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<document id="EFC572849657F47C65C246B53088724F" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B3FFFC9FF79F2BAFEEAFA74" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena perpunctata Perkins 2011, new species" docType="treatment" docVersion="6" lastPageNumber="91" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="91" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E11B473AE1AAC13E8EE797EDA5AE354C">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="B7C6B295BAC89BE05C1DE7E6526EBAF6">Perkins, Philip D.</mods:namePart>
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<mods:date id="3ED23E3AA92A43943A4F940EA5E709F4">2011</mods:date>
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<treatment id="03B087E55B3FFFC9FF79F2BAFEEAFA74" ID-DOI="http://doi.org/10.5281/zenodo.5291654" ID-Zenodo-Dep="5291654" LSID="urn:lsid:plazi:treatment:03B087E55B3FFFC9FF79F2BAFEEAFA74" httpUri="http://treatment.plazi.org/id/03B087E55B3FFFC9FF79F2BAFEEAFA74" lastPageNumber="91" pageId="90" pageNumber="91">
<subSubSection id="C30365785B3FFFC9FF79F2BAFDF3FF27" box="[151,581,151,177]" pageId="90" pageNumber="91" type="nomenclature">
<paragraph id="8BA636F35B3FFFC9FF79F2BAFDF3FF27" blockId="90.[151,581,151,210]" box="[151,581,151,177]" pageId="90" pageNumber="91">
<heading id="D0EE819F5B3FFFC9FF79F2BAFDF3FF27" bold="true" box="[151,581,151,177]" fontSize="11" level="1" pageId="90" pageNumber="91" reason="1">
<emphasis id="B96DEAE15B3FFFC9FF79F2BAFDF3FF27" bold="true" box="[151,581,151,177]" pageId="90" pageNumber="91">
<taxonomicName id="4C194D705B3FFFC9FF79F2BAFE10FF27" ID-CoL="3N3FD" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,422,151,177]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="perpunctata" status="sp. nov.">
<emphasis id="B96DEAE15B3FFFC9FF79F2BAFE10FF27" bold="true" box="[151,422,151,177]" italics="true" pageId="90" pageNumber="91">Hydraena perpunctata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B3FFFC9FE5BF2BAFDF3FF27" box="[437,581,151,177]" pageId="90" pageNumber="91" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B3FFFC9FF79F297FE34FF44" box="[151,386,186,210]" pageId="90" pageNumber="91" type="description">
<paragraph id="8BA636F35B3FFFC9FF79F297FE34FF44" blockId="90.[151,581,151,210]" box="[151,386,186,210]" pageId="90" pageNumber="91">
(
<figureCitation id="13222A765B3FFFC9FF71F297FEBDFF44" box="[159,267,186,210]" captionStart="FIGURE 155" captionStartId="261.[151,250,1753,1776]" captionTargetBox="[260,1328,216,1713]" captionTargetId="figure-26@261.[245,1342,193,1732]" captionTargetPageId="261" captionText="FIGURE 155. Hydraena perpuncta, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283706" httpUri="https://zenodo.org/record/5283706/files/figure.png" pageId="90" pageNumber="91">Figs. 155</figureCitation>
,
<figureCitation id="13222A765B3FFFC9FEF8F297FEF4FF44" box="[278,322,186,210]" captionStart="FIGURES 156157" captionStartId="262.[151,264,1649,1672]" captionTargetBox="[355,1215,202,1617]" captionTargetId="figure-32@262.[349,1238,193,1628]" captionTargetPageId="262" captionText="FIGURES 156157. Aedeagi of holotypes. 156. H. pertransversa. 157. H. perpuncta." figureDoi="http://doi.org/10.5281/zenodo.5283708" httpUri="https://zenodo.org/record/5283708/files/figure.png" pageId="90" pageNumber="91">157</figureCitation>
,
<figureCitation id="13222A765B3FFFC9FEA0F297FECFFF44" box="[334,377,186,210]" captionStart="FIGURES 499502" captionStartId="400.[151,264,1650,1673]" captionTargetBox="[165,1421,275,1591]" captionTargetId="figure-23@400.[165,1421,193,1629]" captionTargetPageId="400" captionText="FIGURES 499502. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5299474" httpUri="https://zenodo.org/record/5299474/files/figure.png" pageId="90" pageNumber="91">500</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B3FFFC9FF79F32EFAAFFEF4" pageId="90" pageNumber="91" type="materials_examined">
<paragraph id="8BA636F35B3FFFC9FF79F32EFAAFFEF4" blockId="90.[151,1436,258,1506]" pageId="90" pageNumber="91">
Type Material.
<materialsCitation id="3B713CAE5B3FFFC9FEBEF32EFAAFFEF4" collectingDate="1988-07-01" collectingDateMax="1988-08-30" collectingDateMin="1988-07-01" collectionCode="NMW" collectorName="D. Dudgeon" elevation="160" latitude="-5.55" location="Koje Creek" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" pageId="90" pageNumber="91" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<typeStatus id="54A288515B3FFFC9FEBEF32EFE0BFE8D" box="[336,445,259,283]" pageId="90" pageNumber="91" type="holotype">
<emphasis id="B96DEAE15B3FFFC9FEBEF32EFE0BFE8D" bold="true" box="[336,445,259,283]" pageId="90" pageNumber="91">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B3FFFC9FDCBF32EFCB5FE8D" bold="true" box="[549,771,258,283]" pageId="90" pageNumber="91">
<collectingRegion id="49DDF8115B3FFFC9FDCBF32EFD48FE8D" box="[549,766,258,283]" country="Papua New Guinea" name="Madang" pageId="90" pageNumber="91">Madang Province</collectingRegion>
:
</emphasis>
<collectingRegion id="49DDF8115B3FFFC9FCFAF32FFCE5FE8C" box="[788,851,258,282]" country="Papua New Guinea" name="West Sepik" pageId="90" pageNumber="91">Sepik</collectingRegion>
<collectingMunicipality id="6BC2AC895B3FFFC9FCB5F32EFC5FFE8C" box="[859,1001,258,282]" pageId="90" pageNumber="91">Ramu Basin</collectingMunicipality>
,
<location id="8EC660285B3FFFC9FC1BF32FFBCFFE8C" LSID="urn:lsid:plazi:treatment:03B087E55B3FFFC9FF79F2BAFEEAFA74:8EC660285B3FFFC9FC1BF32FFBCFFE8C" box="[1013,1145,258,282]" latitude="-5.55" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" name="Koje Creek" pageId="90" pageNumber="91" stateProvince="Madang">Kojé Creek</location>
, unshaded creek, small to medium cobbles, gravel and fine sand (the latter in areas of slack water), filamentous algae on stony substrates, small accumulations of leaf litter,
<quantity id="4CE19B165B3FFFC9FDFAF367FDEBFEF4" box="[532,605,330,354]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="90" pageNumber="91" unit="m" value="160.0">
<elevation id="0034D1C05B3FFFC9FDFAF367FDEBFEF4" box="[532,605,330,354]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="90" pageNumber="91" unit="m" value="160.0">160 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B3FFFC9FD87F367FD74FEF4" box="[617,706,330,354]" degrees="5" direction="south" minutes="33" orientation="latitude" pageId="90" pageNumber="91" precision="925" value="-5.55">5° 33' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B3FFFC9FD21F367FCF0FEF4" box="[719,838,330,354]" degrees="145" direction="east" minutes="23" orientation="longitude" pageId="90" pageNumber="91" precision="925" value="145.38333">145° 23' E</geoCoordinate>
,
<date id="FFA710335B3FFFC9FCBCF367FBAAFEF4" box="[850,1052,330,354]" pageId="90" pageNumber="91" value="1988-07-01" valueMax="1988-08-30" valueMin="1988-07-01">
<collectingDate id="EFE3E9DB5B3FFFC9FCBCF367FBAAFEF4" box="[850,1052,330,354]" pageId="90" pageNumber="91" value="1988-07-01" valueMax="1988-08-30" valueMin="1988-07-01">1 vii30 viii 1988</collectingDate>
</date>
,
<collectorName id="26EC53255B3FFFC9FBC6F366FB07FEF4" box="[1064,1201,330,354]" pageId="90" pageNumber="91">D. Dudgeon</collectorName>
(
<collectionCode id="ED08AE365B3FFFC9FB2EF367FAB8FEF4" box="[1216,1294,330,354]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="90" pageNumber="91">NMW</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B3FFFC9FF29F343FD90FDAD" pageId="90" pageNumber="91" type="diagnosis">
<paragraph id="8BA636F35B3FFFC9FF29F343FD90FDAD" blockId="90.[151,1436,258,1506]" pageId="90" pageNumber="91">
<emphasis id="B96DEAE15B3FFFC9FF29F343FE64FE11" bold="true" box="[199,466,366,391]" pageId="90" pageNumber="91">Differential Diagnosis.</emphasis>
The coarse and dense dorsal punctation is similar to, though even coarser than, that of
<taxonomicName id="4C194D705B3FFFC9FF79F3BEFE82FE3D" authorityName="Perkins" authorityYear="2011" box="[151,308,403,427]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="austrobesa">
<emphasis id="B96DEAE15B3FFFC9FF79F3BEFF1DFE3C" box="[151,171,403,426]" italics="true" pageId="90" pageNumber="91">H</emphasis>
.
<emphasis id="B96DEAE15B3FFFC9FF54F3BEFE82FE3D" box="[186,308,403,427]" italics="true" pageId="90" pageNumber="91">austrobesa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D705B3FFFC9FE82F3BEFDB4FE3D" authorityName="Perkins" authorityYear="2011" box="[364,514,403,427]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="limbobesa">
<emphasis id="B96DEAE15B3FFFC9FE82F3BEFE36FE3C" box="[364,384,403,426]" italics="true" pageId="90" pageNumber="91">H</emphasis>
.
<emphasis id="B96DEAE15B3FFFC9FE61F3BEFDB4FE3D" box="[399,514,403,427]" italics="true" pageId="90" pageNumber="91">limbobesa</emphasis>
</taxonomicName>
(
<figureCitation id="13222A765B3FFFC9FDFCF3BFFDC9FE3C" box="[530,639,402,426]" captionStart="FIGURE 150" captionStartId="257.[151,250,1753,1776]" captionTargetBox="[211,1374,225,1718]" captionTargetId="figure-26@257.[184,1403,193,1732]" captionTargetPageId="257" captionText="FIGURE 150. Hydraena austrobesa, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283694" httpUri="https://zenodo.org/record/5283694/files/figure.png" pageId="90" pageNumber="91">Figs. 150</figureCitation>
,
<figureCitation id="13222A765B3FFFC9FD62F3BFFD0EFE3C" box="[652,696,402,426]" captionStart="FIGURE 151" captionStartId="258.[151,250,1753,1776]" captionTargetBox="[255,1335,209,1718]" captionTargetId="figure-29@258.[233,1354,193,1732]" captionTargetPageId="258" captionText="FIGURE 151. Hydraena limbobesa, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283700" httpUri="https://zenodo.org/record/5283700/files/figure.png" pageId="90" pageNumber="91">151</figureCitation>
,
<figureCitation id="13222A765B3FFFC9FD2BF3BFFD45FE3C" box="[709,755,402,426]" captionStart="FIGURE 155" captionStartId="261.[151,250,1753,1776]" captionTargetBox="[260,1328,216,1713]" captionTargetId="figure-26@261.[245,1342,193,1732]" captionTargetPageId="261" captionText="FIGURE 155. Hydraena perpuncta, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283706" httpUri="https://zenodo.org/record/5283706/files/figure.png" pageId="90" pageNumber="91">155</figureCitation>
). In addition to the coarser punctation,
<taxonomicName id="4C194D705B3FFFC9FB5EF3BEFAECFE3C" authorityName="Perkins" authorityYear="2011" box="[1200,1370,403,426]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="perpunctata">
<emphasis id="B96DEAE15B3FFFC9FB5EF3BEFB72FE3C" box="[1200,1220,403,426]" italics="true" pageId="90" pageNumber="91">H</emphasis>
.
<emphasis id="B96DEAE15B3FFFC9FB3DF3BEFAECFE3C" box="[1235,1370,403,426]" italics="true" pageId="90" pageNumber="91">perpunctata</emphasis>
</taxonomicName>
additionally differs from the compared species by the more angulate sides of the pronotum, the wider explanate margin of the elytra, and the narrower and more widely separated metavental plaques (ratios ca. 6/1/4/10). The aedeagus of
<taxonomicName id="4C194D705B3FFFC9FF79F3D2FEF7FD80" authorityName="Perkins" authorityYear="2011" box="[151,321,511,534]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="perpunctata">
<emphasis id="B96DEAE15B3FFFC9FF79F3D2FF1DFD80" box="[151,171,511,534]" italics="true" pageId="90" pageNumber="91">H</emphasis>
.
<emphasis id="B96DEAE15B3FFFC9FF54F3D2FEF7FD80" box="[186,321,511,534]" italics="true" pageId="90" pageNumber="91">perpunctata</emphasis>
</taxonomicName>
(
<figureCitation id="13222A765B3FFFC9FEBCF3D3FE05FD80" box="[338,435,510,534]" captionStart="FIGURES 156157" captionStartId="262.[151,264,1649,1672]" captionTargetBox="[355,1215,202,1617]" captionTargetId="figure-32@262.[349,1238,193,1628]" captionTargetPageId="262" captionText="FIGURES 156157. Aedeagi of holotypes. 156. H. pertransversa. 157. H. perpuncta." figureDoi="http://doi.org/10.5281/zenodo.5283708" httpUri="https://zenodo.org/record/5283708/files/figure.png" pageId="90" pageNumber="91">Fig. 157</figureCitation>
) is abundantly distinct from that of other members of the
<emphasis id="B96DEAE15B3FFFC9FBA6F3D3FB0DFD80" box="[1096,1211,510,534]" italics="true" pageId="90" pageNumber="91">Spinobesa</emphasis>
group, and perhaps represents a separate species group.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B3FFFC9FF29F06BFA84FAE0" pageId="90" pageNumber="91" type="description">
<paragraph id="8BA636F35B3FFFC9FF29F06BFB3CFD5C" blockId="90.[151,1436,258,1506]" pageId="90" pageNumber="91">
<emphasis id="B96DEAE15B3FFFC9FF29F06BFEEEFDC9" bold="true" box="[199,344,582,607]" inLineHeading="true" pageId="90" pageNumber="91" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B3FFFC9FE42F06BFDB8FDC8" box="[428,526,582,606]" pageId="90" pageNumber="91" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.37/0.71; head 0.25/0.37; pronotum 0.38/0.52, PA 0.41, PB 0.41; elytra 0.87/0.71. Dorsum of head piceous; pronotum light brown to testaceous in front of and behind piceous fascia with ill-defined margins, ratios of color bands, as measured in midline, ca. 7/12/6; elytra and legs brown; maxillary palpi light brown to testaceous, tip not darker.
</paragraph>
<paragraph id="8BA636F35B3FFFC9FF29F0FAFA8CFC50" blockId="90.[151,1436,258,1506]" pageId="90" pageNumber="91">Frons coarsely densely punctate, punctures ca. 34xef, slightly larger and denser near eyes than medially; interstices microreticulate and dull laterally, effacedly microreticulate and weakly shining medially, narrow walls to 1xpd. Clypeus microreticulate laterally, moderately coarsely punctate medially. Mentum and postmentum very sparsely very finely punctulate, shining. Genae raised, shining, without posterior ridge. Pronotum moderately transverse (PL/PW ca. 0.73), ca. median 3/5 of anterior margin slightly emarginate; very coarsely, densely punctate, punctures on disc ca. 3xpd frons punctures, interstices on disc shining, narrow walls to 1xpd, punctures larger and denser at anterior and posterior; PF1 shallow, obsolete; PF2 moderately deep; PF3 deep; PF4 shallow.</paragraph>
<paragraph id="8BA636F35B3FFFC9FF29F1FFFE2BFBEC" blockId="90.[151,1436,258,1506]" pageId="90" pageNumber="91">Elytra with summit of posterior declivity slightly anterior to midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, weakly shining, on disc ca. 1xpd, as are interstices between punctures of a row. Apices in dorsal aspect weakly separately rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="8BA636F35B3FFFC9FF29F6ABFA84FAE0" blockId="90.[151,1436,258,1506]" pageId="90" pageNumber="91">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 6/1/4/10. P1 laminate; median carina sinuate in profile. P2 very narrow, raised slightly, l/w ca. 2/1, sides slightly converging toward apex. Plaques minute, widely separated, carinate lines at sides of median depression that continues shallowly to tip of P2. Metaventrite without midlongitudinal ridge. AIS width at straight posterior margin slightly wider than P2. All legs rather short, femora moderately robust. Profemur (male) without tubercle next to trochanter; protibia widest at distal 1/3 where produced on medial margin. Mesotibia straight, lateral margin slightly arcuate and bearing row of short stout spines. Metatibia missing in
<typeStatus id="54A288515B3FFFC9FF5DF773FEAEFAE0" box="[179,280,1374,1398]" pageId="90" pageNumber="91" type="holotype">holotype</typeStatus>
. Abdominal apex symmetrical; last tergite (male) notched. Aedeagus as illustrated (
<figureCitation id="13222A765B3FFFC9FB2DF773FA90FAE0" box="[1219,1318,1374,1398]" captionStart="FIGURES 156157" captionStartId="262.[151,264,1649,1672]" captionTargetBox="[355,1215,202,1617]" captionTargetId="figure-32@262.[349,1238,193,1628]" captionTargetPageId="262" captionText="FIGURES 156157. Aedeagi of holotypes. 156. H. pertransversa. 157. H. perpuncta." figureDoi="http://doi.org/10.5281/zenodo.5283708" httpUri="https://zenodo.org/record/5283708/files/figure.png" pageId="90" pageNumber="91">Fig. 157</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B3FFFC9FF29F7AEFC78FA0C" box="[199,974,1410,1435]" pageId="90" pageNumber="91" type="etymology">
<paragraph id="8BA636F35B3FFFC9FF29F7AEFC78FA0C" blockId="90.[151,1436,258,1506]" box="[199,974,1410,1435]" pageId="90" pageNumber="91">
<emphasis id="B96DEAE15B3FFFC9FF29F7AEFEFBFA0D" bold="true" box="[199,333,1411,1435]" inLineHeading="true" pageId="90" pageNumber="91" reason="1">Etymology.</emphasis>
Named in reference to the very coarse dorsal punctation.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B3FFFC9FF29F78BFEEAFA74" pageId="90" pageNumber="91" type="distribution">
<paragraph id="8BA636F35B3FFFC9FF29F78BFEEAFA74" blockId="90.[151,1436,258,1506]" pageId="90" pageNumber="91">
<emphasis id="B96DEAE15B3FFFC9FF29F78BFED6FA29" bold="true" box="[199,352,1446,1471]" inLineHeading="true" pageId="90" pageNumber="91" reason="1">Distribution.</emphasis>
Currently known only from the
<typeStatus id="54A288515B3FFFC9FD25F78AFD4DFA29" box="[715,763,1447,1471]" pageId="90" pageNumber="91">type</typeStatus>
locality, Kojé Creek, in the eastern arm of Area 3; elevation
<quantity id="4CE19B165B3FFFC9FF79F7E7FF68FA74" box="[151,222,1482,1506]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="90" pageNumber="91" unit="m" value="160.0">160 m</quantity>
(
<figureCitation id="13222A765B3FFFC9FF03F7E7FEF9FA74" box="[237,335,1482,1506]" captionStart="FIGURES 499502" captionStartId="400.[151,264,1650,1673]" captionTargetBox="[165,1421,275,1591]" captionTargetId="figure-23@400.[165,1421,193,1629]" captionTargetPageId="400" captionText="FIGURES 499502. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5299474" httpUri="https://zenodo.org/record/5299474/files/figure.png" pageId="90" pageNumber="91">Fig. 500</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>