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<document id="C74FB406077BFF27FC0894C9813E4E0F" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55BE6FF17FF79F04BFB25FC82" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena bifunda Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="133" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="132" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="BC54766F7262989A880CCBE5BC54E266" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9995C052F5A6CCAF48C0F45682566606">
<mods:title id="D332860C4B21F8E8BCBCC2DECA6D22E0">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
</mods:titleInfo>
<mods:name id="013563AD4FBF82E47977909F2B7A9CA5" type="personal">
<mods:role id="7DEF2FFF7D107F470D87E55C02FDED41">
<mods:roleTerm id="970D39AC073650D730B86530213C98F5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3C823DBB6A5815D293ADA31F1D6DBF5B">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="AF790CFABBE9A9CA6BEF641F51E542A9">text</mods:typeOfResource>
<mods:relatedItem id="F2EA4406855EBAABCC97F086C56E7B02" type="host">
<mods:titleInfo id="A69270004732EFA9F6F92BEDF45F4FB0">
<mods:title id="846078220452693612014E03810D1C85">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EF9A2AC75F42F14A222540D250B09DD3">
<mods:date id="E4455A98E3A00B98AF1ADB7A7ED839A7">2011</mods:date>
<mods:detail id="1CA10EEF0FD6B5713FB16A67DF086149" type="pubDate">
<mods:number id="71B9E8A394240D80B791387A7917721B">2011-07-08</mods:number>
</mods:detail>
<mods:detail id="535902CF053E4B94ADAD16B98A66715D" type="volume">
<mods:number id="B2EE4545F2426929059F32691B1B3A85">2944</mods:number>
</mods:detail>
<mods:detail id="E42054BCE6B69B657162E3BF23D33C0D" type="issue">
<mods:number id="C52B0AFA62C847AD03FC8181959FAB04">1</mods:number>
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<mods:start id="1685B71532A68D9328CF15C14F403A36">1</mods:start>
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<mods:location id="3E4DDFBD967DECA0468DB4BEDE048DCB">
<mods:url id="2C3202653A66C5565B4666D800DDE12C">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
</mods:location>
<mods:classification id="092C258EE6FD53F817D2E950B6F8C990">journal article</mods:classification>
<mods:identifier id="2438B9AFA7D0BC806E44A55F1A53749A" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
<mods:identifier id="0DFD11E0F71CD770AB27862D40E3060B" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="1EEC68970AC1B6FCB745F168A07CA5A9" type="Zenodo-Dep">5283420</mods:identifier>
</mods:mods>
<treatment id="03B087E55BE6FF17FF79F04BFB25FC82" ID-DOI="http://doi.org/10.5281/zenodo.5291809" ID-Zenodo-Dep="5291809" LSID="urn:lsid:plazi:treatment:03B087E55BE6FF17FF79F04BFB25FC82" httpUri="http://treatment.plazi.org/id/03B087E55BE6FF17FF79F04BFB25FC82" lastPageId="132" lastPageNumber="133" pageId="131" pageNumber="132">
<subSubSection id="C30365785BE6FF10FF79F04BFDA5FD16" box="[151,531,614,640]" pageId="131" pageNumber="132" type="nomenclature">
<paragraph id="8BA636F35BE6FF10FF79F04BFDA5FD16" blockId="131.[151,531,614,673]" box="[151,531,614,640]" pageId="131" pageNumber="132">
<heading id="D0EE819F5BE6FF10FF79F04BFDA5FD16" bold="true" box="[151,531,614,640]" fontSize="11" level="1" pageId="131" pageNumber="132" reason="1">
<emphasis id="B96DEAE15BE6FF10FF79F04BFDA5FD16" bold="true" box="[151,531,614,640]" pageId="131" pageNumber="132">
<taxonomicName id="4C194D705BE6FF10FF79F04BFEC2FD16" ID-CoL="6MHF6" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,372,614,640]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="bifunda" status="sp. nov.">
<emphasis id="B96DEAE15BE6FF10FF79F04BFEC2FD16" bold="true" box="[151,372,614,640]" italics="true" pageId="131" pageNumber="132">Hydraena bifunda</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5BE6FF10FE6DF04BFDA5FD16" box="[387,531,614,640]" pageId="131" pageNumber="132" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785BE6FF10FF79F0A4FE44FD37" box="[151,498,649,673]" pageId="131" pageNumber="132" type="description">
<paragraph id="8BA636F35BE6FF10FF79F0A4FE44FD37" blockId="131.[151,531,614,673]" box="[151,498,649,673]" pageId="131" pageNumber="132">
(
<figureCitation id="13222A765BE6FF10FF71F0A4FEBDFD37" box="[159,267,649,673]" captionStart="FIGURE 243" captionStartId="327.[151,250,1753,1776]" captionTargetBox="[280,1344,239,1706]" captionTargetId="figure-25@327.[224,1363,193,1732]" captionTargetPageId="327" captionText="FIGURE 243. Hydraena bifunda, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283890" httpUri="https://zenodo.org/record/5283890/files/figure.png" pageId="131" pageNumber="132">Figs. 243</figureCitation>
,
<figureCitation id="13222A765BE6FF10FEF9F0A4FEF5FD37" box="[279,323,649,673]" captionStart="FIGURES 244–245" captionStartId="328.[151,264,1650,1673]" captionText="FIGURES 244–245. Aedeagi of holotypes. 244. H. fundextra. 245. H. bifunda." pageId="131" pageNumber="132">245</figureCitation>
,
<figureCitation id="13222A765BE6FF10FEA0F0A4FE05FD37" box="[334,435,649,673]" captionStart="FIGURES 382–387" captionStartId="368.[151,264,1650,1673]" captionTargetBox="[184,1402,196,1625]" captionTargetId="figure-28@368.[181,1405,193,1629]" captionTargetPageId="368" captionText="FIGURES 382–387. Scanning electron micrographs of H. bifunda (382–386) and H. thola (387)." figureDoi="http://doi.org/10.5281/zenodo.5284019" httpUri="https://zenodo.org/record/5284019/files/figure.png" pageId="131" pageNumber="132">382–386</figureCitation>
,
<figureCitation id="13222A765BE6FF10FE50F0A4FE5FFD37" box="[446,489,649,673]" captionStart="FIGURES 543–546" captionStartId="411.[151,264,1650,1673]" captionTargetBox="[165,1421,226,1610]" captionTargetId="figure-20@411.[165,1421,193,1629]" captionTargetPageId="411" captionText="FIGURES 543–546. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284143" httpUri="https://zenodo.org/record/5284143/files/figure.png" pageId="131" pageNumber="132">543</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785BE6FF10FF79F0E1FF5FFA96" pageId="131" pageNumber="132" type="materials_examined">
<paragraph id="8BA636F35BE6FF10FF79F0E1FF5FFA96" blockId="131.[151,1436,715,2036]" pageId="131" pageNumber="132">
Type Material.
<materialsCitation id="3B713CAE5BE6FF10FEA0F0E1FAFBFC91" collectingDate="1964-12-30" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="20" latitude="-6.6833334" location="Lae - Bulolo" longLatPrecision="1304" longitude="146.9" pageId="131" pageNumber="132" specimenCount="1" stateProvince="Morobe" typeStatus="holotype">
<typeStatus id="54A288515BE6FF10FEA0F0E1FE0DFD72" box="[334,443,716,740]" pageId="131" pageNumber="132" type="holotype">
<emphasis id="B96DEAE15BE6FF10FEA0F0E1FE0DFD72" bold="true" box="[334,443,716,740]" pageId="131" pageNumber="132">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15BE6FF10FDF1F0E1FD41FD72" bold="true" box="[543,759,715,740]" pageId="131" pageNumber="132">
<collectingRegion id="49DDF8115BE6FF10FDF1F0E1FD47FD72" box="[543,753,715,740]" country="Papua New Guinea" name="Morobe" pageId="131" pageNumber="132">Morobe Province</collectingRegion>
:
</emphasis>
c.
<quantity id="4CE19B165BE6FF10FCF0F0E1FCE1FD72" box="[798,855,716,740]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="131" pageNumber="132" unit="mi" value="7.0">7 mi.</quantity>
<location id="8EC660285BE6FF10FCB0F0E1FC57FD72" LSID="urn:lsid:plazi:treatment:03B087E55BE6FF17FF79F04BFB25FC82:8EC660285BE6FF10FCB0F0E1FC57FD72" box="[862,993,716,740]" latitude="-6.6833334" longLatPrecision="1304" longitude="146.9" name="Lae - Bulolo" pageId="131" pageNumber="132" stateProvince="Morobe">Lae–Bulolo</location>
road, ex gravel and grass roots on bank of small stream, [GE est.]
<quantity id="4CE19B165BE6FF10FE2BF0C2FD9DFC91" box="[453,555,751,775]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="131" pageNumber="132" unit="m" value="20.0" valueMax="30.0" valueMin="10.0">
<elevation id="0034D1C05BE6FF10FE2BF0C2FD9DFC91" box="[453,555,751,775]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="131" pageNumber="132" unit="m" value="20.0" valueMax="30.0" valueMin="10.0">10–30 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345BE6FF10FDD6F0C2FD22FC91" box="[568,660,751,775]" degrees="6" direction="south" minutes="41" orientation="latitude" pageId="131" pageNumber="132" precision="925" value="-6.6833334">6° 41' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345BE6FF10FD4CF0C2FCAAFC91" box="[674,796,751,775]" degrees="146" direction="east" minutes="54" orientation="longitude" pageId="131" pageNumber="132" precision="925" value="146.9">146° 54' E</geoCoordinate>
,
<date id="FFA710335BE6FF10FCC4F0C2FC19FC91" box="[810,943,751,775]" pageId="131" pageNumber="132" value="1964-12-30">
<collectingDate id="EFE3E9DB5BE6FF10FCC4F0C2FC19FC91" box="[810,943,751,775]" pageId="131" pageNumber="132" value="1964-12-30">30 xii 1964</collectingDate>
</date>
,
<collectorName id="26EC53255BE6FF10FC52F0DDFBD5FC91" box="[956,1123,751,775]" pageId="131" pageNumber="132">M. E. Bacchus</collectorName>
(MEB 120) (
<collectionCode id="ED08AE365BE6FF10FB10F0C2FAF3FC91" box="[1278,1349,751,775]" pageId="131" pageNumber="132">NHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CAE5BE6FF10FAB7F0DDFE75FCD9" collectingDate="1991-03-11" collectionCode="MCZ" collectorName="D. Larson" elevation="35" latitude="-4.9666667" location="5 km N Kawe" longLatPrecision="1306" longitude="145.78334" pageId="131" pageNumber="132" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<emphasis id="B96DEAE15BE6FF10FAB7F0DDFF63FCBA" bold="true" pageId="131" pageNumber="132">
<typeStatus id="54A288515BE6FF10FAB7F0DDFF63FCBA" pageId="131" pageNumber="132" type="paratype">Paratypes</typeStatus>
</emphasis>
(52):
<emphasis id="B96DEAE15BE6FF10FEC8F139FDB2FCBA" bold="true" box="[294,516,787,812]" pageId="131" pageNumber="132">
<collectingRegion id="49DDF8115BE6FF10FEC8F139FE49FCBA" box="[294,511,787,812]" country="Papua New Guinea" name="Madang" pageId="131" pageNumber="132">Madang Province</collectingRegion>
:
</emphasis>
<location id="8EC660285BE6FF10FDF8F139FD0CFCBD" LSID="urn:lsid:plazi:treatment:03B087E55BE6FF17FF79F04BFB25FC82:8EC660285BE6FF10FDF8F139FD0CFCBD" box="[534,698,788,812]" latitude="-4.9666667" longLatPrecision="1306" longitude="145.78334" name="5 km N Kawe" pageId="131" pageNumber="132" stateProvince="Madang">
<quantity id="4CE19B165BE6FF10FDF8F139FDE6FCBA" box="[534,592,788,812]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="131" pageNumber="132" unit="km" value="5.0">5 km</quantity>
N Kawe
</location>
, plantation drain, [GE est.]
<quantity id="4CE19B165BE6FF10FC17F139FBE9FCBD" box="[1017,1119,788,812]" metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" pageId="131" pageNumber="132" unit="m" value="35.0" valueMax="60.0" valueMin="10.0">
<elevation id="0034D1C05BE6FF10FC17F139FBE9FCBD" box="[1017,1119,788,812]" metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" pageId="131" pageNumber="132" unit="m" value="35.0" valueMax="60.0" valueMin="10.0">10–60 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345BE6FF10FB83F139FB7CFCBA" box="[1133,1226,788,812]" degrees="4" direction="south" minutes="58" orientation="latitude" pageId="131" pageNumber="132" precision="925" value="-4.9666667">4° 58' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345BE6FF10FB36F139FAE5FCBD" box="[1240,1363,788,812]" degrees="145" direction="east" minutes="47" orientation="longitude" pageId="131" pageNumber="132" precision="925" value="145.78334">145° 47' E</geoCoordinate>
,
<date id="FFA710335BE6FF10FA8FF139FF67FCD9" pageId="131" pageNumber="132" value="1991-03-11">
<collectingDate id="EFE3E9DB5BE6FF10FA8FF139FF67FCD9" pageId="131" pageNumber="132" value="1991-03-11">11 iii 1991</collectingDate>
</date>
,
<collectorName id="26EC53255BE6FF10FF31F115FEE7FCD9" box="[223,337,824,847]" pageId="131" pageNumber="132">D. Larson</collectorName>
(2
<collectionCode id="ED08AE365BE6FF10FE97F11AFE0CFCD9" box="[377,442,823,847]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="131" pageNumber="132" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5BE6FF10FE3EF11AFE0DFCE2" collectingDate="1991-04-10" collectionCode="MCZ" collectorName="D. Larson" elevation="400" latitude="-5.7" location="Gogal Valley" longLatPrecision="1305" longitude="146.53334" pageId="131" pageNumber="132" specimenCount="1" stateProvince="Madang Province" typeStatus="paratype">
<location id="8EC660285BE6FF10FE3EF11AFDD0FCD9" LSID="urn:lsid:plazi:treatment:03B087E55BE6FF17FF79F04BFB25FC82:8EC660285BE6FF10FE3EF11AFDD0FCD9" box="[464,614,823,847]" latitude="-5.7" longLatPrecision="1305" longitude="146.53334" name="Gogal Valley" pageId="131" pageNumber="132" stateProvince="Madang Province">Gogal Valley</location>
,
<locationDeviation id="9A142FD15BE6FF10FD9CF11AFCD6FCD9" box="[626,864,823,847]" location="Baru Village" pageId="131" pageNumber="132">
<quantity id="4CE19B165BE6FF10FD9CF11AFD1BFCD9" box="[626,685,823,847]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="131" pageNumber="132" unit="km" value="4.0">4 km</quantity>
S Baru Village
</locationDeviation>
, [GE est.]
<quantity id="4CE19B165BE6FF10FC33F11AFBD6FCD9" box="[989,1120,823,847]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="131" pageNumber="132" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">
<elevation id="0034D1C05BE6FF10FC33F11AFBD6FCD9" box="[989,1120,823,847]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="131" pageNumber="132" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">300–500 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345BE6FF10FB83F11AFB7CFCD9" box="[1133,1226,823,847]" degrees="5" direction="south" minutes="42" orientation="latitude" pageId="131" pageNumber="132" precision="925" value="-5.7">5° 42' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345BE6FF10FB37F11AFAE5FCD9" box="[1241,1363,823,847]" degrees="146" direction="east" minutes="32" orientation="longitude" pageId="131" pageNumber="132" precision="925" value="146.53334">146° 32' E</geoCoordinate>
,
<date id="FFA710335BE6FF10FA8FF11AFF67FCE2" pageId="131" pageNumber="132" value="1991-04-10">
<collectingDate id="EFE3E9DB5BE6FF10FA8FF11AFF67FCE2" pageId="131" pageNumber="132" value="1991-04-10">10 iv 1991</collectingDate>
</date>
,
<collectorName id="26EC53255BE6FF10FF32F171FEFAFCE5" box="[220,332,860,883]" pageId="131" pageNumber="132">D. Larson</collectorName>
(2
<collectionCode id="ED08AE365BE6FF10FE9FF171FE04FCE2" box="[369,434,860,884]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="131" pageNumber="132" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5BE6FF10FE28F171FEC2FC01" collectingDate="1991-04-18" collectionCode="MCZ" collectorName="D. Larson" elevation="140" latitude="-5.7166667" location="Ramu Valley" longLatPrecision="1305" longitude="145.36667" pageId="131" pageNumber="132" specimenCount="1" stateProvince="Madang Province" typeStatus="paratype">
<location id="8EC660285BE6FF10FE28F171FDEEFCE2" LSID="urn:lsid:plazi:treatment:03B087E55BE6FF17FF79F04BFB25FC82:8EC660285BE6FF10FE28F171FDEEFCE2" box="[454,600,860,884]" latitude="-5.7166667" longLatPrecision="1305" longitude="145.36667" name="Ramu Valley" pageId="131" pageNumber="132" stateProvince="Madang Province">Ramu Valley</location>
,
<locationDeviation id="9A142FD15BE6FF10FD8DF171FC85FCE2" box="[611,819,860,884]" location="Brahman" pageId="131" pageNumber="132">
<quantity id="4CE19B165BE6FF10FD8DF171FD2DFCE2" box="[611,667,860,884]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="131" pageNumber="132" unit="km" value="2.0">2 km</quantity>
SE Brahman
</locationDeviation>
, [GE est.]
<quantity id="4CE19B165BE6FF10FC44F171FB9CFCE5" box="[938,1066,860,884]" metricMagnitude="2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="131" pageNumber="132" unit="m" value="140.0" valueMax="150.0" valueMin="130.0">
<elevation id="0034D1C05BE6FF10FC44F171FB9CFCE5" box="[938,1066,860,884]" metricMagnitude="2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="131" pageNumber="132" unit="m" value="140.0" valueMax="150.0" valueMin="130.0">130–150 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345BE6FF10FBD8F171FB39FCE2" box="[1078,1167,860,884]" degrees="5" direction="south" minutes="43" orientation="latitude" pageId="131" pageNumber="132" precision="925" value="-5.7166667">5° 43' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345BE6FF10FB75F171FAA4FCE5" box="[1179,1298,860,884]" degrees="145" direction="east" minutes="22" orientation="longitude" pageId="131" pageNumber="132" precision="925" value="145.36667">145° 22' E</geoCoordinate>
,
<date id="FFA710335BE6FF10FAF3F171FA21FCE2" box="[1309,1431,860,884]" pageId="131" pageNumber="132" value="1991-04-18">
<collectingDate id="EFE3E9DB5BE6FF10FAF3F171FA21FCE2" box="[1309,1431,860,884]" pageId="131" pageNumber="132" value="1991-04-18">18 iv 1991</collectingDate>
</date>
,
<collectorName id="26EC53255BE6FF10FF79F1ADFEB0FC01" box="[151,262,896,919]" pageId="131" pageNumber="132">D. Larson</collectorName>
(1
<collectionCode id="ED08AE365BE6FF10FEC5F152FEDAFC01" box="[299,364,895,919]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="131" pageNumber="132" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5BE6FF10FE91F1ADFEE5FC2A" collectingDate="1991-04-15" collectionCode="MCZ" collectorName="D. Larson" elevation="500" latitude="-5.633333" location="Ramu Valley" longLatPrecision="1305" longitude="145.51666" pageId="131" pageNumber="132" specimenCount="1" stateProvince="Madang Province" typeStatus="paratype">
<location id="8EC660285BE6FF10FE91F1ADFDA7FC01" LSID="urn:lsid:plazi:treatment:03B087E55BE6FF17FF79F04BFB25FC82:8EC660285BE6FF10FE91F1ADFDA7FC01" box="[383,529,895,919]" latitude="-5.633333" longLatPrecision="1305" longitude="145.51666" name="Ramu Valley" pageId="131" pageNumber="132" stateProvince="Madang Province">Ramu Valley</location>
,
<locationDeviation id="9A142FD15BE6FF10FDF5F152FCA3FC01" box="[539,789,895,919]" location="Yagumbu" pageId="131" pageNumber="132">
<quantity id="4CE19B165BE6FF10FDF5F152FDC6FC01" box="[539,624,895,919]" metricMagnitude="3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="131" pageNumber="132" unit="km" value="4.5" valueMax="5.0" valueMin="4.0">4–5 km</quantity>
SW Yagumbu
</locationDeviation>
, [GE est.]
<quantity id="4CE19B165BE6FF10FC65F152FBBDFC01" box="[907,1035,895,919]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="131" pageNumber="132" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">
<elevation id="0034D1C05BE6FF10FC65F152FBBDFC01" box="[907,1035,895,919]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="131" pageNumber="132" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400–600 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345BE6FF10FBF8F152FBD9FC01" box="[1046,1135,895,919]" degrees="5" direction="south" minutes="38" orientation="latitude" pageId="131" pageNumber="132" precision="925" value="-5.633333">5° 38' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345BE6FF10FB95F152FB47FC01" box="[1147,1265,895,919]" degrees="145" direction="east" minutes="31" orientation="longitude" pageId="131" pageNumber="132" precision="925" value="145.51666">145° 31' E</geoCoordinate>
,
<date id="FFA710335BE6FF10FB12F152FAC3FC01" box="[1276,1397,895,919]" pageId="131" pageNumber="132" value="1991-04-15">
<collectingDate id="EFE3E9DB5BE6FF10FB12F152FAC3FC01" box="[1276,1397,895,919]" pageId="131" pageNumber="132" value="1991-04-15">15 iv 1991</collectingDate>
</date>
,
<collectorName id="26EC53255BE6FF10FA6FF1ADFF53FC2D" pageId="131" pageNumber="132">D. Larson</collectorName>
(3
<collectionCode id="ED08AE365BE6FF10FEE7F189FEFCFC2A" box="[265,330,932,956]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="131" pageNumber="132" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5BE6FF10FEB0F189FE84FC49" collectingDate="1991-04-15" collectionCode="MCZ" collectorName="D. Larson" elevation="600" latitude="-5.65" location="Ramu Valley" longLatPrecision="1305" longitude="145.53334" pageId="131" pageNumber="132" specimenCount="1" stateProvince="Madang Province" typeStatus="paratype">
<location id="8EC660285BE6FF10FEB0F189FE46FC2A" LSID="urn:lsid:plazi:treatment:03B087E55BE6FF17FF79F04BFB25FC82:8EC660285BE6FF10FEB0F189FE46FC2A" box="[350,496,932,956]" latitude="-5.65" longLatPrecision="1305" longitude="145.53334" name="Ramu Valley" pageId="131" pageNumber="132" stateProvince="Madang Province">Ramu Valley</location>
,
<locationDeviation id="9A142FD15BE6FF10FE15F189FD6FFC2A" box="[507,729,932,956]" location="Yagumbu" pageId="131" pageNumber="132">
<quantity id="4CE19B165BE6FF10FE15F189FD82FC2A" box="[507,564,932,956]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="131" pageNumber="132" unit="km" value="6.0">6 km</quantity>
SW Yagumbu
</locationDeviation>
, [GE est.]
<quantity id="4CE19B165BE6FF10FCBEF189FC67FC2D" box="[848,977,932,956]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="131" pageNumber="132" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">
<elevation id="0034D1C05BE6FF10FCBEF189FC67FC2D" box="[848,977,932,956]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="131" pageNumber="132" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500–700 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345BE6FF10FC33F189FB83FC2A" box="[989,1077,932,956]" degrees="5" direction="south" minutes="39" orientation="latitude" pageId="131" pageNumber="132" precision="925" value="-5.65">5° 39' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345BE6FF10FBACF189FB0FFC2D" box="[1090,1209,932,956]" degrees="145" direction="east" minutes="32" orientation="longitude" pageId="131" pageNumber="132" precision="925" value="145.53334">145° 32' E</geoCoordinate>
,
<date id="FFA710335BE6FF10FB2BF189FA88FC2A" box="[1221,1342,932,956]" pageId="131" pageNumber="132" value="1991-04-15">
<collectingDate id="EFE3E9DB5BE6FF10FB2BF189FA88FC2A" box="[1221,1342,932,956]" pageId="131" pageNumber="132" value="1991-04-15">15 iv 1991</collectingDate>
</date>
,
<collectorName id="26EC53255BE6FF10FAA4F189FF08FC49" pageId="131" pageNumber="132">D. Larson</collectorName>
(3
<collectionCode id="ED08AE365BE6FF10FF06F1EAFE9FFC49" box="[232,297,967,991]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="131" pageNumber="132" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5BE6FF10FED1F1E5FC1EFC49" box="[319,936,967,992]" collectingDate="1964-12-30" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="20" latitude="-6.6833334" location="Same" longLatPrecision="1304" longitude="146.9" pageId="131" pageNumber="132" specimenCount="1" stateProvince="Morobe" typeStatus="paratype">
<emphasis id="B96DEAE15BE6FF10FED1F1E5FDACFC76" bold="true" box="[319,538,967,992]" pageId="131" pageNumber="132">
<collectingRegion id="49DDF8115BE6FF10FED1F1E5FDA3FC76" box="[319,533,967,992]" country="Papua New Guinea" name="Morobe" pageId="131" pageNumber="132">Morobe Province</collectingRegion>
:
</emphasis>
<location id="8EC660285BE6FF10FDC3F1EAFDDDFC49" LSID="urn:lsid:plazi:treatment:03B087E55BE6FF17FF79F04BFB25FC82:8EC660285BE6FF10FDC3F1EAFDDDFC49" box="[557,619,967,991]" latitude="-6.6833334" longLatPrecision="1304" longitude="146.9" name="Same" pageId="131" pageNumber="132" stateProvince="Morobe">Same</location>
data as holotype (6
<collectionCode id="ED08AE365BE6FF10FCB4F1EAFC16FC49" box="[858,928,967,991]" pageId="131" pageNumber="132">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5BE6FF10FC5BF1E5FEFBFBBE" collectingDate="1964-12-30" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="20" latitude="-6.6833334" location="Bulolo" longLatPrecision="1304" longitude="146.9" pageId="131" pageNumber="132" specimenCount="1" stateProvince="Morobe Province" typeStatus="paratype">
c.
<locationDeviation id="9A142FD15BE6FF10FC3DF1E5FBECFC49" box="[979,1114,967,991]" location="Lae" pageId="131" pageNumber="132">
<quantity id="4CE19B165BE6FF10FC3DF1E5FB91FC49" box="[979,1063,967,991]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="131" pageNumber="132" unit="mi" value="7.0">7 miles</quantity>
Lae
</locationDeviation>
–
<location id="8EC660285BE6FF10FB87F1EAFB02FC49" LSID="urn:lsid:plazi:treatment:03B087E55BE6FF17FF79F04BFB25FC82:8EC660285BE6FF10FB87F1EAFB02FC49" box="[1129,1204,967,991]" latitude="-6.6833334" longLatPrecision="1304" longitude="146.9" name="Bulolo" pageId="131" pageNumber="132" stateProvince="Morobe Province">Bulolo</location>
road, ex gravel and grass roots on bank of small stream, [GE est.]
<quantity id="4CE19B165BE6FF10FD42F1C1FCA5FB95" box="[684,787,1004,1028]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="131" pageNumber="132" unit="m" value="20.0" valueMax="30.0" valueMin="10.0">
<elevation id="0034D1C05BE6FF10FD42F1C1FCA5FB95" box="[684,787,1004,1028]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="131" pageNumber="132" unit="m" value="20.0" valueMax="30.0" valueMin="10.0">10–30 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345BE6FF10FCCEF1C1FCCBFB92" box="[800,893,1004,1028]" degrees="6" direction="south" minutes="41" orientation="latitude" pageId="131" pageNumber="132" precision="925" value="-6.6833334">6° 41' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345BE6FF10FC65F1C1FBB3FB95" box="[907,1029,1004,1028]" degrees="146" direction="east" minutes="54" orientation="longitude" pageId="131" pageNumber="132" precision="925" value="146.9">146° 54' E</geoCoordinate>
,
<date id="FFA710335BE6FF10FBFDF1C1FB2EFB92" box="[1043,1176,1004,1028]" pageId="131" pageNumber="132" value="1964-12-30">
<collectingDate id="EFE3E9DB5BE6FF10FBFDF1C1FB2EFB92" box="[1043,1176,1004,1028]" pageId="131" pageNumber="132" value="1964-12-30">30 xii 1964</collectingDate>
</date>
,
<collectorName id="26EC53255BE6FF10FB4BF1C1FAFBFB92" box="[1189,1357,1004,1028]" pageId="131" pageNumber="132">M. E. Bacchus</collectorName>
(MEB 120) (18
<collectionCode id="ED08AE365BE6FF10FF11F63DFEF3FBBE" box="[255,325,1040,1064]" pageId="131" pageNumber="132">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5BE6FF10FEB7F63DFD2BFBDA" collectingDate="2006-03-31" collectionCode="MCZ, NHM, NMW, ZSM" collectorName="Balke &amp; Sagata" country="Papua New Guinea" elevation="250" latitude="-6.5045" location="Kwapsanek" longLatPrecision="9" longitude="146.99301" municipality="Huon Pen." pageId="131" pageNumber="132" specimenCount="1" stateProvince="Morobe Province" typeStatus="paratype">
<collectingMunicipality id="6BC2AC895BE6FF10FEB7F63DFE65FBB1" box="[345,467,1040,1063]" pageId="131" pageNumber="132">Huon Pen.</collectingMunicipality>
, road to
<location id="8EC660285BE6FF10FDD8F63DFD0AFBBE" LSID="urn:lsid:plazi:treatment:03B087E55BE6FF17FF79F04BFB25FC82:8EC660285BE6FF10FDD8F63DFD0AFBBE" box="[566,700,1040,1064]" country="Papua New Guinea" latitude="-6.5045" longLatPrecision="9" longitude="146.99301" municipality="Huon Pen." name="Kwapsanek" pageId="131" pageNumber="132" stateProvince="Morobe Province">Kwapsanek</location>
,
<quantity id="4CE19B165BE6FF10FD26F63DFCA4FBB1" box="[712,786,1040,1064]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="131" pageNumber="132" unit="m" value="250.0">
<elevation id="0034D1C05BE6FF10FD26F63DFCA4FBB1" box="[712,786,1040,1064]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="131" pageNumber="132" unit="m" value="250.0">250 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345BE6FF10FCF1F63DFC2AFBBE" box="[799,924,1040,1064]" degrees="6" direction="south" minutes="30.27" orientation="latitude" pageId="131" pageNumber="132" precision="9" value="-6.5045">6° 30.27' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345BE6FF10FC44F63DFBE2FBB1" box="[938,1108,1040,1064]" degrees="146" direction="east" minutes="59.581" orientation="longitude" pageId="131" pageNumber="132" precision="1" value="146.99301">146° 59.581' E</geoCoordinate>
,
<date id="FFA710335BE6FF10FB8FF63DFB69FBBE" box="[1121,1247,1040,1064]" pageId="131" pageNumber="132" value="2006-03-31">
<collectingDate id="EFE3E9DB5BE6FF10FB8FF63DFB69FBBE" box="[1121,1247,1040,1064]" pageId="131" pageNumber="132" value="2006-03-31">31 iii 2006</collectingDate>
</date>
,
<collectorName id="26EC53255BE6FF10FB05F63DFA9AFBBE" box="[1259,1324,1040,1064]" pageId="131" pageNumber="132">Balke</collectorName>
&amp;
<collectorName id="26EC53255BE6FF10FABFF63DFA2DFBBE" box="[1361,1435,1040,1064]" pageId="131" pageNumber="132">Sagata</collectorName>
(
<collectingCountry id="F30E76635BE6FF10FF70F619FF6EFBDA" box="[158,216,1076,1100]" name="Papua New Guinea" pageId="131" pageNumber="132">PNG</collectingCountry>
24) (7
<collectionCode id="ED08AE365BE6FF10FEC7F619FEDEFBDA" box="[297,360,1076,1100]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="131" pageNumber="132" type="Museum">MCZ</collectionCode>
,
<collectionCode id="ED08AE365BE6FF10FE9DF619FE01FBDD" box="[371,439,1076,1099]" pageId="131" pageNumber="132">NHM</collectionCode>
,
<collectionCode id="ED08AE365BE6FF10FE2CF619FDBCFBDD" box="[450,522,1076,1099]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="131" pageNumber="132">NMW</collectionCode>
,
<collectingCountry id="F30E76635BE6FF10FDFBF619FDFBFBDA" box="[533,589,1076,1100]" name="Papua New Guinea" pageId="131" pageNumber="132">PNG</collectingCountry>
,
<collectionCode id="ED08AE365BE6FF10FDB9F619FD23FBDA" box="[599,661,1076,1100]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="131" pageNumber="132" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5BE6FF10FD47F619FBE2FBE6" collectingDate="1964-12-12" collectionCode="NHM" collectorName="M. E. Bacchus" country="Papua New Guinea" elevation="25" latitude="-6.7166667" location="Botanic Garden" longLatPrecision="1304" longitude="147.0" municipality="Lae" pageId="131" pageNumber="132" specimenCount="1" stateProvince="Morobe Province" typeStatus="paratype">
<collectingMunicipality id="6BC2AC895BE6FF10FD47F619FD63FBDD" box="[681,725,1076,1099]" pageId="131" pageNumber="132">Lae</collectingMunicipality>
, ex muddy gravel bank of small stream in
<location id="8EC660285BE6FF10FB41F619FAD4FBDA" LSID="urn:lsid:plazi:treatment:03B087E55BE6FF17FF79F04BFB25FC82:8EC660285BE6FF10FB41F619FAD4FBDA" box="[1199,1378,1076,1100]" country="Papua New Guinea" latitude="-6.7166667" longLatPrecision="1304" longitude="147.0" municipality="Lae" name="Botanic Garden" pageId="131" pageNumber="132" stateProvince="Morobe Province">Botanic Garden</location>
, [GE est.]
<quantity id="4CE19B165BE6FF10FF20F675FE82FBF9" box="[206,308,1112,1136]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="131" pageNumber="132" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">
<elevation id="0034D1C05BE6FF10FF20F675FE82FBF9" box="[206,308,1112,1136]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="131" pageNumber="132" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">20–30 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345BE6FF10FEAEF675FE2DFBE6" box="[320,411,1112,1136]" degrees="6" direction="south" minutes="43" orientation="latitude" pageId="131" pageNumber="132" precision="925" value="-6.7166667">6° 43' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345BE6FF10FE47F675FDA2FBF9" box="[425,532,1112,1136]" degrees="147" direction="east" minutes="0" orientation="longitude" pageId="131" pageNumber="132" precision="925" value="147.0">147° 0' E</geoCoordinate>
,
<date id="FFA710335BE6FF10FDCEF675FD12FBE6" box="[544,676,1112,1136]" pageId="131" pageNumber="132" value="1964-12-12">
<collectingDate id="EFE3E9DB5BE6FF10FDCEF675FD12FBE6" box="[544,676,1112,1136]" pageId="131" pageNumber="132" value="1964-12-12">12 xii 1964</collectingDate>
</date>
,
<collectorName id="26EC53255BE6FF10FD5EF675FCE1FBE6" box="[688,855,1112,1136]" pageId="131" pageNumber="132">M. E. Bacchus</collectorName>
(MEB 118) (2
<collectionCode id="ED08AE365BE6FF10FBE8F675FBFAFBE6" box="[1030,1100,1112,1136]" pageId="131" pageNumber="132">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5BE6FF10FB8EF675FE73FB2E" collectingDate="1965-01-27" collectingDateMax="1965-01-30" collectingDateMin="1965-01-27" collectionCode="NHM" collectorName="M. E. Bacchus" country="Papua New Guinea" elevation="305" latitude="-5.9833336" location="Gusap" longLatPrecision="1305" longitude="146.08333" municipality="Markham Valley" pageId="131" pageNumber="132" specimenCount="1" stateProvince="Morobe Province" typeStatus="paratype">
<collectingMunicipality id="6BC2AC895BE6FF10FB8EF675FAABFBE6" box="[1120,1309,1112,1136]" pageId="131" pageNumber="132">Markham Valley</collectingMunicipality>
,
<location id="8EC660285BE6FF10FAC7F675FAC5FBE6" LSID="urn:lsid:plazi:treatment:03B087E55BE6FF17FF79F04BFB25FC82:8EC660285BE6FF10FAC7F675FAC5FBE6" box="[1321,1395,1112,1136]" country="Papua New Guinea" latitude="-5.9833336" longLatPrecision="1305" longitude="146.08333" municipality="Markham Valley" name="Gusap" pageId="131" pageNumber="132" stateProvince="Morobe Province">Gusap</location>
,
<locationDeviation id="9A142FD15BE6FF10FA6EF675FEEAFB05" location="Lae" pageId="131" pageNumber="132">
<quantity id="4CE19B165BE6FF10FA6EF675FF62FB02" metricMagnitude="5" metricUnit="m" metricValue="1.4484096" pageId="131" pageNumber="132" unit="mi" value="90.0">90 miles</quantity>
NW of Lae
</locationDeviation>
, ex dead leaves on mud wetted by seepage,
<quantity id="4CE19B165BE6FF10FCA5F651FC23FB05" box="[843,917,1148,1172]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="131" pageNumber="132" unit="m" value="305.0">
<elevation id="0034D1C05BE6FF10FCA5F651FC23FB05" box="[843,917,1148,1172]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="131" pageNumber="132" unit="m" value="305.0">305 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345BE6FF10FC4FF651FC4DFB02" box="[929,1019,1148,1172]" degrees="5" direction="south" minutes="59" orientation="latitude" pageId="131" pageNumber="132" precision="925" value="-5.9833336">5° 59' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345BE6FF10FBE9F651FBC4FB05" box="[1031,1138,1148,1172]" degrees="146" direction="east" minutes="5" orientation="longitude" pageId="131" pageNumber="132" precision="925" value="146.08333">146° 5' E</geoCoordinate>
,
<date id="FFA710335BE6FF10FB90F651FAA2FB02" box="[1150,1300,1148,1172]" pageId="131" pageNumber="132" value="1965-01-27" valueMax="1965-01-30" valueMin="1965-01-27">
<collectingDate id="EFE3E9DB5BE6FF10FB90F651FAA2FB02" box="[1150,1300,1148,1172]" pageId="131" pageNumber="132" value="1965-01-27" valueMax="1965-01-30" valueMin="1965-01-27">27–30 i 1965</collectingDate>
</date>
,
<collectorName id="26EC53255BE6FF10FACEF651FF7CFB2E" pageId="131" pageNumber="132">M. E. Bacchus</collectorName>
(MEB 163) (5
<collectionCode id="ED08AE365BE6FF10FE99F68DFE0BFB2E" box="[375,445,1184,1208]" pageId="131" pageNumber="132">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5BE6FF10FE3FF68DFF53FA96" collectingDate="1965-01-27" collectingDateMax="1965-01-30" collectingDateMin="1965-01-27" collectionCode="NHM" collectorName="M. E. Bacchus" country="Papua New Guinea" elevation="305" latitude="-5.9833336" location="Gusap" longLatPrecision="1305" longitude="146.08333" municipality="Markham Valley" pageId="131" pageNumber="132" specimenCount="1" stateProvince="Morobe Province" typeStatus="paratype">
<collectingMunicipality id="6BC2AC895BE6FF10FE3FF68DFD3AFB2E" box="[465,652,1184,1208]" pageId="131" pageNumber="132">Markham Valley</collectingMunicipality>
,
<location id="8EC660285BE6FF10FD76F68DFD54FB2E" LSID="urn:lsid:plazi:treatment:03B087E55BE6FF17FF79F04BFB25FC82:8EC660285BE6FF10FD76F68DFD54FB2E" box="[664,738,1184,1208]" country="Papua New Guinea" latitude="-5.9833336" longLatPrecision="1305" longitude="146.08333" municipality="Markham Valley" name="Gusap" pageId="131" pageNumber="132" stateProvince="Morobe Province">Gusap</location>
, c.
<locationDeviation id="9A142FD15BE6FF10FCE6F68DFC46FB21" box="[776,1008,1184,1208]" location="Lae" pageId="131" pageNumber="132">
<quantity id="4CE19B165BE6FF10FCE6F68DFCDEFB2E" box="[776,872,1184,1208]" metricMagnitude="5" metricUnit="m" metricValue="1.4484096" pageId="131" pageNumber="132" unit="mi" value="90.0">90 miles</quantity>
NW of Lae
</locationDeviation>
, large pond, muddy bottom with thick layer of dead leaves, dense
<taxonomicName id="4C194D705BE6FF10FE24F6E9FDC4FB4A" box="[458,626,1220,1244]" class="Magnoliopsida" family="Ceratophyllaceae" genus="Ceratophyllum" kingdom="Plantae" order="Ceratophyllales" pageId="131" pageNumber="132" phylum="Tracheophyta" rank="genus">
<emphasis id="B96DEAE15BE6FF10FE24F6E9FDC4FB4A" box="[458,626,1220,1244]" italics="true" pageId="131" pageNumber="132">Ceratophyllum</emphasis>
</taxonomicName>
,
<quantity id="4CE19B165BE6FF10FD6EF6E9FD7FFB4D" box="[640,713,1220,1244]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="131" pageNumber="132" unit="m" value="305.0">
<elevation id="0034D1C05BE6FF10FD6EF6E9FD7FFB4D" box="[640,713,1220,1244]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="131" pageNumber="132" unit="m" value="305.0">305 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345BE6FF10FD3BF6E9FC98FB4A" box="[725,814,1220,1244]" degrees="5" direction="south" minutes="59" orientation="latitude" pageId="131" pageNumber="132" precision="925" value="-5.9833336">5° 59' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345BE6FF10FCD5F6E9FC12FB4D" box="[827,932,1220,1244]" degrees="146" direction="east" minutes="5" orientation="longitude" pageId="131" pageNumber="132" precision="925" value="146.08333">146° 5' E</geoCoordinate>
,
<date id="FFA710335BE6FF10FC5EF6E9FBF3FB4A" box="[944,1093,1220,1244]" pageId="131" pageNumber="132" value="1965-01-27" valueMax="1965-01-30" valueMin="1965-01-27">
<collectingDate id="EFE3E9DB5BE6FF10FC5EF6E9FBF3FB4A" box="[944,1093,1220,1244]" pageId="131" pageNumber="132" value="1965-01-27" valueMax="1965-01-30" valueMin="1965-01-27">27–30 i 1965</collectingDate>
</date>
,
<collectorName id="26EC53255BE6FF10FBBFF6E9FB40FB4A" box="[1105,1270,1220,1244]" pageId="131" pageNumber="132">M. E. Bacchus</collectorName>
(MEB 162) (3
<collectionCode id="ED08AE365BE6FF10FF79F6C5FF6BFA96" box="[151,221,1256,1280]" pageId="131" pageNumber="132">NHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785BE6FF10FF29F726FD1BF9B6" pageId="131" pageNumber="132" type="diagnosis">
<paragraph id="8BA636F35BE6FF10FF29F726FD1BF9B6" blockId="131.[151,1436,715,2036]" pageId="131" pageNumber="132">
<emphasis id="B96DEAE15BE6FF10FF29F726FE64FAB2" bold="true" box="[199,466,1291,1316]" pageId="131" pageNumber="132">Differential Diagnosis.</emphasis>
Among members of the
<emphasis id="B96DEAE15BE6FF10FD05F721FCE9FAB2" box="[747,863,1292,1316]" italics="true" pageId="131" pageNumber="132">Fundextra</emphasis>
group similar to
<taxonomicName id="4C194D705BE6FF10FBCCF720FBCBFAB2" authorityName="Perkins" authorityYear="2011" box="[1058,1149,1292,1316]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="thola">
<emphasis id="B96DEAE15BE6FF10FBCCF720FB80FAB2" box="[1058,1078,1293,1316]" italics="true" pageId="131" pageNumber="132">H</emphasis>
.
<emphasis id="B96DEAE15BE6FF10FBAAF721FBCBFAB2" box="[1092,1149,1292,1316]" italics="true" pageId="131" pageNumber="132">thola</emphasis>
</taxonomicName>
in the plaque ratios, having P2 slightly wider than the plaque separation (ratios ca. 5/1/6/4 and 5/1/8/4 respectively) (
<figureCitation id="13222A765BE6FF10FB0CF71DFAE4FADE" box="[1250,1362,1328,1352]" captionStart="FIGURE 243" captionStartId="327.[151,250,1753,1776]" captionTargetBox="[280,1344,239,1706]" captionTargetId="figure-25@327.[224,1363,193,1732]" captionTargetPageId="327" captionText="FIGURE 243. Hydraena bifunda, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283890" httpUri="https://zenodo.org/record/5283890/files/figure.png" pageId="131" pageNumber="132">Figs. 243</figureCitation>
,
<figureCitation id="13222A765BE6FF10FA8CF71DFA3BFADE" box="[1378,1421,1328,1352]" captionStart="FIGURE 247" captionStartId="330.[151,250,1739,1762]" captionTargetBox="[201,1382,205,1708]" captionTargetId="figure-29@330.[186,1401,193,1718]" captionTargetPageId="330" captionText="FIGURE 247. Hydraena thola, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283896" httpUri="https://zenodo.org/record/5283896/files/figure.png" pageId="131" pageNumber="132">247</figureCitation>
).
<taxonomicName id="4C194D705BE6FF10FF79F779FED3FAFA" authorityName="Perkins" authorityYear="2011" box="[151,357,1364,1388]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="bifunda">
<emphasis id="B96DEAE15BE6FF10FF79F779FED3FAFA" box="[151,357,1364,1388]" italics="true" pageId="131" pageNumber="132">Hydraena bifunda</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C194D705BE6FF10FE12F778FDEEFAFA" authorityName="Perkins" authorityYear="2011" box="[508,600,1364,1388]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="thola">
<emphasis id="B96DEAE15BE6FF10FE12F778FDA6FAFA" box="[508,528,1365,1388]" italics="true" pageId="131" pageNumber="132">H</emphasis>
.
<emphasis id="B96DEAE15BE6FF10FDF1F779FDEEFAFA" box="[543,600,1364,1388]" italics="true" pageId="131" pageNumber="132">thola</emphasis>
</taxonomicName>
and all other members of the
<emphasis id="B96DEAE15BE6FF10FC41F779FB95FAFA" box="[943,1059,1364,1388]" italics="true" pageId="131" pageNumber="132">Fundextra</emphasis>
group in the degree to which the posterior part of P1 is raised.
<taxonomicName id="4C194D705BE6FF10FE1CF755FD74FA06" authorityName="Perkins" authorityYear="2011" box="[498,706,1400,1424]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="bifunda">
<emphasis id="B96DEAE15BE6FF10FE1CF755FD74FA06" box="[498,706,1400,1424]" italics="true" pageId="131" pageNumber="132">Hydraena bifunda</emphasis>
</taxonomicName>
differs dorsally from
<taxonomicName id="4C194D705BE6FF10FC2CF754FB96FA06" authorityName="Perkins" authorityYear="2011" box="[962,1056,1400,1424]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="thola">
<emphasis id="B96DEAE15BE6FF10FC2CF754FC60FA06" box="[962,982,1401,1424]" italics="true" pageId="131" pageNumber="132">H</emphasis>
.
<emphasis id="B96DEAE15BE6FF10FC09F755FB96FA06" box="[999,1056,1400,1424]" italics="true" pageId="131" pageNumber="132">thola</emphasis>
</taxonomicName>
in the following characters: the frons is only weakly microreticulate, the pronotal punctures are only slightly larger than the frons punctures, the elytra have a few subserial discal punctures, and the pronotum has a fascia. The male genitalia of
<taxonomicName id="4C194D705BE6FF10FB1FF7ECFADCFA4E" authorityName="Perkins" authorityYear="2011" box="[1265,1386,1472,1496]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="bifunda">
<emphasis id="B96DEAE15BE6FF10FB1FF7ECFAB3FA4E" box="[1265,1285,1473,1496]" italics="true" pageId="131" pageNumber="132">H</emphasis>
.
<emphasis id="B96DEAE15BE6FF10FAFAF7EDFADCFA4E" box="[1300,1386,1472,1496]" italics="true" pageId="131" pageNumber="132">bifunda</emphasis>
</taxonomicName>
distinctively differ from those of other members of the group (
<figureCitation id="13222A765BE6FF10FCC4F7C9FC3AFA6A" box="[810,908,1508,1532]" captionStart="FIGURES 244–245" captionStartId="328.[151,264,1650,1673]" captionText="FIGURES 244–245. Aedeagi of holotypes. 244. H. fundextra. 245. H. bifunda." pageId="131" pageNumber="132">Fig. 245</figureCitation>
). The metaventrite and complicated leg chaetotaxy are illustrated with SEMs (
<figureCitation id="13222A765BE6FF10FE14F425FD17F9B6" box="[506,673,1544,1568]" captionStart="FIGURES 382–387" captionStartId="368.[151,264,1650,1673]" captionTargetBox="[184,1402,196,1625]" captionTargetId="figure-28@368.[181,1405,193,1629]" captionTargetPageId="368" captionText="FIGURES 382–387. Scanning electron micrographs of H. bifunda (382–386) and H. thola (387)." figureDoi="http://doi.org/10.5281/zenodo.5284019" httpUri="https://zenodo.org/record/5284019/files/figure.png" pageId="131" pageNumber="132">Figs. 382–386</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785BE6FF17FF29F406FDE2FD12" lastPageId="132" lastPageNumber="133" pageId="131" pageNumber="132" type="description">
<paragraph id="8BA636F35BE6FF10FF29F406FB2CF926" blockId="131.[151,1436,715,2036]" pageId="131" pageNumber="132">
<emphasis id="B96DEAE15BE6FF10FF29F406FEEEF9D2" bold="true" box="[199,344,1579,1604]" inLineHeading="true" pageId="131" pageNumber="132" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515BE6FF10FE42F401FDB8F9D2" box="[428,526,1580,1604]" pageId="131" pageNumber="132" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.33/0.60; head 0.22/0.33; pronotum 0.33/0.47, PA 0.37, PB 0.44; elytra 0.81/0.60. Dorsum of head dark brown; pronotum light brown to testaceous around brown, diffusely margined, fascia, ratios of color bands, as measured in midline, ca. 6/10/5; elytra brown; legs light brown; maxillary palpi testaceous, tip not darker. Dorsum weakly shining.
</paragraph>
<paragraph id="8BA636F35BE6FF10FF29F491FEDDF862" blockId="131.[151,1436,715,2036]" pageId="131" pageNumber="132">Frons punctures ca. 1xef, slightly smaller and sparser medially; interstices effacedly microreticulate, weakly shining, 0.5–2xpd. Clypeus effacedly microreticulate, very finely sparsely punctulate medially. Mentum very sparsely very finely punctulate, shining. Postmentum effacedly microreticulate in shallow median depression, otherwise finely sparsely punctulate. Genae very slightly raised, shining, with shallow, obsolete median impression, without posterior ridge. Pronotum transverse, ca. median 3/4 of anterior margin slightly arcuate to posterior, sides arcuate, slightly emarginate between midlength and posterior angle; punctures on disc ca. 1xpd of largest frons punctures, interstices strongly shining, ca. 2–3xpd, punctures posteriorly slightly larger than those on disc; PF1 and PF4 absent; PF2 very shallow, completely shallowly confluent in midline to form U-shaped impression; PF3 moderately deep, wide.</paragraph>
<paragraph id="8BA636F35BE1FF17FF29F2BAFEEFFEA9" blockId="132.[151,1437,151,788]" pageId="132" pageNumber="133">Elytra with posterior declivity very gradual, summit at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming angle with one another.</paragraph>
<paragraph id="8BA636F35BE1FF17FF29F361FDE2FD12" blockId="132.[151,1437,151,788]" pageId="132" pageNumber="133">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 5/1/6/4. P1 very slightly wider than laminate; in profile, median carina straight in front of coxae, behind coxae carina raised, arcuate in profile. P2 markedly concave, length/width ca. 6/5, sides parallel or very slightly converging toward blunt apex. Plaques carinate lines, converging slightly anteriorly, at sides of densely punctate median depression. Metaventrite between mesocoxae slightly sloping, confluent with tip of P2. AIS width at straight posterior margin ca. equal P2. All legs of moderate length. Profemur (male) with minute tubercle on ventral margin near basal 1/3; protibia very slightly arcuate, gradually increasing in width from base to apex, ventral margin with minute carina near midlength. Meso- and metatibia straight, slender. Abdominal apex asymmetrical; last tergite (male) with deep concavity near apex, offset to left side. Aedeagus as illustrated (
<figureCitation id="13222A765BE1FF17FE0BF041FDF1FD12" box="[485,583,620,644]" captionStart="FIGURES 244–245" captionStartId="328.[151,264,1650,1673]" captionText="FIGURES 244–245. Aedeagi of holotypes. 244. H. fundextra. 245. H. bifunda." pageId="132" pageNumber="133">Fig. 245</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785BE1FF17FF29F0BDFA36FD31" box="[199,1408,655,680]" pageId="132" pageNumber="133" type="etymology">
<paragraph id="8BA636F35BE1FF17FF29F0BDFA36FD31" blockId="132.[151,1437,151,788]" box="[199,1408,655,680]" pageId="132" pageNumber="133">
<emphasis id="B96DEAE15BE1FF17FF29F0BDFEFBFD3E" bold="true" box="[199,333,656,680]" inLineHeading="true" pageId="132" pageNumber="133" reason="1">Etymology.</emphasis>
Named in reference to the paired processes of the aedeagus, and to the relationship to
<taxonomicName id="4C194D705BE1FF17FAF9F0BCFACFFD3E" authorityName="Perkins" authorityYear="2011" box="[1303,1401,656,680]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="funda">
<emphasis id="B96DEAE15BE1FF17FAF9F0BCFA9DFD3E" box="[1303,1323,657,680]" italics="true" pageId="132" pageNumber="133">H</emphasis>
.
<emphasis id="B96DEAE15BE1FF17FAD7F0BDFACFFD3E" box="[1337,1401,656,680]" italics="true" pageId="132" pageNumber="133">funda</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785BE1FF17FF29F09EFB25FC82" pageId="132" pageNumber="133" type="distribution">
<paragraph id="8BA636F35BE1FF17FF29F09EFD25FD79" blockId="132.[151,1437,151,788]" pageId="132" pageNumber="133">
<emphasis id="B96DEAE15BE1FF17FF29F09EFED6FD5A" bold="true" box="[199,352,691,716]" inLineHeading="true" pageId="132" pageNumber="133" reason="1">Distribution.</emphasis>
Currently known from the eastern parts of Areas 2 and 3, and both eastern and western parts of Area 5; elevation range
<quantity id="4CE19B165BE1FF17FE4AF0FAFDA3FD79" box="[420,533,727,751]" metricMagnitude="2" metricUnit="m" metricValue="3.55" metricValueMax="7.0" metricValueMin="0.1" pageId="132" pageNumber="133" unit="m" value="355.0" valueMax="700.0" valueMin="10.0">10–700 m</quantity>
(
<figureCitation id="13222A765BE1FF17FDCAF0FAFD30FD79" box="[548,646,727,751]" captionStart="FIGURES 543–546" captionStartId="411.[151,264,1650,1673]" captionTargetBox="[165,1421,226,1610]" captionTargetId="figure-20@411.[165,1421,193,1629]" captionTargetPageId="411" captionText="FIGURES 543–546. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284143" httpUri="https://zenodo.org/record/5284143/files/figure.png" pageId="132" pageNumber="133">Fig. 543</figureCitation>
).
</paragraph>
<paragraph id="8BA636F35BE1FF17FF29F0D1FB25FC82" blockId="132.[151,1437,151,788]" box="[199,1171,764,788]" pageId="132" pageNumber="133">
<emphasis id="B96DEAE15BE1FF17FF29F0D1FEB0FC82" bold="true" box="[199,262,764,788]" pageId="132" pageNumber="133">Note.</emphasis>
The
<typeStatus id="54A288515BE1FF17FED1F0D1FE17FC82" box="[319,417,764,788]" pageId="132" pageNumber="133" type="holotype">holotype</typeStatus>
is slightly teneral; mature specimens have a darker pronotal fascia.
</paragraph>
</subSubSection>
</treatment>
</document>
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