159 lines
20 KiB
XML
159 lines
20 KiB
XML
<document id="E20BF43D27ABC8E8E27FC50DDEC5922D" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B08FFFDFF79F477FD5FFBA2" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena confluenta Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="111" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="110" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="08405ADC24149273E30748CDEC804331">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="A56CEA58C843170516CBB7CCB138C512">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="937306088F61DC1A0349B782A6967314">text</mods:typeOfResource>
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<mods:title id="3F108116F2853B3AC8491F2DFE698668">Zootaxa</mods:title>
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<mods:part id="798031578B0A4476FAED3749AA597A81">
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<mods:date id="FCCFC83A6333D4FB78ED1F83FFCB3EEE">2011</mods:date>
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<mods:number id="B2510470D5052869E1882B3520B3916F">2011-07-08</mods:number>
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<mods:number id="2B1BB1B96AABFC8E422EB477E4E8B42F">2944</mods:number>
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<mods:url id="32A540D8CFF6E677069D0F6CE6D10163">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
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<mods:classification id="45F9BBE78C9B00A96D75444BA8BDE4D2">journal article</mods:classification>
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<mods:identifier id="F15AF60019CB7FA056A861F45F8AB86C" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
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<mods:identifier id="DEEF5977CD92EED772B9B4615AC0D6CD" type="ISSN">11755334</mods:identifier>
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<treatment id="03B087E55B08FFFDFF79F477FD5FFBA2" ID-DOI="http://doi.org/10.5281/zenodo.5291733" ID-Zenodo-Dep="5291733" LSID="urn:lsid:plazi:treatment:03B087E55B08FFFDFF79F477FD5FFBA2" httpUri="http://treatment.plazi.org/id/03B087E55B08FFFDFF79F477FD5FFBA2" lastPageId="110" lastPageNumber="111" pageId="109" pageNumber="110">
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<subSubSection id="C30365785B08FFFEFF79F477FD81F9E2" box="[151,567,1626,1652]" pageId="109" pageNumber="110" type="nomenclature">
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<paragraph id="8BA636F35B08FFFEFF79F477FD81F9E2" blockId="109.[151,567,1626,1685]" box="[151,567,1626,1652]" pageId="109" pageNumber="110">
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<heading id="D0EE819F5B08FFFEFF79F477FD81F9E2" bold="true" box="[151,567,1626,1652]" fontSize="11" level="1" pageId="109" pageNumber="110" reason="1">
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<emphasis id="B96DEAE15B08FFFEFF79F477FD81F9E2" bold="true" box="[151,567,1626,1652]" pageId="109" pageNumber="110">
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<taxonomicName id="4C194D705B08FFFEFF79F477FE2EF9E2" ID-CoL="3N382" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,408,1626,1652]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="confluenta" status="sp. nov.">
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<emphasis id="B96DEAE15B08FFFEFF79F477FE2EF9E2" bold="true" box="[151,408,1626,1652]" italics="true" pageId="109" pageNumber="110">Hydraena confluenta</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25E579A5B08FFFEFE49F477FD81F9E2" box="[423,567,1626,1652]" pageId="109" pageNumber="110" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B08FFFEFF79F450FE34F903" box="[151,386,1661,1685]" pageId="109" pageNumber="110" type="description">
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<paragraph id="8BA636F35B08FFFEFF79F450FE34F903" blockId="109.[151,567,1626,1685]" box="[151,386,1661,1685]" pageId="109" pageNumber="110">
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(
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<figureCitation id="13222A765B08FFFEFF71F450FEBDF903" box="[159,267,1661,1685]" captionStart="FIGURE 195" captionStartId="291.[151,250,1705,1728]" captionTargetBox="[263,1387,220,1679]" captionTargetId="figure-25@291.[186,1401,193,1684]" captionTargetPageId="291" captionText="FIGURE 195. Hydraena confluenta, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283790" httpUri="https://zenodo.org/record/5283790/files/figure.png" pageId="109" pageNumber="110">Figs. 195</figureCitation>
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,
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<figureCitation id="13222A765B08FFFEFEF8F450FEF4F903" box="[278,322,1661,1685]" captionStart="FIGURES 196–197" captionStartId="292.[151,264,1650,1673]" captionTargetBox="[310,1275,212,1616]" captionTargetId="figure-32@292.[264,1323,193,1629]" captionTargetPageId="292" captionText="FIGURES 196–197. Aedeagi of holotypes. 196. H. pacificica. 197. H. confluenta." figureDoi="http://doi.org/10.5281/zenodo.5283792" httpUri="https://zenodo.org/record/5283792/files/figure.png" pageId="109" pageNumber="110">197</figureCitation>
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,
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<figureCitation id="13222A765B08FFFEFEA0F450FECFF903" box="[334,377,1661,1685]" captionStart="FIGURES 523–526" captionStartId="406.[151,264,1650,1673]" captionTargetBox="[165,1421,227,1566]" captionTargetId="figure-23@406.[165,1421,193,1629]" captionTargetPageId="406" captionText="FIGURES 523–526. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284133" httpUri="https://zenodo.org/record/5284133/files/figure.png" pageId="109" pageNumber="110">526</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B08FFFEFF79F4E8FBBBF897" pageId="109" pageNumber="110" type="materials_examined">
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<paragraph id="8BA636F35B08FFFEFF79F4E8FBBBF897" blockId="109.[151,1436,1733,2009]" pageId="109" pageNumber="110">
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Type Material.
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<materialsCitation id="3B713CAE5B08FFFEFEA1F4E8FBBBF897" collectingDate="1980-03-01" collectingDateMax="1980-03-30" collectingDateMin="1980-03-01" collectionCode="MHNG" collectorName="W. G. Ullrich" elevation="1600" latitude="-6.2833333" location="Onerunka" longLatPrecision="1304" longitude="145.86667" municipality="Kainantu" pageId="109" pageNumber="110" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="holotype">
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<typeStatus id="54A288515B08FFFEFEA1F4E8FE0AF94B" box="[335,444,1733,1757]" pageId="109" pageNumber="110" type="holotype">
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<emphasis id="B96DEAE15B08FFFEFEA1F4E8FE0AF94B" bold="true" box="[335,444,1733,1757]" pageId="109" pageNumber="110">Holotype</emphasis>
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</typeStatus>
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(male):
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<emphasis id="B96DEAE15B08FFFEFDCAF4E8FCCBF948" bold="true" box="[548,893,1733,1758]" pageId="109" pageNumber="110">
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<collectingRegion id="49DDF8115B08FFFEFDCAF4E8FCCFF948" box="[548,889,1733,1758]" country="Papua New Guinea" name="Eastern Highlands" pageId="109" pageNumber="110">Eastern Highlands Province</collectingRegion>
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||
:
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</emphasis>
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Umg. [=environs of]
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<collectingMunicipality id="6BC2AC895B08FFFEFB92F4E8FB53F94B" box="[1148,1253,1733,1757]" pageId="109" pageNumber="110">Kainantu</collectingMunicipality>
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||
,
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||
<location id="8EC660285B08FFFEFB1EF4E8FAD7F94B" LSID="urn:lsid:plazi:treatment:03B087E55B08FFFDFF79F477FD5FFBA2:8EC660285B08FFFEFB1EF4E8FAD7F94B" box="[1264,1377,1733,1757]" latitude="-6.2833333" longLatPrecision="1304" longitude="145.86667" municipality="Kainantu" name="Onerunka" pageId="109" pageNumber="110" stateProvince="Eastern Highlands">Onerunka</location>
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||
, [GE est.]
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<quantity id="4CE19B165B08FFFEFF23F4C4FEDCF897" box="[205,362,1769,1793]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="109" pageNumber="110" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">
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<elevation id="0034D1C05B08FFFEFF23F4C4FEDCF897" box="[205,362,1769,1793]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="109" pageNumber="110" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">1500–1700 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE2D50345B08FFFEFE98F4C4FE79F897" box="[374,463,1769,1793]" degrees="6" direction="south" minutes="17" orientation="latitude" pageId="109" pageNumber="110" precision="925" value="-6.2833333">6° 17' S</geoCoordinate>
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,
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<geoCoordinate id="EE2D50345B08FFFEFE32F4C4FDE5F897" box="[476,595,1769,1793]" degrees="145" direction="east" minutes="52" orientation="longitude" pageId="109" pageNumber="110" precision="925" value="145.86667">145° 52' E</geoCoordinate>
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,
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<date id="FFA710335B08FFFEFDB1F4C4FD40F897" box="[607,758,1769,1793]" pageId="109" pageNumber="110" value="1980-03-01" valueMax="1980-03-30" valueMin="1980-03-01">
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<collectingDate id="EFE3E9DB5B08FFFEFDB1F4C4FD40F897" box="[607,758,1769,1793]" pageId="109" pageNumber="110" value="1980-03-01" valueMax="1980-03-30" valueMin="1980-03-01">1–30 iii 1980</collectingDate>
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</date>
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,
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||
<collectorName id="26EC53255B08FFFEFCECF4C7FC20F897" box="[770,918,1769,1793]" pageId="109" pageNumber="110">W. G. Ullrich</collectorName>
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||
(
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||
<collectionCode id="ED08AE365B08FFFEFC4BF4C4FBB7F897" box="[933,1025,1769,1793]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="109" pageNumber="110" type="Museum">MHNG</collectionCode>
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||
).
|
||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30365785B08FFFEFF29F520FE48F84F" pageId="109" pageNumber="110" type="diagnosis">
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||
<paragraph id="8BA636F35B08FFFEFF29F520FE48F84F" blockId="109.[151,1436,1733,2009]" pageId="109" pageNumber="110">
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||
<emphasis id="B96DEAE15B08FFFEFF29F520FE65F8B0" bold="true" box="[199,467,1805,1830]" pageId="109" pageNumber="110">Differential Diagnosis.</emphasis>
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Similar in color and dorsal sculpture to
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||
<taxonomicName id="4C194D705B08FFFEFC70F523FB92F8B3" authorityName="Perkins" authorityYear="2011" box="[926,1060,1806,1829]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="processa">
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||
<emphasis id="B96DEAE15B08FFFEFC70F523FC04F8B3" box="[926,946,1806,1829]" italics="true" pageId="109" pageNumber="110">H</emphasis>
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||
.
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<emphasis id="B96DEAE15B08FFFEFC2CF523FB92F8B3" box="[962,1060,1806,1829]" italics="true" pageId="109" pageNumber="110">processa</emphasis>
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</taxonomicName>
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||
(
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||
<figureCitation id="13222A765B08FFFEFBDAF520FB17F8B3" box="[1076,1185,1805,1829]" captionStart="FIGURE 195" captionStartId="291.[151,250,1705,1728]" captionTargetBox="[263,1387,220,1679]" captionTargetId="figure-25@291.[186,1401,193,1684]" captionTargetPageId="291" captionText="FIGURE 195. Hydraena confluenta, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283790" httpUri="https://zenodo.org/record/5283790/files/figure.png" pageId="109" pageNumber="110">Figs. 195</figureCitation>
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,
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<figureCitation id="13222A765B08FFFEFB40F520FB6AF8B3" box="[1198,1244,1805,1829]" captionStart="FIGURE 198" captionStartId="293.[151,250,1753,1776]" captionTargetBox="[288,1306,226,1718]" captionTargetId="figure-25@293.[261,1325,193,1732]" captionTargetPageId="293" captionText="FIGURE 198. Hydraena processa, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283798" httpUri="https://zenodo.org/record/5283798/files/figure.png" pageId="109" pageNumber="110">198</figureCitation>
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); differing therefrom by the larger size (TL ca. 1.63 vs.
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<quantity id="4CE19B165B08FFFEFDB6F51CFD76F8DF" box="[600,704,1841,1865]" metricMagnitude="-3" metricUnit="m" metricValue="1.48" pageId="109" pageNumber="110" unit="mm" value="1.48">1.48 mm</quantity>
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), the slightly narrower body form (EW/TL ca. 0.44 vs. 0.47), the wider mesoventral intercoxal process (P2), the more widely spaced plaques (plaque ratios ca. 6/1/5/8 vs. 5/1/5/7), and the confluently concave AIS and first abdominal ventrite. The general plan of the male genitalia of the two species also suggests a relationship, but the genitalia distinctively differ in details, and should be examined for reliable determinations (
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||
<figureCitation id="13222A765B08FFFEFEA2F5ECFE0EF84F" box="[332,440,1985,2009]" captionStart="FIGURES 196–197" captionStartId="292.[151,264,1650,1673]" captionTargetBox="[310,1275,212,1616]" captionTargetId="figure-32@292.[264,1323,193,1629]" captionTargetPageId="292" captionText="FIGURES 196–197. Aedeagi of holotypes. 196. H. pacificica. 197. H. confluenta." figureDoi="http://doi.org/10.5281/zenodo.5283792" httpUri="https://zenodo.org/record/5283792/files/figure.png" pageId="109" pageNumber="110">Figs. 197</figureCitation>
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||
,
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<figureCitation id="13222A765B08FFFEFE2AF5ECFE44F84F" box="[452,498,1985,2009]" captionStart="FIGURES 200–201" captionStartId="295.[151,264,1650,1673]" captionText="FIGURES 200–201. Aedeagi of holotypes. 200. H. processa. 201. H. bicarinova." pageId="109" pageNumber="110">200</figureCitation>
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).
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785B0BFFFDFF29F2BAFDC9FC51" pageId="110" pageNumber="111" type="description">
|
||
<paragraph id="8BA636F35B0BFFFDFF29F2BAFBB4FF61" blockId="110.[151,1436,151,1076]" pageId="110" pageNumber="111">
|
||
<emphasis id="B96DEAE15B0BFFFDFF29F2BAFEEEFF26" bold="true" box="[199,344,151,176]" inLineHeading="true" pageId="110" pageNumber="111" reason="1">Description.</emphasis>
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||
Size:
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||
<typeStatus id="54A288515B0BFFFDFE42F2BAFDB8FF39" box="[428,526,151,175]" pageId="110" pageNumber="111" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.63/0.72; head 0.28/0.37; pronotum 0.38/0.52, PA 0.40, PB 0.44; elytra 0.97/0.72. Dorsum dark reddish brown, frons darkest, clypeus and labrum lightest; legs reddish brown; maxillary palpi testaceous, tip not darker.
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</paragraph>
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<paragraph id="8BA636F35B0BFFFDFF29F329FF42FE62" blockId="110.[151,1436,151,1076]" pageId="110" pageNumber="111">Frons punctures ca. 1–2xef, larger and denser near eyes than medially; interstices shining, 1–3xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum and postmentum very sparsely very finely punctulate, shining. Genae raised, shining, without posterior ridge. Pronotum subcordiform, ca. median 3/4 of anterior margin arcuate to posterior; punctures on disc slightly smaller than smallest frons punctures, interstices shining, 2–4xpd, punctures larger and denser at anterior and posterior; disc slightly longitudinally convex; PF1 very shallow, obsolete; PF2 shallow, medially confluent, forming U-shaped impression; PF3 deep; PF4 absent or very shallow.</paragraph>
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<paragraph id="8BA636F35B0BFFFDFF29F3D2FF47FD31" blockId="110.[151,1436,151,1076]" pageId="110" pageNumber="111">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins rather wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 2–3xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
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||
<paragraph id="8BA636F35B0BFFFDFF29F099FDC9FC51" blockId="110.[151,1436,151,1076]" pageId="110" pageNumber="111">
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||
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 6/1/5/8. P1 ca. 2/3 P2; median carina bisinuate in profile. P2 transversely slightly concave, l/w ca. 5/6, sides slightly diverging toward blunt apex. Plaques very narrow, carinate, converging slightly anteriorly, at sides of deep median depression. Metaventrite concave between P2 and plaques. AIS conjointly concave with anteromedial area of first ventrite, width at straight posterior margin ca. 1.3x P2. All legs moderately long and slender. Profemur (male) with small tubercle next to trochanter; protibia slightly arcuate, anterior margin very slightly widened subapically, medial margin flat except small sharp tubercle subapically on posteromedial margin. Meso- and metatibia straight. Abdominal apex symmetrical; last tergite (male) deeply notched. Aedeagus as illustrated (
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||
<figureCitation id="13222A765B0BFFFDFDFFF182FDC5FC51" box="[529,627,943,967]" captionStart="FIGURES 196–197" captionStartId="292.[151,264,1650,1673]" captionTargetBox="[310,1275,212,1616]" captionTargetId="figure-32@292.[264,1323,193,1629]" captionTargetPageId="292" captionText="FIGURES 196–197. Aedeagi of holotypes. 196. H. pacificica. 197. H. confluenta." figureDoi="http://doi.org/10.5281/zenodo.5283792" httpUri="https://zenodo.org/record/5283792/files/figure.png" pageId="110" pageNumber="111">Fig. 197</figureCitation>
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||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785B0BFFFDFF29F1F9FB6FFC7A" box="[199,1241,980,1004]" pageId="110" pageNumber="111" type="etymology">
|
||
<paragraph id="8BA636F35B0BFFFDFF29F1F9FB6FFC7A" blockId="110.[151,1436,151,1076]" box="[199,1241,980,1004]" pageId="110" pageNumber="111">
|
||
<emphasis id="B96DEAE15B0BFFFDFF29F1F9FEFBFC7A" bold="true" box="[199,333,980,1004]" inLineHeading="true" pageId="110" pageNumber="111" reason="1">Etymology.</emphasis>
|
||
Named in reference to the confluently concave AIS and first abdominal ventrite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785B0BFFFDFF29F1DAFD5FFBA2" pageId="110" pageNumber="111" type="distribution">
|
||
<paragraph id="8BA636F35B0BFFFDFF29F1DAFD5FFBA2" blockId="110.[151,1436,151,1076]" pageId="110" pageNumber="111">
|
||
<emphasis id="B96DEAE15B0BFFFDFF29F1DAFED6FB86" bold="true" box="[199,352,1015,1040]" inLineHeading="true" pageId="110" pageNumber="111" reason="1">Distribution.</emphasis>
|
||
Currently known only from the
|
||
<typeStatus id="54A288515B0BFFFDFD22F1D5FD4AFB86" box="[716,764,1016,1040]" pageId="110" pageNumber="111">type</typeStatus>
|
||
locality near Onerunka, in the eastern part of Area 1, where it borders Area 2; elevation
|
||
<quantity id="4CE19B165B0BFFFDFE3EF631FDDDFBA5" box="[464,619,1052,1076]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="110" pageNumber="111" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">1500–1700 m</quantity>
|
||
(
|
||
<figureCitation id="13222A765B0BFFFDFD94F631FD6CFBA2" box="[634,730,1052,1076]" captionStart="FIGURES 523–526" captionStartId="406.[151,264,1650,1673]" captionTargetBox="[165,1421,227,1566]" captionTargetId="figure-23@406.[165,1421,193,1629]" captionTargetPageId="406" captionText="FIGURES 523–526. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284133" httpUri="https://zenodo.org/record/5284133/files/figure.png" pageId="110" pageNumber="111">Fig. 526</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |