496 lines
61 KiB
XML
496 lines
61 KiB
XML
<document id="B8DF5C466A27E8084EC0BBAD6DF738F6" ID-CLB-Dataset="30217" ID-DOI="10.1093/zoolinnean/zlx112" ID-GBIF-Dataset="99b3b009-f006-4146-b335-cbfdc2fcd60b" ID-Zenodo-Dep="1226487" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1524310177419" checkinUser="plazi" docAuthor="David Prötzel, Miguel Vences, Oliver Hawlitschek, Mark D. Scherz, Fanomezana M. Ratsoavina & Frank Glaw" docDate="2018" docId="03B487F21646FFCB398EFE6FFB430777" docLanguage="en" docName="zjls.2018.20.1-28.pdf" docOrigin="Zoological Journal of the Linnean Society 2018 (20)" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.1:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="1" docTitle="Calumma lefona David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw 2018, SP. NOV." docType="treatment" docUuid="99AF4F23-EBDF-4276-AE41-3C04E7443FD8" docUuidSource="ZooBank" docVersion="8" lastPageNumber="15" masterDocId="FF8DFF8A164DFFC53A42FFABFFE90568" masterDocTitle="Endangered beauties: micro-CT cranial osteology, molecular genetics and external morphology reveal three new species of chameleons in the Calumma boettgeri complex (Squamata: Chamaeleonidae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="12" updateTime="1698540677137" updateUser="ExternalLinkService">
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<mods:title id="CE5CADDFD9FAC59C31BB84C7ED0677C6">Endangered beauties: micro-CT cranial osteology, molecular genetics and external morphology reveal three new species of chameleons in the Calumma boettgeri complex (Squamata: Chamaeleonidae)</mods:title>
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<mods:roleTerm id="23D14AC8AAA4BDC657A1387B3E91D7D8">Author</mods:roleTerm>
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<mods:namePart id="82753944D7E148E420EE3FFAEC7B74E3">David Prötzel</mods:namePart>
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<mods:roleTerm id="FB23973D70DA9732547AB217378B8B87">Author</mods:roleTerm>
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<mods:namePart id="49DEF3441219926C2543EAF4751B860F">Miguel Vences</mods:namePart>
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<mods:role id="497737A84D8C257D3D52851B00BCB909">
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<mods:roleTerm id="659BBA672F9A7F4686770482981C33D1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A59C133E5AB47224F0F5EE108061B25B">Oliver Hawlitschek</mods:namePart>
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</mods:name>
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<mods:name id="D890C380628F95A5ECBC3B612783ECD3" type="personal">
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<mods:roleTerm id="AEFC9E6EEC8AB55F09199F9AE7A53369">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C219A75B44516A5C164003C6962454CE">Mark D. Scherz</mods:namePart>
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</mods:name>
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<mods:roleTerm id="AFB72CBC9F2A30C3562413647EEE21DF">Author</mods:roleTerm>
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<mods:namePart id="35D17EF8F9D2A5F0C6B1272C25F103B8">Fanomezana M. Ratsoavina</mods:namePart>
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<mods:roleTerm id="4553CF1CB853EB979AAA414ECD4293B0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5CF6EE047B16EF81FF6EEB7B95638901">Frank Glaw</mods:namePart>
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<mods:title id="D21EE965BEB729A1A27EBC9083816B7F">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="9099B776DC3330ABCDA608FF51599200">
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<mods:date id="7ADF6BDF807693805F1E55D5150C89CD">2018</mods:date>
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<mods:number id="83EEFC5538BFC80AFD16886BC1128944">2018-04-09</mods:number>
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<mods:detail id="80B5F59F727A21C1B5343D68471579EB" type="volume">
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<mods:number id="68B35228367BC6129BD54810B8033F03">2018</mods:number>
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<mods:classification id="C94F34D2FEC9B48646432B3F2BA2B816">journal article</mods:classification>
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<mods:identifier id="229DF738BBC675E37C93040F59E71AFB" type="DOI">10.1093/zoolinnean/zlx112</mods:identifier>
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<treatment id="03B487F21646FFCB398EFE6FFB430777" ID-DOI="http://doi.org/10.5281/zenodo.5949496" ID-GBIF-Taxon="143625132" ID-Zenodo-Dep="5949496" LSID="urn:lsid:zoobank.org:act:99AF4F23-EBDF-4276-AE41-3C04E7443FD8" httpUri="http://treatment.plazi.org/id/03B487F21646FFCB398EFE6FFB430777" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
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<subSubSection id="C307656F1646FFCE398EFE6FFC050759" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="8BA236E41646FFCE398EFE6FFAF804B4" blockId="11.[872,1397,374,476]" box="[972,1297,452,476]" pageId="11" pageNumber="12">
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<taxonomicName id="4C1D4D671646FFCE398EFE6FFB4E04B2" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[972,1191,452,476]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="28" phylum="Chordata" rank="species" species="lefona" status="sp. nov.">CALUMMA LEFONA</taxonomicName>
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<emphasis id="B969EAF61646FFCE3EECFE6CFAF804B4" bold="true" box="[1198,1297,452,476]" pageId="11" pageNumber="12">
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<taxonomicNameLabel id="A25A578D1646FFCE3EECFE6CFAF804B4" box="[1198,1297,452,476]" pageId="11" pageNumber="12">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="8BA236E41646FFCE3979FE56FC050759" blockId="11.[827,1443,509,561]" pageId="11" pageNumber="12">urn:lsid:zoobank.org:act:99AF4F23-EBDF-4276-AE41- 3C04E7443FD8</paragraph>
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<subSubSection id="C307656F1646FFCE3979FDE0FBC50717" pageId="11" pageNumber="12" type="reference_group">
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<emphasis id="B969EAF61646FFCE3979FDE0FC740708" box="[827,925,587,608]" italics="true" pageId="11" pageNumber="12">Remark:</emphasis>
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This species was considered as clade FII by Gehring
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<emphasis id="B969EAF61646FFCE39DDFDC1FC3E0717" box="[927,983,618,639]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
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(
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<date id="FFA310241646FFCE39A7FDC1FBF407E8" box="[997,1053,618,640]" pageId="11" pageNumber="12" value="2012">2012</date>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C307656F1646FFCE3979FD32FBEE06FF" pageId="11" pageNumber="12" type="materials_examined">
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<paragraph id="8BA236E41646FFCE3979FD32FB120600" blockId="11.[827,1443,665,872]" pageId="11" pageNumber="12">
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<materialsCitation id="3B753CB91646FFCE3979FD32FB120600" ID-GBIF-Occurrence="1838289362" collectingDate="2010-06-21" collectionCode="ZSM" collectorName="F. M. Ratsoavina & F. Randrianasola." country="Madagascar" elevation="1717" latitude="-14.3464" location="Andrevorevo" longLatPrecision="7" longitude="49.1028" municipality="Tsaratanana Massif" pageId="11" pageNumber="12" specimenCode="ZSM 2" specimenCount="1" stateProvince="Mahajanga Province" typeStatus="holotype">
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<emphasis id="B969EAF61646FFCE3979FD32FC4607C6" box="[827,943,665,686]" italics="true" pageId="11" pageNumber="12">Holotype</emphasis>
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</typeStatus>
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:
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<specimenCode id="DBBB9E9F1646FFCE398AFD32FBC907C6" box="[968,1056,665,686]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="11" pageNumber="12">
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<collectionCode id="ED0CAE211646FFCE398AFD32FBEB07C6" box="[968,1026,665,686]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="11" pageNumber="12">ZSM</collectionCode>
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8 4 9/
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(DRV
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7), adult male with incompletely everted hemipenes, left hemipenis removed for micro-CT scanning, collected in
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<location id="8EC2603F1646FFCE391FFD5DFC120664" LSID="urn:lsid:plazi:treatment:03B487F21646FFCB398EFE6FFB430777:8EC2603F1646FFCE391FFD5DFC120664" box="[861,1019,758,780]" country="Madagascar" latitude="-14.3464" longLatPrecision="7" longitude="49.1028" municipality="Tsaratanana Massif" name="Andrevorevo" pageId="11" pageNumber="12" stateProvince="Mahajanga Province">Andrevorevo</location>
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||
southeast of
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<collectingMunicipality id="6BC6AC9E1646FFCE3EE5FD5DFA4A0664" box="[1191,1443,758,780]" pageId="11" pageNumber="12">Tsaratanana Massif</collectingMunicipality>
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(
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<geoCoordinate id="EE2950231646FFCE3901FCBFFC2B0642" box="[835,962,788,810]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="-14.3464">
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14.3464°
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<collectionCode id="ED0CAE211646FFCE39F1FCBFFC2B0642" box="[947,962,788,810]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="11" pageNumber="12">S</collectionCode>
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</geoCoordinate>
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,
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<geoCoordinate id="EE2950231646FFCE3992FCBEFBB80642" box="[976,1105,789,810]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="49.1028">
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49.1028°
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<collectionCode id="ED0CAE211646FFCE3E7CFCBEFBB80642" box="[1086,1105,789,810]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="11" pageNumber="12">E</collectionCode>
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</geoCoordinate>
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,
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a.s.l.
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</elevation>
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),
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<collectingRegion id="49D9F8061646FFCE3F5BFCBFFC4D0621" country="Madagascar" name="Mahajanga" pageId="11" pageNumber="12">Mahajanga Province</collectingRegion>
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, northern
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<collectingCountry id="F30A76741646FFCE3E63FC98FB590621" box="[1057,1200,819,841]" name="Madagascar" pageId="11" pageNumber="12">Madagascar</collectingCountry>
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, on
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<date id="FFA310241646FFCE3EA3FC9FFA690621" box="[1249,1408,820,841]" pageId="11" pageNumber="12" value="2010-06-21">
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<collectingDate id="EFE7E9CC1646FFCE3EA3FC9FFA690621" box="[1249,1408,820,841]" pageId="11" pageNumber="12" value="2010-06-21">21 June 2010</collectingDate>
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</date>
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||
by
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<collectorName id="26E853321646FFCE3979FCF8FC100600" box="[827,1017,850,872]" pageId="11" pageNumber="12">
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<collectionCode id="ED0CAE211646FFCE3979FCF8FCA50600" box="[827,844,851,872]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="11" pageNumber="12">F</collectionCode>
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.
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||
<collectionCode id="ED0CAE211646FFCE3917FCF8FC860600" box="[853,879,851,872]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="11" pageNumber="12">M</collectionCode>
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||
. Ratsoavina
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||
</collectorName>
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and
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||
<collectorName id="26E853321646FFCE3E70FCF8FB120600" box="[1074,1275,850,872]" pageId="11" pageNumber="12">
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<collectionCode id="ED0CAE211646FFCE3E70FCF8FBAA0600" box="[1074,1091,851,872]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="11" pageNumber="12">F</collectionCode>
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||
. Randrianasola.
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||
</collectorName>
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</materialsCitation>
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</paragraph>
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<paragraph id="8BA236E41646FFCE3979FC29FC5806FF" blockId="11.[827,945,898,919]" box="[827,945,898,919]" pageId="11" pageNumber="12">
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<heading id="D0EA81881646FFCE3979FC29FC5806FF" box="[827,945,898,919]" centered="true" fontSize="9" level="6" pageId="11" pageNumber="12" reason="4">
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<materialsCitation id="3B753CB91646FFCE3979FC29FC5806FF" ID-GBIF-Occurrence="1838289363" box="[827,945,898,919]" pageId="11" pageNumber="12" typeStatus="paratype">
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<typeStatus id="54A688461646FFCE3979FC29FC4306FF" box="[827,938,898,919]" pageId="11" pageNumber="12" type="paratype">
|
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<emphasis id="B969EAF61646FFCE3979FC29FC4306FF" box="[827,938,898,919]" italics="true" pageId="11" pageNumber="12">Paratypes</emphasis>
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</typeStatus>
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:
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</materialsCitation>
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</heading>
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</paragraph>
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<paragraph id="8BA236E41646FFCE3986FC29FBEE06FF" blockId="11.[964,1031,898,919]" box="[964,1031,898,919]" pageId="11" pageNumber="12">
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<heading id="D0EA81881646FFCE3986FC29FBEE06FF" box="[964,1031,898,919]" centered="true" fontSize="9" level="6" pageId="11" pageNumber="12" reason="4">None.</heading>
|
||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C307656F1646FFC93979FC1AFE940629" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="diagnosis">
|
||
<paragraph id="8BA236E41646FFCE3979FC1AFB650052" blockId="11.[827,1443,944,1896]" pageId="11" pageNumber="12">
|
||
<emphasis id="B969EAF61646FFCE3979FC1AFC5D06AE" box="[827,948,945,966]" italics="true" pageId="11" pageNumber="12">Diagnosis</emphasis>
|
||
:
|
||
<taxonomicName id="4C1D4D671646FFCE3990FC1BFB7406AD" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[978,1181,944,965]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="lefona" status="sp. nov.">
|
||
<emphasis id="B969EAF61646FFCE3990FC1BFB7406AD" box="[978,1181,944,965]" italics="true" pageId="11" pageNumber="12">Calumma lefona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1646FFCE3EE5FC1AFAEB06AE" box="[1191,1282,945,966]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is a member of the phenetic
|
||
<emphasis id="B969EAF61646FFCE39B9FC7BFB6F068D" box="[1019,1158,976,997]" italics="true" pageId="11" pageNumber="12">C. nasutum</emphasis>
|
||
species group (
|
||
<bibRefCitation id="EF8C4B151646FFCE3F04FC64FB76016C" author="Protzel D & Ruthensteiner B & Glaw F." journalOrPublisher="Zoosystematics and Evolution" pageId="11" pageNumber="12" pagination="13 - 21" part="92" refId="ref18271" refString="Protzel D, Ruthensteiner B, Glaw F. 2016. No longer single! Description of female Calumma vatosoa (Squamata, Chamaeleonidae) including a review of the species and its systematic position. Zoosystematics and Evolution 92: 13 - 21." title="No longer single! Description of female Calumma vatosoa (Squamata, Chamaeleonidae) including a review of the species and its systematic position" type="journal article" year="2016">
|
||
Prötzel, Ruthensteiner & Glaw,
|
||
<date id="FFA310241646FFCE3E26FC45FB76016C" box="[1124,1183,1006,1028]" pageId="11" pageNumber="12" value="2016">2016</date>
|
||
</bibRefCitation>
|
||
), on the basis of the presence of a soft, dermal, unpaired rostral appendage, absence of gular or ventral crests, and heterogeneous scalation on the lower arm, consisting mostly of tubercles of ~
|
||
<quantity id="4CE59B011646FFCE3996FBC1FBC201E8" box="[980,1067,1130,1152]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="mm" value="0.5">0.5 mm</quantity>
|
||
diameter. Within the genus, it is a small sized chameleon (SVL
|
||
<quantity id="4CE59B011646FFCE3ED7FB22FB1001F7" box="[1173,1273,1161,1183]" metricMagnitude="-2" metricUnit="m" metricValue="5.13" pageId="11" pageNumber="12" unit="mm" value="51.3">51.3 mm</quantity>
|
||
, TL
|
||
<quantity id="4CE59B011646FFCE3F68FB22FA7501F7" box="[1322,1436,1161,1183]" metricMagnitude="-1" metricUnit="m" metricValue="1.137" pageId="11" pageNumber="12" unit="mm" value="113.7">113.7 mm</quantity>
|
||
) that is characterized by a long and pointed rostral appendage, occipital lobes that are widely notched and completely separated, a distinctly elevated rostral crest, a dorsal and caudal crest, absence of axillary pits and unique skull morphology.
|
||
</paragraph>
|
||
<paragraph id="8BA236E41646FFCE3911FAE8FB2F03A5" blockId="11.[827,1443,944,1896]" pageId="11" pageNumber="12">
|
||
<taxonomicName id="4C1D4D671646FFCE3911FAE8FBCA0030" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[851,1059,1347,1368]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="lefona" status="sp. nov.">
|
||
<emphasis id="B969EAF61646FFCE3911FAE8FBCA0030" box="[851,1059,1347,1368]" italics="true" pageId="11" pageNumber="12">Calumma lefona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1646FFCE3E6EFAEFFB600031" box="[1068,1161,1348,1369]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C1D4D671646FFCE3F70FAEFFA740030" baseAuthorityName="Mocquard" baseAuthorityYear="1900" box="[1330,1437,1347,1369]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="fallax">
|
||
<emphasis id="B969EAF61646FFCE3F70FAEFFA740030" box="[1330,1437,1347,1369]" italics="true" pageId="11" pageNumber="12">C. fallax</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B969EAF61646FFCE3979FAC8FC4D001F" box="[827,932,1378,1400]" italics="true" pageId="11" pageNumber="12">C. gallus</emphasis>
|
||
,
|
||
<emphasis id="B969EAF61646FFCE39F0FAC8FBD20010" box="[946,1083,1379,1400]" italics="true" pageId="11" pageNumber="12">C. nasutum</emphasis>
|
||
,
|
||
<taxonomicName id="4C1D4D671646FFCE3E08FAC8FB360010" baseAuthorityName="Brygoo, Blanc & Domergue" baseAuthorityYear="1974" box="[1098,1247,1379,1400]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="peyrierasi">
|
||
<emphasis id="B969EAF61646FFCE3E08FAC8FB360010" box="[1098,1247,1379,1400]" italics="true" pageId="11" pageNumber="12">C. peyrierasi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B969EAF61646FFCE3EACFAC8FA870010" box="[1262,1390,1379,1400]" italics="true" pageId="11" pageNumber="12">C. vatosoa,</emphasis>
|
||
and
|
||
<emphasis id="B969EAF61646FFCE3979FA29FC5700FE" box="[827,958,1409,1431]" italics="true" pageId="11" pageNumber="12">C. vohibola</emphasis>
|
||
of the
|
||
<emphasis id="B969EAF61646FFCE3E53FA29FB7100FF" box="[1041,1176,1410,1431]" italics="true" pageId="11" pageNumber="12">C. nasutum</emphasis>
|
||
group by the presence of occipital lobes; from
|
||
<emphasis id="B969EAF61646FFCE3E1BFA0BFB0A00DD" box="[1113,1251,1440,1461]" italics="true" pageId="11" pageNumber="12">C. boettgeri</emphasis>
|
||
and
|
||
<emphasis id="B969EAF61646FFCE3F61FA0BFA4A00DD" box="[1315,1443,1440,1461]" italics="true" pageId="11" pageNumber="12">C. linotum</emphasis>
|
||
in the clearly notched occipital lobes with a depth of
|
||
<quantity id="4CE59B011646FFCE3979FA75FC7D009C" box="[827,916,1502,1524]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="12" unit="mm" value="1.8">1.8 mm</quantity>
|
||
(vs. not or slightly notched with
|
||
<quantity id="4CE59B011646FFCE3F5DFA75FA7F009C" box="[1311,1430,1502,1524]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="7.0" metricValueMin="0.0" pageId="11" pageNumber="12" unit="mm" value="0.35" valueMax="0.7" valueMin="0.0">0–0.7 mm</quantity>
|
||
), presence of a frontoparietal fenestra with a width of
|
||
<quantity id="4CE59B011646FFCE3979F9B7FC7A035A" box="[827,915,1564,1586]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="12" unit="mm" value="1.9">1.9 mm</quantity>
|
||
(14.7% of skull length; vs. completely closed brain case), prefrontal, frontal and parietal with many tubercles (vs. smooth or only a few tubercles); additionally, from
|
||
<emphasis id="B969EAF61646FFCE39E8F9D2FBC103E6" box="[938,1064,1657,1678]" italics="true" pageId="11" pageNumber="12">C. boettgeri</emphasis>
|
||
by the higher number (
|
||
<quantity id="4CE59B011646FFCE3F73F9D2FA8303E7" box="[1329,1386,1657,1679]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="11" pageNumber="12" unit="in" value="21.0">21 in</quantity>
|
||
line) of large, juxtaposed tubercle scales on the extremities (vs. seven to 14 isolated tubercles).
|
||
</paragraph>
|
||
<paragraph id="8BA236E41646FFC93911F97DFE940629" blockId="11.[827,1443,944,1896]" lastBlockId="12.[145,762,198,834]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
From the other three taxa with notched occipital lobes,
|
||
<emphasis id="B969EAF61646FFCE39EEF95EFBC70262" box="[940,1070,1781,1802]" italics="true" pageId="11" pageNumber="12">C. gehringi</emphasis>
|
||
,
|
||
<emphasis id="B969EAF61646FFCE3E7EF95EFB430262" box="[1084,1194,1781,1802]" italics="true" pageId="11" pageNumber="12">C. guibei,</emphasis>
|
||
and
|
||
<taxonomicName id="4C1D4D671646FFCE3EA5F95EFAD70263" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[1255,1342,1781,1803]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="uetzi" status="sp. nov.">
|
||
<emphasis id="B969EAF61646FFCE3EA5F95EFAD70263" box="[1255,1342,1781,1803]" italics="true" pageId="11" pageNumber="12">C. uetzi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1646FFCE3F04F95EFA760262" box="[1350,1439,1781,1802]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C1D4D671646FFCE3979F8BFFC4F0241" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[827,934,1812,1833]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="lefona" status="sp. nov.">
|
||
<emphasis id="B969EAF61646FFCE3979F8BFFC4F0241" box="[827,934,1812,1833]" italics="true" pageId="11" pageNumber="12">C. lefona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1646FFCE39EDF8BFFBE30241" box="[943,1034,1812,1833]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs by the long (
|
||
<quantity id="4CE59B011646FFCE3F45F8BFFA8C0241" box="[1287,1381,1812,1834]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="11" pageNumber="12" unit="mm" value="5.6">5.6 mm</quantity>
|
||
) and pointed rostral appendage (vs.
|
||
<quantity id="4CE59B011646FFCE3EE5F898FAD80220" box="[1191,1329,1843,1865]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="5.4" metricValueMin="3.1" pageId="11" pageNumber="12" unit="mm" value="4.25" valueMax="5.4" valueMin="3.1">3.1–5.4 mm</quantity>
|
||
in males, rounded), with 60 large scales (diameter>
|
||
<quantity id="4CE59B011646FFCE3F03F8F9FA72020F" box="[1345,1435,1874,1895]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="12" unit="mm" value="0.3">0.3 mm</quantity>
|
||
) on the right side of the rostral appendage (vs.
|
||
<quantity id="4CE59B011641FFC938F0FF6CFF400593" metricMagnitude="-1" metricUnit="m" metricValue="7.874" metricValueMax="10.668" metricValueMin="5.08" pageId="12" pageNumber="13" unit="in" value="31.0" valueMax="42.0" valueMin="20.0">20–42 in</quantity>
|
||
males), dorsal crest of 23 small conical scales (vs. absence or 5–15 large conical scales), clearly and widely separated occipital lobes with a notch of
|
||
<quantity id="4CE59B011641FFC938E7FE88FD100450" box="[677,761,291,312]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="mm" value="1.8">1.8 mm</quantity>
|
||
(vs. tightly separated or connected with a notch of
|
||
<quantity id="4CE59B011641FFC93AD3FECBFEFE041D" box="[145,279,352,374]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="12" pageNumber="13" unit="mm" value="1.0" valueMax="1.5" valueMin="0.5">0.5–1.5 mm</quantity>
|
||
in males); furthermore, in skull morphology by a narrower frontal, e.g. at the border to the postorbitofrontal with 30.5% of SkL (vs. 35.5–39.7%) and strongly raised maxillae; from
|
||
<emphasis id="B969EAF61641FFC9387DFE17FD2D04B9" box="[575,708,444,465]" italics="true" pageId="12" pageNumber="13">C. gehringi</emphasis>
|
||
and
|
||
<emphasis id="B969EAF61641FFC93AD3FE70FF1A0487" box="[145,243,474,496]" italics="true" pageId="12" pageNumber="13">C. guibei</emphasis>
|
||
in a broader parietal at midpoint with 18.3% of SkL (vs. 9.9–15.3%); from
|
||
<taxonomicName id="4C1D4D671641FFC93B96FE52FDCE0767" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[468,551,505,527]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="uetzi" status="sp. nov.">
|
||
<emphasis id="B969EAF61641FFC93B96FE52FDCE0767" box="[468,551,505,527]" italics="true" pageId="12" pageNumber="13">C. uetzi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1641FFC9386CFE52FD690766" box="[558,640,505,526]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by possession of a frontoparietal fenestra (vs. completely closed brain case) and a narrower head (e.g. RFWPo of 30.5 vs. 35.6–39.4%); from
|
||
<emphasis id="B969EAF61641FFC93BC4FDFEFE0E0702" box="[390,487,597,618]" italics="true" pageId="12" pageNumber="13">C. guibei</emphasis>
|
||
by a smaller frontoparietal fenestra with 14.7% of SkL (vs. 21.0%), prefrontal fontanelle and naris separated by contact of prefrontal with maxilla (vs. fused), thick squamosal (vs. thin) in broad dorsal contact with the parietal (vs. not meeting parietal or only in weak contact); and from
|
||
<emphasis id="B969EAF61641FFC9383EFD44FD10066B" box="[636,761,750,772]" italics="true" pageId="12" pageNumber="13">C. gehringi</emphasis>
|
||
by possession of a larger frontoparietal fenestra (21.0% of SkL vs. 5.8–9.1%).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307656F1641FFCB3AD3FCC0FD88013F" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
|
||
<paragraph id="8BA236E41641FFC93AD3FCC0FC0E015F" blockId="12.[145,761,874,1080]" lastBlockId="12.[809,1425,198,1079]" pageId="12" pageNumber="13">
|
||
<emphasis id="B969EAF61641FFC93AD3FCC0FDD306E8" box="[145,570,874,896]" italics="true" pageId="12" pageNumber="13">
|
||
Description of the
|
||
<typeStatus id="54A688461641FFC93B28FCC1FE250617" box="[362,460,874,895]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13262A611641FFC93B9FFCC0FDDB0617" box="[477,562,874,896]" captionStart="Figure 3" captionStartId="12.[145,223,1850,1872]" captionTargetBox="[385,1185,1135,1809]" captionTargetId="figure@12.[385,1185,1135,1809]" captionTargetPageId="12" captionText="Figure 3. Holotypes of the three new Calumma species as preserved specimens.A, C. uetzi sp. nov. (ZSM 1688/2012). B, C. lefona sp. nov. (ZSM 2849/2010).C, C. juliae sp. nov. (ZSM 143/2016).Scale bar:20 mm." httpUri="https://zenodo.org/record/1226493/files/figure.png" pageId="12" pageNumber="13">Fig. 3B</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: Adult male, in a good state of preservation, left forelimb removed for DNA analysis, left hemipenis removed for micro-CT scanning; mouth slightly opened with tongue between the jaws; originally both hemipenes incompletely everted (
|
||
<figureCitation id="13262A611641FFC93BE7FBAFFE170171" box="[421,510,1027,1050]" captionStart="Figure 5" captionStartId="15.[163,242,696,718]" captionTargetBox="[227,1379,195,655]" captionTargetId="figure@15.[227,1379,195,655]" captionTargetPageId="15" captionText="Figure 5. DiceCT scans of hemipenes.A, right hemipenes of Calumma uetzi sp. nov. in asulcal (left) and sulcal (right) view.B, left hemipenes (incompletely everted) of C. lefona sp. nov. in asulcal (left) and sulcal (right) view; note that characters can be seen only in completely everted hemipenes. Scale bars:1.0 mm." httpUri="https://zenodo.org/record/1226497/files/figure.png" pageId="12" pageNumber="13">Fig. 5B</figureCitation>
|
||
), but left hemipenis cut off for micro-CT scanning and stored in a separate Eppendorf tube alongside the specimen; SVL
|
||
<quantity id="4CE59B011641FFC93927FF4EFC380592" box="[869,977,229,251]" metricMagnitude="-2" metricUnit="m" metricValue="5.13" pageId="12" pageNumber="13" unit="mm" value="51.3">51.3 mm</quantity>
|
||
, tail length
|
||
<quantity id="4CE59B011641FFC93E25FF4EFB3A0592" box="[1127,1235,229,251]" metricMagnitude="-2" metricUnit="m" metricValue="6.24" pageId="12" pageNumber="13" unit="mm" value="62.4">62.4 mm</quantity>
|
||
; see
|
||
<tableCitation id="C69F035F1641FFC93F4FFF4EFA810592" box="[1293,1384,229,251]" captionStart="Table 1" captionStartId="2.[9,73,551,572]" captionText="Table 1. Morphological measurements of the Calumma boettgeri complex" pageId="12" pageNumber="13">Table 1</tableCitation>
|
||
for other measurements; distinct and elevated rostral ridges that form a concave cup on the snout and fuse on the anterior snout at the base of a tapering, laterally compressed dermal rostral appendage that projects straight forward over a length of
|
||
<quantity id="4CE59B011641FFC93F77FED5FA7804FC" box="[1333,1425,382,404]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="12" pageNumber="13" unit="mm" value="5.6">5.6 mm</quantity>
|
||
with a diameter of 2.0 mm, pointed distally; 12 infralabial and 11 supralabial scales; supralabials dorsally serrated; no supra-orbital crest; distinct lateral crest running horizontally; short temporal crest consisting of two tubercles per side; indistinct parietal crest; occipital lobes clearly developed and completely separated by a deep, ‘U’-shaped notch (
|
||
<quantity id="4CE59B011641FFC93939FDDFFC3E07E1" box="[891,983,628,649]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="mm" value="1.8">1.8 mm</quantity>
|
||
); casque raised; dorsal crest present, starting
|
||
<quantity id="4CE59B011641FFC939D2FD39FC0007C0" box="[912,1001,658,680]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="13" unit="mm" value="0.8">0.8 mm</quantity>
|
||
from the base of the notch between the occipital lobes, consisting of a row of 23 separated, small conical scales spaced at irregular intervals from
|
||
<quantity id="4CE59B011641FFC9392EFD45FBF4066C" box="[876,1053,750,772]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="9.0" metricValueMin="1.0" pageId="12" pageNumber="13" unit="mm" value="0.5" valueMax="0.9" valueMin="0.1">0.1 to 0.9 mm</quantity>
|
||
, and several more on the tail decreasing in size towards the tip; no traces of gular or ventral crest. Body laterally compressed, with fine homogeneous scalation with the exception of the extremities and head region; limbs with rounded tubercle scales with a maximal diameter of
|
||
<quantity id="4CE59B011641FFC93F70FC2CFA6406F5" box="[1330,1421,903,925]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="mm" value="0.5">0.5 mm</quantity>
|
||
; heterogeneous scalation on the head, with largest scale on temporal region with diameter of
|
||
<quantity id="4CE59B011641FFC93F74FC6EFA7806B2" box="[1334,1425,965,987]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="13" unit="mm" value="0.6">0.6 mm</quantity>
|
||
and 60 oval tubercle scales (diameter>
|
||
<quantity id="4CE59B011641FFC93F48FC4FFA8C0691" box="[1290,1381,996,1017]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="13" unit="mm" value="0.3">0.3 mm</quantity>
|
||
) on the right side of the rostral appendage; no axillary or inguinal pits.
|
||
</paragraph>
|
||
<caption id="DF62666C1641FFC93AD3F891FB8F0205" httpUri="https://zenodo.org/record/1226493/files/figure.png" pageId="12" pageNumber="13" startId="12.[145,223,1850,1872]" targetBox="[385,1185,1135,1809]" targetPageId="12">
|
||
<paragraph id="8BA236E41641FFC93AD3F891FB8F0205" blockId="12.[145,1425,1850,1901]" pageId="12" pageNumber="13">
|
||
<emphasis id="B969EAF61641FFC93AD3F891FF100227" bold="true" box="[145,249,1850,1872]" pageId="12" pageNumber="13">Figure 3.</emphasis>
|
||
Holotypes of the three new
|
||
<taxonomicName id="4C1D4D671641FFC93867F891FD640227" box="[549,653,1850,1871]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
||
<emphasis id="B969EAF61641FFC93867F891FD640227" box="[549,653,1850,1871]" italics="true" pageId="12" pageNumber="13">Calumma</emphasis>
|
||
</taxonomicName>
|
||
species as preserved specimens. A,
|
||
<emphasis id="B969EAF61641FFC93E40F891FB5E0227" bold="true" box="[1026,1207,1850,1871]" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C1D4D671641FFC93E40F891FBB00227" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[1026,1113,1850,1871]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="uetzi" status="sp. nov.">
|
||
<emphasis id="B969EAF61641FFC93E40F891FBB00227" bold="true" box="[1026,1113,1850,1871]" italics="true" pageId="12" pageNumber="13">C. uetzi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1641FFC93E22F891FB5E0227" box="[1120,1207,1850,1871]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(ZSM 1688/2012). B,
|
||
<emphasis id="B969EAF61641FFC93AD3F8FCFEBC0204" bold="true" box="[145,341,1879,1901]" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C1D4D671641FFC93AD3F8FCFF110205" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[145,248,1879,1901]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="lefona" status="sp. nov.">
|
||
<emphasis id="B969EAF61641FFC93AD3F8FCFF110205" bold="true" box="[145,248,1879,1901]" italics="true" pageId="12" pageNumber="13">C. lefona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1641FFC93ABCF8FCFEBC0204" box="[254,341,1879,1900]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(ZSM 2849/2010). C,
|
||
<emphasis id="B969EAF61641FFC93876F8FCFD1A0204" bold="true" box="[564,755,1879,1901]" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C1D4D671641FFC93876F8FCFD7F0205" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[564,662,1879,1901]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="juliae" status="sp. nov.">
|
||
<emphasis id="B969EAF61641FFC93876F8FCFD7F0205" bold="true" box="[564,662,1879,1901]" italics="true" pageId="12" pageNumber="13">C. juliae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1641FFC938DFF8FCFD1A0204" box="[669,755,1879,1900]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(ZSM 143/2016). Scale bar: 20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF62666C1640FFC83AE1F9ECFC4E03BA" httpUri="https://zenodo.org/record/1226495/files/figure.png" pageId="13" pageNumber="14" startId="13.[163,245,1607,1629]" targetBox="[323,1283,195,1566]" targetPageId="13">
|
||
<paragraph id="8BA236E41640FFC83AE1F9ECFC4E03BA" blockId="13.[163,1443,1607,1746]" pageId="13" pageNumber="14">
|
||
<emphasis id="B969EAF61640FFC83AE1F9ECFEF90334" bold="true" box="[163,272,1607,1629]" pageId="13" pageNumber="14">Figure 4.</emphasis>
|
||
Micro-computed tomography scans of the skulls of the three holotypes in dorsal and lateral view. A, male
|
||
<emphasis id="B969EAF61640FFC83AE1F9CFFE530311" bold="true" box="[163,442,1636,1658]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C1D4D671640FFC83AE1F9CFFEB30311" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[163,346,1636,1658]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="uetzi" status="sp. nov.">
|
||
<emphasis id="B969EAF61640FFC83AE1F9CFFEB30311" bold="true" box="[163,346,1636,1658]" italics="true" pageId="13" pageNumber="14">Calumma uetzi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1640FFC83B22F9CFFE530311" box="[352,442,1636,1657]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(ZSM 1688/2012). B, male
|
||
<emphasis id="B969EAF61640FFC8389EF9CFFC4E0311" bold="true" box="[732,935,1636,1658]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C1D4D671640FFC8389EF9CFFCAE0312" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[732,839,1636,1658]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="lefona" status="sp. nov.">
|
||
<emphasis id="B969EAF61640FFC8389EF9CFFCAE0312" bold="true" box="[732,839,1636,1658]" italics="true" pageId="13" pageNumber="14">C. lefona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1640FFC8390CF9CFFC4E0311" box="[846,935,1636,1657]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(ZSM 2849/2010). C, female
|
||
<emphasis id="B969EAF61640FFC83E9FF9CFFA4A0311" bold="true" box="[1245,1443,1636,1658]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C1D4D671640FFC83E9FF9CFFAAB0312" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[1245,1346,1636,1658]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="juliae" status="sp. nov.">
|
||
<emphasis id="B969EAF61640FFC83E9FF9CFFAAB0312" bold="true" box="[1245,1346,1636,1658]" italics="true" pageId="13" pageNumber="14">C. juliae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1640FFC83F08F9CFFA4A0311" box="[1354,1443,1636,1657]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(ZSM 143/2016). D, male
|
||
<emphasis id="B969EAF61640FFC83BE8F929FDF603FF" box="[426,543,1666,1687]" italics="true" pageId="13" pageNumber="14">C. boettgeri</emphasis>
|
||
(ZSM 440/2000). E, male
|
||
<emphasis id="B969EAF61640FFC8396EF929FC7003FF" box="[812,921,1666,1687]" italics="true" pageId="13" pageNumber="14">C. linotum</emphasis>
|
||
holotype. F, female
|
||
<emphasis id="B969EAF61640FFC83E2AF929FB3C03FF" box="[1128,1237,1666,1687]" italics="true" pageId="13" pageNumber="14">C. linotum</emphasis>
|
||
(ZSM 551/2001). G, male
|
||
<emphasis id="B969EAF61640FFC83A9EF934FEA703DC" box="[220,334,1695,1716]" italics="true" pageId="13" pageNumber="14">C. gehringi</emphasis>
|
||
holotype (ZSM 2851/2010). H, male
|
||
<emphasis id="B969EAF61640FFC83888F934FCCC03DC" box="[714,805,1695,1716]" italics="true" pageId="13" pageNumber="14">C. guibei</emphasis>
|
||
(ZSM 2855/2010). Diagnostic characters are encircled in red. Abbreviations are given in the Material and Methods. Scale bars: 2.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA236E41640FFCB3AE1F955FDCB07E1" blockId="13.[163,779,1790,1874]" lastBlockId="14.[145,761,198,650]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
<emphasis id="B969EAF61640FFC83AE1F955FE22025A" italics="true" pageId="13" pageNumber="14">
|
||
Skull osteology of the
|
||
<typeStatus id="54A688461640FFC83BE9F955FDE7027B" box="[427,526,1790,1811]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13262A611640FFC83862F954FD9F027C" box="[544,630,1791,1812]" captionStart="Figure 4" captionStartId="13.[163,245,1607,1629]" captionTargetBox="[323,1283,195,1566]" captionTargetId="figure@13.[323,1283,195,1566]" captionTargetPageId="13" captionText="Figure 4. Micro-computed tomography scans of the skulls of the three holotypes in dorsal and lateral view. A, male Calumma uetzi sp. nov. (ZSM 1688/2012). B, male C. lefona sp. nov. (ZSM 2849/2010). C, female C. juliae sp. nov. (ZSM 143/2016). D, male C.boettgeri (ZSM 440/2000).E, male C.linotum holotype.F, female C.linotum (ZSM 551/2001).G, male C.gehringi holotype (ZSM 2851/2010).H, male C.guibei (ZSM 2855/2010).Diagnostic characters are encircled in red. Abbreviations are given in the Material and Methods.Scale bars:2.0 mm." httpUri="https://zenodo.org/record/1226495/files/figure.png" pageId="13" pageNumber="14">Fig. 4B</figureCitation>
|
||
, Supporting information, Video S3)
|
||
</emphasis>
|
||
: Skull length
|
||
<quantity id="4CE59B011640FFC838D1F8B6FCEF025B" box="[659,774,1821,1843]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="13" pageNumber="14" unit="mm" value="13.1">13.1 mm</quantity>
|
||
; snout–casque length
|
||
<quantity id="4CE59B011640FFC83BE1F897FDE60239" box="[419,527,1852,1874]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="13" pageNumber="14" unit="mm" value="15.9">15.9 mm</quantity>
|
||
; broad paired nasals meeting anteriorly, anterior tip of frontal exceeding more than half of the naris; prefrontal fontanelle and naris separated by contact of prefrontal with maxilla; elevated maxillae building a rectangular edge at anterior margin; prominent prefrontals with laterally raised tubercles; frontal and parietal with several tubercles, five on the midline forming a parietal crest; frontal with a width of
|
||
<quantity id="4CE59B011643FFCB3BAFFEEAFDAC043F" box="[493,581,321,343]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="14" pageNumber="15" unit="mm" value="2.9">2.9 mm</quantity>
|
||
(22.1% of SkL) at border to prefrontal extending to 4.0 mm (30.5% of SkL) at border to postorbitofrontal; frontoparietal fenestra with transverse diameter of
|
||
<quantity id="4CE59B011643FFCB3808FE36FD4A04DA" box="[586,675,413,434]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="15" unit="mm" value="1.9">1.9 mm</quantity>
|
||
(14.7% of SkL); parietal tapering more or less constantly, with a width of
|
||
<quantity id="4CE59B011643FFCB3B0EFE71FE4D0498" box="[332,420,474,496]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="14" pageNumber="15" unit="mm" value="3.9">3.9 mm</quantity>
|
||
(29.8% of SkL) at the border to frontal and
|
||
<quantity id="4CE59B011643FFCB3B7AFE52FE670766" box="[312,398,505,526]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="14" pageNumber="15" unit="mm" value="2.4">2.4 mm</quantity>
|
||
(18.3% of SkL) at its midpoint, and then tapering rather strongly to the posterior tip; posterodorsally directed parietal meets the squamosal laterally; squamosal thick, with several tubercles. For further measurements, see
|
||
<tableCitation id="C69F035F1643FFCB3B8AFDDFFDF507E1" box="[456,540,628,650]" captionStart="Table 2" captionStartId="5.[8,72,637,658]" captionText="Table 2. Osteological measurements of the Calumma boettgeri complex" pageId="14" pageNumber="15">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA236E41643FFCB3AD3FD09FE3406E7" blockId="14.[145,761,674,911]" pageId="14" pageNumber="15">
|
||
<emphasis id="B969EAF61643FFCB3AD3FD09FD5007D0" box="[145,697,674,696]" italics="true" pageId="14" pageNumber="15">
|
||
Coloration of the
|
||
<typeStatus id="54A688461643FFCB3B0CFD09FE4507DF" box="[334,428,674,695]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
in preservative (
|
||
<figureCitation id="13262A611643FFCB3821FD08FD5B07DF" box="[611,690,674,696]" captionStart="Figure 3" captionStartId="12.[145,223,1850,1872]" captionTargetBox="[385,1185,1135,1809]" captionTargetId="figure@12.[385,1185,1135,1809]" captionTargetPageId="12" captionText="Figure 3. Holotypes of the three new Calumma species as preserved specimens.A, C. uetzi sp. nov. (ZSM 1688/2012). B, C. lefona sp. nov. (ZSM 2849/2010).C, C. juliae sp. nov. (ZSM 143/2016).Scale bar:20 mm." httpUri="https://zenodo.org/record/1226493/files/figure.png" pageId="14" pageNumber="15">Fig. 3B</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: The body of the
|
||
<typeStatus id="54A688461643FFCB3B5AFD6AFE9207BF" box="[280,379,705,727]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
in preservative is of a grey–blue colour, with an indistinct beige lateral stripe; ventral and temporal regions and throat also beige; dark blue line from dorsal part of rostral appendage crossing the eyes and the lateral crest, ending in occipital lobes; extremities speckled with bluish tubercle scales. The coloration in life is unknown.
|
||
</paragraph>
|
||
<paragraph id="8BA236E41643FFCB3AD3FC03FD88013F" blockId="14.[145,761,936,1111]" pageId="14" pageNumber="15">
|
||
<emphasis id="B969EAF61643FFCB3AD3FC03FD5B06D5" box="[145,690,936,957]" italics="true" pageId="14" pageNumber="15">Hemipenial morphology based on diceCT scans</emphasis>
|
||
: The hemipenis of the
|
||
<typeStatus id="54A688461643FFCB3B1FFC6DFE2806B4" box="[349,449,966,988]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
was incompletely everted, and a diceCT scan resulted in an inadequate illustration (
|
||
<figureCitation id="13262A611643FFCB3ADBFBAFFF050171" box="[153,236,1027,1050]" captionStart="Figure 5" captionStartId="15.[163,242,696,718]" captionTargetBox="[227,1379,195,655]" captionTargetId="figure@15.[227,1379,195,655]" captionTargetPageId="15" captionText="Figure 5. DiceCT scans of hemipenes.A, right hemipenes of Calumma uetzi sp. nov. in asulcal (left) and sulcal (right) view.B, left hemipenes (incompletely everted) of C. lefona sp. nov. in asulcal (left) and sulcal (right) view; note that characters can be seen only in completely everted hemipenes. Scale bars:1.0 mm." httpUri="https://zenodo.org/record/1226497/files/figure.png" pageId="14" pageNumber="15">Fig. 5B</figureCitation>
|
||
). Apparently, it showed deep calyces and two pairs of rotulae; changing the threshold revealed two pairs of cornucula that were not everted.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307656F1643FFCB3AD3FBDBFEDE018A" pageId="14" pageNumber="15" type="diagnosis">
|
||
<paragraph id="8BA236E41643FFCB3AD3FBDBFEDE018A" blockId="14.[145,761,1136,1250]" pageId="14" pageNumber="15">
|
||
<emphasis id="B969EAF61643FFCB3AD3FBDBFEBA01ED" box="[145,339,1136,1157]" italics="true" pageId="14" pageNumber="15">Available names</emphasis>
|
||
: Apart from
|
||
<emphasis id="B969EAF61643FFCB3BB8FBDBFD9401ED" box="[506,637,1136,1157]" italics="true" pageId="14" pageNumber="15">C. gehringi</emphasis>
|
||
,
|
||
<emphasis id="B969EAF61643FFCB38C8FBDBFD1001ED" box="[650,761,1136,1157]" italics="true" pageId="14" pageNumber="15">C. guibei,</emphasis>
|
||
and
|
||
<taxonomicName id="4C1D4D671643FFCB3A81FB24FEF001CC" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[195,281,1167,1188]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="uetzi" status="sp. nov.">
|
||
<emphasis id="B969EAF61643FFCB3A81FB24FEF001CC" box="[195,281,1167,1188]" italics="true" pageId="14" pageNumber="15">C. uetzi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1643FFCB3B62FB24FE9F01CC" box="[288,374,1167,1188]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, there is no other valid species or synonym in the
|
||
<emphasis id="B969EAF61643FFCB3B04FB06FE2E01AB" box="[326,455,1197,1219]" italics="true" pageId="14" pageNumber="15">C. nasutum</emphasis>
|
||
group with deeply notched occipital lobes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307656F1643FFCB3AD3FB51FEC8008E" pageId="14" pageNumber="15" type="etymology">
|
||
<paragraph id="8BA236E41643FFCB3AD3FB51FEC8008E" blockId="14.[145,761,1274,1510]" pageId="14" pageNumber="15">
|
||
<emphasis id="B969EAF61643FFCB3AD3FB51FEFA0067" box="[145,275,1274,1295]" italics="true" pageId="14" pageNumber="15">Etymology</emphasis>
|
||
:
|
||
<taxonomicName id="4C1D4D671643FFCB3B70FB51FDEB0067" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[306,514,1274,1295]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="lefona" status="sp. nov.">
|
||
<emphasis id="B969EAF61643FFCB3B70FB51FDEB0067" box="[306,514,1274,1295]" italics="true" pageId="14" pageNumber="15">Calumma lefona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1643FFCB384EFB50FD800078" box="[524,617,1275,1296]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is the only species in the
|
||
<emphasis id="B969EAF61643FFCB3B70FAB2FE5C0047" box="[306,437,1305,1327]" italics="true" pageId="14" pageNumber="15">C. nasutum</emphasis>
|
||
group with a relatively long and pointed/constantly tapering rostral appendage (with the exception of
|
||
<emphasis id="B969EAF61643FFCB3BD4FAFCFE140003" box="[406,509,1366,1388]" italics="true" pageId="14" pageNumber="15">C. gallus</emphasis>
|
||
). This shape reminds of a spearhead; accordingly, we chose the Malagasy word ‘lefona’ (meaning ‘spear’) as its species epithet. It is used as an invariable noun in apposition to the genus name.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307656F1643FFCB3AD3FA54FA840450" pageId="14" pageNumber="15" type="distribution">
|
||
<paragraph id="8BA236E41643FFCB3AD3FA54FD740384" blockId="14.[145,761,1535,1772]" pageId="14" pageNumber="15">
|
||
<emphasis id="B969EAF61643FFCB3AD3FA54FEF5037C" box="[145,284,1535,1556]" italics="true" pageId="14" pageNumber="15">Distribution</emphasis>
|
||
:
|
||
<taxonomicName id="4C1D4D671643FFCB3B76FA54FE19037C" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[308,496,1535,1556]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="lefona" status="sp. nov.">
|
||
<emphasis id="B969EAF61643FFCB3B76FA54FE19037C" box="[308,496,1535,1556]" italics="true" pageId="14" pageNumber="15">Calumma lefona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1643FFCB3BB6F9ABFDAC037D" box="[500,581,1536,1557]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is, so far, known from only a single location in northern
|
||
<collectingCountry id="F30A76741643FFCB3824F9B5FD1C035C" box="[614,757,1566,1588]" name="Madagascar" pageId="14" pageNumber="15">Madagascar</collectingCountry>
|
||
, southeast of the Tsaratanana Massif, called Andrevorevo, at
|
||
<quantity id="4CE59B011643FFCB3B0DF9F7FE4D0319" box="[335,420,1628,1649]" metricMagnitude="3" metricUnit="m" metricValue="1.7169999999999999" pageId="14" pageNumber="15" unit="m" value="1717.0">1717 m</quantity>
|
||
a.s.l. (
|
||
<figureCitation id="13262A611643FFCB3BB3F9F0FDD90319" box="[497,560,1627,1649]" captionStart="Figure6" captionStartId="16.[145,223,1318,1340]" captionTargetBox="[385,1185,206,1277]" captionTargetId="figure@16.[385,1185,195,1277]" captionTargetPageId="16" captionText="Figure6. DistributionmapofsevenspeciesoftheCalummaboettgericomplex inMadagascar.The typelocalityisunknown for Calumma linotum. Contour lines indicate steps of 200 m elevation. The record from Nosy Komba is based on Hyde Roberts & Daly (2014). Coordinates for Calumma boettgeri, Maromiandra (Nagy et al., 2012) in Prötzel et al. (2015) are corrected to 13.9965°S, 48.2177°E." httpUri="https://zenodo.org/record/1226499/files/figure.png" pageId="14" pageNumber="15">Fig. 6</figureCitation>
|
||
; for geographical coordinates see the ‘holotype’ details above). It lives within the lower elevational range of its sister taxon
|
||
<emphasis id="B969EAF61643FFCB3AD3F913FF1703A4" box="[145,254,1719,1741]" italics="true" pageId="14" pageNumber="15">C. guibei</emphasis>
|
||
(from
|
||
<quantity id="4CE59B011643FFCB3B16F91CFE1103A5" box="[340,504,1719,1741]" metricMagnitude="3" metricUnit="m" metricValue="1.92" metricValueMax="2.25" metricValueMin="1.59" pageId="14" pageNumber="15" unit="m" value="1920.0" valueMax="2250.0" valueMin="1590.0">1590–2250 m</quantity>
|
||
a.s.l.) and at higher elevation than
|
||
<emphasis id="B969EAF61643FFCB3B7EF97DFE510383" box="[316,440,1750,1771]" italics="true" pageId="14" pageNumber="15">C. gehringi</emphasis>
|
||
(
|
||
<quantity id="4CE59B011643FFCB3B85F97DFDBB0384" box="[455,594,1750,1772]" metricMagnitude="3" metricUnit="m" metricValue="1.135" metricValueMax="1.54" metricValueMin="0.73" pageId="14" pageNumber="15" unit="m" value="1135.0" valueMax="1540.0" valueMin="730.0">730–1540 m</quantity>
|
||
a.s.l.).
|
||
</paragraph>
|
||
<paragraph id="8BA236E41643FFCB3AD3F8AEFA840450" blockId="14.[145,762,1797,1880]" lastBlockId="14.[809,1425,198,312]" pageId="14" pageNumber="15">
|
||
<emphasis id="B969EAF61643FFCB3AD3F8AEFE270272" box="[145,462,1797,1818]" italics="true" pageId="14" pageNumber="15">Natural history and ecology</emphasis>
|
||
:
|
||
<taxonomicName id="4C1D4D671643FFCB3BA3F8AEFD760272" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[481,671,1797,1818]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="lefona" status="sp. nov.">
|
||
<emphasis id="B969EAF61643FFCB3BA3F8AEFD760272" box="[481,671,1797,1818]" italics="true" pageId="14" pageNumber="15">Calumma lefona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1643FFCB38E5F8AEFD130272" box="[679,762,1797,1818]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was found roosting at night on tree branches ~
|
||
<quantity id="4CE59B011643FFCB38C9F88FFD5D0251" box="[651,692,1828,1849]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="m" value="2.0">2 m</quantity>
|
||
above the ground. The habitat consisted of primary forest with closed canopy cover and small streams. During the nocturnal observations on
|
||
<date id="FFA310241643FFCB3E3AFF4EFAB70592" box="[1144,1374,229,251]" pageId="14" pageNumber="15" value="2010-06-20" valueMax="2010-06-21" valueMin="2010-06-20">20 and 21 June 2010</date>
|
||
, two adults and two juveniles were found along the forest path, but only one adult (the
|
||
<typeStatus id="54A688461643FFCB3E2BFE89FB210450" box="[1129,1224,290,312]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
) was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307656F1643FFCB396BFEFAFB430777" pageId="14" pageNumber="15" type="conservation">
|
||
<paragraph id="8BA236E41643FFCB396BFEFAFB430777" blockId="14.[809,1426,337,543]" pageId="14" pageNumber="15">
|
||
<emphasis id="B969EAF61643FFCB396BFEFAFB6E040E" box="[809,1159,337,358]" italics="true" pageId="14" pageNumber="15">Recommended IUCN status</emphasis>
|
||
: Practically no data are available on the distribution and condition of
|
||
<taxonomicName id="4C1D4D671643FFCB396BFE25FC7A04CB" authority="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw, 2018" authorityName="David Prötzel & Miguel Vences & Oliver Hawlitschek & Mark D. Scherz & Fanomezana M. Ratsoavina & Frank Glaw" authorityYear="2018" box="[809,915,398,419]" class="Reptilia" family="Chamaeleonidae" genus="Calumma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="lefona" status="sp. nov.">
|
||
<emphasis id="B969EAF61643FFCB396BFE25FC7A04CB" box="[809,915,398,419]" italics="true" pageId="14" pageNumber="15">C. lefona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A578D1643FFCB39DEFE24FC1004CC" box="[924,1017,399,420]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and the species is known from a single specimen and a few additional observations. To avoid inflation of perceived risk, we recommend that this species be considered Data Deficient by the IUCN until more data become available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |