString="Mateu, J. (1986) Lebiinae and Brachininae of Saudi Arabia (Coleoptera: Carabidae). Fauna of Saudi Arabia, 8, 198 - 217." type="journal article" year="1986">Mateu 1986</bibRefCitation>
;
<bibRefCitation id="EF8087823D30FFB604F9CADBF7BAFD05" author="El-Hawagry" box="[151,417,649,674]" pageId="11" pageNumber="168" refString="El-Hawagry, M. S., Khalil, M. W., Sharaf, M. R., Fadl, H. H. &amp; Aldawood, A. S. (2013) A preliminary study on the insect fauna of Al-Baha Province, Saudi Arabia, with descriptions of two new species. ZooKeys, 274, 1 - 88." type="journal article" year="2013">
El-Hawagry
<emphasis id="B96526613D30FFB60549CAD8F742FD06" box="[295,345,649,673]" italics="true" pageId="11" pageNumber="168">et al</emphasis>
. 2013
</bibRefCitation>
) and
<taxonomicName id="4C1181F03D30FFB6058ECAD8F25EFD05" authority="Lucas, 1846 (Basilewsky 1979)" authorityName="Lucas, 1846 (Basilewsky" authorityYear="1979" box="[480,1093,649,674]" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="species" species="laevistriata">
<emphasis id="B96526613D30FFB6058ECAD8F7E1FD06" box="[480,506,650,673]" italics="true" pageId="11" pageNumber="168">C.</emphasis>
(
<emphasis id="B96526613D30FFB60665CAD8F45DFD06" box="[523,582,650,673]" italics="true" pageId="11" pageNumber="168">s. str.</emphasis>
)
<emphasis id="B96526613D30FFB60638CADBF4CEFD06" box="[598,725,649,673]" italics="true" pageId="11" pageNumber="168">laevistriata</emphasis>
Lucas, 1846 (
<bibRefCitation id="EF8087823D30FFB60718CADBF227FD05" author="Basilewsky" box="[886,1084,649,674]" pageId="11" pageNumber="168" refString="Basilewsky, P. (1979) Insects of Saudi Arabia, Coleoptera: Fam. Carabidae. Fauna of Saudi Arabia, 1, 141 - 146." type="journal article" year="1979">Basilewsky 1979</bibRefCitation>
)
</taxonomicName>
. During the present work only
<taxonomicName id="4C1181F03D30FFB604F9CAFCF709FD62" ID-CoL="8MGK5" box="[151,274,685,709]" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="species" species="andreae">
<emphasis id="B96526613D30FFB604F9CAFCF709FD62" box="[151,274,685,709]" italics="true" pageId="11" pageNumber="168">C. andreae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1181F03D30FFB60527CAFCF7D0FD62" ID-CoL="33757" box="[329,459,685,709]" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="168" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B96526613D30FFB60527CAFCF7D0FD62" box="[329,459,685,709]" italics="true" pageId="11" pageNumber="168">C. suturalis</emphasis>
</taxonomicName>
were collected in the southwestern
<collectingCountry id="F306BAE33D30FFB60733CAFFF5E8FD61" box="[861,1011,685,710]" name="Saudi Arabia" pageId="11" pageNumber="168">Saudi Arabia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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