165 lines
20 KiB
XML
165 lines
20 KiB
XML
<document id="E00517BC0FA65F97FEB73C53AAD1D80C" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B0CFFFAFF79F30AFE25F9BD" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena impala Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="106" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="106" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="164BB01B670CF5FB508FF6DD359A0A11">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="4723F6CC3F8238315D00584DF664E889">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="DA15A81AB2C641310FCCA13AE9053C2E">text</mods:typeOfResource>
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<mods:title id="C57CA3B995C80402D4F05336090B8A0D">Zootaxa</mods:title>
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<mods:part id="FB11A3C127F7129EE280C3F31E4E0B48">
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<mods:date id="5C92238F42D91D9B92A9BB6AFFB98699">2011</mods:date>
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<mods:number id="A6F0D1EB697802103E6F52B40F3E48E6">2011-07-08</mods:number>
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<mods:number id="98A50E1EB71CE8C1DFD3469F2AFFB4F8">2944</mods:number>
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<mods:url id="4EB82C34687D5D0050C4379EF60D35AE">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
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<mods:classification id="E9E6C53E28C902254222F0CC82A3FD71">journal article</mods:classification>
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<mods:identifier id="246C04986489DAF031029E7B78521D97" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
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<mods:identifier id="FB77119D1AC9B83A1A21705E335C5EB6" type="ISSN">11755334</mods:identifier>
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<treatment id="03B087E55B0CFFFAFF79F30AFE25F9BD" ID-DOI="http://doi.org/10.5281/zenodo.5291708" ID-Zenodo-Dep="5291708" LSID="urn:lsid:plazi:treatment:03B087E55B0CFFFAFF79F30AFE25F9BD" httpUri="http://treatment.plazi.org/id/03B087E55B0CFFFAFF79F30AFE25F9BD" lastPageNumber="106" pageId="105" pageNumber="106">
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<subSubSection id="C30365785B0CFFFAFF79F30AFDBEFED7" box="[151,520,295,321]" pageId="105" pageNumber="106" type="nomenclature">
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<paragraph id="8BA636F35B0CFFFAFF79F30AFDBEFED7" blockId="105.[151,520,295,354]" box="[151,520,295,321]" pageId="105" pageNumber="106">
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<heading id="D0EE819F5B0CFFFAFF79F30AFDBEFED7" bold="true" box="[151,520,295,321]" fontSize="11" level="1" pageId="105" pageNumber="106" reason="1">
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<emphasis id="B96DEAE15B0CFFFAFF79F30AFDBEFED7" bold="true" box="[151,520,295,321]" pageId="105" pageNumber="106">
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<taxonomicName id="4C194D705B0CFFFAFF79F30AFEDFFED7" ID-CoL="6MJBF" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,361,295,321]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="impala" status="sp. nov.">
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<emphasis id="B96DEAE15B0CFFFAFF79F30AFEDFFED7" bold="true" box="[151,361,295,321]" italics="true" pageId="105" pageNumber="106">Hydraena impala</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25E579A5B0CFFFAFE96F30AFDBEFED7" box="[376,520,295,321]" pageId="105" pageNumber="106" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B0CFFFAFF79F367FE34FEF4" box="[151,386,330,354]" pageId="105" pageNumber="106" type="description">
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<paragraph id="8BA636F35B0CFFFAFF79F367FE34FEF4" blockId="105.[151,520,295,354]" box="[151,386,330,354]" pageId="105" pageNumber="106">
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(
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<figureCitation id="13222A765B0CFFFAFF71F367FEBDFEF4" box="[159,267,330,354]" captionStart="FIGURE 183" captionStartId="282.[151,250,1753,1776]" captionTargetBox="[281,1312,209,1718]" captionTargetId="figure-29@282.[259,1327,193,1732]" captionTargetPageId="282" captionText="FIGURE 183. Hydraena impala, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283758" httpUri="https://zenodo.org/record/5283758/files/figure.png" pageId="105" pageNumber="106">Figs. 183</figureCitation>
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,
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<figureCitation id="13222A765B0CFFFAFEF8F367FEF4FEF4" box="[278,322,330,354]" captionStart="FIGURES 184–185" captionStartId="283.[151,264,1650,1673]" captionTargetBox="[290,1288,210,1618]" captionTargetId="figure-29@283.[279,1307,193,1629]" captionTargetPageId="283" captionText="FIGURES 184–185. Aedeagi of holotypes. 184. H. buloba. 185. H. impala." figureDoi="http://doi.org/10.5281/zenodo.5283760" httpUri="https://zenodo.org/record/5283760/files/figure.png" pageId="105" pageNumber="106">185</figureCitation>
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,
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<figureCitation id="13222A765B0CFFFAFEA0F367FECFFEF4" box="[334,377,330,354]" captionStart="FIGURES 531–534" captionStartId="408.[151,264,1650,1673]" captionTargetBox="[165,1421,234,1616]" captionTargetId="figure-23@408.[165,1421,193,1629]" captionTargetPageId="408" captionText="FIGURES 531–534. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284137" httpUri="https://zenodo.org/record/5284137/files/figure.png" pageId="105" pageNumber="106">534</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C30365785B0CFFFAFF79F3BEFA81FE58" pageId="105" pageNumber="106" type="materials_examined">
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<paragraph id="8BA636F35B0CFFFAFF79F3BEFA81FE58" blockId="105.[151,1437,402,1579]" pageId="105" pageNumber="106">
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Type Material.
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<materialsCitation id="3B713CAE5B0CFFFAFEB9F3BEFA81FE58" collectingDate="1965-03-16" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="150" latitude="-9.766666" location="Musgrave River" longLatPrecision="1299" longitude="147.68333" municipality="Sogeri Plateau" pageId="105" pageNumber="106" specimenCount="1" stateProvince="Central" typeStatus="holotype">
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<typeStatus id="54A288515B0CFFFAFEB9F3BEFE72FE3D" box="[343,452,403,427]" pageId="105" pageNumber="106" type="holotype">
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<emphasis id="B96DEAE15B0CFFFAFEB9F3BEFE72FE3D" bold="true" box="[343,452,403,427]" pageId="105" pageNumber="106">Holotype</emphasis>
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</typeStatus>
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(male):
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<emphasis id="B96DEAE15B0CFFFAFDD9F3BFFCA7FE3D" bold="true" box="[567,785,402,427]" pageId="105" pageNumber="106">
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<collectingRegion id="49DDF8115B0CFFFAFDD9F3BFFCBAFE3D" box="[567,780,402,427]" country="Papua New Guinea" name="Central" pageId="105" pageNumber="106">Central Province</collectingRegion>
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:
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</emphasis>
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nr.
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<collectingRegion id="49DDF8115B0CFFFAFCA1F3BEFC59FE3C" box="[847,1007,402,426]" country="Papua New Guinea" name="National Capital" pageId="105" pageNumber="106">Port Moresby</collectingRegion>
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,
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<collectingMunicipality id="6BC2AC895B0CFFFAFC10F3BFFB11FE3C" box="[1022,1191,402,426]" pageId="105" pageNumber="106">Sogeri Plateau</collectingMunicipality>
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,
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<location id="8EC660285B0CFFFAFB59F3BEFAC4FE3C" LSID="urn:lsid:plazi:treatment:03B087E55B0CFFFAFF79F30AFE25F9BD:8EC660285B0CFFFAFB59F3BEFAC4FE3C" box="[1207,1394,402,427]" latitude="-9.766666" longLatPrecision="1299" longitude="147.68333" municipality="Sogeri Plateau" name="Musgrave River" pageId="105" pageNumber="106" stateProvince="Central">Musgrave River</location>
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, ex gravel banks, [GE est.]
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<quantity id="4CE19B165B0CFFFAFE71F39BFD96FE58" box="[415,544,438,462]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="105" pageNumber="106" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
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<elevation id="0034D1C05B0CFFFAFE71F39BFD96FE58" box="[415,544,438,462]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="105" pageNumber="106" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100–200 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE2D50345B0CFFFAFDC2F39BFD33FE58" box="[556,645,438,462]" degrees="9" direction="south" minutes="46" orientation="latitude" pageId="105" pageNumber="106" precision="925" value="-9.766666">9° 46' S</geoCoordinate>
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,
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<geoCoordinate id="EE2D50345B0CFFFAFD7CF39BFCBFFE58" box="[658,777,438,462]" degrees="147" direction="east" minutes="41" orientation="longitude" pageId="105" pageNumber="106" precision="925" value="147.68333">147° 41' E</geoCoordinate>
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,
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<date id="FFA710335B0CFFFAFCFBF39BFC26FE58" box="[789,912,438,462]" pageId="105" pageNumber="106" value="1965-03-16">
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<collectingDate id="EFE3E9DB5B0CFFFAFCFBF39BFC26FE58" box="[789,912,438,462]" pageId="105" pageNumber="106" value="1965-03-16">16 iii 1965</collectingDate>
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</date>
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,
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<collectorName id="26EC53255B0CFFFAFC72F39AFBF7FE58" box="[924,1089,438,462]" pageId="105" pageNumber="106">M. E. Bacchus</collectorName>
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(MEB 200a) (
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<collectionCode id="ED08AE365B0CFFFAFB0DF39BFA9DFE58" box="[1251,1323,438,462]" pageId="105" pageNumber="106">NHM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B0CFFFAFF29F3F7FD37FD14" pageId="105" pageNumber="106" type="diagnosis">
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<paragraph id="8BA636F35B0CFFFAFF29F3F7FD37FD14" blockId="105.[151,1437,402,1579]" pageId="105" pageNumber="106">
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<emphasis id="B96DEAE15B0CFFFAFF29F3F7FE64FE65" bold="true" box="[199,466,474,499]" pageId="105" pageNumber="106">Differential Diagnosis.</emphasis>
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Similar in color and dorsal sculpture to
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<taxonomicName id="4C194D705B0CFFFAFC77F3F6FC4CFE65" authorityName="Perkins" authorityYear="2011" box="[921,1018,475,499]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="galea">
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<emphasis id="B96DEAE15B0CFFFAFC77F3F6FC1BFE64" box="[921,941,475,498]" italics="true" pageId="105" pageNumber="106">H</emphasis>
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.
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<emphasis id="B96DEAE15B0CFFFAFC52F3F6FC4CFE65" box="[956,1018,475,499]" italics="true" pageId="105" pageNumber="106">galea</emphasis>
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</taxonomicName>
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(
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<figureCitation id="13222A765B0CFFFAFBE4F3F7FBC0FE64" box="[1034,1142,474,498]" captionStart="FIGURE 183" captionStartId="282.[151,250,1753,1776]" captionTargetBox="[281,1312,209,1718]" captionTargetId="figure-29@282.[259,1327,193,1732]" captionTargetPageId="282" captionText="FIGURE 183. Hydraena impala, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283758" httpUri="https://zenodo.org/record/5283758/files/figure.png" pageId="105" pageNumber="106">Figs. 183</figureCitation>
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,
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<figureCitation id="13222A765B0CFFFAFB6CF3F7FB06FE64" box="[1154,1200,474,498]" captionStart="FIGURE 186" captionStartId="284.[151,250,1753,1776]" captionTargetBox="[224,1366,215,1724]" captionTargetId="figure-29@284.[208,1379,193,1732]" captionTargetPageId="284" captionText="FIGURE 186. Hydraena galea, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283766" httpUri="https://zenodo.org/record/5283766/files/figure.png" pageId="105" pageNumber="106">186</figureCitation>
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); differing therefrom by the slightly narrower body form (EW/TL ca. 0.40 vs. 0.45), the narrower mesoventral intercoxal process (P2) (plaque ratios ca. 4/1/5/5 vs. 5/1/5/5), and the micropunctulate mentum. The general plan of the male genitalia of the two species also suggests a relationship, but the genitalia distinctively differ in details, and should be examined for reliable determinations (
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<figureCitation id="13222A765B0CFFFAFE21F047FD8CFD14" box="[463,570,618,642]" captionStart="FIGURES 184–185" captionStartId="283.[151,264,1650,1673]" captionTargetBox="[290,1288,210,1618]" captionTargetId="figure-29@283.[279,1307,193,1629]" captionTargetPageId="283" captionText="FIGURES 184–185. Aedeagi of holotypes. 184. H. buloba. 185. H. impala." figureDoi="http://doi.org/10.5281/zenodo.5283760" httpUri="https://zenodo.org/record/5283760/files/figure.png" pageId="105" pageNumber="106">Figs. 185</figureCitation>
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,
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<figureCitation id="13222A765B0CFFFAFDA8F047FDC2FD14" box="[582,628,618,642]" captionStart="FIGURES 188–189" captionStartId="286.[151,264,1650,1673]" captionTargetBox="[305,1277,206,1620]" captionTargetId="figure-32@286.[288,1298,193,1629]" captionTargetPageId="286" captionText="FIGURES 188–189. Aedeagi of holotypes. 188. H. galea. 189. H. inhalista." figureDoi="http://doi.org/10.5281/zenodo.5283770" httpUri="https://zenodo.org/record/5283770/files/figure.png" pageId="105" pageNumber="106">188</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C30365785B0CFFFAFF29F0A3FF67FA0C" pageId="105" pageNumber="106" type="description">
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||
<paragraph id="8BA636F35B0CFFFAFF29F0A3FAF5FC84" blockId="105.[151,1437,402,1579]" pageId="105" pageNumber="106">
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||
<emphasis id="B96DEAE15B0CFFFAFF29F0A3FEEEFD31" bold="true" box="[199,344,654,679]" inLineHeading="true" pageId="105" pageNumber="106" reason="1">Description.</emphasis>
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Size:
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<typeStatus id="54A288515B0CFFFAFE42F0A3FDB8FD30" box="[428,526,654,678]" pageId="105" pageNumber="106" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.36/0.55; head 0.23/0.31; pronotum 0.32/0.45, PA 0.35, PB 0.37; elytra 0.83/0.55. Dorsum of head with frons dark brown to reddish brown, clypeus slightly lighter, labrum testaceous; pronotum brown around ill-defined dark brown macula on disc; elytra dark brown to reddish brown; legs brown to light reddish brown; maxillary palpi testaceous, tip not darker.
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</paragraph>
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<paragraph id="8BA636F35B0CFFFAFF29F132FEFDFC7C" blockId="105.[151,1437,402,1579]" pageId="105" pageNumber="106">Frons punctures ca. 1xef, slightly larger near eyes than medially; interstices shining, 1–3xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum and postmentum micropunctulate, dull. Genae raised, shining, without posterior ridge. Pronotum subcordiform, ca. median 3/4 of anterior margin arcuate to posterior; punctures on disc ca. 1xpd frons punctures, interstices shining, 1–3xpd, punctures slightly larger and denser at anterior and posterior; PF1 very shallow, obsolete; PF2 very shallow, obsolete; PF3 moderately deep; PF4 absent or very shallow.</paragraph>
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<paragraph id="8BA636F35B0CFFFAFF29F1DBFF47FB08" blockId="105.[151,1437,402,1579]" pageId="105" pageNumber="106">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
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<paragraph id="8BA636F35B0CFFFAFF29F687FF67FA0C" blockId="105.[151,1437,402,1579]" pageId="105" pageNumber="106">
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Ratios of P2 width and plaque shape (P2/w/l/s) ca. 4/1/5/5. P1 ca. 1/2 P2; median carina very low, bisinuate in profile. P2 transversely concave, l/w ca. equal, sides slightly diverging toward blunt apex. Plaques very narrow, carinate, anteriorly converging very slightly, at sides of deep median depression. Metaventrite concave between P2 and plaques. AIS width at straight posterior margin ca. 1.7x P2. All legs long and slender. Profemur (male) without tubercle next to trochanter; protibia slightly arcuate, anterior margin very slightly widened subapically. Meso- and metatibia straight. Abdominal apex symmetrical; last tergite (male) deeply notched. Aedeagus as illustrated (
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<figureCitation id="13222A765B0CFFFAFA81F773FF73FA0C" captionStart="FIGURES 184–185" captionStartId="283.[151,264,1650,1673]" captionTargetBox="[290,1288,210,1618]" captionTargetId="figure-29@283.[279,1307,193,1629]" captionTargetPageId="283" captionText="FIGURES 184–185. Aedeagi of holotypes. 184. H. buloba. 185. H. impala." figureDoi="http://doi.org/10.5281/zenodo.5283760" httpUri="https://zenodo.org/record/5283760/files/figure.png" pageId="105" pageNumber="106">Fig. 185</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B0CFFFAFF29F78AFE23FA74" pageId="105" pageNumber="106" type="etymology">
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<paragraph id="8BA636F35B0CFFFAFF29F78AFE23FA74" blockId="105.[151,1437,402,1579]" pageId="105" pageNumber="106">
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<emphasis id="B96DEAE15B0CFFFAFF29F78AFEFBFA29" bold="true" box="[199,333,1447,1471]" inLineHeading="true" pageId="105" pageNumber="106" reason="1">Etymology.</emphasis>
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Named in reference to the relationship to
|
||
<taxonomicName id="4C194D705B0CFFFAFCC6F78AFCCBFA28" authorityName="Perkins" authorityYear="2011" box="[808,893,1446,1470]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="pala">
|
||
<emphasis id="B96DEAE15B0CFFFAFCC6F78AFC8AFA28" box="[808,828,1447,1470]" italics="true" pageId="105" pageNumber="106">H</emphasis>
|
||
.
|
||
<emphasis id="B96DEAE15B0CFFFAFCA5F78BFCCBFA28" box="[843,893,1446,1470]" italics="true" pageId="105" pageNumber="106">pala</emphasis>
|
||
</taxonomicName>
|
||
, but the absence of the prosternal fovea ("pala") present in that species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785B0CFFFAFF29F7C3FE25F9BD" pageId="105" pageNumber="106" type="distribution">
|
||
<paragraph id="8BA636F35B0CFFFAFF29F7C3FE25F9BD" blockId="105.[151,1437,402,1579]" pageId="105" pageNumber="106">
|
||
<emphasis id="B96DEAE15B0CFFFAFF29F7C3FED6F991" bold="true" box="[199,352,1518,1543]" inLineHeading="true" pageId="105" pageNumber="106" reason="1">Distribution.</emphasis>
|
||
Currently known only from the
|
||
<typeStatus id="54A288515B0CFFFAFD3CF7C2FCB4F991" box="[722,770,1519,1543]" pageId="105" pageNumber="106">type</typeStatus>
|
||
locality, the Musgrave River, in central Area 11; elevation
|
||
<quantity id="4CE19B165B0CFFFAFF79F43FFEA0F9BC" box="[151,278,1554,1578]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="105" pageNumber="106" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100–200 m</quantity>
|
||
(
|
||
<figureCitation id="13222A765B0CFFFAFECBF43FFE32F9BC" box="[293,388,1554,1578]" captionStart="FIGURES 531–534" captionStartId="408.[151,264,1650,1673]" captionTargetBox="[165,1421,234,1616]" captionTargetId="figure-23@408.[165,1421,193,1629]" captionTargetPageId="408" captionText="FIGURES 531–534. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284137" httpUri="https://zenodo.org/record/5284137/files/figure.png" pageId="105" pageNumber="106">Fig. 534</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |