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<mods:title id="6BF1409835F683116E1EB8773D23F478">New species and records of Chasmogenus Sharp, 1882 from the southwestern margin of the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae)</mods:title>
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<mods:namePart id="32EA42C097E888F4D22F04C9A39F4A1C">Glynn, Rachel D.</mods:namePart>
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<paragraph id="8BA236B67F14FFC0FF3016D56A80A036" blockId="7.[151,1437,1019,1082]" pageId="7" pageNumber="442">
<heading id="D0EA81DA7F14FFC0FF3016D56A80A036" bold="true" centered="true" fontSize="11" level="1" pageId="7" pageNumber="442" reason="1">
<emphasis id="B969EAA47F14FFC0FF3016D56A80A036" bold="true" pageId="7" pageNumber="442">
Key to the species of
<taxonomicName id="4C1D4D357F14FFC0FE3916D269FAA01A" ID-CoL="8VW66" authority="Sharp" authorityName="Sharp" authorityYear="1882" box="[414,669,1019,1046]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAA47F14FFC0FE3916D2692EA01A" bold="true" box="[414,585,1019,1046]" italics="true" pageId="7" pageNumber="442">Chasmogenus</emphasis>
Sharp
</taxonomicName>
of
<collectingCountry id="F30A76267F14FFC0FD6316D56823A01A" box="[708,836,1020,1046]" name="Venezuela" pageId="7" pageNumber="442">Venezuela</collectingCountry>
,
<collectingCountry id="F30A76267F14FFC0FCF616D568AAA01A" box="[849,973,1020,1046]" name="Suriname" pageId="7" pageNumber="442">Suriname</collectingCountry>
,
<collectingCountry id="F30A76267F14FFC0FC7D16D56F58A01A" box="[986,1087,1020,1046]" name="Guyana" pageId="7" pageNumber="442">Guyana</collectingCountry>
, French Guiana, and
<collectingCountry id="F30A76267F14FFC0FAF716D56EFAA01A" box="[1360,1437,1020,1046]" name="Brazil" pageId="7" pageNumber="442">Brazil</collectingCountry>
north of the Amazon River
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C307653D7F14FFC0FF30114069AAA0E2" pageId="7" pageNumber="442" type="discussion">
<paragraph id="8BA236B67F14FFC0FF30114069AAA0E2" blockId="7.[151,1436,1128,1263]" pageId="7" pageNumber="442">
We have modified the key presented in
<bibRefCitation id="EF8C4B477F14FFC0FDFB11416832A08E" author="Smith, R. R. &amp; Short, A. E. Z." box="[604,853,1128,1154]" pageId="7" pageNumber="442" pagination="25 - 79" refId="ref5625" refString="Smith, R. R. &amp; Short, A. E. Z. (2020) Review of the genus Chasmogenus Sharp, 1882 of northeastern South America with an emphasis on Venezuela, Suriname, and Guyana (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys, 934, 25 - 79. https: // doi. org / 10.3897 / zookeys. 934.49359" type="journal article" year="2020">Smith &amp; Short (2020)</bibRefCitation>
to insert the three species described here. All three species would key to couplet
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the original key, so we have not repeated those here. We emphasize that examination of the genitalia is crucial to accurate identifications and unassociated females of the three species described here cannot be identified with precision.
</paragraph>
</subSubSection>
<subSubSection id="C307653D7F14FFCFFF3010306EFBA669" lastPageId="8" lastPageNumber="443" pageId="7" pageNumber="442" type="key">
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<paragraph id="8BA236B67F14FFC0FF3010306EFBA147" blockId="7.[151,1437,1305,2031]" pageId="7" pageNumber="442">
<keyLead id="30EC95837F14FFC0FF3010306EFBA147" pageId="7" pageNumber="442">
13 Aedeagus with apex of median lobe long, distinctly extending beyond the apex of the parameres; basal piece short, one-third the total length of the parameres or less (e.g.
<bibRefCitation id="EF8C4B477F14FFC0FD35101F683FA147" author="Smith, R. R. &amp; Short, A. E. Z." box="[658,856,1334,1356]" pageId="7" pageNumber="442" pagination="25 - 79" refId="ref5625" refString="Smith, R. R. &amp; Short, A. E. Z. (2020) Review of the genus Chasmogenus Sharp, 1882 of northeastern South America with an emphasis on Venezuela, Suriname, and Guyana (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys, 934, 25 - 79. https: // doi. org / 10.3897 / zookeys. 934.49359" type="journal article" year="2020">Smith &amp; Short 2020</bibRefCitation>
: fig. 15AE, 15GI)............................. 14
</keyLead>
</paragraph>
<paragraph id="8BA236B67F14FFC0FF30107D6EFBA18A" blockId="7.[151,1437,1305,2031]" pageId="7" pageNumber="442">
<keyLead id="30EC95837F14FFC0FF30107D6EFBA18A" pageId="7" pageNumber="442">- Aedeagus with apex of median lobe shorter or not extending beyond the apex of the parameres; basal piece long, up to one-half the length of the entire aedeagus........................................................................ 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E92E137F14FFC0FF3010856EFBA1F0" pageId="7" pageNumber="442">
<paragraph id="8BA236B67F14FFC0FF3010856EFBA1D3" blockId="7.[151,1437,1305,2031]" pageId="7" pageNumber="442">
<keyLead id="30EC95837F14FFC0FF3010856EFBA1D3" pageId="7" pageNumber="442">
14 Parameres of aedeagus smooth along outer edges, not sinuate or undulate, apex of median lobe acuminates at the apex (
<bibRefCitation id="EF8C4B477F14FFC0FAC610856EFBA1CE" author="Smith, R. R. &amp; Short, A. E. Z." box="[1377,1436,1452,1474]" pageId="7" pageNumber="442" pagination="25 - 79" refId="ref5625" refString="Smith, R. R. &amp; Short, A. E. Z. (2020) Review of the genus Chasmogenus Sharp, 1882 of northeastern South America with an emphasis on Venezuela, Suriname, and Guyana (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys, 934, 25 - 79. https: // doi. org / 10.3897 / zookeys. 934.49359" type="journal article" year="2020">Smith</bibRefCitation>
&amp; Short 2020: fig. 15H, I); gonopore situated toward the apex of the median lobe............
<taxonomicName id="4C1D4D357F14FFC0FBDD10E06EFBA1D3" ID-CoL="8XGFN" authority="Smith &amp; Short" authorityName="Smith &amp; Short" authorityYear="2020" box="[1146,1436,1481,1503]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="species" species="acuminatus">
<emphasis id="B969EAA47F14FFC0FBDD10E06E6DA1D2" bold="true" box="[1146,1290,1481,1502]" italics="true" pageId="7" pageNumber="442">C. acuminatus</emphasis>
Smith &amp; Short
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA236B67F14FFC0FF3010CE6EFBA1F0" blockId="7.[151,1437,1305,2031]" box="[151,1436,1510,1532]" pageId="7" pageNumber="442">
<keyLead id="30EC95837F14FFC0FF3010CE6EFBA1F0" box="[151,1436,1510,1532]" pageId="7" pageNumber="442">- Parameres moderately sinuate along outer margins; gonopore situated about in the middle of the median lobe............15</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8BA236B67F14FFC0FF3013086EFBA258" blockId="7.[151,1437,1305,2031]" pageId="7" pageNumber="442">
<keyLead id="30EC95837F14FFC0FF3013086EFBA258" pageId="7" pageNumber="442">
15 Median lobe of aedeagus terminating in a broadly rounded apex past the apex of the parameres (e.g.
<bibRefCitation id="EF8C4B477F14FFC0FB0B13086E14A23A" author="Smith, R. R. &amp; Short, A. E. Z." box="[1196,1395,1569,1591]" pageId="7" pageNumber="442" pagination="25 - 79" refId="ref5625" refString="Smith, R. R. &amp; Short, A. E. Z. (2020) Review of the genus Chasmogenus Sharp, 1882 of northeastern South America with an emphasis on Venezuela, Suriname, and Guyana (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys, 934, 25 - 79. https: // doi. org / 10.3897 / zookeys. 934.49359" type="journal article" year="2020">Smith &amp; Short 2020</bibRefCitation>
: fig. 15E);..........................................................................
<taxonomicName id="4C1D4D357F14FFC0FB3013176EFBA258" ID-CoL="8XGFZ" authority="Smith &amp; Short" authorityName="Smith &amp; Short" authorityYear="2020" box="[1175,1436,1598,1620]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="B969EAA47F14FFC0FB3013176E6DA258" bold="true" box="[1175,1290,1598,1620]" italics="true" pageId="7" pageNumber="442">C. ligulatus</emphasis>
Smith &amp; Short
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA236B67F14FFC0FF3013756EFBA27E" blockId="7.[151,1437,1305,2031]" box="[151,1436,1628,1650]" pageId="7" pageNumber="442">
<keyLead id="30EC95837F14FFC0FF3013756EFBA27E" box="[151,1436,1628,1650]" pageId="7" pageNumber="442">- Median lobe narrowing to a moderately to acutely pointed apex past the apex of the parameres...................... 16a</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E92E137F14FFC0FF3013BF6EFBA39A" pageId="7" pageNumber="442">
<paragraph id="8BA236B67F14FFC0FF3013BF6EFAA2C6" blockId="7.[151,1437,1305,2031]" pageId="7" pageNumber="442">
<keyLead id="30EC95837F14FFC0FF3013BF6EFAA2C6" pageId="7" pageNumber="442">
16a Median lobe with apex acuminate (e.g.
<bibRefCitation id="EF8C4B477F14FFC0FDC413BF684BA2A7" author="Smith, R. R. &amp; Short, A. E. Z." box="[611,812,1686,1708]" pageId="7" pageNumber="442" pagination="25 - 79" refId="ref5625" refString="Smith, R. R. &amp; Short, A. E. Z. (2020) Review of the genus Chasmogenus Sharp, 1882 of northeastern South America with an emphasis on Venezuela, Suriname, and Guyana (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys, 934, 25 - 79. https: // doi. org / 10.3897 / zookeys. 934.49359" type="journal article" year="2020">Smith &amp; Short 2020</bibRefCitation>
: fig. 15AD); emargination of the clypeus narrow (e.g.
<bibRefCitation id="EF8C4B477F14FFC0FAEE13BF6A37A2C5" author="Smith, R. R. &amp; Short, A. E. Z." pageId="7" pageNumber="442" pagination="25 - 79" refId="ref5625" refString="Smith, R. R. &amp; Short, A. E. Z. (2020) Review of the genus Chasmogenus Sharp, 1882 of northeastern South America with an emphasis on Venezuela, Suriname, and Guyana (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys, 934, 25 - 79. https: // doi. org / 10.3897 / zookeys. 934.49359" type="journal article" year="2020">Smith &amp; Short 2020</bibRefCitation>
: figs 5DF, 10D).......................................................................... 16b
</keyLead>
</paragraph>
<paragraph id="8BA236B67F14FFC0FF3013F86EFBA308" blockId="7.[151,1437,1305,2031]" pageId="7" pageNumber="442">
<keyLead id="30EC95837F14FFC0FF3013F86EFBA308" pageId="7" pageNumber="442">
- Median lobe with apex rounded (
<bibRefCitation id="EF8C4B477F14FFC0FD8013F86994A2EA" author="Smith, R. R. &amp; Short, A. E. Z." box="[551,755,1745,1767]" pageId="7" pageNumber="442" pagination="25 - 79" refId="ref5625" refString="Smith, R. R. &amp; Short, A. E. Z. (2020) Review of the genus Chasmogenus Sharp, 1882 of northeastern South America with an emphasis on Venezuela, Suriname, and Guyana (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys, 934, 25 - 79. https: // doi. org / 10.3897 / zookeys. 934.49359" type="journal article" year="2020">Smith &amp; Short 2020</bibRefCitation>
: fig. 15G); dorsal coloration dark brown, emargination of the clypeus rounded to angulate.............................................................
<taxonomicName id="4C1D4D357F14FFC0FB2D13C76EFBA308" ID-CoL="8XGG6" authority="Smith &amp; Short" authorityName="Smith &amp; Short" authorityYear="2020" box="[1162,1436,1774,1796]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B969EAA47F14FFC0FB2D13C76E6DA308" bold="true" box="[1162,1290,1774,1796]" italics="true" pageId="7" pageNumber="442">C. undulatus</emphasis>
Smith &amp; Short
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA236B67F14FFC0FF3012006EFBA350" blockId="7.[151,1437,1305,2031]" pageId="7" pageNumber="442">
<keyLead id="30EC95837F14FFC0FF3012006EFBA350" pageId="7" pageNumber="442">
<emphasis id="B969EAA47F14FFC0FF3012006BDBA332" bold="true" box="[151,188,1833,1854]" pageId="7" pageNumber="442">16b</emphasis>
Outer margins of parameres with two constrictions (bisinuate) (e.g.
<bibRefCitation id="EF8C4B477F14FFC0FCCC12006F27A332" author="Smith, R. R. &amp; Short, A. E. Z." box="[875,1088,1833,1855]" pageId="7" pageNumber="442" pagination="25 - 79" refId="ref5625" refString="Smith, R. R. &amp; Short, A. E. Z. (2020) Review of the genus Chasmogenus Sharp, 1882 of northeastern South America with an emphasis on Venezuela, Suriname, and Guyana (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys, 934, 25 - 79. https: // doi. org / 10.3897 / zookeys. 934.49359" type="journal article" year="2020">Smith and Short 2020</bibRefCitation>
: fig. 15AD); dorsal coloration dark reddish brown and translucent (e.g.
<bibRefCitation id="EF8C4B477F14FFC0FD9D126F6877A357" author="Smith, R. R. &amp; Short, A. E. Z." box="[570,784,1862,1884]" pageId="7" pageNumber="442" pagination="25 - 79" refId="ref5625" refString="Smith, R. R. &amp; Short, A. E. Z. (2020) Review of the genus Chasmogenus Sharp, 1882 of northeastern South America with an emphasis on Venezuela, Suriname, and Guyana (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys, 934, 25 - 79. https: // doi. org / 10.3897 / zookeys. 934.49359" type="journal article" year="2020">Smith and Short 2020</bibRefCitation>
: figs. 5DF, 10D)...................
<taxonomicName id="4C1D4D357F14FFC0FB01126F6EFBA350" ID-CoL="8XGG3" authority="Smith &amp; Short" authorityName="Smith &amp; Short" authorityYear="2020" box="[1190,1436,1862,1884]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="species" species="pandus">
<emphasis id="B969EAA47F14FFC0FB01126F6E6DA350" bold="true" box="[1190,1290,1862,1884]" italics="true" pageId="7" pageNumber="442">C. pandus</emphasis>
Smith &amp; Short
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA236B67F14FFC0FF30124D6EFBA39A" blockId="7.[151,1437,1305,2031]" pageId="7" pageNumber="442">
<keyLead id="30EC95837F14FFC0FF30124D6EFBA39A" pageId="7" pageNumber="442">
- Outer margins of parameres with a single constriction (sinuate) (fig. 4AD); dorsal coloration uniformly dark brown (fig. 2A, B)....................................................................................
<emphasis id="B969EAA47F14FFC0FB5212A86EFBA39A" bold="true" box="[1269,1436,1920,1942]" pageId="7" pageNumber="442">
<taxonomicName id="4C1D4D357F14FFC0FB5212A86E07A39A" ID-CoL="8XXDD" authority="Glynn &amp; Short, 2021" authorityName="Glynn &amp; Short" authorityYear="2021" box="[1269,1376,1920,1942]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="species" species="clinatus" status="sp. nov.">
<emphasis id="B969EAA47F14FFC0FB5212A86E07A39A" bold="true" box="[1269,1376,1920,1942]" italics="true" pageId="7" pageNumber="442">C. clinatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57DF7F14FFC0FAC112A86EFBA39A" box="[1382,1436,1921,1942]" pageId="7" pageNumber="442" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E92E137F14FFC0FF3012956EFBA3E2" pageId="7" pageNumber="442">
<paragraph id="8BA236B67F14FFC0FF3012956EFBA3DD" blockId="7.[151,1437,1305,2031]" box="[151,1436,1980,2002]" pageId="7" pageNumber="442">
<keyLead id="30EC95837F14FFC0FF3012956EFBA3DD" box="[151,1436,1980,2002]" pageId="7" pageNumber="442">17 Median lobe of aedeagus shorter than length of parameres, aedeagus symmetrical................................. 18</keyLead>
</paragraph>
<paragraph id="8BA236B67F14FFC0FF3012F06EFBA3E2" blockId="7.[151,1437,1305,2031]" box="[151,1436,2009,2031]" pageId="7" pageNumber="442">
<keyLead id="30EC95837F14FFC0FF3012F06EFBA3E2" box="[151,1436,2009,2031]" pageId="7" pageNumber="442">- Apex of median lobe of aedeagus approximately even with apex of parameres.................................... 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E92E137F1BFFCFFF3015B16EFBA4C7" pageId="8" pageNumber="443">
<paragraph id="8BA236B67F1BFFCFFF3015B16EFBA4A1" blockId="8.[151,1436,151,613]" box="[151,1436,151,174]" pageId="8" pageNumber="443">
<keyLead id="30EC95837F1BFFCFFF3015B16EFBA4A1" box="[151,1436,151,174]" pageId="8" pageNumber="443">
18 Apex of parameres bluntly squared (
<bibRefCitation id="EF8C4B477F1BFFCFFDE715BE6860A4A1" author="Smith, R. R. &amp; Short, A. E. Z." box="[576,775,151,173]" pageId="8" pageNumber="443" pagination="25 - 79" refId="ref5625" refString="Smith, R. R. &amp; Short, A. E. Z. (2020) Review of the genus Chasmogenus Sharp, 1882 of northeastern South America with an emphasis on Venezuela, Suriname, and Guyana (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys, 934, 25 - 79. https: // doi. org / 10.3897 / zookeys. 934.49359" type="journal article" year="2020">Smith &amp; Short 2020</bibRefCitation>
: fig. 14F)..........................
<taxonomicName id="4C1D4D357F1BFFCFFB0315BE6EFBA4A1" ID-CoL="8XGG4" authority="Smith &amp; Short" authorityName="Smith &amp; Short" authorityYear="2020" box="[1188,1436,151,173]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="species" species="schmits">
<emphasis id="B969EAA47F1BFFCFFB0315BE6E6CA4A1" bold="true" box="[1188,1291,151,173]" italics="true" pageId="8" pageNumber="443">C. schmits</emphasis>
Smith &amp; Short
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA236B67F1BFFCFFF30159C6EFBA4C7" blockId="8.[151,1436,151,613]" box="[151,1436,181,203]" pageId="8" pageNumber="443">
<keyLead id="30EC95837F1BFFCFFF30159C6EFBA4C7" box="[151,1436,181,203]" pageId="8" pageNumber="443">
- Apex of parameres rounded (
<bibRefCitation id="EF8C4B477F1BFFCFFE5D159C69A6A4C6" author="Smith, R. R. &amp; Short, A. E. Z." box="[506,705,181,203]" pageId="8" pageNumber="443" pagination="25 - 79" refId="ref5625" refString="Smith, R. R. &amp; Short, A. E. Z. (2020) Review of the genus Chasmogenus Sharp, 1882 of northeastern South America with an emphasis on Venezuela, Suriname, and Guyana (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys, 934, 25 - 79. https: // doi. org / 10.3897 / zookeys. 934.49359" type="journal article" year="2020">Smith &amp; Short 2020</bibRefCitation>
: fig. 14G, H)..........................
<taxonomicName id="4C1D4D357F1BFFCFFB24159C6EFBA4C7" ID-CoL="8XGFX" authority="Smith &amp; Short" authorityName="Smith &amp; Short" authorityYear="2020" box="[1155,1436,181,203]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis id="B969EAA47F1BFFCFFB24159C6E6DA4C6" bold="true" box="[1155,1290,181,202]" italics="true" pageId="8" pageNumber="443">C. guianensis</emphasis>
Smith &amp; Short
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E92E137F1BFFCFFF3015D96EFBA533" pageId="8" pageNumber="443">
<paragraph id="8BA236B67F1BFFCFFF3015D96EFBA52F" blockId="8.[151,1436,151,613]" pageId="8" pageNumber="443">
<keyLead id="30EC95837F1BFFCFFF3015D96EFBA52F" pageId="8" pageNumber="443">
19 Distinct dark patches on mesal portion of clypeus and labrum (
<bibRefCitation id="EF8C4B477F1BFFCFFC9B15C66F63A509" author="Smith, R. R. &amp; Short, A. E. Z." box="[828,1028,239,261]" pageId="8" pageNumber="443" pagination="25 - 79" refId="ref5625" refString="Smith, R. R. &amp; Short, A. E. Z. (2020) Review of the genus Chasmogenus Sharp, 1882 of northeastern South America with an emphasis on Venezuela, Suriname, and Guyana (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys, 934, 25 - 79. https: // doi. org / 10.3897 / zookeys. 934.49359" type="journal article" year="2020">Smith &amp; Short 2020</bibRefCitation>
: fig.10A)..................................................................................................
<taxonomicName id="4C1D4D357F1BFFCFFBEE14246EFBA52F" ID-CoL="8XGFR" authority="Smith &amp; Short" authorityName="Smith &amp; Short" authorityYear="2020" box="[1097,1436,268,291]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="species" species="brownsbergensis">
<emphasis id="B969EAA47F1BFFCFFBEE14246E6DA52E" bold="true" box="[1097,1290,268,290]" italics="true" pageId="8" pageNumber="443">C. brownsbergensis</emphasis>
Smith &amp; Short
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA236B67F1BFFCFFF3014026EFBA533" blockId="8.[151,1436,151,613]" box="[151,1436,298,320]" pageId="8" pageNumber="443">
<keyLead id="30EC95837F1BFFCFFF3014026EFBA533" box="[151,1436,298,320]" pageId="8" pageNumber="443">- Clypeus and labrum without distinct dark patches........................................................... 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E92E137F1BFFCFFF30144C6EFBA5B9" pageId="8" pageNumber="443">
<paragraph id="8BA236B67F1BFFCFFF30144C6EFBA576" blockId="8.[151,1436,151,613]" box="[151,1436,357,379]" pageId="8" pageNumber="443">
<keyLead id="30EC95837F1BFFCFFF30144C6EFBA576" box="[151,1436,357,379]" pageId="8" pageNumber="443">20 Aedeagus with parameres straight along outer margins, only curving inward past the position of the gonopore........... 21</keyLead>
</paragraph>
<paragraph id="8BA236B67F1BFFCFFF3014AA6EFBA5B9" blockId="8.[151,1436,151,613]" pageId="8" pageNumber="443">
<keyLead id="30EC95837F1BFFCFFF3014AA6EFBA5B9" pageId="8" pageNumber="443">
- Aedeagus with parameres that curve slightly inward along the outer margins throughout the entire length (figs. 5A, B)...............................................................................................
<emphasis id="B969EAA47F1BFFCFFABF14B66EFBA5B9" bold="true" box="[1304,1436,415,437]" pageId="8" pageNumber="443">
<taxonomicName id="4C1D4D357F1BFFCFFABF14B66E07A5B8" ID-CoL="8XXDG" authority="Glynn &amp; Short, 2021" authorityName="Glynn &amp; Short" authorityYear="2021" box="[1304,1376,415,436]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="species" species="inpa" status="sp. nov.">
<emphasis id="B969EAA47F1BFFCFFABF14B66E07A5B8" bold="true" box="[1304,1376,415,436]" italics="true" pageId="8" pageNumber="443">C. inpa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57DF7F1BFFCFFAC114896EFBA5B9" box="[1382,1436,416,437]" pageId="8" pageNumber="443" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E92E137F1BFFCFFF3014F36EFBA601" pageId="8" pageNumber="443">
<paragraph id="8BA236B67F1BFFCFFF3014F36EFBA5E3" blockId="8.[151,1436,151,613]" box="[151,1436,474,496]" pageId="8" pageNumber="443">
<keyLead id="30EC95837F1BFFCFFF3014F36EFBA5E3" box="[151,1436,474,496]" pageId="8" pageNumber="443">
21 Apex of median lobe rounded triangular in shape (figs. 5C, D)....................................
<emphasis id="B969EAA47F1BFFCFFB5F14F36EFBA5E3" bold="true" box="[1272,1436,474,495]" pageId="8" pageNumber="443">
<taxonomicName id="4C1D4D357F1BFFCFFB5F14F36E07A5E3" ID-CoL="8XXDF" authority="Glynn &amp; Short, 2021" authorityName="Glynn &amp; Short" authorityYear="2021" box="[1272,1376,474,495]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="species" species="gironae" status="sp. nov.">
<emphasis id="B969EAA47F1BFFCFFB5F14F36E07A5E3" bold="true" box="[1272,1376,474,495]" italics="true" pageId="8" pageNumber="443">C. gironae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57DF7F1BFFCFFAC114F36EFBA5E3" box="[1382,1436,474,495]" pageId="8" pageNumber="443" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BA236B67F1BFFCFFF3014D16EFBA601" blockId="8.[151,1436,151,613]" box="[151,1436,503,525]" pageId="8" pageNumber="443">
<keyLead id="30EC95837F1BFFCFFF3014D16EFBA601" box="[151,1436,503,525]" pageId="8" pageNumber="443">- Apex of median lobe acutely triangular in shape............................................................ 22</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E92E137F1BFFCFFF30171B6EFBA669" pageId="8" pageNumber="443">
<paragraph id="8BA236B67F1BFFCFFF30171B6EFBA644" blockId="8.[151,1436,151,613]" box="[151,1436,562,584]" pageId="8" pageNumber="443">
<keyLead id="30EC95837F1BFFCFFF30171B6EFBA644" box="[151,1436,562,584]" pageId="8" pageNumber="443">
22 Total body size ≤
<quantity id="4CE59B537F1BFFCFFE2A171B6ABEA644" box="[397,473,562,584]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="443" unit="mm" value="3.8">3.8 mm</quantity>
, dorsal coloration very dark brown................................. ..
<taxonomicName id="4C1D4D357F1BFFCFFB63171B6EFBA644" ID-CoL="8XGFW" authority="Smith &amp; Short" authorityName="Smith &amp; Short" authorityYear="2020" box="[1220,1436,562,584]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="species" species="gato">
<emphasis id="B969EAA47F1BFFCFFB63171B6E6CA64B" bold="true" box="[1220,1291,562,583]" italics="true" pageId="8" pageNumber="443">C. gato</emphasis>
Smith &amp; Short
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA236B67F1BFFCFFF3017796EFBA669" blockId="8.[151,1436,151,613]" box="[151,1436,591,613]" pageId="8" pageNumber="443">
<keyLead id="30EC95837F1BFFCFFF3017796EFBA669" box="[151,1436,591,613]" pageId="8" pageNumber="443">
- Total body length ≥
<quantity id="4CE59B537F1BFFCFFE0317796A97A669" box="[420,496,592,613]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="443" unit="mm" value="3.8">3.8 mm</quantity>
, dorsal coloration pale orange-brown.....................
<taxonomicName id="4C1D4D357F1BFFCFFBF517666E2FA669" ID-CoL="8XGFS" authority="Smith &amp; Short" authorityName="Smith &amp; Short" authorityYear="2020" box="[1106,1352,591,613]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="species" species="clavijoi">
<emphasis id="B969EAA47F1BFFCFFBF517666FD1A669" bold="true" box="[1106,1206,591,613]" italics="true" pageId="8" pageNumber="443">C. clavijoi</emphasis>
Smith &amp; Short
</taxonomicName>
(in part)
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</subSection>
</document>