181 lines
23 KiB
XML
181 lines
23 KiB
XML
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<mods:title id="91482E49C7C6C86EC252B4AD7B122D46">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="1FBDA8DEA52E15700C39A35E040CE508">Perkins, Philip D.</mods:namePart>
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<mods:date id="17CA7F30C3E584F49A7B9E06FBBC5A85">2011</mods:date>
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<taxonomicName id="4C194D705B0CFFFAFF79F453FEE1F90E" ID-CoL="6MHG7" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,343,1662,1688]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="galea" status="sp. nov.">
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<emphasis id="B96DEAE15B0CFFFAFF79F453FEE1F90E" bold="true" box="[151,343,1662,1688]" italics="true" pageId="105" pageNumber="106">Hydraena galea</emphasis>
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,
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(
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<figureCitation id="13222A765B0CFFFAFF71F48CFEBDF92F" box="[159,267,1697,1721]" captionStart="FIGURE 186" captionStartId="284.[151,250,1753,1776]" captionTargetBox="[224,1366,215,1724]" captionTargetId="figure-29@284.[208,1379,193,1732]" captionTargetPageId="284" captionText="FIGURE 186. Hydraena galea, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283766" httpUri="https://zenodo.org/record/5283766/files/figure.png" pageId="105" pageNumber="106">Figs. 186</figureCitation>
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,
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<figureCitation id="13222A765B0CFFFAFEF8F48CFEF4F92F" box="[278,322,1697,1721]" captionStart="FIGURES 188–189" captionStartId="286.[151,264,1650,1673]" captionTargetBox="[305,1277,206,1620]" captionTargetId="figure-32@286.[288,1298,193,1629]" captionTargetPageId="286" captionText="FIGURES 188–189. Aedeagi of holotypes. 188. H. galea. 189. H. inhalista." figureDoi="http://doi.org/10.5281/zenodo.5283770" httpUri="https://zenodo.org/record/5283770/files/figure.png" pageId="105" pageNumber="106">188</figureCitation>
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,
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<figureCitation id="13222A765B0CFFFAFEA0F48CFECFF92F" box="[334,377,1697,1721]" captionStart="FIGURES 523–526" captionStartId="406.[151,264,1650,1673]" captionTargetBox="[165,1421,227,1566]" captionTargetId="figure-23@406.[165,1421,193,1629]" captionTargetPageId="406" captionText="FIGURES 523–526. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284133" httpUri="https://zenodo.org/record/5284133/files/figure.png" pageId="105" pageNumber="106">523</figureCitation>
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)
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<paragraph id="8BA636F35B0CFFFAFF79F4C7FB77F8FB" blockId="105.[151,1436,1769,2009]" pageId="105" pageNumber="106">
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Type Material.
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<typeStatus id="54A288515B0CFFFAFEA6F4C4FE06F897" box="[328,432,1769,1793]" pageId="105" pageNumber="106" type="holotype">Holotype</typeStatus>
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(male):
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:
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<location id="8EC660285B0CFFFAFC9EF4C4FBFDF897" LSID="urn:lsid:plazi:treatment:03B087E55B0CFFF9FF79F453FD59FBCE:8EC660285B0CFFFAFC9EF4C4FBFDF897" box="[880,1099,1769,1793]" country="Papua New Guinea" latitude="-5.8792334" longLatPrecision="1" longitude="145.38681" municipality="Bismarck Range" name="Akameku-Brahmin" pageId="105" pageNumber="106" stateProvince="Eastern Highlands">Akameku-Brahmin</location>
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,
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<collectingMunicipality id="6BC2AC895B0CFFFAFBB4F4C4FAAFF894" box="[1114,1305,1769,1794]" pageId="105" pageNumber="106">Bismarck Range</collectingMunicipality>
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,
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,
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<date id="FFA710335B0CFFFAFE2EF520FD8CF8B3" box="[448,570,1805,1829]" pageId="105" pageNumber="106" value="2006-11-24">
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<collectingDate id="EFE3E9DB5B0CFFFAFE2EF520FD8CF8B3" box="[448,570,1805,1829]" pageId="105" pageNumber="106" value="2006-11-24">24 xi 2006</collectingDate>
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,
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<collectorName id="26EC53255B0CFFFAFDABF520FD30F8B3" box="[581,646,1805,1829]" pageId="105" pageNumber="106">Balke</collectorName>
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&
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<collectorName id="26EC53255B0CFFFAFD44F520FD48F8B3" box="[682,766,1805,1829]" pageId="105" pageNumber="106">Kinibel</collectorName>
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(
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<collectingCountry id="F30E76635B0CFFFAFCE2F520FCF0F8B3" box="[780,838,1805,1829]" name="Papua New Guinea" pageId="105" pageNumber="106">PNG</collectingCountry>
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109) (
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<collectionCode id="ED08AE365B0CFFFAFC60F520FC7BF8B3" box="[910,973,1805,1829]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="105" pageNumber="106" type="Museum">ZSM</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B713CAE5B0CFFFAFC0EF523FB0BF8FB" collectingDate="1988-07-01" collectingDateMax="1988-08-30" collectingDateMin="1988-07-01" collectionCode="NMW" collectorName="D. Dudgeon" country="Papua New Guinea" elevation="1540" latitude="-6.3" location="River Basin" longLatPrecision="1304" longitude="145.91667" pageId="105" pageNumber="106" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="paratype">
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<typeStatus id="54A288515B0CFFFAFC0EF523FBFBF8B0" box="[992,1101,1806,1830]" pageId="105" pageNumber="106" type="paratype">Paratypes</typeStatus>
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(2):
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||
<collectingRegion id="49DDF8115B0CFFFAFB67F523FF7CF8DF" country="Papua New Guinea" name="Eastern Highlands" pageId="105" pageNumber="106">Eastern Highlands Province</collectingRegion>
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:
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<collectingRegion id="49DDF8115B0CFFFAFF36F51CFEA1F8DF" box="[216,279,1841,1865]" country="Papua New Guinea" name="West Sepik" pageId="105" pageNumber="106">Sepik</collectingRegion>
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<location id="8EC660285B0CFFFAFECCF51CFE1BF8DF" LSID="urn:lsid:plazi:treatment:03B087E55B0CFFF9FF79F453FD59FBCE:8EC660285B0CFFFAFECCF51CFE1BF8DF" box="[290,429,1841,1865]" country="Papua New Guinea" latitude="-6.3" longLatPrecision="1304" longitude="145.91667" name="River Basin" pageId="105" pageNumber="106" stateProvince="Eastern Highlands">River Basin</location>
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, stream beside milestone labelled G-99, unshaded stream with angular boulders, some coated with java moss,
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</quantity>
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,
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<geoCoordinate id="EE2D50345B0CFFFAFDEEF578FDEFF8FB" box="[512,601,1877,1901]" degrees="6" direction="south" minutes="18" orientation="latitude" pageId="105" pageNumber="106" precision="925" value="-6.3">6° 18' S</geoCoordinate>
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,
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<geoCoordinate id="EE2D50345B0CFFFAFD88F578FD6BF8FB" box="[614,733,1877,1901]" degrees="145" direction="east" minutes="55" orientation="longitude" pageId="105" pageNumber="106" precision="925" value="145.91667">145° 55' E</geoCoordinate>
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,
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<date id="FFA710335B0CFFFAFD07F578FC18F8FB" box="[745,942,1877,1901]" pageId="105" pageNumber="106" value="1988-07-01" valueMax="1988-08-30" valueMin="1988-07-01">
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<collectingDate id="EFE3E9DB5B0CFFFAFD07F578FC18F8FB" box="[745,942,1877,1901]" pageId="105" pageNumber="106" value="1988-07-01" valueMax="1988-08-30" valueMin="1988-07-01">1 vii-30 viii 1988</collectingDate>
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</date>
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,
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<collectorName id="26EC53255B0CFFFAFC54F57BFBF5F8FB" box="[954,1091,1877,1901]" pageId="105" pageNumber="106">D. Dudgeon</collectorName>
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(2
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<collectionCode id="ED08AE365B0CFFFAFB86F578FB03F8FB" box="[1128,1205,1877,1901]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="105" pageNumber="106">NMW</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B0CFFF9FF79F554FDEFFF42" lastPageId="106" lastPageNumber="107" pageId="105" pageNumber="106" type="diagnosis">
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<paragraph id="8BA636F35B0CFFF9FF79F554FDEFFF42" blockId="105.[151,1436,1769,2009]" lastBlockId="106.[151,1436,151,1112]" lastPageId="106" lastPageNumber="107" pageId="105" pageNumber="106">
|
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<emphasis id="B96DEAE15B0CFFFAFF79F554FE14F804" bold="true" box="[151,418,1913,1938]" pageId="105" pageNumber="106">Differential Diagnosis.</emphasis>
|
||
Similar in color and dorsal sculpture to
|
||
<taxonomicName id="4C194D705B0CFFFAFC89F557FC6EF807" authorityName="Perkins" authorityYear="2011" box="[871,984,1913,1937]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="impala">
|
||
<emphasis id="B96DEAE15B0CFFFAFC89F557FCCDF807" box="[871,891,1914,1937]" italics="true" pageId="105" pageNumber="106">H</emphasis>
|
||
.
|
||
<emphasis id="B96DEAE15B0CFFFAFC64F554FC6EF807" box="[906,984,1913,1937]" italics="true" pageId="105" pageNumber="106">impala</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13222A765B0CFFFAFC09F554FBE5F807" box="[999,1107,1913,1937]" captionStart="FIGURE 183" captionStartId="282.[151,250,1753,1776]" captionTargetBox="[281,1312,209,1718]" captionTargetId="figure-29@282.[259,1327,193,1732]" captionTargetPageId="282" captionText="FIGURE 183. Hydraena impala, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283758" httpUri="https://zenodo.org/record/5283758/files/figure.png" pageId="105" pageNumber="106">Figs. 183</figureCitation>
|
||
,
|
||
<figureCitation id="13222A765B0CFFFAFBB1F554FB3BF807" box="[1119,1165,1913,1937]" captionStart="FIGURE 186" captionStartId="284.[151,250,1753,1776]" captionTargetBox="[224,1366,215,1724]" captionTargetId="figure-29@284.[208,1379,193,1732]" captionTargetPageId="284" captionText="FIGURE 186. Hydraena galea, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283766" httpUri="https://zenodo.org/record/5283766/files/figure.png" pageId="105" pageNumber="106">186</figureCitation>
|
||
); differing therefrom by the slightly wider body form (EW/TL ca. 0.45 vs. 0.40), the wider mesoventral intercoxal process (P2) (plaque ratios ca. 5/1/5/5 vs. 4/1/5/5), and the non-micropunctulate mentum. The general plan of the male genitalia of the two species also suggests a relationship, but the genitalia distinctively differ in details, and should be examined for reliable determinations (
|
||
<figureCitation id="13222A765B0FFFF9FE49F291FDA5FF42" box="[423,531,188,212]" captionStart="FIGURES 184–185" captionStartId="283.[151,264,1650,1673]" captionTargetBox="[290,1288,210,1618]" captionTargetId="figure-29@283.[279,1307,193,1629]" captionTargetPageId="283" captionText="FIGURES 184–185. Aedeagi of holotypes. 184. H. buloba. 185. H. impala." figureDoi="http://doi.org/10.5281/zenodo.5283760" httpUri="https://zenodo.org/record/5283760/files/figure.png" pageId="106" pageNumber="107">Figs. 185</figureCitation>
|
||
,
|
||
<figureCitation id="13222A765B0FFFF9FDF1F291FDFBFF42" box="[543,589,188,212]" captionStart="FIGURES 188–189" captionStartId="286.[151,264,1650,1673]" captionTargetBox="[305,1277,206,1620]" captionTargetId="figure-32@286.[288,1298,193,1629]" captionTargetPageId="286" captionText="FIGURES 188–189. Aedeagi of holotypes. 188. H. galea. 189. H. inhalista." figureDoi="http://doi.org/10.5281/zenodo.5283770" httpUri="https://zenodo.org/record/5283770/files/figure.png" pageId="106" pageNumber="107">188</figureCitation>
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||
).
|
||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="C30365785B0FFFF9FF29F2F2FC6BFC7A" pageId="106" pageNumber="107" type="description">
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<paragraph id="8BA636F35B0FFFF9FF29F2F2FACAFEF2" blockId="106.[151,1436,151,1112]" pageId="106" pageNumber="107">
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<emphasis id="B96DEAE15B0FFFF9FF29F2F2FEEEFF6E" bold="true" box="[199,344,223,248]" inLineHeading="true" pageId="106" pageNumber="107" reason="1">Description.</emphasis>
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Size:
|
||
<typeStatus id="54A288515B0FFFF9FE42F2F2FDB8FF61" box="[428,526,223,247]" pageId="106" pageNumber="107" type="holotype">holotype</typeStatus>
|
||
(length/width, mm): body (length to elytral apices) 1.45/0.65; head 0.19/0.34; pronotum 0.35/0.49, PA 0.35, PB 0.38; elytra 0.92/0.65. Dorsum of head with frons dark brown to dark reddish brown, clypeus lighter, brown, labrum testaceous; pronotum brown around ill-defined dark reddish brown macula on disc; elytra dark brown to dark reddish brown; legs reddish brown; maxillary palpi testaceous, tip not darker.
|
||
</paragraph>
|
||
<paragraph id="8BA636F35B0FFFF9FF29F35DFDA7FDAA" blockId="106.[151,1436,151,1112]" pageId="106" pageNumber="107">Frons punctures ca. 1–2xef, larger and denser near eyes than medially; interstices shining, 1–3xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum very sparsely very finely punctulate, shining; postmentum microreticulate. Genae raised, shining, without posterior ridge. Pronotum subcordiform, ca. median 3/4 of anterior margin arcuate to posterior; punctures on disc ca. 1xpd frons punctures, interstices shining, 1–3xpd, punctures larger and denser at anterior and posterior; PF1 and PF2 very shallow, obsolete; PF3 moderately deep; PF4 absent or very shallow.</paragraph>
|
||
<paragraph id="8BA636F35B0FFFF9FF29F06AFE78FD79" blockId="106.[151,1436,151,1112]" pageId="106" pageNumber="107">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins rather wide in anterior 1/2; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 2–3xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
|
||
<paragraph id="8BA636F35B0FFFF9FF29F0D1FC6BFC7A" blockId="106.[151,1436,151,1112]" pageId="106" pageNumber="107">
|
||
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 5/1/5/5. P1 ca. 2/5 P2; median carina bisinuate in profile. P2 l/w ca. 4/5, sides slightly diverging toward blunt apex. Plaques very narrow, carinate, converging slightly anteriorly, at sides of deep median depression. Metaventrite concave between P2 and plaques. AIS width at straight posterior margin ca. 1.4x P2. All legs moderately long and slender. Profemur (male) with small tubercle next to trochanter; protibia slightly arcuate, anterior margin very slightly widened subapically, medial margin flat except small sharp tubercle subapically on posteromedial margin. Meso- and metatibia straight. Abdominal apex symmetrical; last tergite (male) deeply notched. Aedeagus as illustrated (
|
||
<figureCitation id="13222A765B0FFFF9FC80F1F9FC67FC7A" box="[878,977,980,1004]" captionStart="FIGURES 188–189" captionStartId="286.[151,264,1650,1673]" captionTargetBox="[305,1277,206,1620]" captionTargetId="figure-32@286.[288,1298,193,1629]" captionTargetPageId="286" captionText="FIGURES 188–189. Aedeagi of holotypes. 188. H. galea. 189. H. inhalista." figureDoi="http://doi.org/10.5281/zenodo.5283770" httpUri="https://zenodo.org/record/5283770/files/figure.png" pageId="106" pageNumber="107">Fig. 188</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785B0FFFF9FF29F1D5FB2DFB99" box="[199,1179,1015,1040]" pageId="106" pageNumber="107" type="etymology">
|
||
<paragraph id="8BA636F35B0FFFF9FF29F1D5FB2DFB99" blockId="106.[151,1436,151,1112]" box="[199,1179,1015,1040]" pageId="106" pageNumber="107">
|
||
<emphasis id="B96DEAE15B0FFFF9FF29F1D5FEFBFB86" bold="true" box="[199,333,1016,1040]" inLineHeading="true" pageId="106" pageNumber="107" reason="1">Etymology.</emphasis>
|
||
Named in reference to the helmet shape of the distal piece of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785B0FFFF9FF29F636FD59FBCE" pageId="106" pageNumber="107" type="distribution">
|
||
<paragraph id="8BA636F35B0FFFF9FF29F636FD59FBCE" blockId="106.[151,1436,151,1112]" pageId="106" pageNumber="107">
|
||
<emphasis id="B96DEAE15B0FFFF9FF29F636FED6FBA2" bold="true" box="[199,352,1051,1076]" inLineHeading="true" pageId="106" pageNumber="107" reason="1">Distribution.</emphasis>
|
||
Currently known from two localities at the border of the eastern parts of Areas 1 and 2, at very different elevations of
|
||
<quantity id="4CE19B165B0FFFF9FE7AF66DFE6DFBC1" box="[404,475,1088,1112]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="106" pageNumber="107" unit="m" value="700.0">700 m</quantity>
|
||
and
|
||
<quantity id="4CE19B165B0FFFF9FDFFF66DFDD0FBC1" box="[529,614,1088,1112]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="106" pageNumber="107" unit="m" value="1540.0">1540 m</quantity>
|
||
(
|
||
<figureCitation id="13222A765B0FFFF9FD98F66DFD56FBCE" box="[630,736,1088,1112]" captionStart="FIGURES 523–526" captionStartId="406.[151,264,1650,1673]" captionTargetBox="[165,1421,227,1566]" captionTargetId="figure-23@406.[165,1421,193,1629]" captionTargetPageId="406" captionText="FIGURES 523–526. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284133" httpUri="https://zenodo.org/record/5284133/files/figure.png" pageId="106" pageNumber="107">Figs. 523</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |