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<document id="C5390EB394D1CBF59B50981F583C96B4" ID-CLB-Dataset="278359" ID-DOI="10.11646/zootaxa.5361.1.6" ID-GBIF-Dataset="969de1fd-69d2-497e-9a19-c39ecea9b64d" ID-ISSN="1175-5326" ID-Zenodo-Dep="10150590" ID-ZooBank="33F819C3-F34D-431B-B262-C8FDBFD5FF4F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1700247603070" checkinUser="plazi" docAuthor="Yang, Qiao-Zhi, Geshev, Georgi &amp; Pathomwattananurak, Wuttipon" docDate="2023" docId="03B8222866745502FF75FA20FBE5F8E5" docLanguage="en" docName="zootaxa.5361.1.6.pdf" docOrigin="Zootaxa 5361 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudohomonyx continentalis Yang &amp; Pathomwattananurak 2023, new species" docType="treatment" docVersion="9" lastPageNumber="117" masterDocId="FF815A5066755501FFE2FFDBFFD6FFBF" masterDocTitle="A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake &amp; Yamaya, 1997 male and a catalogue of the genus" masterLastPageNumber="124" masterPageNumber="114" pageNumber="115" updateTime="1701279501061" updateUser="julia" zenodo-license-document="CLOSED">
<mods:mods id="7D75E133FE1E08DCA6BC387D08257C67" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="86B8EAD4D1A0E67DD7D90A2356F4F17E">
<mods:title id="3DFBAE47BB02CF0E69553E8DD7D7BAD4">A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake &amp; Yamaya, 1997 male and a catalogue of the genus</mods:title>
</mods:titleInfo>
<mods:name id="172747099C7BC14BEB5AFAAE167224DB" type="personal">
<mods:role id="98024656FAF548D3C076C8D7B9EE7510">
<mods:roleTerm id="E992B83481C2979188537F1529053FD2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8F2EEF1E20C8DC1E71DC3844713007F7">Yang, Qiao-Zhi</mods:namePart>
</mods:name>
<mods:name id="72D05129492AD60EADA63D2FF32F1CA1" type="personal">
<mods:role id="FAB430B18A9C2A23C1AB3E6FB93EFF16">
<mods:roleTerm id="A86931C41A0061D7CFAAEAF0C599577F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D5C0DCEF97F7D2CFF7BFC24D1A40A4E2">Geshev, Georgi</mods:namePart>
<mods:affiliation id="8063C71EC474E412E101FD66B50B61BC">3 B, Pingguoshequ, Baiziwan Road, Chaoyang Dist., Beijing, 100022, P. R. China</mods:affiliation>
</mods:name>
<mods:name id="B8E125B0AA8C48181A408AD43D764C01" type="personal">
<mods:role id="368EBCF4360ACC15752C683DABD2DC78">
<mods:roleTerm id="C578D6A812904E19B67B148DC3A7BEA9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C055553F2A2D44CEF5E672A9D58147E6">Pathomwattananurak, Wuttipon</mods:namePart>
<mods:affiliation id="8C2C07F617CD050BD0FF371DDE7A810F">1051 / 38 Wiang, Mueang Dist., Chiang Rai Province, Thailand 57000</mods:affiliation>
</mods:name>
<mods:typeOfResource id="18161B0467E79A7D2445E6CEC620542B">text</mods:typeOfResource>
<mods:relatedItem id="B762DE7190419A0C55FC7FC62DB2D3EF" type="host">
<mods:titleInfo id="48F725571DBDCDD05E9A2091FDB160CD">
<mods:title id="82B82EA2B576FBBCA2E965D2A14CA426">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="64495F8E2CB251426419C0EAA928F7B6">
<mods:date id="48841775296B2FAC026EBDFF6182D45E">2023</mods:date>
<mods:detail id="C9668BAF73662DDA8F430A9FC1BB50CE" type="pubDate">
<mods:number id="AE59E99C5C2F25A8338B11D4FDB48DD1">2023-10-31</mods:number>
</mods:detail>
<mods:detail id="6916F40D6A630C37C0122D77EBA02B23" type="volume">
<mods:number id="398DF0ABC31D25C95511BFF2632F8520">5361</mods:number>
</mods:detail>
<mods:detail id="ED7E011AA285DEF66EF204E5B52688AE" type="issue">
<mods:number id="54C7F2E092B53667CE4D0A4F5CDF603A">1</mods:number>
</mods:detail>
<mods:extent id="EF85286BF68D2B65A1710F20173819A1" unit="page">
<mods:start id="0F8B7CA5EC2016D80E0A77729CE4AA5C">114</mods:start>
<mods:end id="A2CEE9F980D32D93B9BDA1B558406454">124</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:location id="20D04CC5AF58AC0538C52B9A91030324">
<mods:url id="A127D088A62054AA59E2A087B274F6BE">https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164</mods:url>
</mods:location>
<mods:classification id="F599765362F0FBD21F341ED8830775B2">journal article</mods:classification>
<mods:identifier id="BE87AA4DC6A9D4A2D3CC3C475E7F2E85" type="CLB-Dataset">278359</mods:identifier>
<mods:identifier id="D48B0D48E281145C16095527C5FD24E5" type="DOI">10.11646/zootaxa.5361.1.6</mods:identifier>
<mods:identifier id="0E896EC4662D4A4DDF0F7D65E39564A2" type="GBIF-Dataset">969de1fd-69d2-497e-9a19-c39ecea9b64d</mods:identifier>
<mods:identifier id="5F9472D2C4C07C0887F67D6CF157B9AC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="846F86BADC3FAEAB8D6EA065CDE58999" type="Zenodo-Dep">10150590</mods:identifier>
<mods:identifier id="FC879658AA31076A5B009C6F6AF64060" type="ZooBank">33F819C3-F34D-431B-B262-C8FDBFD5FF4F</mods:identifier>
</mods:mods>
<treatment id="03B8222866745502FF75FA20FBE5F8E5" ID-GBIF-Taxon="217755405" LSID="urn:lsid:plazi:treatment:03B8222866745502FF75FA20FBE5F8E5" httpUri="http://treatment.plazi.org/id/03B8222866745502FF75FA20FBE5F8E5" lastPageId="3" lastPageNumber="117" pageId="1" pageNumber="115">
<subSubSection id="C30BC0B566745500FF75FA20FBF5F9AA" box="[151,1059,1531,1558]" pageId="1" pageNumber="115" type="nomenclature">
<paragraph id="8BAE933E66745500FF75FA20FBF5F9AA" blockId="1.[151,1059,1531,1600]" box="[151,1059,1531,1558]" pageId="1" pageNumber="115">
<heading id="D0E6245266745500FF75FA20FBF5F9AA" bold="true" box="[151,1059,1531,1558]" fontSize="11" level="1" pageId="1" pageNumber="115" reason="1">
<emphasis id="B9654F2C66745500FF75FA20FBF5F9AA" bold="true" box="[151,1059,1531,1558]" pageId="1" pageNumber="115">
<taxonomicName id="4C11E8BD66745500FF75FA20FC50F9AA" ID-CoL="BV63M" authority="Yang &amp; Pathomwattananurak" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2023" box="[151,902,1531,1558]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="115" phylum="Arthropoda" rank="species" species="continentalis" status="sp. nov.">
<emphasis id="B9654F2C66745500FF75FA20FDD5F9A9" bold="true" box="[151,515,1531,1558]" italics="true" pageId="1" pageNumber="115">Pseudohomonyx continentalis</emphasis>
Yang &amp; Pathomwattananurak
</taxonomicName>
,
<taxonomicNameLabel id="A256F25766745500FC76FA20FBF5F9AA" box="[916,1059,1531,1557]" pageId="1" pageNumber="115" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30BC0B566745500FF75F9FDFEEEF9FF" box="[151,312,1574,1600]" pageId="1" pageNumber="115" type="description">
<paragraph id="8BAE933E66745500FF75F9FDFEEEF9FF" blockId="1.[151,1059,1531,1600]" box="[151,312,1574,1600]" pageId="1" pageNumber="115">
(
<figureCitation id="132A8FBB66745500FF7DF9FDFEF9F9FF" box="[159,303,1574,1600]" captionStart="FIGURE 1" captionStartId="2.[151,250,1975,1999]" captionTargetBox="[215,1370,718,1935]" captionTargetId="figure-254@2.[215,1373,709,1942]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Pseudohomonyx continentalis Yang &amp; Pathomwattananurak, new species.A, habitus (dorsal view); B, habitus (ventral view); C, head and pronotum; D, parameres (caudal view); E, aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10150592" httpUri="https://zenodo.org/record/10150592/files/figure.png" pageId="1" pageNumber="115">Figs. 1A–1E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30BC0B566745500FF75F9AEFAB7F944" pageId="1" pageNumber="115" type="materials_examined">
<paragraph id="8BAE933E66745500FF75F9AEFAB7F944" blockId="1.[151,1437,1652,2039]" pageId="1" pageNumber="115">
<emphasis id="B9654F2C66745500FF75F9AEFEEBF931" bold="true" box="[151,317,1652,1679]" pageId="1" pageNumber="115">Type material</emphasis>
(2♁♁,
<specimenCount id="9D1758B766745500FE70F9AEFE62F930" box="[402,436,1653,1679]" count="1" pageId="1" pageNumber="115" type="female">1♀</specimenCount>
).
<materialsCitation id="3B79996366745500FE2EF9AEFEA5F90D" ID-GBIF-Occurrence="4440434320" collectingDate="2022-04" collectionCode="CAU" country="Malaysia" location="Malaysia" municipality="Genting Highlands" pageId="1" pageNumber="115" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
<emphasis id="B9654F2C66745500FE2EF9AEFD69F931" bold="true" box="[460,703,1652,1679]" pageId="1" pageNumber="115">
<typeStatus id="54AA2D9C66745500FE2EF9AEFDEBF930" box="[460,573,1653,1679]" pageId="1" pageNumber="115" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F306D3AE66745500FDABF9AFFD6CF931" box="[585,698,1652,1678]" name="Malaysia" pageId="1" pageNumber="115">Malaysia</collectingCountry>
:
</emphasis>
♁ (
<collectionCode id="ED000BFB66745500FD0BF9AEFCF0F930" box="[745,806,1653,1679]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="1" pageNumber="115">CAU</collectionCode>
<emphasis id="B9654F2C66745500FCC7F9AFFCF8F931" bold="true" box="[805,814,1652,1678]" pageId="1" pageNumber="115">)</emphasis>
, “
<collectingMunicipality id="6BCA094466745500FCA4F9AFFBC9F931" box="[838,1055,1652,1678]" pageId="1" pageNumber="115">Genting Highlands</collectingMunicipality>
,
<collectingRegion id="49D55DDC66745500FBC8F9AFFB68F930" box="[1066,1214,1652,1679]" country="Malaysia" name="Pahang" pageId="1" pageNumber="115">Pahang State</collectingRegion>
,
<location id="8ECEC5E566745500FB2AF9AFFAE7F931" LSID="urn:lsid:plazi:treatment:03B8222866745502FF75FA20FBE5F8E5:8ECEC5E566745500FB2AF9AFFAE7F931" box="[1224,1329,1652,1678]" country="Malaysia" municipality="Genting Highlands" name="Malaysia" pageId="1" pageNumber="115" stateProvince="Pahang">Malaysia</location>
,
<date id="FFAFB5FE66745500FADEF9AEFA4EF930" box="[1340,1432,1653,1679]" pageId="1" pageNumber="115" value="2022-04">
<collectingDate id="EFEB4C1666745500FADEF9AEFA4EF930" box="[1340,1432,1653,1679]" pageId="1" pageNumber="115" value="2022-04">IV.2022</collectingDate>
</date>
, local collector leg.”
</materialsCitation>
<materialsCitation id="3B79996366745500FE98F942FBD0F968" ID-GBIF-Occurrence="4440434325" collectingDate="1986-04-15" collectingDateMax="2023-05-19" collectingDateMin="1986-04-15" collectionCode="THNHM" collectorName="J. Nabhitabhata &amp; Chayaphol Photaworn" country="Thailand" county="Ban Na Sarn Dist." location="Nakhon Sri Thammarat Prov." municipality="Nopphitam Dist." pageId="1" pageNumber="115" specimenCount="1" specimenCount-female="1" stateProvince="Surat Thani" typeStatus="paratype">
<emphasis id="B9654F2C66745500FE98F942FDAEF90D" bold="true" box="[378,632,1688,1715]" pageId="1" pageNumber="115">
<typeStatus id="54AA2D9C66745500FE98F942FE20F90C" box="[378,502,1689,1715]" pageId="1" pageNumber="115" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F306D3AE66745500FDE0F943FDA5F90D" box="[514,627,1688,1714]" name="Thailand" pageId="1" pageNumber="115">Thailand</collectingCountry>
:
</emphasis>
1♁ (
<collectionCode id="ED000BFB66745500FD53F942FCC8F90C" box="[689,798,1689,1715]" pageId="1" pageNumber="115">THNHM</collectionCode>
), “S Thailand:
<collectingRegion id="49D55DDC66745500FC27F942FB5AF90C" box="[965,1164,1688,1715]" country="Thailand" name="Surat Thani" pageId="1" pageNumber="115">Surat Thani Prov.</collectingRegion>
,
<collectingCounty id="62CFEBB266745500FB75F942FAB0F90D" box="[1175,1382,1688,1715]" pageId="1" pageNumber="115">Ban Na Sarn Dist.</collectingCounty>
, Plai Nam Subdist.,
<date id="FFAFB5FE66745500FEA6F967FDC5F969" box="[324,531,1724,1751]" pageId="1" pageNumber="115" value="1986-04-15" valueMax="1986-04-28" valueMin="1986-04-15">
<collectingDate id="EFEB4C1666745500FEA6F967FDC5F969" box="[324,531,1724,1751]" pageId="1" pageNumber="115" value="1986-04-15" valueMax="1986-04-28" valueMin="1986-04-15">15–28 April 1986</collectingDate>
</date>
,
<collectorName id="26E4F6E866745500FDC3F966FD04F969" box="[545,722,1724,1751]" pageId="1" pageNumber="115">J. Nabhitabhata</collectorName>
leg. // THNHM-I-10375”;
</materialsCitation>
<materialsCitation id="3B79996366745500FBF3F966FAB7F944" ID-GBIF-Occurrence="4440434306" collectingDate="1986-04-15" collectingDateMax="2023-05-19" collectingDateMin="1986-04-15" collectionCode="THNHM" collectorName="J. Nabhitabhata &amp; Chayaphol Photaworn" country="Thailand" county="Ban Na Sarn Dist." location="Nakhon Sri Thammarat Prov." municipality="Nopphitam Dist." pageId="1" pageNumber="115" specimenCount="1" specimenCount-female="1" stateProvince="Surat Thani" typeStatus="paratype">
<specimenCount id="9D1758B766745500FBF3F966FBE2F968" box="[1041,1076,1725,1751]" count="1" pageId="1" pageNumber="115" type="female">1♀</specimenCount>
(
<collectionCode id="ED000BFB66745500FBA4F966FB65F968" box="[1094,1203,1725,1751]" pageId="1" pageNumber="115">THNHM</collectionCode>
), “larva collected in 2023,
<collectingMunicipality id="6BCA094466745500FF3FF93BFE4DF945" box="[221,411,1760,1786]" pageId="1" pageNumber="115">Nopphitam Dist.</collectingMunicipality>
,
<location id="8ECEC5E566745500FE44F93BFD24F944" LSID="urn:lsid:plazi:treatment:03B8222866745502FF75FA20FBE5F8E5:8ECEC5E566745500FE44F93BFD24F944" box="[422,754,1760,1787]" country="Thailand" county="Ban Na Sarn Dist." municipality="Nopphitam Dist." name="Nakhon Sri Thammarat Prov." pageId="1" pageNumber="115" stateProvince="Surat Thani">Nakhon Sri Thammarat Prov.</location>
,
<collectorName id="26E4F6E866745500FD1FF93BFC20F945" box="[765,1014,1760,1786]" pageId="1" pageNumber="115">Chayaphol Photaworn</collectorName>
leg., adult emerged
<date id="FFAFB5FE66745500FB38F93AFA84F944" box="[1242,1362,1761,1787]" pageId="1" pageNumber="115" value="2023-05-19">
<collectingDate id="EFEB4C1666745500FB38F93AFA84F944" box="[1242,1362,1761,1787]" pageId="1" pageNumber="115" value="2023-05-19">19.V.2023</collectingDate>
</date>
.”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30BC0B566745502FF25F8DFFD3CFB48" lastPageId="3" lastPageNumber="117" pageId="1" pageNumber="115" type="description">
<paragraph id="8BAE933E66745503FF25F8DFFC29FD16" blockId="1.[151,1437,1652,2039]" lastBlockId="2.[151,1437,151,681]" lastPageId="2" lastPageNumber="116" pageId="1" pageNumber="115">
<emphasis id="B9654F2C66745500FF25F8DFFE87F8A1" bold="true" box="[199,337,1796,1822]" pageId="1" pageNumber="115">Description</emphasis>
(
<typeStatus id="54AA2D9C66745500FE82F8DFFE12F8A1" box="[352,452,1796,1822]" pageId="1" pageNumber="115" type="holotype">holotype</typeStatus>
male).
<emphasis id="B9654F2C66745500FDFAF8DFFDAEF8A1" bold="true" box="[536,632,1796,1822]" pageId="1" pageNumber="115">General</emphasis>
(
<figureCitation id="132A8FBB66745500FD6AF8DFFCCFF8A0" box="[648,793,1796,1823]" captionStart="FIGURE 1" captionStartId="2.[151,250,1975,1999]" captionTargetBox="[215,1370,718,1935]" captionTargetId="figure-254@2.[215,1373,709,1942]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Pseudohomonyx continentalis Yang &amp; Pathomwattananurak, new species.A, habitus (dorsal view); B, habitus (ventral view); C, head and pronotum; D, parameres (caudal view); E, aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10150592" httpUri="https://zenodo.org/record/10150592/files/figure.png" pageId="1" pageNumber="115">Figs. 1A–1B</figureCitation>
): Body dark brown, surface mildly shiny; ventral surface covered with dense, long reddish-brown setae.
<emphasis id="B9654F2C66745500FD51F8F2FD25F8FC" bold="true" box="[691,755,1833,1859]" pageId="1" pageNumber="115">Head</emphasis>
(
<figureCitation id="132A8FBB66745500FCE6F8F3FC8DF8FC" box="[772,859,1832,1859]" captionStart="FIGURE 1" captionStartId="2.[151,250,1975,1999]" captionTargetBox="[215,1370,718,1935]" captionTargetId="figure-254@2.[215,1373,709,1942]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Pseudohomonyx continentalis Yang &amp; Pathomwattananurak, new species.A, habitus (dorsal view); B, habitus (ventral view); C, head and pronotum; D, parameres (caudal view); E, aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10150592" httpUri="https://zenodo.org/record/10150592/files/figure.png" pageId="1" pageNumber="115">Fig. 1C</figureCitation>
): Surface mostly dark brown. Clypeus with dense transverse rugopunctures throughout; clypeal apex almost flat; lateral and apical margins of clypeus bordered. Frons with transverse rugopunctures throughout. Transverse carina separating frons and clypeus indicated by an obtuse tubercle at each side. Mandible at outer margin rounded near apex, almost straight at middle.
<emphasis id="B9654F2C66745500FB01F84EFA8AF810" bold="true" box="[1251,1372,1941,1967]" pageId="1" pageNumber="115">Pronotum</emphasis>
(
<figureCitation id="132A8FBB66745500FA8DF84FFF6EF86C" captionStart="FIGURE 1" captionStartId="2.[151,250,1975,1999]" captionTargetBox="[215,1370,718,1935]" captionTargetId="figure-254@2.[215,1373,709,1942]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Pseudohomonyx continentalis Yang &amp; Pathomwattananurak, new species.A, habitus (dorsal view); B, habitus (ventral view); C, head and pronotum; D, parameres (caudal view); E, aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10150592" httpUri="https://zenodo.org/record/10150592/files/figure.png" pageId="1" pageNumber="115">Fig. 1C</figureCitation>
): Surface dark brown; sparsely micropunctate throughout, only with a few slightly larger punctures sparsely distributed near lateral margin. impression absent. Side roundly curved, widest near posterior one-fourth; all margins emarginate except narrowly interrupted in lateral margin; anterior margin without any tubercle at middle. Anterior and posterior angles rounded.
<emphasis id="B9654F2C66775503FE0AFF61FDBFFF6B" bold="true" box="[488,617,186,212]" pageId="2" pageNumber="116">Scutellum:</emphasis>
Surface dark brown; sparsely punctate near base.
<emphasis id="B9654F2C66775503FB7AFF60FB3BFF6A" bold="true" box="[1176,1261,187,213]" pageId="2" pageNumber="116">Elytra:</emphasis>
Surface mostly dark brown, except dark reddish sutural ridge; with sparse micropunctures and dense, round, large punctures in striae. Sutural striae narrow, deeply sulcate; other striae wider, even intervals slightly wider than their adjacent odd intervals. Intervals micropunctate, with dense, larger punctures on fourth and sixth intervals posteriorly. Apex densely punctured. Humeral and apical calli obtuse. Epipleuron of elytron moderately dilated horizontally from base to posterior one-fifth, widest behind middle.
<emphasis id="B9654F2C66775503FD74FEB4FC97FE36" bold="true" box="[662,833,367,393]" pageId="2" pageNumber="116">Metasternum:</emphasis>
Surface with dense setae; punctures small and dense close to sides; median region with sparser and smaller, with a longitudinal depression.
<emphasis id="B9654F2C66775503FB8DFE49FB3BFE13" bold="true" box="[1135,1261,402,428]" pageId="2" pageNumber="116">Abdomen:</emphasis>
Pygidium with dark brown surface, slightly convex, with very dense punctures throughout. Abdominal ventrites with dark brown surface, with reddish anterior margin; with sparse micropunctures, larger punctures near base and longitudinal midline, and a row of punctures with long setae at sides.
<emphasis id="B9654F2C66775503FCE8FE24FC9AFDA6" bold="true" box="[778,844,511,537]" pageId="2" pageNumber="116">Legs:</emphasis>
All legs dark brown, slightly more reddish than other parts of the body. Protibia tridentate, protarsi very strongly thickened, inner protibial claw very strongly enlarged and elongated.
<emphasis id="B9654F2C66775503FEDDFD9CFE16FDDE" bold="true" box="[319,448,583,609]" pageId="2" pageNumber="116">Parameres</emphasis>
(
<figureCitation id="132A8FBB66775503FE2CFD9CFD8AFDDE" box="[462,604,583,609]" captionStart="FIGURE 1" captionStartId="2.[151,250,1975,1999]" captionTargetBox="[215,1370,718,1935]" captionTargetId="figure-254@2.[215,1373,709,1942]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Pseudohomonyx continentalis Yang &amp; Pathomwattananurak, new species.A, habitus (dorsal view); B, habitus (ventral view); C, head and pronotum; D, parameres (caudal view); E, aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10150592" httpUri="https://zenodo.org/record/10150592/files/figure.png" pageId="2" pageNumber="116">Figs. 1D–1E</figureCitation>
): In caudal view (
<figureCitation id="132A8FBB66775503FCC7FD9CFCA1FDDE" box="[805,887,583,609]" captionStart="FIGURE 1" captionStartId="2.[151,250,1975,1999]" captionTargetBox="[215,1370,718,1935]" captionTargetId="figure-254@2.[215,1373,709,1942]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Pseudohomonyx continentalis Yang &amp; Pathomwattananurak, new species.A, habitus (dorsal view); B, habitus (ventral view); C, head and pronotum; D, parameres (caudal view); E, aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10150592" httpUri="https://zenodo.org/record/10150592/files/figure.png" pageId="2" pageNumber="116">Fig.1D</figureCitation>
), slender and symmetric; without any setae; outer sides with a pair of short, acute, outward and strongly backward processes at posterior one-sixth; inner sides simply curved; apex rounded; in lateral view (
<figureCitation id="132A8FBB66775503FDAAFD54FD4BFD16" box="[584,669,655,681]" captionStart="FIGURE 1" captionStartId="2.[151,250,1975,1999]" captionTargetBox="[215,1370,718,1935]" captionTargetId="figure-254@2.[215,1373,709,1942]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Pseudohomonyx continentalis Yang &amp; Pathomwattananurak, new species.A, habitus (dorsal view); B, habitus (ventral view); C, head and pronotum; D, parameres (caudal view); E, aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10150592" httpUri="https://zenodo.org/record/10150592/files/figure.png" pageId="2" pageNumber="116">Fig. 1E</figureCitation>
), apex slightly bent downward.
</paragraph>
<caption id="DF6EC3B666775503FF75F86CFB73F84C" ID-DOI="http://doi.org/10.5281/zenodo.10150592" ID-Zenodo-Dep="10150592" httpUri="https://zenodo.org/record/10150592/files/figure.png" pageId="2" pageNumber="116" startId="2.[151,250,1975,1999]" targetBox="[215,1370,718,1935]" targetPageId="2" targetType="figure">
<paragraph id="8BAE933E66775503FF75F86CFB73F84C" blockId="2.[151,1437,1975,2035]" pageId="2" pageNumber="116">
<emphasis id="B9654F2C66775503FF75F86CFEC5F870" bold="true" box="[151,275,1975,1999]" pageId="2" pageNumber="116">FIGURE 1.</emphasis>
Holotype of
<taxonomicName id="4C11E8BD66775503FE75F86CFC2EF870" authority="Yang &amp; Pathomwattananurak" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2023" box="[407,1016,1975,1999]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="116" phylum="Arthropoda" rank="species" species="continentalis" status="sp. nov.">
<emphasis id="B9654F2C66775503FE75F86CFD10F870" box="[407,710,1975,1999]" italics="true" pageId="2" pageNumber="116">Pseudohomonyx continentalis</emphasis>
Yang &amp; Pathomwattananurak
</taxonomicName>
,
<taxonomicNameLabel id="A256F25766775503FBE3F86CFBABF870" box="[1025,1149,1975,1999]" pageId="2" pageNumber="116" rank="species">new species</taxonomicNameLabel>
. A, habitus (dorsal view); B, habitus (ventral view); C, head and pronotum; D, parameres (caudal view); E, aedeagus (lateral view).
</paragraph>
</caption>
<paragraph id="8BAE933E66765502FF25FF4CFED1FF6A" blockId="3.[151,1437,151,357]" pageId="3" pageNumber="117">
<emphasis id="B9654F2C66765502FF25FF4CFEE2FF0E" bold="true" box="[199,308,151,177]" pageId="3" pageNumber="117">
<typeStatus id="54AA2D9C66765502FF25FF4CFEE2FF0E" box="[199,308,151,177]" pageId="3" pageNumber="117" type="paratype">Paratype</typeStatus>
</emphasis>
(male). Characters stable, except punctures on fourth and sixth intervals sparser, lesser in number (
<figureCitation id="132A8FBB66765502FF7DFF60FF2CFF6A" box="[159,250,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1066,1090]" captionTargetBox="[174,1411,393,1024]" captionTargetId="figure-96@3.[165,1422,387,1035]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of paratypes of Pseudohomonyx continentalis Yang &amp; Pathomwattananurak, new species.A, male paratype (dorsal view, ©THNHM/Tatsanai Jeenthong); B–C, female paratype (dorsal and ventral views, ©Thitipong Hongsuwong)." figureDoi="http://doi.org/10.5281/zenodo.10150594" httpUri="https://zenodo.org/record/10150594/files/figure.png" pageId="3" pageNumber="117">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="8BAE933E66765502FF25FF04FDBFFEDA" blockId="3.[151,1437,151,357]" pageId="3" pageNumber="117">
<emphasis id="B9654F2C66765502FF25FF04FEE2FF46" bold="true" box="[199,308,223,249]" pageId="3" pageNumber="117">
<typeStatus id="54AA2D9C66765502FF25FF04FEE2FF46" box="[199,308,223,249]" pageId="3" pageNumber="117" type="paratype">Paratype</typeStatus>
</emphasis>
(female). Characters mainly stable, except color much more reddish (it died two days after emerging before full melanization, thus the color is likely not its natural color); transverse carina separating frons and clypeus absent; punctures on fourth and sixth intervals sparser, lesser in number; protarsi not thickened, inner protibial claw not enlarged and elongated (
<figureCitation id="132A8FBB66765502FE36FE90FD8BFEDA" box="[468,605,331,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1066,1090]" captionTargetBox="[174,1411,393,1024]" captionTargetId="figure-96@3.[165,1422,387,1035]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of paratypes of Pseudohomonyx continentalis Yang &amp; Pathomwattananurak, new species.A, male paratype (dorsal view, ©THNHM/Tatsanai Jeenthong); B–C, female paratype (dorsal and ventral views, ©Thitipong Hongsuwong)." figureDoi="http://doi.org/10.5281/zenodo.10150594" httpUri="https://zenodo.org/record/10150594/files/figure.png" pageId="3" pageNumber="117">Fig. 2B–2C</figureCitation>
).
</paragraph>
<caption id="DF6EC3B666765502FF75FBF1FAA5FBD9" ID-DOI="http://doi.org/10.5281/zenodo.10150594" ID-Zenodo-Dep="10150594" httpUri="https://zenodo.org/record/10150594/files/figure.png" pageId="3" pageNumber="117" startId="3.[151,250,1066,1090]" targetBox="[174,1411,393,1024]" targetPageId="3" targetType="figure">
<paragraph id="8BAE933E66765502FF75FBF1FAA5FBD9" blockId="3.[151,1436,1066,1128]" pageId="3" pageNumber="117">
<emphasis id="B9654F2C66765502FF75FBF1FEC4FBFD" bold="true" box="[151,274,1066,1090]" pageId="3" pageNumber="117">FIGURE 2.</emphasis>
Habitus of paratypes of
<taxonomicName id="4C11E8BD66765502FDE4FBF1FBB3FBFD" authority="Yang &amp; Pathomwattananurak" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2023" box="[518,1125,1066,1090]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="species" species="continentalis" status="sp. nov.">
<emphasis id="B9654F2C66765502FDE4FBF1FCE3FBFD" box="[518,821,1066,1090]" italics="true" pageId="3" pageNumber="117">Pseudohomonyx continentalis</emphasis>
Yang &amp; Pathomwattananurak
</taxonomicName>
,
<taxonomicNameLabel id="A256F25766765502FB8CFBF1FB3FFBFD" box="[1134,1257,1066,1090]" pageId="3" pageNumber="117" rank="species">new species</taxonomicNameLabel>
. A, male paratype (dorsal view, ©THNHM/Tatsanai Jeenthong); B–C, female paratype (dorsal and ventral views, ©Thitipong Hongsuwong).
</paragraph>
</caption>
<paragraph id="8BAE933E66765502FF25FB63FD3CFB48" blockId="3.[151,1437,1208,1883]" pageId="3" pageNumber="117">
<emphasis id="B9654F2C66765502FF25FB63FEA8FB6D" bold="true" box="[199,382,1208,1234]" pageId="3" pageNumber="117">Measurements.</emphasis>
Body length: male 19.5–20.0 mm (holotype 20.0 mm), female 20.7 mm; body width: male 10.3–11.0 mm (holotype 10.3 mm), female 11.6 mm.
</paragraph>
</subSubSection>
<subSubSection id="C30BC0B566765502FF25FADBFCF8FA14" pageId="3" pageNumber="117" type="diagnosis">
<paragraph id="8BAE933E66765502FF25FADBFCF8FA14" blockId="3.[151,1437,1208,1883]" pageId="3" pageNumber="117">
<emphasis id="B9654F2C66765502FF25FADBFE94FAA5" bold="true" box="[199,322,1280,1306]" pageId="3" pageNumber="117">Diagnosis.</emphasis>
This species is relatively unique in its appearances and can be easily distinguished from all other species of the genus by absent of the frontal impression and frontal tubercle on the pronotum in both sexes, the combination of the following characters can also distinguish this species from all other species: male with very strongly thickened protarsi and very strongly enlarged and elongated inner protibial claw; fourth and sixth intervals with punctures posteriorly; and the shape of the parameres.
</paragraph>
</subSubSection>
<subSubSection id="C30BC0B566765502FF25FA6FFCF5FA4D" pageId="3" pageNumber="117" type="etymology">
<paragraph id="8BAE933E66765502FF25FA6FFCF5FA4D" blockId="3.[151,1437,1208,1883]" pageId="3" pageNumber="117">
<emphasis id="B9654F2C66765502FF25FA6FFE98FA71" bold="true" box="[199,334,1460,1486]" pageId="3" pageNumber="117">Etymology.</emphasis>
This species is named for its limited distribution on the continent, which is currently the only species known exclusively from mainland Southeast Asia.
</paragraph>
</subSubSection>
<subSubSection id="C30BC0B566765502FF25FA27FCC4F9A9" box="[199,786,1532,1558]" pageId="3" pageNumber="117" type="distribution">
<paragraph id="8BAE933E66765502FF25FA27FCC4F9A9" blockId="3.[151,1437,1208,1883]" box="[199,786,1532,1558]" pageId="3" pageNumber="117">
<emphasis id="B9654F2C66765502FF25FA27FEB7F9A9" bold="true" box="[199,353,1532,1558]" pageId="3" pageNumber="117">Distribution.</emphasis>
Malaysia: Malay Peninsula; Thailand.
</paragraph>
</subSubSection>
<subSubSection id="C30BC0B566765502FF25F9FBFBE5F8E5" pageId="3" pageNumber="117" type="discussion">
<paragraph id="8BAE933E66765502FF25F9FBFB01F919" blockId="3.[151,1437,1208,1883]" pageId="3" pageNumber="117">
<emphasis id="B9654F2C66765502FF25F9FBFEEDF985" bold="true" box="[199,315,1568,1594]" pageId="3" pageNumber="117">Remarks.</emphasis>
This species belongs to a group who lacks setation on the parameres (the small white dots on the photographs of the parameres (
<figureCitation id="132A8FBB66765502FE17F99FFD53F9E1" box="[501,645,1604,1630]" captionStart="FIGURE 1" captionStartId="2.[151,250,1975,1999]" captionTargetBox="[215,1370,718,1935]" captionTargetId="figure-254@2.[215,1373,709,1942]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Pseudohomonyx continentalis Yang &amp; Pathomwattananurak, new species.A, habitus (dorsal view); B, habitus (ventral view); C, head and pronotum; D, parameres (caudal view); E, aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10150592" httpUri="https://zenodo.org/record/10150592/files/figure.png" pageId="3" pageNumber="117">Figs. 1D–1E</figureCitation>
) are small pieces of dust instead of setae). Other species of this group are:
<taxonomicName id="4C11E8BD66765502FF28F9B3FC9FF93D" authority="Miyake &amp; Yamaya, 1997" authorityName="Miyake &amp; Yamaya" authorityYear="1997" box="[202,841,1640,1667]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="species" species="belitungensis">
<emphasis id="B9654F2C66765502FF28F9B3FDC9F93D" box="[202,543,1640,1666]" italics="true" pageId="3" pageNumber="117">Pseudohomonyx belitungensis</emphasis>
<bibRefCitation id="EF80EECF66765502FDC5F9B3FC9FF93D" author="Miyake, Y. &amp; Yamaya, S." box="[551,841,1640,1667]" pageId="3" pageNumber="117" pagination="5 - 20" refId="ref5570" refString="Miyake, Y. &amp; Yamaya, S. (1997) New or little known Scarabaeid beetles from Southeast Asia, preserved in Nagaoka Municipal Science Museum. Bulletin of the Nagaoka municipal Science Museum, 32, 5 - 20" type="journal article" year="1997">Miyake &amp; Yamaya, 1997</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C11E8BD66765502FCB4F9B2FB50F93D" authority="Arrow, 1908" authorityName="Arrow" authorityYear="1908" box="[854,1158,1640,1666]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="B9654F2C66765502FCB4F9B2FC3AF93D" box="[854,1004,1640,1666]" italics="true" pageId="3" pageNumber="117">P. borneensis</emphasis>
<bibRefCitation id="EF80EECF66765502FC16F9B3FB50F93D" author="Arrow, G. J." box="[1012,1158,1640,1666]" pageId="3" pageNumber="117" pagination="321 - 358" refId="ref5162" refString="Arrow, G. J. (1908) A contribution to the classification of the Coleopterous family Dynastidae. Transactions of the Entomological Society of London, 56 (2), 321 - 358. https: // doi. org / 10.1111 / j. 1365 - 2311.1908. tb 02151. x" type="journal article" year="1908">Arrow, 1908</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C11E8BD66765502FB71F9B2FF0DF919" authority="(Burmeister, 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B9654F2C66765502FB71F9B2FADEF93D" box="[1171,1288,1641,1666]" italics="true" pageId="3" pageNumber="117">P. javanus</emphasis>
(
<bibRefCitation id="EF80EECF66765502FAFAF9B3FF05F919" author="Burmeister, H. C. C." pageId="3" pageNumber="117" refId="ref5318" refString="Burmeister, H. C. C. (1847) Handbuch der Entomologie. Funfter Band. Besondere Entomologie. Forsetzung. Coleoptera Lamellicornia Xvlophila et Pectinicornia. G. Reimer, Reimer, 584 pp. https: // doi. org / 10.5962 / bhl. title. 8135" type="book" year="1847">Burmeister, 1847</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C11E8BD66765502FF05F956FDA6F919" authority="Jakl &amp; Zidek, 2017" authorityName="Jakl &amp; Zidek" authorityYear="2017" box="[231,624,1676,1702]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="species" species="loksadoensis">
<emphasis id="B9654F2C66765502FF05F956FE47F919" box="[231,401,1676,1702]" italics="true" pageId="3" pageNumber="117">P. loksadoensis</emphasis>
<bibRefCitation id="EF80EECF66765502FE7AF957FDA6F919" author="Jakl, S. &amp; Zidek, J." box="[408,624,1676,1702]" pageId="3" pageNumber="117" pagination="19 - 30" refId="ref5477" refString="Jakl, S. &amp; Zidek, J. (2017) A contribution to the knowledge of Pseudohomonyx Arrow, 1908. Folia Heyrovskyana, Series A, 25 (2), 19 - 30." type="journal article" year="2017">Jákl &amp; Zídek, 2017</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C11E8BD66765502FD45F956FC32F919" authority="Jakl &amp; Zidek, 2017" authorityName="Jakl &amp; Zidek" authorityYear="2017" box="[679,996,1676,1702]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B9654F2C66765502FD45F956FCD3F919" box="[679,773,1677,1702]" italics="true" pageId="3" pageNumber="117">P. minor</emphasis>
<bibRefCitation id="EF80EECF66765502FCEEF957FC32F919" author="Jakl, S. &amp; Zidek, J." box="[780,996,1676,1702]" pageId="3" pageNumber="117" pagination="19 - 30" refId="ref5477" refString="Jakl, S. &amp; Zidek, J. (2017) A contribution to the knowledge of Pseudohomonyx Arrow, 1908. Folia Heyrovskyana, Series A, 25 (2), 19 - 30." type="journal article" year="2017">Jákl &amp; Zídek, 2017</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EF80EECF66765502FC11F957FB1DF919" author="Jakl, S. &amp; Zidek, J." box="[1011,1227,1676,1702]" pageId="3" pageNumber="117" pagination="19 - 30" refId="ref5477" refString="Jakl, S. &amp; Zidek, J. (2017) A contribution to the knowledge of Pseudohomonyx Arrow, 1908. Folia Heyrovskyana, Series A, 25 (2), 19 - 30." type="journal article" year="2017">Jákl &amp; Zídek 2017</bibRefCitation>
).
</paragraph>
<paragraph id="8BAE933E66765502FF25F96BFC50F8AD" blockId="3.[151,1437,1208,1883]" pageId="3" pageNumber="117">
<taxonomicName id="4C11E8BD66765502FF25F96BFCAEF975" authority="Yang &amp; Pathomwattananurak" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2023" box="[199,888,1712,1739]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="species" species="continentalis" status="sp. nov.">
<emphasis id="B9654F2C66765502FF25F96BFDCCF975" box="[199,538,1712,1738]" italics="true" pageId="3" pageNumber="117">Pseudohomonyx continentalis</emphasis>
Yang &amp; Pathomwattananurak
</taxonomicName>
,
<emphasis id="B9654F2C66765502FC65F96BFBC2F975" bold="true" box="[903,1044,1712,1738]" pageId="3" pageNumber="117">
<taxonomicNameLabel id="A256F25766765502FC65F96BFBC2F975" box="[903,1044,1712,1738]" pageId="3" pageNumber="117" rank="species">new species</taxonomicNameLabel>
</emphasis>
, alongside with
<taxonomicName id="4C11E8BD66765502FB3BF96AFA99F975" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1241,1359,1713,1738]" class="Insecta" family="Scarabaeidae" genus="Pseudohomonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B9654F2C66765502FB3BF96AFA99F975" box="[1241,1359,1713,1738]" italics="true" pageId="3" pageNumber="117">P. javanus</emphasis>
</taxonomicName>
(more information given in the according section in the catalogue), represents the first records of this genus from mainland Southeast Asia. It is also the only species endemic to the mainland.
</paragraph>
<paragraph id="8BAE933E66765502FF25F8C6FBE5F8E5" blockId="3.[151,1437,1208,1883]" pageId="3" pageNumber="117">It should be noted that the female paratype was assembled from pieces after it died before full melanization. Fortunately, characters can still be clearly visible and compared with the holotype.</paragraph>
</subSubSection>
</treatment>
</document>
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