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<document id="190A9E09BBD230645326CD018C26A083" ID-DOI="10.11646/zootaxa.3926.1.5" ID-GBIF-Dataset="71045880-a3e6-4cbd-930c-fd33408f883f" ID-ISSN="1175-5326" ID-Zenodo-Dep="239465" ID-ZooBank="78FAB512-1F6D-46A4-9D8F-A6B1C84E3C23" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461006333168" checkinUser="plazi" docAuthor="Santos, Jandir C., Castilho, Raphael C., Silva, Edmilson S. &amp; De Moraes, Gilberto J." docDate="2015" docId="03B47964FFC1FF9BFF5BF8C9AA867E82" docLanguage="en" docName="zt03926p121.pdf" docOrigin="Zootaxa 3926 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rykellus" docType="key" docVersion="8" lastPageNumber="120" masterDocId="FF8D011CFFC9FF92FFCCFFA7AF1A7A11" masterDocTitle="Two new species of Rykellus (Acari: Mesostigmata: Ologamasidae) from Brazil and a key to the world species of the genus" masterLastPageNumber="121" masterPageNumber="111" pageNumber="119" updateTime="1698565632399" updateUser="plazi">
<mods:mods id="F898E53C3872D27D1DB677016FDA68DC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5712CC99134DA197B7B1FABB34ECF54C">Two new species of Rykellus (Acari: Mesostigmata: Ologamasidae) from Brazil and a key to the world species of the genus</mods:title>
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<mods:name id="D2C473D22B141895B8FCB7177FB8CCAC" type="personal">
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<mods:namePart id="3BBA68C3665F5055E8C673D8B1676CE4">Santos, Jandir C.</mods:namePart>
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<mods:namePart id="97A3567C324DBF083CCE71F06ADCC644">Castilho, Raphael C.</mods:namePart>
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<mods:namePart id="46D07643C3AEC20F1974BE4EA8C670B6">Silva, Edmilson S.</mods:namePart>
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<mods:namePart id="CE73F0D72DE9BCAFF3E42DBA886FED6C">De Moraes, Gilberto J.</mods:namePart>
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<mods:title id="DD1B8514806B66C16993AA790BCBFE5A">Zootaxa</mods:title>
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<mods:date id="EF616723B9641FCAA3CB3954AD3A7F12">2015</mods:date>
<mods:detail id="74EAABC939452B86C95E39D3DEE1CAAF" type="volume">
<mods:number id="27D96EDC20E15368860867519D4EA605">3926</mods:number>
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<mods:classification id="B15FB46B9DE13A0B7C5E57823EAA19EB">journal article</mods:classification>
<mods:identifier id="8825704676E6B01645E439390AFD46EF" type="DOI">10.11646/zootaxa.3926.1.5</mods:identifier>
<mods:identifier id="20713BE2B209F89BE58427F2E94ABC62" type="GBIF-Dataset">71045880-a3e6-4cbd-930c-fd33408f883f</mods:identifier>
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<treatment id="03B47964FFC1FF9BFF5BF8C9AA867E82" ID-DOI="http://doi.org/10.5281/zenodo.6111917" ID-GBIF-Taxon="119598850" ID-Zenodo-Dep="6111917" LSID="urn:lsid:plazi:treatment:03B47964FFC1FF9BFF5BF8C9AA867E82" httpUri="http://treatment.plazi.org/id/03B47964FFC1FF9BFF5BF8C9AA867E82" lastPageId="9" lastPageNumber="120" pageId="8" pageNumber="119">
<subSubSection id="C3079BF9FFC1FF9BFF5BF8C9AC827BBC" lastPageId="9" lastPageNumber="120" pageId="8" pageNumber="119" type="nomenclature">
<paragraph id="8BA2C872FFC1FF9AFF5BF8C9AC537D99" blockId="8.[151,841,1902,1928]" box="[151,841,1902,1928]" pageId="8" pageNumber="119">
<heading id="D0EA7F1EFFC1FF9AFF5BF8C9AC537D99" bold="true" box="[151,841,1902,1928]" fontSize="11" level="1" pageId="8" pageNumber="119" reason="1">
<emphasis id="B9691460FFC1FF9AFF5BF8C9AC537D99" bold="true" box="[151,841,1902,1928]" pageId="8" pageNumber="119">
Key to world species of
<taxonomicName id="4C1DB3F1FFC1FF9AFE72F8C9AD3C7D99" ID-CoL="63RK8" box="[446,550,1902,1928]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B9691460FFC1FF9AFE72F8C9AD3C7D99" bold="true" box="[446,550,1902,1928]" italics="true" pageId="8" pageNumber="119">Rykellus</emphasis>
</taxonomicName>
based on adult females
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA2C872FFC1FF9AFF5BF809AB827DFA" blockId="8.[151,1436,1966,2027]" pageId="8" pageNumber="119">
<taxonomicName id="4C1DB3F1FFC1FF9AFF5BF809AFEF7DD7" box="[151,245,1966,1990]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B9691460FFC1FF9AFF5BF809AFEF7DD7" box="[151,245,1966,1990]" italics="true" pageId="8" pageNumber="119">Rykellus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C1DB3F1FFC1FF9AFE17F809AC287DD6" authority="Hischmann." authorityName="Hischmann." box="[475,818,1966,1991]" class="Arachnida" family="Ologamasidae" genus="Hydrogamasellus" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B9691460FFC1FF9AFE17F809ADBA7DD7" box="[475,672,1966,1990]" italics="true" pageId="8" pageNumber="119">Hydrogamasellus</emphasis>
Hischmann.
</taxonomicName>
Species of the latter genus also have podonotal and opisthonotal shields fused, but the line of fusion of
<taxonomicName id="4C1DB3F1FFC1FF9AFD1AF874AC807DFA" box="[726,922,2003,2027]" class="Arachnida" family="Ologamasidae" genus="Hydrogamasellus" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B9691460FFC1FF9AFD1AF874AC807DFA" box="[726,922,2003,2027]" italics="true" pageId="8" pageNumber="119">Hydrogamasellus</emphasis>
</taxonomicName>
species is not distinct.
</paragraph>
<paragraph id="8BA2C872FFC0FF9BFF0BFF3FADC77B75" blockId="9.[151,1436,151,429]" pageId="9" pageNumber="120">
Four species were mentioned by
<bibRefCitation id="EF8CB583FFC0FF9BFD8EFF3FAC917AA1" author="Karg" box="[578,907,151,176]" pageId="9" pageNumber="120" refString="Karg, W. &amp; Schorlemmer, A. (2013) Origin of five unique mite-genera in geological periods compared to other group of Gamasina (Acarina, Parasitiformes) and description of two new species of Rykellus Lee and Oloopticus Karg. Zoosystematics and Evolution, 89, 193 - 207. http: // dx. doi. org / 10.1002 / zoos. 201300006" type="journal article" year="2013">Karg &amp; Schorlemmer (2013)</bibRefCitation>
in
<taxonomicName id="4C1DB3F1FFC0FF9BFC7FFF3FAB0B7AA1" box="[947,1041,152,176]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B9691460FFC0FF9BFC7FFF3FAB0B7AA1" box="[947,1041,152,176]" italics="true" pageId="9" pageNumber="120">Rykellus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C1DB3F1FFC0FF9BFBEFFF3EABD57AA1" box="[1059,1231,152,176]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="brevipellitus">
<emphasis id="B9691460FFC0FF9BFBEFFF3EAB2E7AA1" box="[1059,1076,153,176]" italics="true" pageId="9" pageNumber="120">R</emphasis>
.
<emphasis id="B9691460FFC0FF9BFB88FF3FABD57AA1" box="[1092,1231,152,176]" italics="true" pageId="9" pageNumber="120">brevipellitus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1DB3F1FFC0FF9BFB13FF3EAA6E7AA1" box="[1247,1396,152,176]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="darglensis">
<emphasis id="B9691460FFC0FF9BFB13FF3EABEA7AA1" box="[1247,1264,153,176]" italics="true" pageId="9" pageNumber="120">R</emphasis>
.
<emphasis id="B9691460FFC0FF9BFACCFF3FAA6E7AA1" box="[1280,1396,152,176]" italics="true" pageId="9" pageNumber="120">darglensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1DB3F1FFC0FF9BFA48FF3EAE2D7AC5" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="nkandhlaensis">
<emphasis id="B9691460FFC0FF9BFA48FF3EAA8F7AA1" box="[1412,1429,153,176]" italics="true" pageId="9" pageNumber="120">R</emphasis>
.
<emphasis id="B9691460FFC0FF9BFF5BFF1BAE2D7AC5" box="[151,311,188,212]" italics="true" pageId="9" pageNumber="120">nkandhlaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1DB3F1FFC0FF9BFEBAFF1AAEC77AC5" box="[374,477,188,212]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="B9691460FFC0FF9BFEBAFF1AAE9D7AC5" box="[374,391,189,212]" italics="true" pageId="9" pageNumber="120">R</emphasis>
.
<emphasis id="B9691460FFC0FF9BFE56FF1BAEC77AC5" box="[410,477,188,212]" italics="true" pageId="9" pageNumber="120">ovalis</emphasis>
</taxonomicName>
). Two species previously placed in
<taxonomicName id="4C1DB3F1FFC0FF9BFC46FF1BAB557AC5" box="[906,1103,188,212]" class="Arachnida" family="Ologamasidae" genus="Hydrogamasellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B9691460FFC0FF9BFC46FF1BAB557AC5" box="[906,1103,188,212]" italics="true" pageId="9" pageNumber="120">Hydrogamasellus</emphasis>
</taxonomicName>
were recently transferred to
<taxonomicName id="4C1DB3F1FFC0FF9BFF5BFF47AFEF7AE9" box="[151,245,224,248]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B9691460FFC0FF9BFF5BFF47AFEF7AE9" box="[151,245,224,248]" italics="true" pageId="9" pageNumber="120">Rykellus</emphasis>
</taxonomicName>
because they have the typical characteristics of species of this genus, including the distinct, complete and V-shaped line of fusion between podonotal and opisthonotal shields (
<bibRefCitation id="EF8CB583FFC0FF9BFC3CFEA3ABF87B0D" author="Santos" box="[1008,1250,260,284]" pageId="9" pageNumber="120" refString="Santos, J. C., Castilho, R. C., Silva, E. S. &amp; Moraes, G. J. de (2013) A new species of Hydrogamasellus (Acari: Mesostigmata: Ologamasidae) from Brazil, with a key to the world species of the genus. Zootaxa, 3718 (1), 81 - 88. http: // dx. doi. org / 10.11646 / zootaxa. 3718.1.7" type="journal article" year="2013">
Santos
<emphasis id="B9691460FFC0FF9BFB80FEA2AB9D7B0D" box="[1100,1159,260,284]" italics="true" pageId="9" pageNumber="120">et al</emphasis>
., 2013
</bibRefCitation>
). These are
<taxonomicName id="4C1DB3F1FFC0FF9BFA48FEA2AE317B51" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="ubatubaensis">
<emphasis id="B9691460FFC0FF9BFA48FEA2AA8F7B0D" box="[1412,1429,261,284]" italics="true" pageId="9" pageNumber="120">R</emphasis>
.
<emphasis id="B9691460FFC0FF9BFF5BFE8FAE317B51" box="[151,299,296,320]" italics="true" pageId="9" pageNumber="120">ubatubaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1DB3F1FFC0FF9BFEAFFE8EAEE37B51" box="[355,505,296,320]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="longopilus">
<emphasis id="B9691460FFC0FF9BFEAFFE8EAE6E7B51" box="[355,372,297,320]" italics="true" pageId="9" pageNumber="120">R</emphasis>
.
<emphasis id="B9691460FFC0FF9BFE4FFE8FAEE37B51" box="[387,505,296,320]" italics="true" pageId="9" pageNumber="120">longopilus</emphasis>
</taxonomicName>
. The former had been placed in
<taxonomicName id="4C1DB3F1FFC0FF9BFCA0FE8FAB2B7B51" box="[876,1073,296,320]" class="Arachnida" family="Ologamasidae" genus="Hydrogamasellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B9691460FFC0FF9BFCA0FE8FAB2B7B51" box="[876,1073,296,320]" italics="true" pageId="9" pageNumber="120">Hydrogamasellus</emphasis>
</taxonomicName>
for convenience by
<bibRefCitation id="EF8CB583FFC0FF9BFAD4FE8FAA8D7B51" author="Lee" box="[1304,1431,296,320]" pageId="9" pageNumber="120" refString="Lee, D. C. (1970) Rhodacaridae (Acari: Mesostigmata); classification, external morphology and distribution of genera. Records of the South Australian Museum, 16, 1 - 219." type="journal article" year="1970">Lee (1970)</bibRefCitation>
, but the author considered it a species
<emphasis id="B9691460FFC0FF9BFDF0FEEAADCC7B75" box="[572,726,332,356]" italics="true" pageId="9" pageNumber="120">incertae sedis</emphasis>
.
</paragraph>
<paragraph id="8BA2C872FFC0FF9BFF0BFEC8AC827BBC" blockId="9.[151,1436,151,429]" pageId="9" pageNumber="120">Sufficient information is presently available in the literature to allow the recognition of all species attributed to this genus. These can be separated with the use of the following key.</paragraph>
</subSubSection>
<subSubSection id="C3079BF9FFC0FF9BFF5BFE7CAA867E82" pageId="9" pageNumber="120" type="key">
<keyStep id="30E9D0D7FFC0FF9BFF5BFE7CAA81783A" pageId="9" pageNumber="120">
<paragraph id="8BA2C872FFC0FF9BFF5BFE7CAA867BE0" blockId="9.[151,1436,475,1172]" box="[151,1436,475,497]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFE7CAA867BE0" box="[151,1436,475,497]" pageId="9" pageNumber="120">1. Sternal shield fused with section of endopodal shield near coxae IV; exopodal shields totally fused with peritrematic shield. 2</keyLead>
</paragraph>
<paragraph id="8BA2C872FFC0FF9BFF5BFE5EAA81783A" blockId="9.[151,1436,475,1172]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFE5EAA81783A" pageId="9" pageNumber="120">- Sternal shield not fused with section of endopodal shield near coxae IV; only posterior end of exopodal shield fused with peritrematic shield........................................................................................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E9D0D7FFC0FF9BFF5BFD94AA8178B0" pageId="9" pageNumber="120">
<paragraph id="8BA2C872FFC0FF9BFF5BFD94AA817877" blockId="9.[151,1436,475,1172]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFD94AA817877" pageId="9" pageNumber="120">
2. Seta
<emphasis id="B9691460FFC0FF9BFEDCFD93AE077858" box="[272,285,564,585]" italics="true" pageId="9" pageNumber="120">Z</emphasis>
5 about 1.1 times as long as
<emphasis id="B9691460FFC0FF9BFDE1FD93AD227858" box="[557,568,564,585]" italics="true" pageId="9" pageNumber="120">J</emphasis>
5; ventrianal shield with the two most anterior setae in longitudinal line.............................................................................
<taxonomicName id="4C1DB3F1FFC0FF9BFC1BFDF7AA0D7877" ID-CoL="78Z62" authority="Ryke, 1962" authorityName="Ryke" authorityYear="1962" box="[983,1303,592,614]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="darglensis">
<emphasis id="B9691460FFC0FF9BFC1BFDF7AB8B7877" box="[983,1169,592,614]" italics="true" pageId="9" pageNumber="120">Rykellus darglensis</emphasis>
(
<bibRefCitation id="EF8CB583FFC0FF9BFB52FDF7AA157877" author="Ryke" box="[1182,1295,592,614]" pageId="9" pageNumber="120" refString="Ryke, P. A. J. (1962) A revision of the subgenera Cyrtolaelaps Berlese and Gamasellus Berlese of the genus Cyrtolaelaps Berlese and descriptions of new species (Acarina: Rhodacaridae). Memoirs of the Entomological Society of Southern Africa, 7, 1 - 59." type="journal article" year="1962">Ryke, 1962</bibRefCitation>
)
</taxonomicName>
,
<collectingCountry id="F30A88E2FFC0FF9BFAEDFDF7AA817877" box="[1313,1435,592,614]" name="South Africa" pageId="9" pageNumber="120">South Africa</collectingCountry>
</keyLead>
</paragraph>
<paragraph id="8BA2C872FFC0FF9BFF5BFDC9AA8178B0" blockId="9.[151,1436,475,1172]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFDC9AA8178B0" pageId="9" pageNumber="120">
- Seta
<emphasis id="B9691460FFC0FF9BFEDCFDC9AE077892" box="[272,285,622,643]" italics="true" pageId="9" pageNumber="120">Z</emphasis>
5 about 8 times as long as
<emphasis id="B9691460FFC0FF9BFDD7FDC9AD3C7892" box="[539,550,622,643]" italics="true" pageId="9" pageNumber="120">J</emphasis>
5; ventrianal shield with the two most anterior setae in transverse line.............................................................................
<taxonomicName id="4C1DB3F1FFC0FF9BFC7CFD2CAA0D78B0" ID-CoL="4TTXB" authority="Ryke, 1962" authorityName="Ryke" authorityYear="1962" box="[944,1303,651,673]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="nkandhlaensis">
<emphasis id="B9691460FFC0FF9BFC7CFD2CAB8B78B0" box="[944,1169,651,673]" italics="true" pageId="9" pageNumber="120">Rykellus nkandhlaensis</emphasis>
(
<bibRefCitation id="EF8CB583FFC0FF9BFB52FD2CAA1578B0" author="Ryke" box="[1182,1295,651,673]" pageId="9" pageNumber="120" refString="Ryke, P. A. J. (1962) A revision of the subgenera Cyrtolaelaps Berlese and Gamasellus Berlese of the genus Cyrtolaelaps Berlese and descriptions of new species (Acarina: Rhodacaridae). Memoirs of the Entomological Society of Southern Africa, 7, 1 - 59." type="journal article" year="1962">Ryke, 1962</bibRefCitation>
)
</taxonomicName>
,
<collectingCountry id="F30A88E2FFC0FF9BFAEDFD2CAA8178B0" box="[1313,1435,651,673]" name="South Africa" pageId="9" pageNumber="120">South Africa</collectingCountry>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E9D0D7FFC0FF9BFF5BFD0FAA8178CA" pageId="9" pageNumber="120">
<paragraph id="8BA2C872FFC0FF9BFF5BFD0FAA8F78AF" blockId="9.[151,1436,475,1172]" box="[151,1429,680,702]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFD0FAA8F78AF" box="[151,1429,680,702]" pageId="9" pageNumber="120">3. Podonotal region of dorsal shield with 19–20 pairs of setae................................................... 4</keyLead>
</paragraph>
<paragraph id="8BA2C872FFC0FF9BFF5BFD61AA8178CA" blockId="9.[151,1436,475,1172]" box="[151,1435,709,731]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFD61AA8178CA" box="[151,1435,709,731]" pageId="9" pageNumber="120">- Podonotal region of dorsal shield with 22 pairs of setae...................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E9D0D7FFC0FF9BFF5BFD44AA817940" pageId="9" pageNumber="120">
<paragraph id="8BA2C872FFC0FF9BFF5BFD44AA867907" blockId="9.[151,1436,475,1172]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFD44AA867907" pageId="9" pageNumber="120">
4. Podonotal region of dorsal shield with 19 pairs of setae; opisthogaster with seven pairs of setae (
<emphasis id="B9691460FFC0FF9BFB5FFD43ABB278E8" box="[1171,1192,740,761]" italics="true" pageId="9" pageNumber="120">Jv</emphasis>
1–
<emphasis id="B9691460FFC0FF9BFB0CFD43ABCF78E8" box="[1216,1237,740,761]" italics="true" pageId="9" pageNumber="120">Jv</emphasis>
3,
<emphasis id="B9691460FFC0FF9BFB23FD43AA1E78E8" box="[1263,1284,740,761]" italics="true" pageId="9" pageNumber="120">Jv</emphasis>
5 and
<emphasis id="B9691460FFC0FF9BFA8DFD43AA4378E8" box="[1345,1369,740,761]" italics="true" pageId="9" pageNumber="120">Zv</emphasis>
1–
<emphasis id="B9691460FFC0FF9BFABDFD43AA9378E8" box="[1393,1417,740,761]" italics="true" pageId="9" pageNumber="120">Zv</emphasis>
3;
<emphasis id="B9691460FFC0FF9BFF2DFCA6AFEC7907" box="[225,246,769,790]" italics="true" pageId="9" pageNumber="120">Jv</emphasis>
4 absent).............................................
<taxonomicName id="4C1DB3F1FFC0FF9BFCBBFCA7AA4D7907" ID-CoL="4TTX9" authority="Karg &amp; Schorlemmer, 2009" authorityName="Karg &amp; Schorlemmer" authorityYear="2009" box="[887,1367,768,790]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="brevipellitus">
<emphasis id="B9691460FFC0FF9BFCBBFCA7AB5F7907" box="[887,1093,768,790]" italics="true" pageId="9" pageNumber="120">Rykellus brevipellitus</emphasis>
<bibRefCitation id="EF8CB583FFC0FF9BFB87FCA6AA4D7907" author="Karg" box="[1099,1367,768,790]" pageId="9" pageNumber="120" refString="Karg, W. &amp; Schorlemmer, A. (2009) New insights into predatory mites (Acarina, Gamasina) from tropical rain forests with special reference to distribuition and taxonomy. Zoosystematics and Evolution, 85, 57 - 91. http: // dx. doi. org / 10.1002 / zoos. 200800016" type="journal article" year="2009">Karg &amp; Schorlemmer, 2009</bibRefCitation>
</taxonomicName>
,
<collectingCountry id="F30A88E2FFC0FF9BFAADFCA7AA867907" box="[1377,1436,768,790]" name="Brazil" pageId="9" pageNumber="120">Brazil</collectingCountry>
</keyLead>
</paragraph>
<paragraph id="8BA2C872FFC0FF9BFF5BFCB9AA817940" blockId="9.[151,1436,475,1172]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFCB9AA817940" pageId="9" pageNumber="120">
- Podonotal region of dorsal shield with 20 pairs of setae; opisthogaster with eight pairs of setae (
<emphasis id="B9691460FFC0FF9BFBBBFCB9AB967922" box="[1143,1164,798,819]" italics="true" pageId="9" pageNumber="120">Jv</emphasis>
1–
<emphasis id="B9691460FFC0FF9BFB68FCB9ABA37922" box="[1188,1209,798,819]" italics="true" pageId="9" pageNumber="120">Jv</emphasis>
5 and
<emphasis id="B9691460FFC0FF9BFB38FCB9AA167922" box="[1268,1292,798,819]" italics="true" pageId="9" pageNumber="120">Zv</emphasis>
1–
<emphasis id="B9691460FFC0FF9BFAE8FCB9AA267922" box="[1316,1340,798,819]" italics="true" pageId="9" pageNumber="120">Zv</emphasis>
3)..............................................................
<taxonomicName id="4C1DB3F1FFC0FF9BFC5AFC9CAA237940" ID-CoL="9f69b8bd-b7e7-4eef-a374-ebe66022d58d" authority="Karg &amp; Schorlemmer, 2013" authorityName="Karg &amp; Schorlemmer" authorityYear="2013" box="[918,1337,827,849]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="B9691460FFC0FF9BFC5AFC9CAB3C7940" box="[918,1062,827,849]" italics="true" pageId="9" pageNumber="120">Rykellus ovalis</emphasis>
<bibRefCitation id="EF8CB583FFC0FF9BFBE0FC9BAA237940" author="Karg" box="[1068,1337,827,849]" pageId="9" pageNumber="120" refString="Karg, W. &amp; Schorlemmer, A. (2013) Origin of five unique mite-genera in geological periods compared to other group of Gamasina (Acarina, Parasitiformes) and description of two new species of Rykellus Lee and Oloopticus Karg. Zoosystematics and Evolution, 89, 193 - 207. http: // dx. doi. org / 10.1002 / zoos. 201300006" type="journal article" year="2013">Karg &amp; Schorlemmer, 2013</bibRefCitation>
</taxonomicName>
,
<collectingCountry id="F30A88E2FFC0FF9BFA8EFC9BAA817940" box="[1346,1435,828,849]" name="Paraguay" pageId="9" pageNumber="120">Paraguay</collectingCountry>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E9D0D7FFC0FF9BFF5BFCFFAA86799A" pageId="9" pageNumber="120">
<paragraph id="8BA2C872FFC0FF9BFF5BFCFFAA86797F" blockId="9.[151,1436,475,1172]" box="[151,1436,856,878]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFCFFAA86797F" box="[151,1436,856,878]" pageId="9" pageNumber="120">
5. Setae
<emphasis id="B9691460FFC0FF9BFED7FCFEAE3C797F" box="[283,294,857,878]" italics="true" pageId="9" pageNumber="120">J</emphasis>
2,
<emphasis id="B9691460FFC0FF9BFEF1FCFEAE52797F" box="[317,328,857,878]" italics="true" pageId="9" pageNumber="120">J</emphasis>
3 and
<emphasis id="B9691460FFC0FF9BFE4FFCFEAE94797F" box="[387,398,857,878]" italics="true" pageId="9" pageNumber="120">J</emphasis>
4 at least as long as distance between their bases and bases of respective subsequent setae of
<emphasis id="B9691460FFC0FF9BFACBFCFEAA08797F" box="[1287,1298,857,878]" italics="true" pageId="9" pageNumber="120">J</emphasis>
series..... 6
</keyLead>
</paragraph>
<paragraph id="8BA2C872FFC0FF9BFF5BFCD1AA86799A" blockId="9.[151,1436,475,1172]" box="[151,1436,885,907]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFCD1AA86799A" box="[151,1436,885,907]" pageId="9" pageNumber="120">
- Setae
<emphasis id="B9691460FFC0FF9BFED7FCD1AE3C799A" box="[283,294,886,907]" italics="true" pageId="9" pageNumber="120">J</emphasis>
2,
<emphasis id="B9691460FFC0FF9BFEF1FCD1AE52799A" box="[317,328,886,907]" italics="true" pageId="9" pageNumber="120">J</emphasis>
3 and
<emphasis id="B9691460FFC0FF9BFE4FFCD1AE94799A" box="[387,398,886,907]" italics="true" pageId="9" pageNumber="120">J</emphasis>
4 at most half as long as distance between their bases and bases of respective subsequent setae of
<emphasis id="B9691460FFC0FF9BFAF9FCD1AA5A799A" box="[1333,1344,886,907]" italics="true" pageId="9" pageNumber="120">J</emphasis>
series. 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E9D0D7FFC0FF9BFF5BFC34AA867E0F" pageId="9" pageNumber="120">
<paragraph id="8BA2C872FFC0FF9BFF5BFC34AA8679F2" blockId="9.[151,1436,475,1172]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFC34AA8679F2" pageId="9" pageNumber="120">
6. Opisthonotal region of dorsal shield with 21 pairs of setae (
<emphasis id="B9691460FFC0FF9BFCD2FC33AC3379B8" box="[798,809,916,937]" italics="true" pageId="9" pageNumber="120">J</emphasis>
1–
<emphasis id="B9691460FFC0FF9BFC8DFC33AC5679B8" box="[833,844,916,937]" italics="true" pageId="9" pageNumber="120">J</emphasis>
5,
<emphasis id="B9691460FFC0FF9BFCA8FC33AC6B79B8" box="[868,881,916,937]" italics="true" pageId="9" pageNumber="120">Z</emphasis>
1–
<emphasis id="B9691460FFC0FF9BFC45FC33AC8C79B8" box="[905,918,916,937]" italics="true" pageId="9" pageNumber="120">Z</emphasis>
5,
<emphasis id="B9691460FFC0FF9BFC63FC34ACA179B8" box="[943,955,915,937]" italics="true" pageId="9" pageNumber="120">S</emphasis>
1–
<emphasis id="B9691460FFC0FF9BFC1FFC34ACC579B8" box="[979,991,915,937]" italics="true" pageId="9" pageNumber="120">S</emphasis>
5,
<emphasis id="B9691460FFC0FF9BFC3BFC33AB1C79B8" box="[1015,1030,916,937]" italics="true" pageId="9" pageNumber="120">R</emphasis>
1–
<emphasis id="B9691460FFC0FF9BFBD2FC33AB3779B8" box="[1054,1069,916,937]" italics="true" pageId="9" pageNumber="120">R</emphasis>
5 and
<emphasis id="B9691460FFC0FF9BFBA4FC33AB9279B8" box="[1128,1160,916,937]" italics="true" pageId="9" pageNumber="120">UR</emphasis>
5); ventrianal shield with six pairs of setae (
<emphasis id="B9691460FFC0FF9BFEA1FC16AE9F79D7" box="[365,389,945,966]" italics="true" pageId="9" pageNumber="120">Zv</emphasis>
3 on unsclerotised integument along margins of the shield).....................................................................................................
<taxonomicName id="4C1DB3F1FFC0FF9BFC78FC6AAA4D79F2" ID-CoL="2bbf07f7-c534-48b9-81df-51a0b1fd0fe3" authority="Hirschmann, 1966" authorityName="Hirschmann" authorityYear="1966" box="[948,1367,973,995]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="ubatubaensis">
<emphasis id="B9691460FFC0FF9BFC78FC6AAB9479F2" box="[948,1166,973,995]" italics="true" pageId="9" pageNumber="120">Rykellus ubatubaensis</emphasis>
(
<bibRefCitation id="EF8CB583FFC0FF9BFB57FC6AAA4A79F2" author="Hirschmann" box="[1179,1360,973,995]" pageId="9" pageNumber="120" refString="Hirschmann, W. (1966) Gangsystematik der Parasitiformes. Teil 11. Die Gattung Hydrogamasus Berlese 1892 nov. comb. und die neuen Untergattungen Hydrogamasus (Austrohydrogamasus nov. subgen.) und Gamasellus (Hydrogamasellus nov." type="journal article" year="1966">Hirschmann, 1966</bibRefCitation>
)
</taxonomicName>
,
<collectingCountry id="F30A88E2FFC0FF9BFAADFC6AAA8679F2" box="[1377,1436,973,995]" name="Brazil" pageId="9" pageNumber="120">Brazil</collectingCountry>
</keyLead>
</paragraph>
<paragraph id="8BA2C872FFC0FF9BFF5BFC4BAA867E0F" blockId="9.[151,1436,475,1172]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFC4BAA867E0F" pageId="9" pageNumber="120">
- Opisthonotal region of dorsal shield with 18 pairs of setae (
<emphasis id="B9691460FFC0FF9BFCE3FC4BAC207E10" box="[815,826,1004,1025]" italics="true" pageId="9" pageNumber="120">J</emphasis>
1–
<emphasis id="B9691460FFC0FF9BFC9EFC4BAC477E10" box="[850,861,1004,1025]" italics="true" pageId="9" pageNumber="120">J</emphasis>
5,
<emphasis id="B9691460FFC0FF9BFCBAFC4BAC997E10" box="[886,899,1004,1025]" italics="true" pageId="9" pageNumber="120">Z</emphasis>
1–
<emphasis id="B9691460FFC0FF9BFC50FC4BACB37E10" box="[924,937,1004,1025]" italics="true" pageId="9" pageNumber="120">Z</emphasis>
5,
<emphasis id="B9691460FFC0FF9BFC0FFC4CACD57E10" box="[963,975,1003,1025]" italics="true" pageId="9" pageNumber="120">S</emphasis>
1–
<emphasis id="B9691460FFC0FF9BFC2BFC4CACE97E10" box="[999,1011,1003,1025]" italics="true" pageId="9" pageNumber="120">S</emphasis>
5 and
<emphasis id="B9691460FFC0FF9BFBFEFC4BAB5A7E10" box="[1074,1088,1004,1025]" italics="true" pageId="9" pageNumber="120">R</emphasis>
2–
<emphasis id="B9691460FFC0FF9BFB94FC4BAB7D7E10" box="[1112,1127,1004,1025]" italics="true" pageId="9" pageNumber="120">R</emphasis>
4); ventrianal shield with seven pairs of setae (
<emphasis id="B9691460FFC0FF9BFEA1FBAEAE9F7E0F" box="[365,389,1033,1054]" italics="true" pageId="9" pageNumber="120">Zv</emphasis>
3 on the shield)..........................................
<taxonomicName id="4C1DB3F1FFC0FF9BFBD2FBAFAA447E0F" ID-CoL="c0d33526-824b-4fa2-b5b4-06c6b9c1db8d" authority="Karg, 1976" authorityName="Karg" authorityYear="1976" box="[1054,1374,1032,1054]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="longopilus">
<emphasis id="B9691460FFC0FF9BFBD2FBAFABC07E0F" box="[1054,1242,1032,1054]" italics="true" pageId="9" pageNumber="120">Rykellus longopilus</emphasis>
(
<bibRefCitation id="EF8CB583FFC0FF9BFB2BFBAFAA4D7E0F" author="Karg" box="[1255,1367,1032,1054]" pageId="9" pageNumber="120" refString="Karg, W. (1976) Revision der Milbengattung Hydrogamasellus Hirschmann, 1966 (Acarina Parasitiformes). Deutsche Entomologische Zeitschrift, 23, 37 - 55. http: // dx. doi. org / 10.1002 / mmnd. 19760230104" type="journal article" year="1976">Karg, 1976</bibRefCitation>
)
</taxonomicName>
,
<collectingCountry id="F30A88E2FFC0FF9BFAA4FBAFAA867E0F" box="[1384,1436,1032,1054]" name="Chile" pageId="9" pageNumber="120">Chile</collectingCountry>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E9D0D7FFC0FF9BFF5BFB81AA867E82" pageId="9" pageNumber="120">
<paragraph id="8BA2C872FFC0FF9BFF5BFB81AA867E48" blockId="9.[151,1436,475,1172]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFB81AA867E48" pageId="9" pageNumber="120">
7. Anterolateral extensions of epistome rounded distally; with a platelet between presternal plates; ventrianal shield with anterolateral reticulation..............................................................
<taxonomicName id="4C1DB3F1FFC0FF9BFBB1FBE4AA037E48" ID-CoL="24267de1-6757-4b8c-a3ce-3c7d2a325cd1" box="[1149,1305,1091,1113]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="anibali" status="sp. nov.">
<emphasis id="B9691460FFC0FF9BFBB1FBE4AA037E48" box="[1149,1305,1091,1113]" italics="true" pageId="9" pageNumber="120">Rykellus anibali</emphasis>
</taxonomicName>
<emphasis id="B9691460FFC0FF9BFAD3FBE4AA4F7E48" bold="true" box="[1311,1365,1091,1113]" pageId="9" pageNumber="120">
<taxonomicNameLabel id="A25AA91BFFC0FF9BFAD3FBE4AA4F7E48" box="[1311,1365,1091,1113]" pageId="9" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<collectingCountry id="F30A88E2FFC0FF9BFAADFBE4AA867E48" box="[1377,1436,1091,1113]" name="Brazil" pageId="9" pageNumber="120">Brazil</collectingCountry>
</keyLead>
</paragraph>
<paragraph id="8BA2C872FFC0FF9BFF5BFBC6AA867E82" blockId="9.[151,1436,475,1172]" pageId="9" pageNumber="120">
<keyLead id="30EC6B47FFC0FF9BFF5BFBC6AA867E82" pageId="9" pageNumber="120">
- Anterolateral extensions of epistome sharp-tipped; without platelet between presternal plates; ventrianal shield without anterolateral reticulation.............................................................
<taxonomicName id="4C1DB3F1FFC0FF9BFBBEFBDAAA037E82" ID-CoL="97e05fa7-9759-4a91-8143-1f77db2deb74" box="[1138,1305,1149,1171]" class="Arachnida" family="Ologamasidae" genus="Rykellus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="mineiroi" status="sp. nov.">
<emphasis id="B9691460FFC0FF9BFBBEFBDAAA037E82" box="[1138,1305,1149,1171]" italics="true" pageId="9" pageNumber="120">Rykellus mineiroi</emphasis>
</taxonomicName>
<emphasis id="B9691460FFC0FF9BFAD3FBD9AA4F7E85" bold="true" box="[1311,1365,1150,1172]" pageId="9" pageNumber="120">
<taxonomicNameLabel id="A25AA91BFFC0FF9BFAD3FBD9AA4F7E85" box="[1311,1365,1150,1172]" pageId="9" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<collectingCountry id="F30A88E2FFC0FF9BFAADFBDAAA867E82" box="[1377,1436,1149,1171]" name="Brazil" pageId="9" pageNumber="120">Brazil</collectingCountry>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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