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<document id="7E7ED7BB8A66EE18F1AFB5F5375458C0" ID-CLB-Dataset="8431" ID-DOI="10.11646/zootaxa.4858.2.1" ID-GBIF-Dataset="985a0e09-5ed8-4596-9047-fc5fe27db488" ID-ISSN="1175-5326" ID-Zenodo-Dep="4411669" ID-ZooBank="A6C6FBA5-8026-46D8-9DA6-57A8BBE0655C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609609713693" checkinUser="plazi" docAuthor="Joshi, Shantanu, Subramanian, K. A., Babu, R., Sawant, Dattaprasad &amp; Kunte, Krushnamegh" docDate="2020" docId="03B087FBFF8DD665FF05D53BB261FB47" docLanguage="en" docName="zootaxa.4858.2.1.pdf" docOrigin="Zootaxa 4858 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protosticta myristicaensis Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte 2020, spec. nov." docType="treatment" docUuid="8E1C4562-9142-42BD-8328-0B6FA733A52B" docUuidSource="ZooBank" docVersion="9" lastPageNumber="169" masterDocId="FF89FF83FF87D677FF92D045B15DFF81" masterDocTitle="Three new species of Protosticta Selys, 1885 (Odonata: Zygoptera: Platystictidae) from the Western Ghats, India, with taxonomic notes on P. mortoni Fraser, 1922 and rediscovery of P. rufostigma Kimmins, 1958" masterLastPageNumber="185" masterPageNumber="151" pageNumber="161" updateTime="1701999446184" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="22C159A668964DB7E3EFF73D74DE0536">Three new species of Protosticta Selys, 1885 (Odonata: Zygoptera: Platystictidae) from the Western Ghats, India, with taxonomic notes on P. mortoni Fraser, 1922 and rediscovery of P. rufostigma Kimmins, 1958</mods:title>
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<mods:namePart id="732D977CA8657E121CCAE59B2E769846">Joshi, Shantanu</mods:namePart>
<mods:affiliation id="5B54160054035DF428ADC4F8F7D813B2">National Centre for Biological Sciences, GKVK Campus, Bellary Rd, Bengaluru, 560065, Karnataka, India</mods:affiliation>
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<mods:namePart id="E784012FCFFD14A3D791BFE3815AC447">Subramanian, K. A.</mods:namePart>
<mods:nameIdentifier id="C11C200D84791664167B7D898BC3DA31" type="ORCID">0000-0003-0872-9771</mods:nameIdentifier>
<mods:affiliation id="06E3A2A79EA56BCD4EA279AB03FBABFB">Zoological Survey of India, Southern Regional Centre, Santhome High Road, Chennai 600028, Tamil Nadu, India &amp; subbuka. zsi @ gmail. com; https: // orcid. org / 0000 - 0003 - 0872 - 9771</mods:affiliation>
<mods:nameIdentifier id="05F93C7E2AD2A4EEF5AB2DD5A8AF06FC" type="email">subbuka.zsi@gmail.com</mods:nameIdentifier>
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<mods:namePart id="348842FBBBCF9E6BAEDFF611FA58BE12">Babu, R.</mods:namePart>
<mods:nameIdentifier id="A0D686089F04C04143F6D3D695781C6C" type="ORCID">0000-0001-9147-4540</mods:nameIdentifier>
<mods:affiliation id="9C634AAFDA415282DF4A1F9CD9B05822">Zoological Survey of India, Southern Regional Centre, Santhome High Road, Chennai 600028, Tamil Nadu, India &amp; baburzsi @ gmail. com; https: // orcid. org / 0000 - 0001 - 9147 - 4540</mods:affiliation>
<mods:nameIdentifier id="FAD78BCDE7FC26DECDB95FD567C79612" type="email">baburzsi@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F4CD4982470D15EE0A7613196FE229C3">Sawant, Dattaprasad</mods:namePart>
<mods:affiliation id="F99EDC8B2781DEACF4E7BE755E970ED6">M. D. Community Medicine, Seth G S Medical College and KEM Hospital, Mumbai, 400 012, Maharashtra, India &amp; dattaprasad. 101 @ gmail. com; http: // orcid. org / 0000 - 0003 - 0533 - 9292</mods:affiliation>
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<mods:namePart id="7FD8D19A0B0A0C27EE69264B8AE3446A">Kunte, Krushnamegh</mods:namePart>
<mods:affiliation id="EBEB33CC27F4F4BFF5633DECDD7BE8EB">National Centre for Biological Sciences, GKVK Campus, Bellary Rd, Bengaluru, 560065, Karnataka, India &amp; krushnamegh @ ncbs. res. in; http: // orcid. org / 0000 - 0002 - 3860 - 6118</mods:affiliation>
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<mods:title id="D0E44C2CFDC4DF0663D5EB3CC88A208E">Zootaxa</mods:title>
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<mods:date id="656B39C9E002B51568316E7BE3617980">2020</mods:date>
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<mods:number id="602CFC853EB50A035609C6958AEFDCF8">4858</mods:number>
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<taxonomicName id="4C194D6EFF8DD67DFF05D53BB091FA19" ID-CoL="9X45W" authority="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte, 2020" authorityName="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte" authorityYear="2020" box="[151,460,1406,1432]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="myristicaensis" status="sp. nov.">
<emphasis id="B96DEAFFFF8DD67DFF05D53BB091FA19" bold="true" box="[151,460,1406,1432]" italics="true" pageId="10" pageNumber="161">Protosticta myristicaensis</emphasis>
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<emphasis id="B96DEAFFFF8DD67DFF05D5E7B01BFA3C" bold="true" box="[151,326,1442,1469]" pageId="10" pageNumber="161">Joshi &amp; Kunte</emphasis>
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urn:lsid:zoobank.org:act:
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(
<figureCitation id="13222A68FF8DD67DFF0DD5AFB045F984" box="[159,280,1514,1541]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStart-4="FIGURE 11" captionStartId-0="12.[151,250,1857,1881]" captionStartId-1="13.[151,250,1657,1681]" captionStartId-2="14.[151,250,1797,1821]" captionStartId-3="15.[151,250,1943,1967]" captionStartId-4="16.[151,250,1947,1971]" captionTargetBox-0="[151,1436,181,1833]" captionTargetBox-1="[151,1436,181,1633]" captionTargetBox-2="[151,1435,181,1773]" captionTargetBox-3="[214,1374,181,1920]" captionTargetBox-4="[151,1436,417,1924]" captionTargetId-0="figure-33@12.[151,1436,181,1833]" captionTargetId-1="figure-17@13.[151,1436,181,1633]" captionTargetId-2="figure-33@14.[151,1435,181,1773]" captionTargetId-3="figure-17@15.[214,1374,181,1920]" captionTargetId-4="figure-139@16.[151,1436,417,1924]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionTargetPageId-4="16" captionText-0="FIGURE 7. Protosticta myristicaensis sp. nov. Holotype male [NCBS-BH110]: (a) habitus, lateral view; (b) face; (c) prothorax, dorsal view; (d) pterostigma of right HW &amp; FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." captionText-1="FIGURE 8. Protosticta myristicaensis sp. nov. Paratype males: (ab) NCBS-BM655, (cf) NCBS-BM654: (a) prothorax, dorsal view; (b) caudal appendages, ventral view; (c) face; (d) pterostigma of right FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." captionText-2="FIGURE 9. Genital ligula of Protosticta myristicaensis sp. nov. [based on NCBS-BH110 &amp; NCBS-BM656]: (a) dorsal view; (b) ventral view; (c) left lateral view." captionText-3="FIGURE 10. Protosticta myristicaensis sp. nov. Paratype female [NCBS-BH111]: (a) habitus, lateral view; (b) face; (c) pterostigma of right FW; (d) prothorax, left lateral view; (e) prothorax, dorsal view; (f) caudal appendages, dorsal view; (g) caudal appendages, lateral view." captionText-4="FIGURE 11. Protosticta myristicaensis sp. nov. Paratype females: (a-e) NCBS-BM657, (f) NCBS-BH112: (a) face; (b) prothorax, dorsal view; (c) pterostigma of right FW; (d) caudal appendages, dorsal view; (e-f) caudal appendages, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.4411683" figureDoi-1="http://doi.org/10.5281/zenodo.4411685" figureDoi-2="http://doi.org/10.5281/zenodo.4411687" figureDoi-3="http://doi.org/10.5281/zenodo.4411689" figureDoi-4="http://doi.org/10.5281/zenodo.4411691" httpUri-0="https://zenodo.org/record/4411683/files/figure.png" httpUri-1="https://zenodo.org/record/4411685/files/figure.png" httpUri-2="https://zenodo.org/record/4411687/files/figure.png" httpUri-3="https://zenodo.org/record/4411689/files/figure.png" httpUri-4="https://zenodo.org/record/4411691/files/figure.png" pageId="10" pageNumber="161">Figs. 711</figureCitation>
,
<figureCitation id="13222A68FF8DD67DFEB6D5AFB062F984" box="[292,319,1514,1541]" captionStart="FIGURE 12" captionStartId="17.[151,250,1965,1989]" captionTargetBox="[189,1399,181,1941]" captionTargetId="figure-17@17.[189,1399,181,1941]" captionTargetPageId="17" captionText="FIGURE 12. Field images of: (ab) Protosticta cyanofemora sp. nov. (a) male; (b) female; (cd) P. myristicaensis sp. nov. (c) male; (d) female. Photos: (a, cd) SJ, &amp; (b) KAS." figureDoi="http://doi.org/10.5281/zenodo.4411695" httpUri="https://zenodo.org/record/4411695/files/figure.png" pageId="10" pageNumber="161">12</figureCitation>
cd)
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.
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(
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),
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,
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,
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(
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,
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; Alt:
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),
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<collectingDate id="EFE3E9C5FF8DD67DFF05D614B052F9EA" box="[151,271,1617,1643]" pageId="10" pageNumber="161" value="2019-04-16">16.iv.2019</collectingDate>
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,
<collectorName id="26EC533BFF8DD67DFE89D614B35EF9ED" box="[283,515,1617,1644]" pageId="10" pageNumber="161">Krushnamegh Kunte</collectorName>
leg.
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<materialsCitation id="3B713CB0FF8DD67DFF55D630B196F935" ID-GBIF-Occurrence="3031208306" collectionCode="NCBS-BH" pageId="10" pageNumber="161" specimenCode="NCBS-BH111, NCBS-BH112" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<emphasis id="B96DEAFFFF8DD67DFF55D630B0D2F913" bold="true" box="[199,399,1653,1682]" pageId="10" pageNumber="161">
<typeStatus id="54A2884FFF8DD67DFF55D630B01CF90E" box="[199,321,1653,1679]" pageId="10" pageNumber="161" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="9D1FFD64FF8DD67DFEDDD630B0D2F913" box="[335,399,1653,1682]" pageId="10" pageNumber="161" type="female">2 ♀♀</specimenCount>
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(
<specimenCode id="DBBF9E96FF8DD67DFE0DD630B31FF90E" box="[415,578,1653,1679]" collectionCode="NCBS-BH" pageId="10" pageNumber="161">NCBS-BH111</specimenCode>
&amp;
<specimenCode id="DBBF9E96FF8DD67DFDFBD630B24DF90E" box="[617,784,1653,1679]" collectionCode="NCBS-BH" pageId="10" pageNumber="161">NCBS-BH112</specimenCode>
), location, date of collection, and collector same as
<typeStatus id="54A2884FFF8DD67DFAF3D630B196F935" pageId="10" pageNumber="161" type="holotype">holotype</typeStatus>
</materialsCitation>
.
<materialsCitation id="3B713CB0FF8DD67DFF47D6DCB415F932" ID-GBIF-Occurrence="3031208303" box="[213,1352,1689,1718]" collectingDate="2019-11-23" collectionCode="BM" collectorName="Krushnamegh Kunte" country="India" elevation="560" latitude="14.2743" location="Kathlekan" municipality="Shivamogga" pageId="10" pageNumber="161" specimenCode="NCBS-BM654, BM657" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Karnataka" typeStatus="paratype">
<emphasis id="B96DEAFFFF8DD67DFF47D6DCB011F937" bold="true" box="[213,332,1689,1718]" pageId="10" pageNumber="161">
<specimenCount id="9D1FFD64FF8DD67DFF47D6DCB049F937" box="[213,276,1689,1718]" pageId="10" pageNumber="161" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D1FFD64FF8DD67DFEB0D6DCB011F937" box="[290,332,1689,1718]" pageId="10" pageNumber="161" type="female">1 ♀</specimenCount>
</emphasis>
(
<specimenCode id="DBBF9E96FF8DD67DFEC9D6DCB355F935" box="[347,520,1689,1716]" collectionCode="NCBS-BM" pageId="10" pageNumber="161">NCBS-BM654</specimenCode>
<specimenCode id="DBBF9E96FF8DD67DFD85D6DCB330F935" box="[535,621,1689,1716]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="10" pageNumber="161">BM657</specimenCode>
), same locality as holotype,
<date id="FFA7102DFF8DD67DFC3BD6DCB57EF932" box="[937,1059,1689,1715]" pageId="10" pageNumber="161" value="2019-11-23">
<collectingDate id="EFE3E9C5FF8DD67DFC3BD6DCB57EF932" box="[937,1059,1689,1715]" pageId="10" pageNumber="161" value="2019-11-23">23.xi.2019</collectingDate>
</date>
,
<collectorName id="26EC533BFF8DD67DFBBDD6DCB44AF935" box="[1071,1303,1689,1716]" pageId="10" pageNumber="161">Krushnamegh Kunte</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3036566FF8DD67DFF55D6F8B57CF97A" pageId="10" pageNumber="161" type="etymology">
<paragraph id="8BA636EDFF8DD67DFF55D6F8B57CF97A" blockId="10.[151,1437,1581,2040]" pageId="10" pageNumber="161">
<emphasis id="B96DEAFFFF8DD67DFF55D6F8B013F956" bold="true" box="[199,334,1725,1751]" pageId="10" pageNumber="161">Etymology.</emphasis>
The specific name is given after
<taxonomicName id="4C194D6EFF8DD67DFD4BD6FBB219F956" box="[729,836,1726,1751]" class="Magnoliopsida" family="Myristicaceae" genus="Myristica" kingdom="Plantae" order="Magnoliales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="genus">
<emphasis id="B96DEAFFFF8DD67DFD4BD6FBB219F956" box="[729,836,1726,1751]" italics="true" pageId="10" pageNumber="161">Myristica</emphasis>
</taxonomicName>
swamps, a unique, biodiverse forest swamp habitat endemic to the Western Ghats, where the
<typeStatus id="54A2884FFF8DD67DFDFBD6A7B3C4F97D" box="[617,665,1762,1788]" pageId="10" pageNumber="161">type</typeStatus>
series was observed and collected.
</paragraph>
</subSubSection>
<subSubSection id="C3036566FF8DD665FF55D740B0B0FF78" lastPageId="18" lastPageNumber="169" pageId="10" pageNumber="161" type="description">
<paragraph id="8BA636EDFF8DD67DFF55D740B01DF82E" blockId="10.[151,1437,1581,2040]" pageId="10" pageNumber="161">
<emphasis id="B96DEAFFFF8DD67DFF55D740B32AF89E" bold="true" box="[199,631,1797,1823]" pageId="10" pageNumber="161">
Description of
<typeStatus id="54A2884FFF8DD67DFEEFD740B0B9F89E" box="[381,484,1797,1823]" pageId="10" pageNumber="161" type="holotype">holotype</typeStatus>
(
<figureCitation id="13222A68FF8DD67DFE64D740B314F89E" box="[502,585,1797,1823]" captionStart="FIGURE 7" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-33@12.[151,1436,181,1833]" captionTargetPageId="12" captionText="FIGURE 7. Protosticta myristicaensis sp. nov. Holotype male [NCBS-BH110]: (a) habitus, lateral view; (b) face; (c) prothorax, dorsal view; (d) pterostigma of right HW &amp; FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411683" httpUri="https://zenodo.org/record/4411683/files/figure.png" pageId="10" pageNumber="161">Figs. 7</figureCitation>
,
<figureCitation id="13222A68FF8DD67DFDCBD740B335F89E" box="[601,616,1797,1823]" captionStart="FIGURE 9" captionStartId="14.[151,250,1797,1821]" captionTargetBox="[151,1435,181,1773]" captionTargetId="figure-33@14.[151,1435,181,1773]" captionTargetPageId="14" captionText="FIGURE 9. Genital ligula of Protosticta myristicaensis sp. nov. [based on NCBS-BH110 &amp; NCBS-BM656]: (a) dorsal view; (b) ventral view; (c) left lateral view." figureDoi="http://doi.org/10.5281/zenodo.4411687" httpUri="https://zenodo.org/record/4411687/files/figure.png" pageId="10" pageNumber="161">9</figureCitation>
).
</emphasis>
<emphasis id="B96DEAFFFF8DD67DFD13D740B3E3F89E" box="[641,702,1797,1823]" italics="true" pageId="10" pageNumber="161">Head</emphasis>
(
<figureCitation id="13222A68FF8DD67DFD42D740B27AF8A1" box="[720,807,1797,1824]" captionStart="FIGURE 7" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-33@12.[151,1436,181,1833]" captionTargetPageId="12" captionText="FIGURE 7. Protosticta myristicaensis sp. nov. Holotype male [NCBS-BH110]: (a) habitus, lateral view; (b) face; (c) prothorax, dorsal view; (d) pterostigma of right HW &amp; FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411683" httpUri="https://zenodo.org/record/4411683/files/figure.png" pageId="10" pageNumber="161">Fig. 7b</figureCitation>
). Labrum, anteclypeus, postclypeus bright blue in live condition, pinkish white in preserved individual; lower 1/3rd of labrum, mandibles and labium brown; antefrons, postfrons black; vertex and postocular lobes black with metallic green sheen; eyes pastel blue in situ, pale pinkish brown post-mortem; both sides of the occipital ridge with a small rounded protuberance; base of antenna white, filament black.
</paragraph>
<paragraph id="8BA636EDFF8DD67CFF55D7FCB422FF78" blockId="10.[151,1437,1581,2040]" lastBlockId="11.[151,1437,150,2013]" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161">
<emphasis id="B96DEAFFFF8DD67DFF55D7FCB07DF852" box="[199,288,1977,2003]" italics="true" pageId="10" pageNumber="161">Thorax.</emphasis>
Prothorax (
<figureCitation id="13222A68FF8DD67DFE31D7FCB0AEF855" box="[419,499,1977,2004]" captionStart="FIGURE 7" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-33@12.[151,1436,181,1833]" captionTargetPageId="12" captionText="FIGURE 7. Protosticta myristicaensis sp. nov. Holotype male [NCBS-BH110]: (a) habitus, lateral view; (b) face; (c) prothorax, dorsal view; (d) pterostigma of right HW &amp; FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411683" httpUri="https://zenodo.org/record/4411683/files/figure.png" pageId="10" pageNumber="161">Fig. 7c</figureCitation>
) dorsally bright turquoise blue, brown laterally, dorsal carina pale yellow. Pterothorax (
<figureCitation id="13222A68FF8DD67DFF5ED798B040F879" box="[204,285,2013,2040]" captionStart="FIGURE 7" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-33@12.[151,1436,181,1833]" captionTargetPageId="12" captionText="FIGURE 7. Protosticta myristicaensis sp. nov. Holotype male [NCBS-BH110]: (a) habitus, lateral view; (b) face; (c) prothorax, dorsal view; (d) pterostigma of right HW &amp; FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411683" httpUri="https://zenodo.org/record/4411683/files/figure.png" pageId="10" pageNumber="161">Fig. 7a</figureCitation>
) black, with copper colored sheen, marked with white: dorsal carina, across mesepimeron occupying dorsal 1/3
<superScript id="7C6C9BA5FF8CD67CFE9AD0D3B04BFF25" attach="none" box="[264,278,150,164]" fontSize="6" pageId="11" pageNumber="162">rd</superScript>
, connected to white metacoxae, and across metepimeron connecting to the white metacoxae laterally, metinfraepisternum slightly white at ventral border adjacent to metacoxae; ventrum of metathorax black.
<emphasis id="B96DEAFFFF8CD67CFAD2D0FEB428FF55" box="[1344,1397,187,212]" italics="true" pageId="11" pageNumber="162">Legs</emphasis>
including coxae white, femur faintly marked black on the posterior face, tarsus dark at apices, tarsal claws yellow.
</paragraph>
<paragraph id="8BA636EDFF8CD67CFF55D146B07FFEE4" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="162">
<emphasis id="B96DEAFFFF8CD67CFF55D146B056FE9D" box="[199,267,259,284]" italics="true" pageId="11" pageNumber="162">Wings</emphasis>
(
<figureCitation id="13222A68FF8CD67CFE89D146B0DFFE9F" box="[283,386,259,286]" captionStart="FIGURE 7" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-33@12.[151,1436,181,1833]" captionTargetPageId="12" captionText="FIGURE 7. Protosticta myristicaensis sp. nov. Holotype male [NCBS-BH110]: (a) habitus, lateral view; (b) face; (c) prothorax, dorsal view; (d) pterostigma of right HW &amp; FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411683" httpUri="https://zenodo.org/record/4411683/files/figure.png" pageId="11" pageNumber="162">Fig. 7a,d</figureCitation>
). Basal wing venation white; Pt dark brown bordered with yellow; two cells between junction of RP
<subScript id="179D34A8FF8CD67CFF4BD17CB1BCFEC6" attach="left" box="[217,225,313,327]" fontSize="6" pageId="11" pageNumber="162">1</subScript>
-RP
<subScript id="179D34A8FF8CD67CFE9FD17CB048FEC6" attach="left" box="[269,277,313,327]" fontSize="6" pageId="11" pageNumber="162">2</subScript>
and origin of IR
<quantity id="4CE19B08FF8CD67CFE46D17CB0A6FEC0" box="[468,507,295,327]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="162" unit="in" value="1.0">
<subScript id="179D34A8FF8CD67CFE46D17CB081FEC6" attach="left" box="[468,476,313,327]" fontSize="6" pageId="11" pageNumber="162">1</subScript>
in
</quantity>
FW and right HW, three in left HW; Ab absent; Ax:
<quantity id="4CE19B08FF8CD67CFBCAD162B5D9FEC0" box="[1112,1156,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="162" unit="in" value="2.0">2 in</quantity>
all wings; Px: FW = 11, HW = 910.
</paragraph>
<paragraph id="8BA636EDFF8CD67CFF55D12AB421FDBC" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="162">
<emphasis id="B96DEAFFFF8CD67CFF55D12AB06CFE08" box="[199,305,367,393]" italics="true" pageId="11" pageNumber="162">Abdomen</emphasis>
(
<figureCitation id="13222A68FF8CD67CFED3D12AB0C9FE08" box="[321,404,367,393]" captionStart="FIGURE 7" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-33@12.[151,1436,181,1833]" captionTargetPageId="12" captionText="FIGURE 7. Protosticta myristicaensis sp. nov. Holotype male [NCBS-BH110]: (a) habitus, lateral view; (b) face; (c) prothorax, dorsal view; (d) pterostigma of right HW &amp; FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411683" httpUri="https://zenodo.org/record/4411683/files/figure.png" pageId="11" pageNumber="162">Fig. 7a</figureCitation>
). Marked with black and white: anterior half of S1 white laterally, S2 with a slanting white marking laterally reaching 2/3
<superScript id="7C6C9BA5FF8CD67CFE7FD1D7B0A6FE21" attach="left" box="[493,507,402,416]" fontSize="6" pageId="11" pageNumber="162">rd</superScript>
of its length; S37 with white annules at anterior border and black annules at the posterior border, white markings reduced on S3, more prominent on subsequent segments; S8 with a broad white basal ring occupying about 3/4
<superScript id="7C6C9BA5FF8CD67CFE61D19FB35DFE69" attach="left" box="[499,512,474,488]" fontSize="6" pageId="11" pageNumber="162">th</superScript>
of the segment, continued ventro-laterally till posterior margin; S910 black. Anterior hamuli small, ovoid, dark brown, covered with thick brown hair at apices. Posterior hamuli saddle-shaped, broader at base curved anteriorly at a 45° angle, base covered with sparse hair, apex covered by a plume of hair.
</paragraph>
<paragraph id="8BA636EDFF8CD67CFF55D202B06DFD70" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="162">
<emphasis id="B96DEAFFFF8CD67CFF55D202B03BFDE0" box="[199,358,583,609]" italics="true" pageId="11" pageNumber="162">Genital ligula</emphasis>
(
<figureCitation id="13222A68FF8CD67CFEEBD202B09CFDE0" box="[377,449,583,609]" captionStart="FIGURE 9" captionStartId="14.[151,250,1797,1821]" captionTargetBox="[151,1435,181,1773]" captionTargetId="figure-33@14.[151,1435,181,1773]" captionTargetPageId="14" captionText="FIGURE 9. Genital ligula of Protosticta myristicaensis sp. nov. [based on NCBS-BH110 &amp; NCBS-BM656]: (a) dorsal view; (b) ventral view; (c) left lateral view." figureDoi="http://doi.org/10.5281/zenodo.4411687" httpUri="https://zenodo.org/record/4411687/files/figure.png" pageId="11" pageNumber="162">Fig. 9</figureCitation>
). Ventral half of first and second segments black, dorsally dark brown; first segment thicker at base, thinner midway and thick at the juncture with second segment; second and third segments attached; third segment complex; dorsally with two inwardly curved ridges on apices; these two ridges connecting at the apices; laterally third segment with conical protuberances on both sides, apices tapering, elongated reaching till the first segment.
</paragraph>
<paragraph id="8BA636EDFF8CD67CFF55D2BEB303FC48" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="162">
<emphasis id="B96DEAFFFF8CD67CFF55D2BEB0FAFC94" box="[199,423,763,789]" italics="true" pageId="11" pageNumber="162">Caudal appendages</emphasis>
(
<figureCitation id="13222A68FF8CD67CFE24D2BEB0A5FC97" box="[438,504,763,790]" captionStart="FIGURE 7" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-33@12.[151,1436,181,1833]" captionTargetPageId="12" captionText="FIGURE 7. Protosticta myristicaensis sp. nov. Holotype male [NCBS-BH110]: (a) habitus, lateral view; (b) face; (c) prothorax, dorsal view; (d) pterostigma of right HW &amp; FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411683" httpUri="https://zenodo.org/record/4411683/files/figure.png" pageId="11" pageNumber="162">Fig. 7</figureCitation>
ef). Cerci black, about 3.5 times of S
<quantity id="4CE19B08FF8CD67CFC36D2BEB283FC94" box="[932,990,763,789]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="162" unit="in" value="10.0">10 in</quantity>
length, curving inwards at apices; cerci with a basal spine, bifurcated at the apices into two arms; inner, ventral arm thick at its base, curved, with bluntly pointed apices, expanded at base and curved downwards, at the base of inner arm a small blunt spine pointing posteriorly; outer dorsal arm much thinner than inner, slightly thinner at base, thicker medially and with blunt thin apices; paraprocts brown, curved ventro-laterally inwards and upwards towards apices, a small inwardly curved basal spine present, broader at base, apices rounded.
</paragraph>
<paragraph id="8BA636EDFF8CD67CFF55D396B46FFC6C" blockId="11.[151,1437,150,2013]" box="[199,1330,979,1005]" pageId="11" pageNumber="162">Measurements: abdomen + caudal appendages = 19.2, FW &amp; HW = 12, cerci = 0.7, paraprocts = 0.6.</paragraph>
<paragraph id="8BA636EDFF8CD67CFF55D3B3B3FFFB44" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="162">
<emphasis id="B96DEAFFFF8CD67CFF55D3B3B352FB91" bold="true" box="[199,527,1014,1041]" pageId="11" pageNumber="162">
Variation in males (
<figureCitation id="13222A68FF8CD67CFE2AD3B3B0A2FB91" box="[440,511,1014,1040]" captionStart="FIGURE 8" captionStartId="13.[151,250,1657,1681]" captionTargetBox="[151,1436,181,1633]" captionTargetId="figure-17@13.[151,1436,181,1633]" captionTargetPageId="13" captionText="FIGURE 8. Protosticta myristicaensis sp. nov. Paratype males: (ab) NCBS-BM655, (cf) NCBS-BM654: (a) prothorax, dorsal view; (b) caudal appendages, ventral view; (c) face; (d) pterostigma of right FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411685" httpUri="https://zenodo.org/record/4411685/files/figure.png" pageId="11" pageNumber="162">Fig. 8</figureCitation>
).
</emphasis>
In NCBS-BM654, bright markings on S27 fainter; marking on S8 slightly irregular on left lateral area, but of same length as
<typeStatus id="54A2884FFF8CD67CFD55D45EB270FBB4" box="[711,813,1051,1077]" pageId="11" pageNumber="162" type="holotype">holotype</typeStatus>
; yellow border of pterostigma less extensive; two cells between junction of RP
<subScript id="179D34A8FF8CD67CFE36D414B0F1FBDE" attach="left" box="[420,428,1105,1119]" fontSize="6" pageId="11" pageNumber="162">1</subScript>
-RP
<subScript id="179D34A8FF8CD67CFE4AD414B0BDFBDE" attach="left" box="[472,480,1105,1119]" fontSize="6" pageId="11" pageNumber="162">2</subScript>
and origin of IR
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<subScript id="179D34A8FF8CD67CFD30D414B3F7FBDE" attach="left" box="[674,682,1105,1119]" fontSize="6" pageId="11" pageNumber="162">1</subScript>
in
</quantity>
FW, three in the HW. In NCBS-BM655, border of anteclypeus and postclypeus bluish, postclypeus brown; three cells between junction of RP
<subScript id="179D34A8FF8CD67CFB98D430B54FFB02" attach="left" box="[1034,1042,1141,1155]" fontSize="6" pageId="11" pageNumber="162">1</subScript>
-RP
<subScript id="179D34A8FF8CD67CFBACD430B51BFB02" attach="left" box="[1086,1094,1141,1155]" fontSize="6" pageId="11" pageNumber="162">2</subScript>
and origin of IR
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in
</quantity>
all wings. In NCBS-BM656, three cells between junction of RP
<subScript id="179D34A8FF8CD67CFD7CD4DCB3ABFB26" attach="right" box="[750,758,1177,1191]" fontSize="6" pageId="11" pageNumber="162">1</subScript>
-RP
<subScript id="179D34A8FF8CD67CFCB3D4DCB274FB26" attach="left" box="[801,809,1177,1191]" fontSize="6" pageId="11" pageNumber="162">2</subScript>
and origin of IR
<quantity id="4CE19B08FF8CD67CFC77D4DCB55AFB20" box="[997,1031,1159,1191]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="162" unit="in" value="1.0">
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in
</quantity>
all wings, except left HW with four cells. Immature males have white pterostigma.
</paragraph>
<paragraph id="8BA636EDFF8CD67CFF55D48AB344FB68" blockId="11.[151,1437,150,2013]" box="[199,537,1231,1257]" pageId="11" pageNumber="162">Px: HW = 910, FW = 1011.</paragraph>
<paragraph id="8BA636EDFF8CD67CFF55D4B7B3A8FA8D" blockId="11.[151,1437,150,2013]" box="[199,757,1266,1292]" pageId="11" pageNumber="162">
<emphasis id="B96DEAFFFF8CD67CFF55D4B7B3A8FA8D" bold="true" box="[199,757,1266,1292]" pageId="11" pageNumber="162">
Description of female (NCBS-BH111) (
<figureCitation id="13222A68FF8CD67CFD00D4B7B3BBFA8D" box="[658,742,1266,1292]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1920]" captionTargetId="figure-17@15.[214,1374,181,1920]" captionTargetPageId="15" captionText="FIGURE 10. Protosticta myristicaensis sp. nov. Paratype female [NCBS-BH111]: (a) habitus, lateral view; (b) face; (c) pterostigma of right FW; (d) prothorax, left lateral view; (e) prothorax, dorsal view; (f) caudal appendages, dorsal view; (g) caudal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4411689" httpUri="https://zenodo.org/record/4411689/files/figure.png" pageId="11" pageNumber="162">Fig. 10</figureCitation>
).
</emphasis>
</paragraph>
<paragraph id="8BA636EDFF8CD67CFF55D552B192FA1C" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="162">
<emphasis id="B96DEAFFFF8CD67CFF55D552B059FAB0" box="[199,260,1303,1329]" italics="true" pageId="11" pageNumber="162">Head</emphasis>
(
<figureCitation id="13222A68FF8CD67CFE87D552B025FAB3" box="[277,376,1303,1330]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1920]" captionTargetId="figure-17@15.[214,1374,181,1920]" captionTargetPageId="15" captionText="FIGURE 10. Protosticta myristicaensis sp. nov. Paratype female [NCBS-BH111]: (a) habitus, lateral view; (b) face; (c) pterostigma of right FW; (d) prothorax, left lateral view; (e) prothorax, dorsal view; (f) caudal appendages, dorsal view; (g) caudal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4411689" httpUri="https://zenodo.org/record/4411689/files/figure.png" pageId="11" pageNumber="162">Fig. 10b</figureCitation>
). Labrum, anteclypeus, bright blue; lower half of labrum, mandibles and labium dark brown; postclypeus, frons, vertex and postocular space black with metallic green sheen; both sides of the occipital ridge with a small rounded protuberance; base of antennae white, rest dark brown; eyes brown centrally largely pale blue.
</paragraph>
<paragraph id="8BA636EDFF8CD67CFF55D5E2B049F960" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="162">
<emphasis id="B96DEAFFFF8CD67CFF55D5E2B07DFA40" box="[199,288,1447,1473]" italics="true" pageId="11" pageNumber="162">Thorax.</emphasis>
Prothorax (
<figureCitation id="13222A68FF8CD67CFE31D5E2B0AFFA43" box="[419,498,1447,1474]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1920]" captionTargetId="figure-17@15.[214,1374,181,1920]" captionTargetPageId="15" captionText="FIGURE 10. Protosticta myristicaensis sp. nov. Paratype female [NCBS-BH111]: (a) habitus, lateral view; (b) face; (c) pterostigma of right FW; (d) prothorax, left lateral view; (e) prothorax, dorsal view; (f) caudal appendages, dorsal view; (g) caudal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4411689" httpUri="https://zenodo.org/record/4411689/files/figure.png" pageId="11" pageNumber="162">Fig. 10</figureCitation>
de) bright blue in situ, laterally median and posterior lobes faint purple, propleuron brown; anterior margin of anterior lobe raised; posterior lobe depressed medially. Pterothorax (
<figureCitation id="13222A68FF8CD67CFB4BD58EB461FA64" box="[1241,1340,1483,1509]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1920]" captionTargetId="figure-17@15.[214,1374,181,1920]" captionTargetPageId="15" captionText="FIGURE 10. Protosticta myristicaensis sp. nov. Paratype female [NCBS-BH111]: (a) habitus, lateral view; (b) face; (c) pterostigma of right FW; (d) prothorax, left lateral view; (e) prothorax, dorsal view; (f) caudal appendages, dorsal view; (g) caudal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4411689" httpUri="https://zenodo.org/record/4411689/files/figure.png" pageId="11" pageNumber="162">Fig. 10a</figureCitation>
). Dorsal carina white, mesepisternum brown with purplish sheen at some viewing angles, dorsal carina pale yellow, mesepimeron brown with a faint metallic blue band adjacent to the ventral border; white band across upper half of metepisternum connecting to mesocoxae; mesinfraepisternum largely brown, white at lower border adjacent to coxa; metepimeron with lower half white; brown marking on lower half of metepisternum and upper half of metepimeron connecting, expanded medially, tapering posteriorly, anteriorly continued on metinfraepisternum; venter of metathorax black with a medial broad white band continuing on S1.
<emphasis id="B96DEAFFFF8CD67CFCF6D6E6B2C4F93D" box="[868,921,1699,1724]" italics="true" pageId="11" pageNumber="162">Legs</emphasis>
including coxae white, junction of femur and tibia black.
</paragraph>
<paragraph id="8BA636EDFF8CD67CFF55D6AEB0B2F8CC" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="162">
<emphasis id="B96DEAFFFF8CD67CFF55D6AEB056F885" box="[199,267,1771,1796]" italics="true" pageId="11" pageNumber="162">Wings</emphasis>
(
<figureCitation id="13222A68FF8CD67CFE89D6AEB0CDF884" box="[283,400,1771,1797]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1920]" captionTargetId="figure-17@15.[214,1374,181,1920]" captionTargetPageId="15" captionText="FIGURE 10. Protosticta myristicaensis sp. nov. Paratype female [NCBS-BH111]: (a) habitus, lateral view; (b) face; (c) pterostigma of right FW; (d) prothorax, left lateral view; (e) prothorax, dorsal view; (f) caudal appendages, dorsal view; (g) caudal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4411689" httpUri="https://zenodo.org/record/4411689/files/figure.png" pageId="11" pageNumber="162">Fig. 10a,c</figureCitation>
). Basal membranules white, venation black; Pt brown post-mortem, slightly longer than one cell; three cells between junction of RP
<subScript id="179D34A8FF8CD67CFDC3D764B304F8AE" attach="left" box="[593,601,1825,1839]" fontSize="6" pageId="11" pageNumber="162">1</subScript>
-RP
<subScript id="179D34A8FF8CD67CFD17D764B3D0F8AE" attach="left" box="[645,653,1825,1839]" fontSize="6" pageId="11" pageNumber="162">2</subScript>
and origin of IR
<quantity id="4CE19B08FF8CD67CFCDBD764B231F8A8" box="[841,876,1807,1839]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="162" unit="in" value="1.0">
<subScript id="179D34A8FF8CD67CFCDBD764B20CF8AE" attach="left" box="[841,849,1825,1839]" fontSize="6" pageId="11" pageNumber="162">1</subScript>
in
</quantity>
all wings; Ab absent; Ax:
<quantity id="4CE19B08FF8CD67CFB05D74AB59FF8A8" box="[1175,1218,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="162" unit="in" value="2.0">2 in</quantity>
all wings; Px: FW: left = 10, right = 11, HW = 10.
</paragraph>
<paragraph id="8BA636EDFF8CD67CFF55D712B3B0F85C" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="162">
<emphasis id="B96DEAFFFF8CD67CFF55D712B06CF8F0" box="[199,305,1879,1905]" italics="true" pageId="11" pageNumber="162">Abdomen</emphasis>
(
<figureCitation id="13222A68FF8CD67CFED3D712B0FEF8F0" box="[321,419,1879,1905]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1920]" captionTargetId="figure-17@15.[214,1374,181,1920]" captionTargetPageId="15" captionText="FIGURE 10. Protosticta myristicaensis sp. nov. Paratype female [NCBS-BH111]: (a) habitus, lateral view; (b) face; (c) pterostigma of right FW; (d) prothorax, left lateral view; (e) prothorax, dorsal view; (f) caudal appendages, dorsal view; (g) caudal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4411689" httpUri="https://zenodo.org/record/4411689/files/figure.png" pageId="11" pageNumber="162">Fig. 10a</figureCitation>
). Anterior half of S1 laterally white; S28 with white annules at the anterior border, more extensive laterally and ventrally; dorsal markings reduced on S2; white marking at the anterior border of S8 not connected dorsally, continuing laterally about ½ the length of the segment; S9 white laterally but not connected dorsally; S10 brown; S27 darker at posterior border.
</paragraph>
<caption id="DF666665FF8BD67BFF05D704B4D2F8FC" ID-DOI="http://doi.org/10.5281/zenodo.4411683" ID-Zenodo-Dep="4411683" httpUri="https://zenodo.org/record/4411683/files/figure.png" pageId="12" pageNumber="163" startId="12.[151,250,1857,1881]" targetBox="[151,1436,181,1833]" targetPageId="12">
<paragraph id="8BA636EDFF8BD67BFF05D704B4D2F8FC" blockId="12.[151,1437,1857,1917]" pageId="12" pageNumber="163">
<emphasis id="B96DEAFFFF8BD67BFF05D704B049F8D8" bold="true" box="[151,276,1857,1881]" pageId="12" pageNumber="163">FIGURE 7.</emphasis>
<taxonomicName id="4C194D6EFF8BD67BFE89D704B37EF8D9" authority="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte, 2020" authorityName="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte" authorityYear="2020" box="[283,547,1857,1880]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="myristicaensis" status="sp. nov.">
<emphasis id="B96DEAFFFF8BD67BFE89D704B37EF8D9" box="[283,547,1857,1880]" italics="true" pageId="12" pageNumber="163">Protosticta myristicaensis</emphasis>
</taxonomicName>
<emphasis id="B96DEAFFFF8BD67BFDB8D704B326F8D8" bold="true" box="[554,635,1857,1881]" pageId="12" pageNumber="163">
<taxonomicNameLabel id="A25E5784FF8BD67BFDB8D704B326F8D8" box="[554,635,1857,1881]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype male [NCBS-BH110]: (a) habitus, lateral view; (b) face; (c) prothorax, dorsal view; (d) pterostigma of right HW &amp; FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view.
</paragraph>
</caption>
<caption id="DF666665FF8AD67AFF05D63CB083F958" ID-DOI="http://doi.org/10.5281/zenodo.4411685" ID-Zenodo-Dep="4411685" httpUri="https://zenodo.org/record/4411685/files/figure.png" pageId="13" pageNumber="164" startId="13.[151,250,1657,1681]" targetBox="[151,1436,181,1633]" targetPageId="13">
<paragraph id="8BA636EDFF8AD67AFF05D63CB083F958" blockId="13.[151,1437,1657,1753]" pageId="13" pageNumber="164">
<emphasis id="B96DEAFFFF8AD67AFF05D63CB04BF910" bold="true" box="[151,278,1657,1681]" pageId="13" pageNumber="164">FIGURE 8.</emphasis>
<taxonomicName id="4C194D6EFF8AD67AFEB2D63FB376F910" authority="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte, 2020" authorityName="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte" authorityYear="2020" box="[288,555,1658,1681]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="myristicaensis" status="sp. nov.">
<emphasis id="B96DEAFFFF8AD67AFEB2D63FB376F910" box="[288,555,1658,1681]" italics="true" pageId="13" pageNumber="164">Protosticta myristicaensis</emphasis>
</taxonomicName>
<emphasis id="B96DEAFFFF8AD67AFDA7D63CB3D4F910" bold="true" box="[565,649,1657,1681]" pageId="13" pageNumber="164">
<taxonomicNameLabel id="A25E5784FF8AD67AFDA7D63CB3D4F910" box="[565,649,1657,1681]" pageId="13" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype males: (ab) NCBS-BM655, (cf) NCBS-BM654: (a) prothorax, dorsal view; (b) caudal appendages, ventral view; (c) face; (d) pterostigma of right FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view.
</paragraph>
</caption>
<caption id="DF666665FF89D679FF05D740B350F8C0" ID-DOI="http://doi.org/10.5281/zenodo.4411687" ID-Zenodo-Dep="4411687" httpUri="https://zenodo.org/record/4411687/files/figure.png" pageId="14" pageNumber="165" startId="14.[151,250,1797,1821]" targetBox="[151,1435,181,1773]" targetPageId="14">
<paragraph id="8BA636EDFF89D679FF05D740B350F8C0" blockId="14.[151,1436,1797,1857]" pageId="14" pageNumber="165">
<emphasis id="B96DEAFFFF89D679FF05D740B04EF89C" bold="true" box="[151,275,1797,1821]" pageId="14" pageNumber="165">FIGURE 9.</emphasis>
Genital ligula of
<taxonomicName id="4C194D6EFF89D679FE5BD743B38DF89C" authority="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte, 2020" authorityName="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte" authorityYear="2020" box="[457,720,1798,1821]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="myristicaensis" status="sp. nov.">
<emphasis id="B96DEAFFFF89D679FE5BD743B38DF89C" box="[457,720,1798,1821]" italics="true" pageId="14" pageNumber="165">Protosticta myristicaensis</emphasis>
</taxonomicName>
<emphasis id="B96DEAFFFF89D679FD45D740B275F89C" bold="true" box="[727,808,1797,1821]" pageId="14" pageNumber="165">
<taxonomicNameLabel id="A25E5784FF89D679FD45D740B275F89C" box="[727,808,1797,1821]" pageId="14" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
[based on NCBS-BH110 &amp; NCBS-BM656]: (a) dorsal view; (b) ventral view; (c) left lateral view.
</paragraph>
</caption>
<caption id="DF666665FF88D678FF05D7D2B0CAF879" ID-DOI="http://doi.org/10.5281/zenodo.4411689" ID-Zenodo-Dep="4411689" httpUri="https://zenodo.org/record/4411689/files/figure.png" pageId="15" pageNumber="166" startId="15.[151,250,1943,1967]" targetBox="[214,1374,181,1920]" targetPageId="15">
<paragraph id="8BA636EDFF88D678FF05D7D2B0CAF879" blockId="15.[151,1436,1943,2040]" pageId="15" pageNumber="166">
<emphasis id="B96DEAFFFF88D678FF05D7D2B07CF82E" bold="true" box="[151,289,1943,1967]" pageId="15" pageNumber="166">FIGURE 10.</emphasis>
<taxonomicName id="4C194D6EFF88D678FEBAD7DDB36DF82E" authority="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte, 2020" authorityName="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte" authorityYear="2020" box="[296,560,1944,1967]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="myristicaensis" status="sp. nov.">
<emphasis id="B96DEAFFFF88D678FEBAD7DDB36DF82E" box="[296,560,1944,1967]" italics="true" pageId="15" pageNumber="166">Protosticta myristicaensis</emphasis>
</taxonomicName>
<emphasis id="B96DEAFFFF88D678FDA5D7DDB3D5F831" bold="true" box="[567,648,1944,1968]" pageId="15" pageNumber="166">
<taxonomicNameLabel id="A25E5784FF88D678FDA5D7DDB3D5F831" box="[567,648,1944,1968]" pageId="15" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype female [NCBS-BH111]: (a) habitus, lateral view; (b) face; (c) pterostigma of right FW; (d) prothorax, left lateral view; (e) prothorax, dorsal view; (f) caudal appendages, dorsal view; (g) caudal appendages, lateral view.
</paragraph>
</caption>
<paragraph id="8BA636EDFF97D667FF55D0D2B277FE9C" blockId="16.[151,1436,151,393]" pageId="16" pageNumber="167">
<emphasis id="B96DEAFFFF97D667FF55D0D2B0F4FF30" box="[199,425,151,177]" italics="true" pageId="16" pageNumber="167">Caudal appendages</emphasis>
(
<figureCitation id="13222A68FF97D667FE2BD0D2B351FF33" box="[441,524,151,178]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1920]" captionTargetId="figure-17@15.[214,1374,181,1920]" captionTargetPageId="15" captionText="FIGURE 10. Protosticta myristicaensis sp. nov. Paratype female [NCBS-BH111]: (a) habitus, lateral view; (b) face; (c) pterostigma of right FW; (d) prothorax, left lateral view; (e) prothorax, dorsal view; (f) caudal appendages, dorsal view; (g) caudal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4411689" httpUri="https://zenodo.org/record/4411689/files/figure.png" pageId="16" pageNumber="167">Fig. 10</figureCitation>
fg). Cerci black, conical, pointed at apices, approximately same length as S10; paraprocts black, rounded, half the length of cerci; ovipositor and valve of ovipositor black, ventrally dark brown, terebra large, about twice the length of cerci, grey-brown; ovipositor dark brown and sharply pointed at apices, ending in long dark brown styles extending well beyond cerci.
</paragraph>
<paragraph id="8BA636EDFF97D667FF55D162B520FEC0" blockId="16.[151,1436,151,393]" box="[199,1149,295,321]" pageId="16" pageNumber="167">Measurements: abdomen + caudal appendages = 20.4, FW &amp; HW = 14, cerci = 0.27.</paragraph>
<paragraph id="8BA636EDFF97D667FF55D10FB4CBFE08" blockId="16.[151,1436,151,393]" pageId="16" pageNumber="167">
<emphasis id="B96DEAFFFF97D667FF55D10FB370FEE5" bold="true" box="[199,557,330,356]" pageId="16" pageNumber="167">
Variation in females (
<figureCitation id="13222A68FF97D667FE58D10FB343FEE5" box="[458,542,330,356]" captionStart="FIGURE 11" captionStartId="16.[151,250,1947,1971]" captionTargetBox="[151,1436,417,1924]" captionTargetId="figure-139@16.[151,1436,417,1924]" captionTargetPageId="16" captionText="FIGURE 11. Protosticta myristicaensis sp. nov. Paratype females: (a-e) NCBS-BM657, (f) NCBS-BH112: (a) face; (b) prothorax, dorsal view; (c) pterostigma of right FW; (d) caudal appendages, dorsal view; (e-f) caudal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4411691" httpUri="https://zenodo.org/record/4411691/files/figure.png" pageId="16" pageNumber="167">Fig. 11</figureCitation>
).
</emphasis>
Wings enfumed brown and basal membranules yellow in NCBS-BH112. Immature females (e.g. NCBS-BM657,
<figureCitation id="13222A68FF97D667FDABD12AB3D8FE08" box="[569,645,367,393]" captionStart="FIGURE 11" captionStartId="16.[151,250,1947,1971]" captionTargetBox="[151,1436,417,1924]" captionTargetId="figure-139@16.[151,1436,417,1924]" captionTargetPageId="16" captionText="FIGURE 11. Protosticta myristicaensis sp. nov. Paratype females: (a-e) NCBS-BM657, (f) NCBS-BH112: (a) face; (b) prothorax, dorsal view; (c) pterostigma of right FW; (d) caudal appendages, dorsal view; (e-f) caudal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4411691" httpUri="https://zenodo.org/record/4411691/files/figure.png" pageId="16" pageNumber="167">Fig. 11</figureCitation>
ae) have white pterostigma and basal membranules, venation brown.
</paragraph>
<caption id="DF666665FF97D667FF05D7DEB404F856" ID-DOI="http://doi.org/10.5281/zenodo.4411691" ID-Zenodo-Dep="4411691" httpUri="https://zenodo.org/record/4411691/files/figure.png" pageId="16" pageNumber="167" startId="16.[151,250,1947,1971]" targetBox="[151,1436,417,1924]" targetPageId="16">
<paragraph id="8BA636EDFF97D667FF05D7DEB404F856" blockId="16.[151,1437,1947,2007]" pageId="16" pageNumber="167">
<emphasis id="B96DEAFFFF97D667FF05D7DEB043F832" bold="true" box="[151,286,1947,1971]" pageId="16" pageNumber="167">FIGURE 11.</emphasis>
<taxonomicName id="4C194D6EFF97D667FEB1D7D9B374F832" authority="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte, 2020" authorityName="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte" authorityYear="2020" box="[291,553,1948,1971]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="myristicaensis" status="sp. nov.">
<emphasis id="B96DEAFFFF97D667FEB1D7D9B374F832" box="[291,553,1948,1971]" italics="true" pageId="16" pageNumber="167">Protosticta myristicaensis</emphasis>
</taxonomicName>
<emphasis id="B96DEAFFFF97D667FDBDD7DEB322F832" bold="true" box="[559,639,1947,1971]" pageId="16" pageNumber="167">
<taxonomicNameLabel id="A25E5784FF97D667FDBDD7DEB322F832" box="[559,639,1947,1971]" pageId="16" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype females: (a-e) NCBS-BM657, (f) NCBS-BH112: (a) face; (b) prothorax, dorsal view; (c) pterostigma of right FW; (d) caudal appendages, dorsal view; (e-f) caudal appendages, lateral view.
</paragraph>
</caption>
<caption id="DF666665FF96D666FF05D7E8B3D3F868" ID-DOI="http://doi.org/10.5281/zenodo.4411695" ID-Zenodo-Dep="4411695" httpUri="https://zenodo.org/record/4411695/files/figure.png" pageId="17" pageNumber="168" startId="17.[151,250,1965,1989]" targetBox="[189,1399,181,1941]" targetPageId="17">
<paragraph id="8BA636EDFF96D666FF05D7E8B3D3F868" blockId="17.[151,1436,1965,2025]" pageId="17" pageNumber="168">
<emphasis id="B96DEAFFFF96D666FF05D7E8B07DF844" bold="true" box="[151,288,1965,1989]" pageId="17" pageNumber="168">FIGURE 12.</emphasis>
Field images of: (ab)
<taxonomicName id="4C194D6EFF96D666FD9FD7EBB25AF844" authority="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte, 2020" authorityName="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte" authorityYear="2020" box="[525,775,1965,1989]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="cyanofemora" status="sp. nov.">
<emphasis id="B96DEAFFFF96D666FD9FD7EBB25AF844" box="[525,775,1965,1989]" italics="true" pageId="17" pageNumber="168">Protosticta cyanofemora</emphasis>
</taxonomicName>
<emphasis id="B96DEAFFFF96D666FC9CD7E8B203F844" bold="true" box="[782,862,1965,1989]" pageId="17" pageNumber="168">
<taxonomicNameLabel id="A25E5784FF96D666FC9CD7E8B203F844" box="[782,862,1965,1989]" pageId="17" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(a) male; (b) female; (cd)
<taxonomicName id="4C194D6EFF96D666FBEAD7EBB47FF844" authority="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte, 2020" authorityName="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte" authorityYear="2020" box="[1144,1314,1966,1989]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="myristicaensis" status="sp. nov.">
<emphasis id="B96DEAFFFF96D666FBEAD7EBB47FF844" box="[1144,1314,1966,1989]" italics="true" pageId="17" pageNumber="168">P. myristicaensis</emphasis>
</taxonomicName>
<emphasis id="B96DEAFFFF96D666FABBD7E8B424F844" bold="true" box="[1321,1401,1965,1989]" pageId="17" pageNumber="168">
<taxonomicNameLabel id="A25E5784FF96D666FABBD7E8B424F844" box="[1321,1401,1965,1989]" pageId="17" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(c) male; (d) female. Photos: (a, cd) SJ, &amp; (b) KAS.
</paragraph>
</caption>
<paragraph id="8BA636EDFF95D665FF55D0D2B004FF54" blockId="18.[151,1437,150,1222]" pageId="18" pageNumber="169">
In NCBS-BM657, three cells between junction of RP
<subScript id="179D34A8FF95D665FCB3D0ECB274FF36" attach="left" box="[801,809,169,183]" fontSize="6" pageId="18" pageNumber="169">1</subScript>
-RP
<subScript id="179D34A8FF95D665FCC7D0ECB200FF36" attach="left" box="[853,861,169,183]" fontSize="6" pageId="18" pageNumber="169">2</subScript>
and origin of IR
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<subScript id="179D34A8FF95D665FB88D0ECB57FFF36" attach="left" box="[1050,1058,169,183]" fontSize="6" pageId="18" pageNumber="169">1</subScript>
in
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left FW and both HW and two cells in right FW.
</paragraph>
<paragraph id="8BA636EDFF95D665FF55D09AB0B0FF78" blockId="18.[151,1437,150,1222]" box="[199,493,223,249]" pageId="18" pageNumber="169">Px: FW = 12, HW: 1011.</paragraph>
</subSubSection>
<subSubSection id="C3036566FF95D665FF55D147B261FB47" pageId="18" pageNumber="169" type="diagnosis">
<paragraph id="8BA636EDFF95D665FF55D147B0DCFE2C" blockId="18.[151,1437,150,1222]" pageId="18" pageNumber="169">
<emphasis id="B96DEAFFFF95D665FF55D147B01FFE9D" bold="true" box="[199,322,258,284]" pageId="18" pageNumber="169">Diagnosis.</emphasis>
This species is clearly the smallest Indian
<taxonomicName id="4C194D6EFF95D665FCBAD146B2FEFE9D" authorityName="Selys" authorityYear="1885" box="[808,931,259,284]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFFFF95D665FCBAD146B2FEFE9D" box="[808,931,259,284]" italics="true" pageId="18" pageNumber="169">Protosticta</emphasis>
</taxonomicName>
. Apart from its small size, males can be distinguished from all other species by the shape of cerci, specifically the small tubercle at the base of the apical fork (
<figureCitation id="13222A68FF95D665FF0DD10EB1A5FEE4" box="[159,248,331,357]" captionStart="FIGURE 7" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-33@12.[151,1436,181,1833]" captionTargetPageId="12" captionText="FIGURE 7. Protosticta myristicaensis sp. nov. Holotype male [NCBS-BH110]: (a) habitus, lateral view; (b) face; (c) prothorax, dorsal view; (d) pterostigma of right HW &amp; FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411683" httpUri="https://zenodo.org/record/4411683/files/figure.png" pageId="18" pageNumber="169">Figs. 7f</figureCitation>
,
<figureCitation id="13222A68FF95D665FE90D10EB06FFEE7" box="[258,306,331,358]" captionStart="FIGURE 8" captionStartId="13.[151,250,1657,1681]" captionTargetBox="[151,1436,181,1633]" captionTargetId="figure-17@13.[151,1436,181,1633]" captionTargetPageId="13" captionText="FIGURE 8. Protosticta myristicaensis sp. nov. Paratype males: (ab) NCBS-BM655, (cf) NCBS-BM654: (a) prothorax, dorsal view; (b) caudal appendages, ventral view; (c) face; (d) pterostigma of right FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411685" httpUri="https://zenodo.org/record/4411685/files/figure.png" pageId="18" pageNumber="169">8b,f</figureCitation>
). It can be further distinguished from congeneric species by the color of eyes (blue,
<figureCitation id="13222A68FF95D665FB43D10EB473FEE4" box="[1233,1326,331,357]" captionStart="FIGURE 12" captionStartId="17.[151,250,1965,1989]" captionTargetBox="[189,1399,181,1941]" captionTargetId="figure-17@17.[189,1399,181,1941]" captionTargetPageId="17" captionText="FIGURE 12. Field images of: (ab) Protosticta cyanofemora sp. nov. (a) male; (b) female; (cd) P. myristicaensis sp. nov. (c) male; (d) female. Photos: (a, cd) SJ, &amp; (b) KAS." figureDoi="http://doi.org/10.5281/zenodo.4411695" httpUri="https://zenodo.org/record/4411695/files/figure.png" pageId="18" pageNumber="169">Fig. 12c</figureCitation>
), uniform coloration of prothorax (purple in live condition) (
<figureCitation id="13222A68FF95D665FD58D12AB27AFE08" box="[714,807,367,393]" captionStart="FIGURE 7" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-33@12.[151,1436,181,1833]" captionTargetPageId="12" captionText="FIGURE 7. Protosticta myristicaensis sp. nov. Holotype male [NCBS-BH110]: (a) habitus, lateral view; (b) face; (c) prothorax, dorsal view; (d) pterostigma of right HW &amp; FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411683" httpUri="https://zenodo.org/record/4411683/files/figure.png" pageId="18" pageNumber="169">Figs. 7c</figureCitation>
,
<figureCitation id="13222A68FF95D665FCA0D12AB212FE08" box="[818,847,367,393]" captionStart="FIGURE 8" captionStartId="13.[151,250,1657,1681]" captionTargetBox="[151,1436,181,1633]" captionTargetId="figure-17@13.[151,1436,181,1633]" captionTargetPageId="13" captionText="FIGURE 8. Protosticta myristicaensis sp. nov. Paratype males: (ab) NCBS-BM655, (cf) NCBS-BM654: (a) prothorax, dorsal view; (b) caudal appendages, ventral view; (c) face; (d) pterostigma of right FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411685" httpUri="https://zenodo.org/record/4411685/files/figure.png" pageId="18" pageNumber="169">8a</figureCitation>
), and bright marking on S8 (anterior 3/4
<superScript id="7C6C9BA5FF95D665FA87D12BB47FFEFD" attach="left" box="[1301,1314,366,380]" fontSize="6" pageId="18" pageNumber="169">th</superScript>
of the segment white,
<figureCitation id="13222A68FF95D665FEB0D1D6B02EFE2C" box="[290,371,403,429]" captionStart="FIGURE 7" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-33@12.[151,1436,181,1833]" captionTargetPageId="12" captionText="FIGURE 7. Protosticta myristicaensis sp. nov. Holotype male [NCBS-BH110]: (a) habitus, lateral view; (b) face; (c) prothorax, dorsal view; (d) pterostigma of right HW &amp; FW; (e) caudal appendages, lateral view; (f) caudal appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411683" httpUri="https://zenodo.org/record/4411683/files/figure.png" pageId="18" pageNumber="169">Fig. 7a</figureCitation>
).
</paragraph>
<paragraph id="8BA636EDFF95D665FF55D1F2B398FD98" blockId="18.[151,1437,150,1222]" pageId="18" pageNumber="169">Female of this species can be differentiated from other species in Western Ghats by its small size, color of prothorax (purple in live condition), and the extent and shape of bright markings on S89. Both males &amp; females can be distinguished by the broad white dorsal carina.</paragraph>
<paragraph id="8BA636EDFF95D665FF55D267B4C6FCDC" blockId="18.[151,1437,150,1222]" pageId="18" pageNumber="169">
<emphasis id="B96DEAFFFF95D665FF55D267B09AFDBC" bold="true" box="[199,455,546,573]" pageId="18" pageNumber="169">Habitat and Ecology.</emphasis>
<taxonomicName id="4C194D6EFF95D665FE42D266B366FDBC" box="[464,571,547,573]" class="Magnoliopsida" family="Myristicaceae" genus="Myristica" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="genus">Myristica</taxonomicName>
swamps are freshwater swamps characterized by trees with stilt roots (
<figureCitation id="13222A68FF95D665FAFDD266B199FDE3" captionStart="FIGURE 18" captionStartId="25.[151,250,1959,1983]" captionTargetBox="[403,1185,181,1935]" captionTargetId="figure-17@25.[403,1185,181,1935]" captionTargetPageId="25" captionText="FIGURE 18. Habitats of (a) Protosticta cyanofemora sp. nov.; (b) P. myristicaensis sp. nov.; (c) P. sholai sp. nov. Photos: (a) Raghu Ram, (b) SJ, &amp; (c) KAS." figureDoi="http://doi.org/10.5281/zenodo.4411707" httpUri="https://zenodo.org/record/4411707/files/figure.png" pageId="18" pageNumber="169">Fig. 18b</figureCitation>
). These are relict habitats thought to have existed over large areas but have become highly fragmented, and now occupy less than 200 hectares (
<bibRefCitation id="EF884B1CFF95D665FDBBD22EB249FD04" author="Kumaran, K. &amp; Limaye, R. B. &amp; Punekar, S. A. &amp; Rajaguru, S. N. &amp; Joshi, S. V. &amp; Karlekar, S. N." box="[553,788,619,645]" pageId="18" pageNumber="169" pagination="3 - 18" refId="ref15409" refString="Kumaran, K., Limaye, R. B., Punekar, S. A., Rajaguru, S. N., Joshi, S. V. &amp; Karlekar, S. N. (2013) Vegetation response to South Asian Monsoon variations in Konkan, western India during the Late Quaternary: Evidence from fluvio-lacustrine archives. Quaternary International, 286, 3 - 18. https: // doi. org / 10.1016 / j. quaint. 2012.03.010" type="journal article" year="2013">
Kumaran
<emphasis id="B96DEAFFFF95D665FD08D22EB38FFD04" box="[666,722,619,645]" italics="true" pageId="18" pageNumber="169">et al.</emphasis>
2013
</bibRefCitation>
,
<bibRefCitation id="EF884B1CFF95D665FC8DD22EB2AAFD07" author="Limaye, R. B. &amp; Padmalal, D. &amp; Kumaran, K." box="[799,1015,619,646]" pageId="18" pageNumber="169" pagination="143 - 154" refId="ref15529" refString="Limaye, R. B., Padmalal, D. &amp; Kumaran, K. (2016) Late Pleistocenee Holocene monsoon variations on climate, landforms and vegetation covering southwestern India: An overview. Quaternary International, 443, 143 - 154. https: // doi. org / 10.1016 / j. quaint. 2016.08.004" type="journal article" year="2016">
Limaye
<emphasis id="B96DEAFFFF95D665FCE9D22EB2EEFD04" box="[891,947,619,645]" italics="true" pageId="18" pageNumber="169">et al.</emphasis>
2016
</bibRefCitation>
). These unique ecosystems are highly threatened from various factors such as land-use changes like conversion into agricultural fields and plantations (
<bibRefCitation id="EF884B1CFF95D665FF0DD2F6B09FFD4C" author="Chandran, M. D. &amp; Mesta, D." box="[159,450,691,717]" pageId="18" pageNumber="169" pagination="1 - 19" refId="ref14402" refString="Chandran, M. D. &amp; Mesta, D. (2001) On the conservation of the Myristica swamps of the Western Ghats. In: Uma Shaanker, R., Ganeshaiah, K. N. &amp; Bawa, K. S. (Eds.), Forest Genetic Resources: Status, Threats, and Conservation Strategies. Oxford &amp; IBH Publishing Co., New Delhi, pp. 1 - 19." type="book chapter" year="2001">Chandran &amp; Mesta 2001</bibRefCitation>
), and excessive water withdrawal (
<bibRefCitation id="EF884B1CFF95D665FCC8D2F6B55EFD4C" author="Ganesan, R." box="[858,1027,691,717]" pageId="18" pageNumber="169" pagination="1351 - 1359" refId="ref14929" refString="Ganesan, R. (2002) Evergreen freshwater swamps and their plant diversity on Kalakkad Mundanthurai Tiger Reserve, south Western Ghats, India. The Indian Forester, 12, 1351 - 1359." type="journal article" year="2002">Ganesan 2002</bibRefCitation>
). We also observed this at the
<typeStatus id="54A2884FFF95D665FAFED2F6B4C1FD4C" box="[1388,1436,691,717]" pageId="18" pageNumber="169">type</typeStatus>
locality of
<taxonomicName id="4C194D6EFF95D665FE86D292B08CFD71" authorityName="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte" authorityYear="2020" box="[276,465,727,752]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="myristicaensis">
<emphasis id="B96DEAFFFF95D665FE86D292B08CFD71" box="[276,465,727,752]" italics="true" pageId="18" pageNumber="169">P. myristicaensis</emphasis>
</taxonomicName>
where on one side of the road
<taxonomicName id="4C194D6EFF95D665FCAAD292B2FEFD70" box="[824,931,727,753]" class="Magnoliopsida" family="Myristicaceae" genus="Myristica" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="genus">Myristica</taxonomicName>
swamps have been converted into areca nut plantations. Of the recorded wildlife in
<taxonomicName id="4C194D6EFF95D665FDC7D2BEB39DFC94" box="[597,704,763,789]" class="Magnoliopsida" family="Myristicaceae" genus="Myristica" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="genus">Myristica</taxonomicName>
swamps 16.3% are endemic to Western Ghats (
<bibRefCitation id="EF884B1CFF95D665FB48D2BEB4CDFC94" author="Jose, J. &amp; Ramachandran, K. K. &amp; Nair, P. V." box="[1242,1424,763,789]" pageId="18" pageNumber="169" pagination="724 - 726" refId="ref15015" refString="Jose, J., Ramachandran, K. K. &amp; Nair, P. V. (2007) Animal diversity of the Myristica swamps forests of Southern Kerala with special reference to herpetofauna. In: Kerala Science Congress &amp; Muthunayagam, A. E. (Eds.), The proceedings of the 19 th Kerala Science Congress. Thiruvananthapuram: Kerala State Council for Science, Technology, and Environment, Government of Kerala, Kerala, pp. 724 - 726." type="book chapter" year="2007">
Jose
<emphasis id="B96DEAFFFF95D665FA86D2BEB410FC94" box="[1300,1357,763,789]" italics="true" pageId="18" pageNumber="169">et al.</emphasis>
2007
</bibRefCitation>
), and many new species exclusive to
<taxonomicName id="4C194D6EFF95D665FDBBD35AB3C9FCB8" box="[553,660,799,825]" class="Magnoliopsida" family="Myristicaceae" genus="Myristica" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="genus">Myristica</taxonomicName>
swamps have been discovered in recent years such as the monotypic
<taxonomicName id="4C194D6EFF95D665FF09D306B03AFCDC" authority="Swamp" authorityName="Swamp" box="[155,359,835,861]" class="Magnoliopsida" family="Myristicaceae" genus="Myristica" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="genus">Myristica Swamp</taxonomicName>
tree frog,
<taxonomicName id="4C194D6EFF95D665FE77D306B4C5FCDC" authority="Abraham, Pyron, Ansil, Zachariah &amp; Zachariah, 2013" authorityName="Abraham, Pyron, Ansil, Zachariah &amp; Zachariah" authorityYear="2013" box="[485,1432,835,861]" class="Amphibia" family="Rhacophoridae" genus="Mercurana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="myristicapalustris">
<emphasis id="B96DEAFFFF95D665FE77D306B26EFCDC" box="[485,819,835,861]" italics="true" pageId="18" pageNumber="169">Mercurana myristicapalustris</emphasis>
Abraham, Pyron, Ansil, Zachariah &amp; Zachariah, 2013
</taxonomicName>
.
</paragraph>
<paragraph id="8BA636EDFF95D665FF55D322B261FB47" blockId="18.[151,1437,150,1222]" pageId="18" pageNumber="169">
<taxonomicName id="4C194D6EFF95D665FF55D322B0B4FC01" authorityName="Joshi &amp; Subramanian &amp; Babu &amp; Sawant &amp; Kunte" authorityYear="2020" box="[199,489,871,896]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="myristicaensis">
<emphasis id="B96DEAFFFF95D665FF55D322B0B4FC01" box="[199,489,871,896]" italics="true" pageId="18" pageNumber="169">Protosticta myristicaensis</emphasis>
</taxonomicName>
was observed exclusively in
<taxonomicName id="4C194D6EFF95D665FCA2D322B2ACFC00" authority="swamp" authorityName="Swamp" box="[816,1009,871,897]" class="Magnoliopsida" family="Myristicaceae" genus="Myristica" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="genus">Myristica swamp</taxonomicName>
patch, along the Bengaluru-Honnavar road at Kathlekan,
<collectingRegion id="49DDF80FFF95D665FEFCD3CEB0BCFC24" box="[366,481,907,933]" country="India" name="Karnataka" pageId="18" pageNumber="169">Karnataka</collectingRegion>
(
<figureCitation id="13222A68FF95D665FE63D3CEB30EFC24" box="[497,595,907,933]" captionStart="FIGURE 23" captionStartId="31.[151,250,1092,1116]" captionTargetBox="[155,1433,185,1066]" captionTargetId="figure-17@31.[151,1436,181,1068]" captionTargetPageId="31" captionText="FIGURE 23. Maps depicting the known distribution of Protosticta spp. in Western Ghats of India. (a) P. cyanofemora sp. nov., P. myristicaensis sp. nov., P. sholai sp. nov.; (b) P. hearseyi, P. ponmudiensis, P. rufostigma; (c) P. antelopoides, P. gravelyi, P. mortoni; (d) P. davenporti, P. monticola, P. sanguinostigma." figureDoi="http://doi.org/10.5281/zenodo.4411717" httpUri="https://zenodo.org/record/4411717/files/figure.png" pageId="18" pageNumber="169">Fig. 23a</figureCitation>
). This site was surveyed twice (
<date id="FFA7102DFF95D665FC28D3CEB5BAFC24" box="[954,1255,907,933]" pageId="18" pageNumber="169" value="2019-04" valueMax="2019-11-31" valueMin="2019-04-00">April and November 2019</date>
) and both times the species was observed (April: four males and two females; November: three males and two females). This species was observed alongside
<taxonomicName id="4C194D6EFF95D665FE4FD396B308FC6C" authorityName="Laidlaw" authorityYear="1915" box="[477,597,979,1005]" class="Insecta" family="Platystictidae" genus="Protositcta" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="gravelyi">
<emphasis id="B96DEAFFFF95D665FE4FD396B308FC6C" box="[477,597,979,1005]" italics="true" pageId="18" pageNumber="169">P. gravelyi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C194D6EFF95D665FDF1D396B512FC6C" authority="Hagen in Selys, 1853" authorityName="Hagen in Selys" authorityYear="1853" box="[611,1103,978,1005]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="bisignata">
<emphasis id="B96DEAFFFF95D665FDF1D396B209FC6C" box="[611,852,979,1005]" italics="true" pageId="18" pageNumber="169">Heliocypha bisignata</emphasis>
Hagen in Selys, 1853
</taxonomicName>
,
<taxonomicName id="4C194D6EFF95D665FBC9D396B186FB90" authority="(Fabricius, 1798)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" class="Insecta" family="Libellulidae" genus="Tholymis" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="tillarga">
<emphasis id="B96DEAFFFF95D665FBC9D396B441FC6C" box="[1115,1308,979,1005]" italics="true" pageId="18" pageNumber="169">Tholymis tillarga</emphasis>
(Fabricius, 1798)
</taxonomicName>
,
<taxonomicName id="4C194D6EFF95D665FF74D3B2B32DFB90" authority="Fraser, 1920" authorityName="Fraser" authorityYear="1920" box="[230,624,1015,1041]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="varralli">
<emphasis id="B96DEAFFFF95D665FF74D3B2B0BCFB90" box="[230,481,1015,1041]" italics="true" pageId="18" pageNumber="169">Mortonagrion varralli</emphasis>
Fraser, 1920
</taxonomicName>
and
<taxonomicName id="4C194D6EFF95D665FD37D3B2B599FB93" authority="(Hagen in Selys, 1860)" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1860" box="[677,1220,1015,1042]" class="Insecta" family="Protoneuridae" genus="Phylloneura" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="westermanii">
<emphasis id="B96DEAFFFF95D665FD37D3B2B2E2FB91" box="[677,959,1015,1041]" italics="true" pageId="18" pageNumber="169">Phylloneura westermanii</emphasis>
(Hagen in Selys, 1860)
</taxonomicName>
(a species endemic to
<taxonomicName id="4C194D6EFF95D665FF27D45EB07DFBB4" box="[181,288,1051,1077]" class="Magnoliopsida" family="Myristicaceae" genus="Myristica" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="genus">Myristica</taxonomicName>
swamps). Unlike other Indian
<taxonomicName id="4C194D6EFF95D665FD13D45EB3A1FBB5" authorityName="Selys" authorityYear="1885" box="[641,764,1051,1076]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAFFFF95D665FD13D45EB3A1FBB5" box="[641,764,1051,1076]" italics="true" pageId="18" pageNumber="169">Protosticta</emphasis>
</taxonomicName>
spp., this species has the unique behavior of perching very low (not flying above the height of
<quantity id="4CE19B08FF95D665FDB9D47AB3C0FBD8" box="[555,669,1087,1113]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="18" pageNumber="169" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">2030 cm</quantity>
) among roots of trees such as
<taxonomicName id="4C194D6EFF95D665FC6AD47AB1B3FBFF" authority="(Bedd.)" baseAuthorityName="Bedd." class="Magnoliopsida" family="Myristicaceae" genus="Myristica" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="variety" species="fatua" variety="magnifica">
<emphasis id="B96DEAFFFF95D665FC6AD47AB5F8FBD8" box="[1016,1189,1087,1113]" italics="true" pageId="18" pageNumber="169">Myristica fatua</emphasis>
Houtt. var.
<emphasis id="B96DEAFFFF95D665FABED47AB4C1FBD8" box="[1324,1436,1087,1113]" italics="true" pageId="18" pageNumber="169">magnifica</emphasis>
(Bedd.)
</taxonomicName>
,
<taxonomicName id="4C194D6EFF95D665FF6ED426B391FBFC" authority="(King) Warb." authorityName="Warb." baseAuthorityName="King" box="[252,716,1123,1150]" class="Magnoliopsida" family="Myristicaceae" genus="Gymnacranthera" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="species" species="canarica">
<emphasis id="B96DEAFFFF95D665FF6ED426B374FBFD" box="[252,553,1123,1149]" italics="true" pageId="18" pageNumber="169">Gymnacranthera canarica</emphasis>
(King) Warb.
</taxonomicName>
,
<taxonomicName id="4C194D6EFF95D665FD4BD426B471FBFC" authority="Dasappa &amp; Swaminath" authorityName="Dasappa &amp; Swaminath" box="[729,1324,1123,1149]" class="Magnoliopsida" family="Anacardiaceae" genus="Semecarpus" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="species" species="kathalekanensis">
<emphasis id="B96DEAFFFF95D665FD4BD426B541FBFC" box="[729,1052,1123,1149]" italics="true" pageId="18" pageNumber="169">Semecarpus kathalekanensis</emphasis>
Dasappa &amp; Swaminath
</taxonomicName>
(all three species endemic to
<taxonomicName id="4C194D6EFF95D665FEE7D4C2B0BDFB20" box="[373,480,1159,1185]" class="Magnoliopsida" family="Myristicaceae" genus="Myristica" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="genus">Myristica</taxonomicName>
swamps),
<taxonomicName id="4C194D6EFF95D665FDC9D4C2B521FB20" authority="(Hook. Fil. &amp; Thomas.) Warb." authorityName="Warb." baseAuthorityName="Hook. Fil. &amp; Thomas." box="[603,1148,1159,1186]" class="Magnoliopsida" family="Myristicaceae" genus="Knema" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="species" species="attenuata">
<emphasis id="B96DEAFFFF95D665FDC9D4C2B241FB21" box="[603,796,1159,1184]" italics="true" pageId="18" pageNumber="169">Knema attenuata</emphasis>
(Hook. Fil. &amp; Thomas.) Warb.
</taxonomicName>
,
<taxonomicName id="4C194D6EFF95D665FB18D4C2B05DFB44" authority="Arn." authorityName="Arn." class="Magnoliopsida" family="Celastraceae" genus="Lophopetalum" kingdom="Plantae" order="Celastrales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="species" species="wrightianum">
<emphasis id="B96DEAFFFF95D665FB18D4C2B19AFB45" italics="true" pageId="18" pageNumber="169">Lophopetalum wrightianum</emphasis>
Arn.
</taxonomicName>
and
<taxonomicName id="4C194D6EFF95D665FEA4D4EEB3E0FB44" authority="(Dennst.) Mabberley" authorityName="Mabberley" authorityYear="1979" baseAuthorityName="Dennst." box="[310,701,1195,1221]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Hopea" kingdom="Plantae" order="Malvales" pageId="18" pageNumber="169" phylum="Tracheophyta" rank="species" species="ponga">
<emphasis id="B96DEAFFFF95D665FEA4D4EEB090FB45" box="[310,461,1195,1220]" italics="true" pageId="18" pageNumber="169">Hopea ponga</emphasis>
(Dennst.) Mabberley
</taxonomicName>
(
<figureCitation id="13222A68FF95D665FD5FD4EEB272FB47" box="[717,815,1195,1222]" captionStart="FIGURE 18" captionStartId="25.[151,250,1959,1983]" captionTargetBox="[403,1185,181,1935]" captionTargetId="figure-17@25.[403,1185,181,1935]" captionTargetPageId="25" captionText="FIGURE 18. Habitats of (a) Protosticta cyanofemora sp. nov.; (b) P. myristicaensis sp. nov.; (c) P. sholai sp. nov. Photos: (a) Raghu Ram, (b) SJ, &amp; (c) KAS." figureDoi="http://doi.org/10.5281/zenodo.4411707" httpUri="https://zenodo.org/record/4411707/files/figure.png" pageId="18" pageNumber="169">Fig. 18b</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>