158 lines
19 KiB
XML
158 lines
19 KiB
XML
<document id="7747F900312CB726447386691140128D" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B12FFEBFF79F73AFE7CFD31" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena fasciolata Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="121" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="120" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="EA36271BCB57E402292D44AA6D567865">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="F649140F8E2E29940DC51A5324340AD1">Perkins, Philip D.</mods:namePart>
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<mods:title id="E81377494695A31FD5FD335527C11AB1">Zootaxa</mods:title>
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<mods:part id="B602251A5814234C4B00FB61E74AE53E">
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<mods:date id="66A92368064B56DB2054C71DB0FEEF41">2011</mods:date>
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<mods:number id="1D9248EAF4AF80220339B68AF95564E6">2011-07-08</mods:number>
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<mods:number id="9274D6CA23EF3E3F8174133D7D64C254">2944</mods:number>
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<mods:url id="3496F9F2A523E7FB308B300B009DE932">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
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<mods:classification id="F3F0FAD15EF812478E55C6CB59925755">journal article</mods:classification>
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<mods:identifier id="8DA22273AA1CF1FFE0555343D5438A29" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
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<treatment id="03B087E55B12FFEBFF79F73AFE7CFD31" ID-DOI="http://doi.org/10.5281/zenodo.5291765" ID-Zenodo-Dep="5291765" LSID="urn:lsid:plazi:treatment:03B087E55B12FFEBFF79F73AFE7CFD31" httpUri="http://treatment.plazi.org/id/03B087E55B12FFEBFF79F73AFE7CFD31" lastPageId="120" lastPageNumber="121" pageId="119" pageNumber="120">
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<subSubSection id="C30365785B12FFE4FF79F73AFD9CFAA7" box="[151,554,1303,1329]" pageId="119" pageNumber="120" type="nomenclature">
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<paragraph id="8BA636F35B12FFE4FF79F73AFD9CFAA7" blockId="119.[151,554,1303,1362]" box="[151,554,1303,1329]" pageId="119" pageNumber="120">
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<heading id="D0EE819F5B12FFE4FF79F73AFD9CFAA7" bold="true" box="[151,554,1303,1329]" fontSize="11" level="1" pageId="119" pageNumber="120" reason="1">
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<emphasis id="B96DEAE15B12FFE4FF79F73AFD9CFAA7" bold="true" box="[151,554,1303,1329]" pageId="119" pageNumber="120">
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<taxonomicName id="4C194D705B12FFE4FF79F73AFE3DFAA7" ID-CoL="6MHH5" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,395,1303,1329]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="fasciolata" status="sp. nov.">
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<emphasis id="B96DEAE15B12FFE4FF79F73AFE3DFAA7" bold="true" box="[151,395,1303,1329]" italics="true" pageId="119" pageNumber="120">Hydraena fasciolata</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25E579A5B12FFE4FE74F73AFD9CFAA7" box="[410,554,1303,1329]" pageId="119" pageNumber="120" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B12FFE4FF79F717FE34FAC4" box="[151,386,1338,1362]" pageId="119" pageNumber="120" type="description">
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<paragraph id="8BA636F35B12FFE4FF79F717FE34FAC4" blockId="119.[151,554,1303,1362]" box="[151,386,1338,1362]" pageId="119" pageNumber="120">
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(
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<figureCitation id="13222A765B12FFE4FF71F717FEBDFAC4" box="[159,267,1338,1362]" captionStart="FIGURE 219" captionStartId="309.[151,250,1753,1776]" captionTargetBox="[191,1371,203,1718]" captionTargetId="figure-26@309.[186,1400,193,1732]" captionTargetPageId="309" captionText="FIGURE 219. Hydraena fasciolata, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283842" httpUri="https://zenodo.org/record/5283842/files/figure.png" pageId="119" pageNumber="120">Figs. 219</figureCitation>
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,
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<figureCitation id="13222A765B12FFE4FEF8F717FEF4FAC4" box="[278,322,1338,1362]" captionStart="FIGURES 220–221" captionStartId="310.[151,264,1650,1673]" captionTargetBox="[263,1294,222,1611]" captionTargetId="figure-32@310.[226,1360,193,1629]" captionTargetPageId="310" captionText="FIGURES 220–221. Aedeagi of holotypes. 220. H. dudgeoni. 221. H. fasciolata." figureDoi="http://doi.org/10.5281/zenodo.5283846" httpUri="https://zenodo.org/record/5283846/files/figure.png" pageId="119" pageNumber="120">221</figureCitation>
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,
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<figureCitation id="13222A765B12FFE4FEA0F717FECFFAC4" box="[334,377,1338,1362]" captionStart="FIGURES 527–530" captionStartId="407.[151,264,1650,1673]" captionTargetBox="[165,1421,224,1616]" captionTargetId="figure-20@407.[165,1421,193,1629]" captionTargetPageId="407" captionText="FIGURES 527–530. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284135" httpUri="https://zenodo.org/record/5284135/files/figure.png" pageId="119" pageNumber="120">530</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C30365785B12FFE4FF79F7AEFD2DFA28" pageId="119" pageNumber="120" type="materials_examined">
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<paragraph id="8BA636F35B12FFE4FF79F7AEFD2DFA28" blockId="119.[151,1437,1410,2010]" pageId="119" pageNumber="120">
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Type Material.
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<materialsCitation id="3B713CAE5B12FFE4FEA0F7AEFD2DFA28" collectingDate="2006-04-30" collectionCode="ZSM" collectorName="Balke & Sagata" country="Papua New Guinea" elevation="160" latitude="-5.23205" location="Ohu Village" longLatPrecision="1" longitude="145.67938" pageId="119" pageNumber="120" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
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<typeStatus id="54A288515B12FFE4FEA0F7AEFE0DFA0D" box="[334,443,1411,1435]" pageId="119" pageNumber="120" type="holotype">
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||
<emphasis id="B96DEAE15B12FFE4FEA0F7AEFE0DFA0D" bold="true" box="[334,443,1411,1435]" pageId="119" pageNumber="120">Holotype</emphasis>
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</typeStatus>
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(male):
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<emphasis id="B96DEAE15B12FFE4FDF1F7AEFD4DFA0D" bold="true" box="[543,763,1410,1435]" pageId="119" pageNumber="120">
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||
<collectingRegion id="49DDF8115B12FFE4FDF1F7AEFD41FA0D" box="[543,759,1410,1435]" country="Papua New Guinea" name="Madang" pageId="119" pageNumber="120">Madang Province</collectingRegion>
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:
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</emphasis>
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<collectingRegion id="49DDF8115B12FFE4FCE6F7AFFCD1FA0C" box="[776,871,1410,1434]" country="Papua New Guinea" name="Madang" pageId="119" pageNumber="120">Madang</collectingRegion>
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,
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<location id="8EC660285B12FFE4FC9CF7AFFC4DFA0C" LSID="urn:lsid:plazi:treatment:03B087E55B12FFEBFF79F73AFE7CFD31:8EC660285B12FFE4FC9CF7AFFC4DFA0C" box="[882,1019,1410,1434]" country="Papua New Guinea" latitude="-5.23205" longLatPrecision="1" longitude="145.67938" name="Ohu Village" pageId="119" pageNumber="120" stateProvince="Madang">Ohu Village</location>
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,
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<quantity id="4CE19B165B12FFE4FBE8F7AFFBF9FA0C" box="[1030,1103,1410,1434]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="119" pageNumber="120" unit="m" value="160.0">
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<elevation id="0034D1C05B12FFE4FBE8F7AFFBF9FA0C" box="[1030,1103,1410,1434]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="119" pageNumber="120" unit="m" value="160.0">160 m</elevation>
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</quantity>
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,
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||
<geoCoordinate id="EE2D50345B12FFE4FBB4F7AFFB52FA0C" box="[1114,1252,1410,1434]" degrees="5" direction="south" minutes="13.923" orientation="latitude" pageId="119" pageNumber="120" precision="1" value="-5.23205">5° 13.923' S</geoCoordinate>
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,
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||
<geoCoordinate id="EE2D50345B12FFE4FB1EF7AFFA21FA0C" box="[1264,1431,1410,1434]" degrees="145" direction="east" minutes="40.763" orientation="longitude" pageId="119" pageNumber="120" precision="1" value="145.67938">145° 40.763' E</geoCoordinate>
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,
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<date id="FFA710335B12FFE4FF79F78BFEA7FA28" box="[151,273,1446,1470]" pageId="119" pageNumber="120" value="2006-04-30">
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<collectingDate id="EFE3E9DB5B12FFE4FF79F78BFEA7FA28" box="[151,273,1446,1470]" pageId="119" pageNumber="120" value="2006-04-30">30 iv 2006</collectingDate>
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</date>
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,
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<collectorName id="26EC53255B12FFE4FEF2F78BFEEBFA28" box="[284,349,1446,1470]" pageId="119" pageNumber="120">Balke</collectorName>
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&
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<collectorName id="26EC53255B12FFE4FE6CF78BFE7AFA28" box="[386,460,1446,1470]" pageId="119" pageNumber="120">Sagata</collectorName>
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(
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<collectingCountry id="F30E76635B12FFE4FE34F78BFDA2FA28" box="[474,532,1446,1470]" name="Papua New Guinea" pageId="119" pageNumber="120">PNG</collectingCountry>
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||
49) (
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<collectionCode id="ED08AE365B12FFE4FDBEF78BFD39FA28" box="[592,655,1446,1470]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="119" pageNumber="120" type="Museum">ZSM</collectionCode>
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).
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30365785B12FFE4FF29F7E7FC4BF9D8" pageId="119" pageNumber="120" type="diagnosis">
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||
<paragraph id="8BA636F35B12FFE4FF29F7E7FC4BF9D8" blockId="119.[151,1437,1410,2010]" pageId="119" pageNumber="120">
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<emphasis id="B96DEAE15B12FFE4FF29F7E7FE64FA75" bold="true" box="[199,466,1482,1507]" pageId="119" pageNumber="120">Differential Diagnosis.</emphasis>
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A distinctively banded, very broad species. Somewhat similar to
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<taxonomicName id="4C194D705B12FFE4FB5AF7E6FA88FA75" authorityName="Perkins" authorityYear="2011" box="[1204,1342,1483,1507]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="dudgeoni">
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<emphasis id="B96DEAE15B12FFE4FB5AF7E6FB7EFA74" box="[1204,1224,1483,1506]" italics="true" pageId="119" pageNumber="120">H</emphasis>
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.
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<emphasis id="B96DEAE15B12FFE4FB38F7E6FA88FA75" box="[1238,1342,1483,1507]" italics="true" pageId="119" pageNumber="120">dudgeoni</emphasis>
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</taxonomicName>
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in habitus, both having a very transverse pronotum (
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<figureCitation id="13222A765B12FFE4FD46F7C3FCA0F990" box="[680,790,1518,1542]" captionStart="FIGURE 218" captionStartId="308.[151,250,1753,1776]" captionTargetBox="[245,1337,231,1721]" captionTargetId="figure-29@308.[218,1369,193,1732]" captionTargetPageId="308" captionText="FIGURE 218. Hydraena dudgeoni, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283838" httpUri="https://zenodo.org/record/5283838/files/figure.png" pageId="119" pageNumber="120">Figs. 218</figureCitation>
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,
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<figureCitation id="13222A765B12FFE4FCCBF7C3FCE5F990" box="[805,851,1518,1542]" captionStart="FIGURE 219" captionStartId="309.[151,250,1753,1776]" captionTargetBox="[191,1371,203,1718]" captionTargetId="figure-26@309.[186,1400,193,1732]" captionTargetPageId="309" captionText="FIGURE 219. Hydraena fasciolata, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283842" httpUri="https://zenodo.org/record/5283842/files/figure.png" pageId="119" pageNumber="120">219</figureCitation>
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), but more markedly color banded, and with more widely spaced coxae (plaque ratios ca. 9/1/6/8 vs. 7/1/6/8). This species has the widest separation of the mesocoxae. The male genitalia of the two species differ markedly (
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<figureCitation id="13222A765B12FFE4FCA5F41BFC47F9D8" box="[843,1009,1590,1614]" captionStart="FIGURES 220–221" captionStartId="310.[151,264,1650,1673]" captionTargetBox="[263,1294,222,1611]" captionTargetId="figure-32@310.[226,1360,193,1629]" captionTargetPageId="310" captionText="FIGURES 220–221. Aedeagi of holotypes. 220. H. dudgeoni. 221. H. fasciolata." figureDoi="http://doi.org/10.5281/zenodo.5283846" httpUri="https://zenodo.org/record/5283846/files/figure.png" pageId="119" pageNumber="120">Figs. 220, 221</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B12FFEBFF29F477FEE3FDAA" lastPageId="120" lastPageNumber="121" pageId="119" pageNumber="120" type="description">
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<paragraph id="8BA636F35B12FFE4FF29F477FE14F894" blockId="119.[151,1437,1410,2010]" pageId="119" pageNumber="120">
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||
<emphasis id="B96DEAE15B12FFE4FF29F477FEEEF9E5" bold="true" box="[199,344,1626,1651]" inLineHeading="true" pageId="119" pageNumber="120" reason="1">Description.</emphasis>
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Size:
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<typeStatus id="54A288515B12FFE4FE42F477FDB8F9E4" box="[428,526,1626,1650]" pageId="119" pageNumber="120" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.44/0.68; head 0.23/0.35; pronotum 0.36/0.55, PA 0.43, PB 0.46; elytra 0.84/0.68. Dorsum very distinctively banded: labrum and clypeus light brown to testaceous; frons dark brown; pronotum testaceous; elytra dark brown with light brown to testaceous fascia at posterior declivity, and similarly colored band across elytral apices; legs and maxillary palpi testaceous, tip of palpus not darker.
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</paragraph>
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<paragraph id="8BA636F35B12FFE4FF29F522FC1EF84C" blockId="119.[151,1437,1410,2010]" pageId="119" pageNumber="120">Frons punctures ca. 1xef, slightly larger near eyes than medially; interstices shining, 2–4xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum and postmentum very sparsely very finely punctulate, shining. Genae raised, shining, without posterior ridge. Pronotum very transverse, sides markedly rounded, ca. median 5/7 of anterior margin arcuate to posterior; punctures on disc ca. 1xpd frons punctures, interstices shining, 1–2xpd, punctures not appreciably larger or denser at anterior and posterior; PF1 absent; PF2 absent or so shallow as to be almost imperceptible; PF3 shallow; PF4 absent.</paragraph>
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<paragraph id="8BA636F35B1DFFEBFF29F2BAFCBCFEA9" blockId="120.[151,1437,151,679]" pageId="120" pageNumber="121">Elytra with summit of posterior declivity anterior to midlength; lateral explanate margins rather wide in anterior 1/2 where sides of elytra gradually sloping; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 2–5xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
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<paragraph id="8BA636F35B1DFFEBFF29F361FEE3FDAA" blockId="120.[151,1437,151,679]" pageId="120" pageNumber="121">
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Ratios of P2 width and plaque shape (P2/w/l/s) ca. 9/1/6/8. P1 ca. 2/3 P2; median carina straight in profile; postcoxal process medially concave. P2 transversely concave, l/w ca. 4/9, sides slightly diverging toward blunt apex. Plaques very narrow, carinate, very slightly arcuate, converging slightly anteriorly, at sides of deep median depression. Metaventrite concave between P2 and plaques. AIS width at straight posterior margin ca. equal P2. All legs rather short, moderately slender. Profemur (male) with small tubercle next to trochanter; protibia arcuate. Meso- and metatibia straight. Abdominal apex symmetrical; last tergite (male) deeply notched. Aedeagus as illustrated (
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<figureCitation id="13222A765B1DFFEBFF08F009FEFEFDAA" box="[230,328,548,572]" captionStart="FIGURES 220–221" captionStartId="310.[151,264,1650,1673]" captionTargetBox="[263,1294,222,1611]" captionTargetId="figure-32@310.[226,1360,193,1629]" captionTargetPageId="310" captionText="FIGURES 220–221. Aedeagi of holotypes. 220. H. dudgeoni. 221. H. fasciolata." figureDoi="http://doi.org/10.5281/zenodo.5283846" httpUri="https://zenodo.org/record/5283846/files/figure.png" pageId="120" pageNumber="121">Fig. 221</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B1DFFEBFF29F065FC66FDC9" box="[199,976,583,608]" pageId="120" pageNumber="121" type="etymology">
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<paragraph id="8BA636F35B1DFFEBFF29F065FC66FDC9" blockId="120.[151,1437,151,679]" box="[199,976,583,608]" pageId="120" pageNumber="121">
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<emphasis id="B96DEAE15B1DFFEBFF29F065FEFBFDF6" bold="true" box="[199,333,584,608]" inLineHeading="true" pageId="120" pageNumber="121" reason="1">Etymology.</emphasis>
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Named in reference to the dorsal fasciae, and wide body.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B1DFFEBFF29F046FE7CFD31" pageId="120" pageNumber="121" type="distribution">
|
||
<paragraph id="8BA636F35B1DFFEBFF29F046FE7CFD31" blockId="120.[151,1437,151,679]" pageId="120" pageNumber="121">
|
||
<emphasis id="B96DEAE15B1DFFEBFF29F046FED6FD12" bold="true" box="[199,352,619,644]" inLineHeading="true" pageId="120" pageNumber="121" reason="1">Distribution.</emphasis>
|
||
Currently known only from the
|
||
<typeStatus id="54A288515B1DFFEBFD36F041FCBEFD12" box="[728,776,620,644]" pageId="120" pageNumber="121">type</typeStatus>
|
||
locality, Ohu Village, in the northwestern part of Area 5; elevation
|
||
<quantity id="4CE19B165B1DFFEBFEEAF0A2FEFDFD31" box="[260,331,655,679]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="120" pageNumber="121" unit="m" value="160.0">160 m</quantity>
|
||
(
|
||
<figureCitation id="13222A765B1DFFEBFEB5F0A2FE08FD31" box="[347,446,655,679]" captionStart="FIGURES 527–530" captionStartId="407.[151,264,1650,1673]" captionTargetBox="[165,1421,224,1616]" captionTargetId="figure-20@407.[165,1421,193,1629]" captionTargetPageId="407" captionText="FIGURES 527–530. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284135" httpUri="https://zenodo.org/record/5284135/files/figure.png" pageId="120" pageNumber="121">Fig. 530</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |