221 lines
32 KiB
XML
221 lines
32 KiB
XML
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<mods:title id="69B468C80B2249FC7A92B0C4756F5A6B">Diadelophora, a new phorid genus from central and western Brazil (Diptera: Phoridae)</mods:title>
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<mods:namePart id="DD3396F065018421A6B3A9EF02676DCA">Ament, Danilo César</mods:namePart>
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<mods:date id="091BB1D61728BA18A73D9088FAE171F6">2024</mods:date>
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<subSubSection id="C3076501FF9E8158FF7ABFCF4D93FD6B" pageId="1" pageNumber="145" type="nomenclature">
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<paragraph id="8BA2368AFF9E8158FF7ABFCF4F9CFD36" blockId="1.[151,1004,643,737]" box="[151,435,643,670]" pageId="1" pageNumber="145">
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<heading id="D0EA81E6FF9E8158FF7ABFCF4F9CFD36" bold="true" box="[151,435,643,670]" fontSize="11" level="1" pageId="1" pageNumber="145" reason="1">
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<emphasis id="B969EA98FF9E8158FF7ABFCF4F9CFD36" bold="true" box="[151,435,643,670]" pageId="1" pageNumber="145">
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<taxonomicName id="4C1D4D09FF9E8158FF7ABFCF4F12FD36" authority="Ament, 2024" authorityName="Ament" authorityYear="2024" box="[151,317,643,670]" class="Insecta" family="Phoridae" genus="Diadelophora" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="145" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B969EA98FF9E8158FF7ABFCF4F12FD36" bold="true" box="[151,317,643,670]" italics="true" pageId="1" pageNumber="145">Diadelophora</emphasis>
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<taxonomicNameLabel id="A25A57E3FF9E8158FEA8BFC84F9CFD36" box="[325,435,644,670]" pageId="1" pageNumber="145" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA2368AFF9E8158FF7ABFE44D93FD6B" blockId="1.[151,1004,643,737]" box="[151,956,680,707]" pageId="1" pageNumber="145">
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<uri id="FF8C3A88FF9E8158FF7ABFE44D93FD6B" box="[151,956,680,707]" pageId="1" pageNumber="145">
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urn:lsid:zoobank.org:act:
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<uuid id="FFBB0C5FFF9E8158FE42BFE44D93FD6B" box="[431,956,680,707]" pageId="1" pageNumber="145">B10B5E22-8A65-4F36-879F-05033C1C6F9A</uuid>
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</uri>
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</paragraph>
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<subSubSection id="C3076501FF9E8158FF7ABF844DC3FD48" box="[151,1004,712,737]" pageId="1" pageNumber="145" type="description">
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<paragraph id="8BA2368AFF9E8158FF7ABF844DC3FD48" blockId="1.[151,1004,643,737]" box="[151,1004,712,737]" pageId="1" pageNumber="145">
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Type-species:
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<taxonomicName id="4C1D4D09FF9E8158FEC5BF844C20FD48" authority="Ament, 2024" authorityName="Ament" authorityYear="2024" box="[296,527,712,736]" class="Insecta" family="Phoridae" genus="Diadelophora" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="145" phylum="Arthropoda" rank="species" species="stilbella" status="sp. nov.">
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<emphasis id="B969EA98FF9E8158FEC5BF844C20FD48" box="[296,527,712,736]" italics="true" pageId="1" pageNumber="145">Diadelophora stilbella</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25A57E3FF9E8158FDF8BF854C4BFD48" box="[533,612,713,736]" pageId="1" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
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Refs:
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<bibRefCitation id="EF8C4B7BFF9E8158FD4EBF854D09FD49" author="Ament, D. C." box="[675,806,713,737]" pageId="1" pageNumber="145" pagination="151 - 188" refId="ref3930" refString="Ament, D. C. (2017) Phylogeny of Phorinae sensu latu (Diptera: Phoridae) inferred from a morphological analysis with comprehensive taxon sampling and an uncommon method of character coding. Zoological Journal of the Linnean Society, 181 (1), 151 - 188. https: // doi. org / 10.1093 / zoolinnean / zlx 038" type="journal article" year="2017">Ament 2017</bibRefCitation>
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:
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<taxonomicName id="4C1D4D09FF9E8158FCC2BF844D80FD48" authority="Ament, 2024" authorityName="Ament" authorityYear="2024" box="[815,943,712,736]" class="Insecta" family="Phoridae" genus="Hypocerides" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="145" phylum="Arthropoda" rank="genus" status="sp. n.">
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<emphasis id="B969EA98FF9E8158FCC2BF844D80FD48" box="[815,943,712,736]" italics="true" pageId="1" pageNumber="145">Hypocerides</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25A57E3FF9E8158FC58BF854DC3FD48" box="[949,1004,713,736]" pageId="1" pageNumber="145" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3076501FF9E8158FF7ABE424FB1FAC1" pageId="1" pageNumber="145" type="diagnosis">
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<paragraph id="8BA2368AFF9E8158FF7ABE424DC1FC3D" blockId="1.[151,1437,782,2033]" pageId="1" pageNumber="145">
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<emphasis id="B969EA98FF9E8158FF7ABE424F25FC80" bold="true" box="[151,266,782,808]" pageId="1" pageNumber="145">Diagnosis</emphasis>
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. This genus differs from other
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<taxonomicName id="4C1D4D09FF9E8158FDBBBE434C95FC81" box="[598,698,783,809]" class="Insecta" family="Phoridae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="145" phylum="Arthropoda" rank="subFamily" subFamily="Phorinae">Phorinae</taxonomicName>
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genera by the combination of characteristics: antennal flagellomere conical-shaped; frontal setae reduced with at least ventral fronto-orbitals lacking; costa long, apically swollen; and tibiae lacking large setae.
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<taxonomicName id="4C1D4D09FF9E8158FE55BE1B4C7AFCD9" authorityName="Ament" authorityYear="2024" box="[440,597,855,881]" class="Insecta" family="Phoridae" genus="Diadelophora" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis id="B969EA98FF9E8158FE55BE1B4C7AFCD9" box="[440,597,855,881]" italics="true" pageId="1" pageNumber="145">Diadelophora</emphasis>
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</taxonomicName>
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also has a characteristic hypopygium with epandrial left and right posterior margins invaginated, continuous with a curved process under the hypoproct.
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</paragraph>
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<paragraph id="8BA2368AFF9E8158FF2ABED24FE3FAED" blockId="1.[151,1437,782,2033]" pageId="1" pageNumber="145">
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<emphasis id="B969EA98FF9E8158FF2ABED24F8BFC10" bold="true" box="[199,420,926,952]" pageId="1" pageNumber="145">Genus description</emphasis>
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. Male. Frontal setae reduced, ventral fronto-orbitals absent, ventral interfrontals absent or present, supra-antennals, when present, almost imperceptible, reclinate (
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<figureCitation id="13262A0FFF9E8158FC15BE8F4A72FC75" box="[1016,1117,963,989]" captionStart="FIGURE 1" captionStartId="2.[151,250,1928,1952]" captionTargetBox="[153,1434,179,1902]" captionTargetId="figure-16@2.[153,1434,179,1902]" captionTargetPageId="2" captionText="FIGURE 1. Diadelophora stilbella sp. nov., male. A, habitus (holotype); B, head in dorsal view; C, palpus in dorsal view (MEV); D, thorax in lateral view (MEV); E, wing; F, hind femur; G, foretibia; H, midtibia; I, hind tibia." figureDoi="http://doi.org/10.5281/zenodo.10692685" httpUri="https://zenodo.org/record/10692685/files/figure.png" pageId="1" pageNumber="145">Figs. 1B</figureCitation>
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,
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<figureCitation id="13262A0FFF9E8158FB81BE8F4ABFFC75" box="[1132,1168,963,989]" captionStart="FIGURE 4" captionStartId="6.[151,250,1737,1761]" captionTargetBox="[151,1435,237,1713]" captionTargetId="figure-46@6.[151,1435,237,1714]" captionTargetPageId="6" captionText="FIGURE 4. Diadelophora inornata sp. nov., male. A, habitus (holotype); B, head in dorsal view; C, microscopic details of the first flagellomere of the antenna (MEV); D, thorax in lateral view; E, wing; F, hind femur; G, foretibia; H, midtibia; I, hind tibia." figureDoi="http://doi.org/10.5281/zenodo.10692694" httpUri="https://zenodo.org/record/10692694/files/figure.png" pageId="1" pageNumber="145">4B</figureCitation>
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). Flagellomere 1 large, conical, with several sensilla of different shapes (
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<figureCitation id="13262A0FFF9E8158FD53BEAB4D3AFBA9" box="[702,789,999,1025]" captionStart="FIGURE 4" captionStartId="6.[151,250,1737,1761]" captionTargetBox="[151,1435,237,1713]" captionTargetId="figure-46@6.[151,1435,237,1714]" captionTargetPageId="6" captionText="FIGURE 4. Diadelophora inornata sp. nov., male. A, habitus (holotype); B, head in dorsal view; C, microscopic details of the first flagellomere of the antenna (MEV); D, thorax in lateral view; E, wing; F, hind femur; G, foretibia; H, midtibia; I, hind tibia." figureDoi="http://doi.org/10.5281/zenodo.10692694" httpUri="https://zenodo.org/record/10692694/files/figure.png" pageId="1" pageNumber="145">Fig. 4C</figureCitation>
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), without subcutaneous pit sensilla (SPS); arista preapical (
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<figureCitation id="13262A0FFF9E8158FF72B9474F2AFB8D" box="[159,261,1035,1061]" captionStart="FIGURE 1" captionStartId="2.[151,250,1928,1952]" captionTargetBox="[153,1434,179,1902]" captionTargetId="figure-16@2.[153,1434,179,1902]" captionTargetPageId="2" captionText="FIGURE 1. Diadelophora stilbella sp. nov., male. A, habitus (holotype); B, head in dorsal view; C, palpus in dorsal view (MEV); D, thorax in lateral view (MEV); E, wing; F, hind femur; G, foretibia; H, midtibia; I, hind tibia." figureDoi="http://doi.org/10.5281/zenodo.10692685" httpUri="https://zenodo.org/record/10692685/files/figure.png" pageId="1" pageNumber="145">Figs. 1A</figureCitation>
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,
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<figureCitation id="13262A0FFF9E8158FEFFB9474F17FB8D" box="[274,312,1035,1061]" captionStart="FIGURE 4" captionStartId="6.[151,250,1737,1761]" captionTargetBox="[151,1435,237,1713]" captionTargetId="figure-46@6.[151,1435,237,1714]" captionTargetPageId="6" captionText="FIGURE 4. Diadelophora inornata sp. nov., male. A, habitus (holotype); B, head in dorsal view; C, microscopic details of the first flagellomere of the antenna (MEV); D, thorax in lateral view; E, wing; F, hind femur; G, foretibia; H, midtibia; I, hind tibia." figureDoi="http://doi.org/10.5281/zenodo.10692694" httpUri="https://zenodo.org/record/10692694/files/figure.png" pageId="1" pageNumber="145">4D</figureCitation>
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). Palpus large, with small ventral setulae (
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<figureCitation id="13262A0FFF9E8158FCF6B9474D9FFB8D" box="[795,944,1035,1061]" captionStart="FIGURE 1" captionStartId="2.[151,250,1928,1952]" captionTargetBox="[153,1434,179,1902]" captionTargetId="figure-16@2.[153,1434,179,1902]" captionTargetPageId="2" captionText="FIGURE 1. Diadelophora stilbella sp. nov., male. A, habitus (holotype); B, head in dorsal view; C, palpus in dorsal view (MEV); D, thorax in lateral view (MEV); E, wing; F, hind femur; G, foretibia; H, midtibia; I, hind tibia." figureDoi="http://doi.org/10.5281/zenodo.10692685" httpUri="https://zenodo.org/record/10692685/files/figure.png" pageId="1" pageNumber="145">Figs. 1A, 1C</figureCitation>
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). Body in general robust, with large thorax and abdomen and relatively thin legs. Anepisternum furrow absent, anterior spiracle large; meron dorsally slightly convex (
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<figureCitation id="13262A0FFF9E8158FF1BB91F4F61FBC5" box="[246,334,1107,1133]" captionStart="FIGURE 1" captionStartId="2.[151,250,1928,1952]" captionTargetBox="[153,1434,179,1902]" captionTargetId="figure-16@2.[153,1434,179,1902]" captionTargetPageId="2" captionText="FIGURE 1. Diadelophora stilbella sp. nov., male. A, habitus (holotype); B, head in dorsal view; C, palpus in dorsal view (MEV); D, thorax in lateral view (MEV); E, wing; F, hind femur; G, foretibia; H, midtibia; I, hind tibia." figureDoi="http://doi.org/10.5281/zenodo.10692685" httpUri="https://zenodo.org/record/10692685/files/figure.png" pageId="1" pageNumber="145">Fig. 1D</figureCitation>
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). Scutellum with two pairs of setae. Tibiae without isolated large setae (
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<figureCitation id="13262A0FFF9E8158FB8AB91F4ACEFBC5" box="[1127,1249,1107,1133]" captionStart="FIGURE 1" captionStartId="2.[151,250,1928,1952]" captionTargetBox="[153,1434,179,1902]" captionTargetId="figure-16@2.[153,1434,179,1902]" captionTargetPageId="2" captionText="FIGURE 1. Diadelophora stilbella sp. nov., male. A, habitus (holotype); B, head in dorsal view; C, palpus in dorsal view (MEV); D, thorax in lateral view (MEV); E, wing; F, hind femur; G, foretibia; H, midtibia; I, hind tibia." figureDoi="http://doi.org/10.5281/zenodo.10692685" httpUri="https://zenodo.org/record/10692685/files/figure.png" pageId="1" pageNumber="145">Figs. 1G–I</figureCitation>
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,
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<figureCitation id="13262A0FFF9E8158FB01B91F4B04FBC5" box="[1260,1323,1107,1133]" captionStart="FIGURE 4" captionStartId="6.[151,250,1737,1761]" captionTargetBox="[151,1435,237,1713]" captionTargetId="figure-46@6.[151,1435,237,1714]" captionTargetPageId="6" captionText="FIGURE 4. Diadelophora inornata sp. nov., male. A, habitus (holotype); B, head in dorsal view; C, microscopic details of the first flagellomere of the antenna (MEV); D, thorax in lateral view; E, wing; F, hind femur; G, foretibia; H, midtibia; I, hind tibia." figureDoi="http://doi.org/10.5281/zenodo.10692694" httpUri="https://zenodo.org/record/10692694/files/figure.png" pageId="1" pageNumber="145">4G–I</figureCitation>
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). Midtibia basal half with or without an anterodorsal and a posterodorsal subtle setal palisades. Hind femur dorso-ventrally narrow, with ventral row of elongated thin setae near base (
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<figureCitation id="13262A0FFF9E8158FCC5B9D74DACFB1D" box="[808,899,1179,1205]" captionStart="FIGURE 1" captionStartId="2.[151,250,1928,1952]" captionTargetBox="[153,1434,179,1902]" captionTargetId="figure-16@2.[153,1434,179,1902]" captionTargetPageId="2" captionText="FIGURE 1. Diadelophora stilbella sp. nov., male. A, habitus (holotype); B, head in dorsal view; C, palpus in dorsal view (MEV); D, thorax in lateral view (MEV); E, wing; F, hind femur; G, foretibia; H, midtibia; I, hind tibia." figureDoi="http://doi.org/10.5281/zenodo.10692685" httpUri="https://zenodo.org/record/10692685/files/figure.png" pageId="1" pageNumber="145">Figs. 1F</figureCitation>
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,
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<figureCitation id="13262A0FFF9E8158FC62B9D74D9FFB1D" box="[911,944,1179,1205]" captionStart="FIGURE 4" captionStartId="6.[151,250,1737,1761]" captionTargetBox="[151,1435,237,1713]" captionTargetId="figure-46@6.[151,1435,237,1714]" captionTargetPageId="6" captionText="FIGURE 4. Diadelophora inornata sp. nov., male. A, habitus (holotype); B, head in dorsal view; C, microscopic details of the first flagellomere of the antenna (MEV); D, thorax in lateral view; E, wing; F, hind femur; G, foretibia; H, midtibia; I, hind tibia." figureDoi="http://doi.org/10.5281/zenodo.10692694" httpUri="https://zenodo.org/record/10692694/files/figure.png" pageId="1" pageNumber="145">4F</figureCitation>
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). Hind tibia with a dorsal palisade of setulae (sometimes subtle, hard to observe). Costa apically swollen, Sc apparently absent, M
|
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<subScript id="179934CFFF9E8158FBD3B99D4A69FB77" attach="left" box="[1086,1094,1233,1247]" fontSize="6" pageId="1" pageNumber="145">1</subScript>
|
||
and M
|
||
<subScript id="179934CFFF9E8158FB7FB99D4AB5FB77" attach="left" box="[1170,1178,1233,1247]" fontSize="6" pageId="1" pageNumber="145">2</subScript>
|
||
sinuous (
|
||
<figureCitation id="13262A0FFF9E8158FAEEB9F34B4DFB71" box="[1283,1378,1215,1241]" captionStart="FIGURE 1" captionStartId="2.[151,250,1928,1952]" captionTargetBox="[153,1434,179,1902]" captionTargetId="figure-16@2.[153,1434,179,1902]" captionTargetPageId="2" captionText="FIGURE 1. Diadelophora stilbella sp. nov., male. A, habitus (holotype); B, head in dorsal view; C, palpus in dorsal view (MEV); D, thorax in lateral view (MEV); E, wing; F, hind femur; G, foretibia; H, midtibia; I, hind tibia." figureDoi="http://doi.org/10.5281/zenodo.10692685" httpUri="https://zenodo.org/record/10692685/files/figure.png" pageId="1" pageNumber="145">Figs. 1E</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0FFF9E8158FA80B9F34BBFFB71" box="[1389,1424,1215,1241]" captionStart="FIGURE 4" captionStartId="6.[151,250,1737,1761]" captionTargetBox="[151,1435,237,1713]" captionTargetId="figure-46@6.[151,1435,237,1714]" captionTargetPageId="6" captionText="FIGURE 4. Diadelophora inornata sp. nov., male. A, habitus (holotype); B, head in dorsal view; C, microscopic details of the first flagellomere of the antenna (MEV); D, thorax in lateral view; E, wing; F, hind femur; G, foretibia; H, midtibia; I, hind tibia." figureDoi="http://doi.org/10.5281/zenodo.10692694" httpUri="https://zenodo.org/record/10692694/files/figure.png" pageId="1" pageNumber="145">4E</figureCitation>
|
||
). Epandrial left and right posterior margins invaginated, continuous with a curved process under the hypoproct (
|
||
<figureCitation id="13262A0FFF9E8158FA82B9AF4E87FA89" captionStart="FIGURE 2" captionStartId="3.[151,250,1909,1933]" captionTargetBox="[167,1415,324,1867]" captionTargetId="figure-75@3.[151,1435,309,1886]" captionTargetPageId="3" captionText="FIGURE 2. Hypopygium of Diadelophora stilbella sp. nov.. A, dorsal; B, ventral; C, left lateral; D, right lateral.Abbreviations: cerc, cercus; ep, epandrium; hyp, hypandrium; hyprct, hypoproct; prj, projection." figureDoi="http://doi.org/10.5281/zenodo.10692688" httpUri="https://zenodo.org/record/10692688/files/figure.png" pageId="1" pageNumber="145">Fig. 2</figureCitation>
|
||
). Distiphallus complex; core plate extending over distiphallus; right arm curved reaching left lateral, connected to a scaled epiphallus (
|
||
<figureCitation id="13262A0FFF9E8158FE97B8674F91FAED" box="[378,446,1323,1349]" captionStart="FIGURE 3" captionStartId="4.[151,250,1706,1730]" captionTargetBox="[189,1398,597,1682]" captionTargetId="figure-144@4.[189,1398,597,1682]" captionTargetPageId="4" captionText="FIGURE 3. A, B, Phallus of Diadelophora stilbella sp. nov. in left lateral (A) and dorsal view (B); C, phallus of Diadelophora inornata sp. nov. in left lateral view. Abbreviations: bs, basiphallus; cp, core plate; epi, epiphallus; ra, right arm." figureDoi="http://doi.org/10.5281/zenodo.10692690" httpUri="https://zenodo.org/record/10692690/files/figure.png" pageId="1" pageNumber="145">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA2368AFF9E8158FF2AB8034FB1FAC1" blockId="1.[151,1437,782,2033]" box="[199,414,1359,1385]" pageId="1" pageNumber="145">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3076501FF9E815AFF2AB83F4FDEFEB5" lastPageId="3" lastPageNumber="147" pageId="1" pageNumber="145" type="etymology">
|
||
<paragraph id="8BA2368AFF9E8158FF2AB83F4C92FA19" blockId="1.[151,1437,782,2033]" pageId="1" pageNumber="145">
|
||
<emphasis id="B969EA98FF9E8158FF2AB83F4F67FA25" bold="true" box="[199,328,1395,1421]" pageId="1" pageNumber="145">Etymology</emphasis>
|
||
. Name derived from the Greek word
|
||
<emphasis id="B969EA98FF9E8158FD01B83F4D64FA25" box="[748,843,1395,1421]" italics="true" pageId="1" pageNumber="145">diadelos</emphasis>
|
||
(distinguishable, distinctive) referring to the unusual general body shape and chaetotaxy of this genus.
|
||
</paragraph>
|
||
<paragraph id="8BA2368AFF9E8158FF2AB8F64A29FA7D" blockId="1.[151,1437,782,2033]" box="[199,1030,1466,1493]" pageId="1" pageNumber="145">
|
||
<emphasis id="B969EA98FF9E8158FF2AB8F64DD0FA7D" bold="true" box="[199,1023,1466,1493]" pageId="1" pageNumber="145">Phylogenetic position and justification of the proposal of a new genus</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA2368AFF9E8158FF2AB8934F3DF905" blockId="1.[151,1437,782,2033]" pageId="1" pageNumber="145">
|
||
<bibRefCitation id="EF8C4B7BFF9E8158FF2AB8934F5DFA51" author="Brown, B. V." box="[199,370,1503,1529]" pageId="1" pageNumber="145" pagination="571 - 581" refId="ref4791" refString="Brown, B. V. (2022 b) Two new genera of phorid flies, Macgrathphora and Aurisetiphora, from Costa Rica (Diptera: Phoridae), with recommendations for naming new genera in the family. Zootaxa, 5115 (4), 571 - 581. https: // doi. org / 10.11646 / zootaxa. 5115.4.7" type="journal article" year="2022">Brown (2022b)</bibRefCitation>
|
||
provided a series of recommendations regarding in which cases new genera should be proposed in the
|
||
<taxonomicName id="4C1D4D09FF9E8158FF0CBB4F4F6AF9B5" box="[225,325,1539,1565]" class="Insecta" family="Phoridae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="145" phylum="Arthropoda" rank="family">Phoridae</taxonomicName>
|
||
family, including ‘…if there is good evidence (i.e. one or more synapomophies) that a unknown specimen is related to a known genus, then unless differences from the known genus are truly exceptional, I would be reluctant to recognize a new genus assignment. This is because there are two possibilities for the situation: the new specimens are part of a sister-taxon to the known genus, or it is something that has diverged extensively within the genus’.
|
||
</paragraph>
|
||
<paragraph id="8BA2368AFF9E8158FF2ABBFB4D3EF859" blockId="1.[151,1437,782,2033]" pageId="1" pageNumber="145">
|
||
Commonly during
|
||
<taxonomicName id="4C1D4D09FF9E8158FE4DBBFB4C2BF979" box="[416,516,1719,1745]" class="Insecta" family="Phoridae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="145" phylum="Arthropoda" rank="family">Phoridae</taxonomicName>
|
||
genera proposition, we do not have information about the phylogenetic position of the putative genus or even reliable information to compare it to closely related genera. Fortunately, in the case of
|
||
<taxonomicName id="4C1D4D09FF9E8158FF55BBB34F7AF8B1" authorityName="Ament" authorityYear="2024" box="[184,341,1791,1817]" class="Insecta" family="Phoridae" genus="Diadelophora" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA98FF9E8158FF55BBB34F7AF8B1" box="[184,341,1791,1817]" italics="true" pageId="1" pageNumber="145">Diadelophora</emphasis>
|
||
</taxonomicName>
|
||
, we have evidence supporting its sister-group relationship with
|
||
<taxonomicName id="4C1D4D09FF9E8158FBD6BBB34AE6F8B1" authorityName="Ament" authorityYear="2024" box="[1083,1225,1791,1817]" class="Insecta" family="Phoridae" genus="Hypocerides" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA98FF9E8158FBD6BBB34AE6F8B1" box="[1083,1225,1791,1817]" italics="true" pageId="1" pageNumber="145">Hypocerides</emphasis>
|
||
</taxonomicName>
|
||
with these genera sharing the possible synapomorphies: ventral fronto-orbitals absent, costa apically swollen, epandrial left and right posterior margins invaginated, continuous with a process under the hypoproct (
|
||
<bibRefCitation id="EF8C4B7BFF9E8158FBF3BA0B4A9DF8C9" author="Ament, D. C." box="[1054,1202,1863,1889]" pageId="1" pageNumber="145" pagination="151 - 188" refId="ref3930" refString="Ament, D. C. (2017) Phylogeny of Phorinae sensu latu (Diptera: Phoridae) inferred from a morphological analysis with comprehensive taxon sampling and an uncommon method of character coding. Zoological Journal of the Linnean Society, 181 (1), 151 - 188. https: // doi. org / 10.1093 / zoolinnean / zlx 038" type="journal article" year="2017">Ament 2017</bibRefCitation>
|
||
). Concomitantly, the differences between these genera may be considered truly exceptional.
|
||
<taxonomicName id="4C1D4D09FF9E8158FC43BA274A64F82D" authorityName="Ament" authorityYear="2024" box="[942,1099,1899,1925]" class="Insecta" family="Phoridae" genus="Diadelophora" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA98FF9E8158FC43BA274A64F82D" box="[942,1099,1899,1925]" italics="true" pageId="1" pageNumber="145">Diadelophora</emphasis>
|
||
</taxonomicName>
|
||
has a much larger and distinct body general shape, conical antenna, palpus large with small ventral setulae, long costa which is also less thickened apically, sinuous wing vein M
|
||
<subScript id="179934CFFF9E8158FE0FBA894FC5F87B" attach="left" box="[482,490,1989,2003]" fontSize="6" pageId="1" pageNumber="145">1</subScript>
|
||
, and epandrium, hypandrium, surstyli and phallus morphology substantially different from
|
||
<taxonomicName id="4C1D4D09FF9E8158FF38BA9B4F4CF859" authorityName="Ament" authorityYear="2024" box="[213,355,2007,2033]" class="Insecta" family="Phoridae" genus="Hypocerides" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA98FF9E8158FF38BA9B4F4CF859" box="[213,355,2007,2033]" italics="true" pageId="1" pageNumber="145">Hypocerides</emphasis>
|
||
</taxonomicName>
|
||
(as described in the previous section).
|
||
</paragraph>
|
||
<caption id="DF626602FF9D815BFF7ABAC44A93F86C" ID-DOI="http://doi.org/10.5281/zenodo.10692685" ID-Zenodo-Dep="10692685" httpUri="https://zenodo.org/record/10692685/files/figure.png" pageId="2" pageNumber="146" startId="2.[151,250,1928,1952]" targetBox="[153,1434,179,1902]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BA2368AFF9D815BFF7ABAC44A93F86C" blockId="2.[151,1436,1928,1988]" pageId="2" pageNumber="146">
|
||
<emphasis id="B969EA98FF9D815BFF7ABAC44F3FF808" bold="true" box="[151,272,1928,1952]" pageId="2" pageNumber="146">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="4C1D4D09FF9D815BFEF2BAC44C27F808" authority="Ament, 2024" authorityName="Ament" authorityYear="2024" box="[287,520,1928,1952]" class="Insecta" family="Phoridae" genus="Diadelophora" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="146" phylum="Arthropoda" rank="species" species="stilbella" status="sp. nov.">
|
||
<emphasis id="B969EA98FF9D815BFEF2BAC44C27F808" box="[287,520,1928,1952]" italics="true" pageId="2" pageNumber="146">Diadelophora stilbella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A57E3FF9D815BFDFDBAC44C4BF837" box="[528,612,1928,1951]" pageId="2" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male.
|
||
<emphasis id="B969EA98FF9D815BFD5DBAC44CEDF808" bold="true" box="[688,706,1928,1952]" pageId="2" pageNumber="146">A</emphasis>
|
||
, habitus (holotype);
|
||
<emphasis id="B969EA98FF9D815BFC70BAC44D81F808" bold="true" box="[925,942,1928,1952]" pageId="2" pageNumber="146">B</emphasis>
|
||
, head in dorsal view;
|
||
<emphasis id="B969EA98FF9D815BFB76BAC44A82F808" bold="true" box="[1179,1197,1928,1952]" pageId="2" pageNumber="146">C</emphasis>
|
||
, palpus in dorsal view (MEV);
|
||
<emphasis id="B969EA98FF9D815BFF03BAE04F2FF86C" bold="true" box="[238,256,1964,1988]" pageId="2" pageNumber="146">D</emphasis>
|
||
, thorax in lateral view (MEV);
|
||
<emphasis id="B969EA98FF9D815BFDAFBAE04C7CF86C" bold="true" box="[578,595,1964,1988]" pageId="2" pageNumber="146">E</emphasis>
|
||
, wing;
|
||
<emphasis id="B969EA98FF9D815BFD72BAE04C81F86C" bold="true" box="[671,686,1964,1988]" pageId="2" pageNumber="146">F</emphasis>
|
||
, hind femur;
|
||
<emphasis id="B969EA98FF9D815BFCD5BAE04D63F86C" bold="true" box="[824,844,1964,1988]" pageId="2" pageNumber="146">G</emphasis>
|
||
, foretibia;
|
||
<emphasis id="B969EA98FF9D815BFC51BAE04DFFF86C" bold="true" box="[956,976,1964,1988]" pageId="2" pageNumber="146">H</emphasis>
|
||
, midtibia;
|
||
<emphasis id="B969EA98FF9D815BFBD3BAE04A67F86C" bold="true" box="[1086,1096,1964,1988]" pageId="2" pageNumber="146">I</emphasis>
|
||
, hind tibia.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA2368AFF9C815AFF2ABDDB4FDEFEB5" blockId="3.[151,1437,151,285]" pageId="3" pageNumber="147">
|
||
Concerning phylogenetic relationships,
|
||
<taxonomicName id="4C1D4D09FF9C815AFD64BDDB4D38FF19" authorityName="Ament" authorityYear="2024" box="[649,791,151,177]" class="Insecta" family="Phoridae" genus="Hypocerides" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="147" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA98FF9C815AFD64BDDB4D38FF19" box="[649,791,151,177]" italics="true" pageId="3" pageNumber="147">Hypocerides</emphasis>
|
||
</taxonomicName>
|
||
is a peculiar genus comprising eight species with almost worldwide distribution but almost no considerable morphological divergence among them (
|
||
<bibRefCitation id="EF8C4B7BFF9C815AFB04BDF74BBEFF7D" author="Ament, D. C." box="[1257,1425,187,213]" pageId="3" pageNumber="147" pagination="113 - 126" refId="ref4066" refString="Ament, D. C. (2021 b) Taxonomy and the unusual morphological similarity among Hypocerides Schmitz species (Diptera: Phoridae). Zootaxa, 4980 (1), 113 - 126. https: // doi. org / 10.11646 / zootaxa. 4980.1.7" type="journal article" year="2021">Ament 2021b</bibRefCitation>
|
||
). Therefore, the monophyly of
|
||
<taxonomicName id="4C1D4D09FF9C815AFE09BD934C5DFF51" authorityName="Ament" authorityYear="2024" box="[484,626,223,249]" class="Insecta" family="Phoridae" genus="Hypocerides" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="147" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA98FF9C815AFE09BD934C5DFF51" box="[484,626,223,249]" italics="true" pageId="3" pageNumber="147">Hypocerides</emphasis>
|
||
</taxonomicName>
|
||
seems to be well supported and the proposition of
|
||
<taxonomicName id="4C1D4D09FF9C815AFB5FBD934B60FF51" authorityName="Ament" authorityYear="2024" box="[1202,1359,223,249]" class="Insecta" family="Phoridae" genus="Diadelophora" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="147" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA98FF9C815AFB5FBD934B60FF51" box="[1202,1359,223,249]" italics="true" pageId="3" pageNumber="147">Diadelophora</emphasis>
|
||
</taxonomicName>
|
||
would unlikely render it paraphyletic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |