treatments-xml/data/03/B0/87/03B087E55B07FFF0FF79F48FFBE3FA9A.xml
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<document id="0C9AAD0B6127A57B87FA0E7EE5EC912F" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B07FFF0FF79F48FFBE3FA9A" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena transvallis Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="100" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="99" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9A699E104DDB6EDEAC60D27D162D29A4">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="33810D7FBC1A19D43D67308BC7CEE2BE">Perkins, Philip D.</mods:namePart>
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<mods:date id="D860C477213A3F010DB2991BF2BCC160">2011</mods:date>
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<mods:classification id="518636338237E7E1968504B425EC3C09">journal article</mods:classification>
<mods:identifier id="C34B72AACAD74AF249FDFEAE15010389" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
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<treatment id="03B087E55B07FFF0FF79F48FFBE3FA9A" LSID="urn:lsid:plazi:treatment:03B087E55B07FFF0FF79F48FFBE3FA9A" httpUri="http://treatment.plazi.org/id/03B087E55B07FFF0FF79F48FFBE3FA9A" lastPageId="99" lastPageNumber="100" pageId="98" pageNumber="99">
<subSubSection id="C30365785B07FFF1FF79F48FFD85F92A" box="[151,563,1698,1724]" pageId="98" pageNumber="99" type="nomenclature">
<paragraph id="8BA636F35B07FFF1FF79F48FFD85F92A" blockId="98.[151,563,1698,1757]" box="[151,563,1698,1724]" pageId="98" pageNumber="99">
<heading id="D0EE819F5B07FFF1FF79F48FFD85F92A" bold="true" box="[151,563,1698,1724]" fontSize="11" level="1" pageId="98" pageNumber="99" reason="1">
<emphasis id="B96DEAE15B07FFF1FF79F48FFD85F92A" bold="true" box="[151,563,1698,1724]" pageId="98" pageNumber="99">
<taxonomicName id="4C194D705B07FFF1FF79F48FFE25F92A" ID-CoL="3N3JW" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,403,1698,1724]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="transvallis" status="sp. nov.">
<emphasis id="B96DEAE15B07FFF1FF79F48FFE25F92A" bold="true" box="[151,403,1698,1724]" italics="true" pageId="98" pageNumber="99">Hydraena transvallis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B07FFF1FE4DF48FFD85F92A" box="[419,563,1698,1724]" pageId="98" pageNumber="99" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B07FFF1FF79F4E8FE34F94B" box="[151,386,1733,1757]" pageId="98" pageNumber="99" type="description">
<paragraph id="8BA636F35B07FFF1FF79F4E8FE34F94B" blockId="98.[151,563,1698,1757]" box="[151,386,1733,1757]" pageId="98" pageNumber="99">
(
<figureCitation id="13222A765B07FFF1FF71F4E8FEBDF94B" box="[159,267,1733,1757]" captionStart="FIGURE 171" captionStartId="273.[151,250,1753,1776]" captionTargetBox="[219,1367,214,1720]" captionTargetId="figure-25@273.[200,1387,193,1732]" captionTargetPageId="273" captionText="FIGURE 171. Hydraena transvallis, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283738" httpUri="https://zenodo.org/record/5283738/files/figure.png" pageId="98" pageNumber="99">Figs. 171</figureCitation>
,
<figureCitation id="13222A765B07FFF1FEF8F4E8FEF4F94B" box="[278,322,1733,1757]" captionStart="FIGURES 172173" captionStartId="274.[151,264,1650,1673]" captionText="FIGURES 172173. Aedeagi of holotypes. 172. H. implacopaca. 173. H. transvallis." pageId="98" pageNumber="99">173</figureCitation>
,
<figureCitation id="13222A765B07FFF1FEA0F4E8FECFF94B" box="[334,377,1733,1757]" captionStart="FIGURES 503506" captionStartId="401.[151,264,1650,1673]" captionTargetBox="[165,1421,222,1609]" captionTargetId="figure-20@401.[165,1421,193,1629]" captionTargetPageId="401" captionText="FIGURES 503506. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5299476" httpUri="https://zenodo.org/record/5299476/files/figure.png" pageId="98" pageNumber="99">506</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B07FFF1FF79F520FB87F8FB" pageId="98" pageNumber="99" type="materials_examined">
<paragraph id="8BA636F35B07FFF1FF79F520FB87F8FB" blockId="98.[151,1436,1805,2009]" pageId="98" pageNumber="99">
Type Material.
<materialsCitation id="3B713CAE5B07FFF1FEBFF520FCD4F8DF" collectingDate="1964-10-02" collectingDateMax="1964-10-07" collectingDateMin="1964-10-02" collectionCode="NHM" collectorName="M. E. Bacchus" county="Finisterre Mts." elevation="1082" latitude="-5.9166665" location="Damanti" longLatPrecision="1305" longitude="145.96666" municipality="Naho River Valley" pageId="98" pageNumber="99" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<typeStatus id="54A288515B07FFF1FEBFF520FE08F8B3" box="[337,446,1805,1829]" pageId="98" pageNumber="99" type="holotype">
<emphasis id="B96DEAE15B07FFF1FEBFF520FE08F8B3" bold="true" box="[337,446,1805,1829]" pageId="98" pageNumber="99">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B07FFF1FDC6F520FCB0F8B0" bold="true" box="[552,774,1805,1830]" pageId="98" pageNumber="99">
<collectingRegion id="49DDF8115B07FFF1FDC6F520FCB7F8B0" box="[552,769,1805,1830]" country="Papua New Guinea" name="Madang" pageId="98" pageNumber="99">Madang Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62C74E7F5B07FFF1FCF9F520FC76F8B3" box="[791,960,1805,1829]" pageId="98" pageNumber="99">Finisterre Mts.</collectingCounty>
,
<collectingMunicipality id="6BC2AC895B07FFF1FC23F520FB17F8B3" box="[973,1185,1805,1829]" pageId="98" pageNumber="99">Naho River Valley</collectingMunicipality>
,
<location id="8EC660285B07FFF1FB43F520FAA6F8B3" LSID="urn:lsid:plazi:treatment:03B087E55B07FFF0FF79F48FFBE3FA9A:8EC660285B07FFF1FB43F520FAA6F8B3" box="[1197,1296,1805,1829]" county="Finisterre Mts." latitude="-5.9166665" longLatPrecision="1305" longitude="145.96666" municipality="Naho River Valley" name="Damanti" pageId="98" pageNumber="99" stateProvince="Madang">Damanti</location>
,
<quantity id="4CE19B165B07FFF1FAF3F520FAC3F8B3" box="[1309,1397,1805,1829]" metricMagnitude="3" metricUnit="m" metricValue="1.082" pageId="98" pageNumber="99" unit="m" value="1082.0">
<elevation id="0034D1C05B07FFF1FAF3F520FAC3F8B3" box="[1309,1397,1805,1829]" metricMagnitude="3" metricUnit="m" metricValue="1.082" pageId="98" pageNumber="99" unit="m" value="1082.0">1082 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B07FFF1FA6DF520FF79F8DF" degrees="5" direction="south" minutes="55" orientation="latitude" pageId="98" pageNumber="99" precision="925" value="-5.9166665">5° 55' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B07FFF1FF35F51CFEE4F8DF" box="[219,338,1841,1865]" degrees="145" direction="east" minutes="58" orientation="longitude" pageId="98" pageNumber="99" precision="925" value="145.96666">145° 58' E</geoCoordinate>
,
<date id="FFA710335B07FFF1FEB3F51CFE6AF8DF" box="[349,476,1841,1865]" pageId="98" pageNumber="99" value="1964-10-02" valueMax="1964-10-07" valueMin="1964-10-02">
<collectingDate id="EFE3E9DB5B07FFF1FEB3F51CFE6AF8DF" box="[349,476,1841,1865]" pageId="98" pageNumber="99" value="1964-10-02" valueMax="1964-10-07" valueMin="1964-10-02">27 x 1964</collectingDate>
</date>
,
<collectorName id="26EC53255B07FFF1FE09F51FFD3AF8DF" box="[487,652,1841,1865]" pageId="98" pageNumber="99">M. E. Bacchus</collectorName>
(MEB 30) (
<collectionCode id="ED08AE365B07FFF1FCFCF51CFCECF8DF" box="[786,858,1841,1865]" pageId="98" pageNumber="99">NHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CAE5B07FFF1FC82F51CFB9BF8FB" collectingDate="1961-06-05" collectionCode="BPBM" collectorName="J. L. &amp; M. Gressitt" elevation="1000" latitude="-6.5" location="Goroka" longLatPrecision="1304" longitude="144.85" municipality="Karimui" pageId="98" pageNumber="99" specimenCount="1" stateProvince="Chimbu" typeStatus="paratype">
<emphasis id="B96DEAE15B07FFF1FC82F51CFC6FF8DF" bold="true" box="[876,985,1841,1865]" pageId="98" pageNumber="99">
<typeStatus id="54A288515B07FFF1FC82F51CFC6FF8DF" box="[876,985,1841,1865]" pageId="98" pageNumber="99" type="paratype">Paratype</typeStatus>
</emphasis>
:
<emphasis id="B96DEAE15B07FFF1FC00F51CFB02F8DC" bold="true" box="[1006,1204,1841,1866]" pageId="98" pageNumber="99">
<collectingRegion id="49DDF8115B07FFF1FC00F51CFB06F8DC" box="[1006,1200,1841,1866]" country="Papua New Guinea" name="Chimbu" pageId="98" pageNumber="99">Simbu Province</collectingRegion>
:
</emphasis>
NE,
<collectingMunicipality id="6BC2AC895B07FFF1FB1AF51CFAE2F8DF" box="[1268,1364,1841,1865]" pageId="98" pageNumber="99">Karimui</collectingMunicipality>
, south of
<location id="8EC660285B07FFF1FF5BF578FEBAF8FB" LSID="urn:lsid:plazi:treatment:03B087E55B07FFF0FF79F48FFBE3FA9A:8EC660285B07FFF1FF5BF578FEBAF8FB" box="[181,268,1877,1901]" latitude="-6.5" longLatPrecision="1304" longitude="144.85" municipality="Karimui" name="Goroka" pageId="98" pageNumber="99" stateProvince="Chimbu">Goroka</location>
,
<quantity id="4CE19B165B07FFF1FEF9F578FED8F8FB" box="[279,366,1877,1901]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="98" pageNumber="99" unit="m" value="1000.0">
<elevation id="0034D1C05B07FFF1FEF9F578FED8F8FB" box="[279,366,1877,1901]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="98" pageNumber="99" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B07FFF1FE94F578FE65F8FB" box="[378,467,1877,1901]" degrees="6" direction="south" minutes="30" orientation="latitude" pageId="98" pageNumber="99" precision="925" value="-6.5">6° 30' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B07FFF1FE0EF578FDE1F8FB" box="[480,599,1877,1901]" degrees="144" direction="east" minutes="51" orientation="longitude" pageId="98" pageNumber="99" precision="925" value="144.85">144° 51' E</geoCoordinate>
,
<date id="FFA710335B07FFF1FD8DF578FD79F8FB" box="[611,719,1877,1901]" pageId="98" pageNumber="99" value="1961-06-05">
<collectingDate id="EFE3E9DB5B07FFF1FD8DF578FD79F8FB" box="[611,719,1877,1901]" pageId="98" pageNumber="99" value="1961-06-05">5 vi 1961</collectingDate>
</date>
,
<collectorName id="26EC53255B07FFF1FD35F57BFC1BF8FB" box="[731,941,1877,1901]" pageId="98" pageNumber="99">J. L. &amp; M. Gressitt</collectorName>
(1
<collectionCode id="ED08AE365B07FFF1FC3DF578FB90F8FB" box="[979,1062,1877,1901]" country="USA" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="98" pageNumber="99" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B07FFF0FF29F554FD1AFF61" lastPageId="99" lastPageNumber="100" pageId="98" pageNumber="99" type="diagnosis">
<paragraph id="8BA636F35B07FFF1FF29F554FA2AF84F" blockId="98.[151,1436,1805,2009]" pageId="98" pageNumber="99">
<emphasis id="B96DEAE15B07FFF1FF29F554FE6FF804" bold="true" box="[199,473,1913,1938]" pageId="98" pageNumber="99">Differential Diagnosis.</emphasis>
Among members of the
<emphasis id="B96DEAE15B07FFF1FCFBF554FC17F807" box="[789,929,1913,1937]" italics="true" pageId="98" pageNumber="99">Cristatigena</emphasis>
group, similar to
<taxonomicName id="4C194D705B07FFF1FB69F557FA8DF807" authorityName="Perkins" authorityYear="2011" box="[1159,1339,1913,1937]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="inplacopaca">
<emphasis id="B96DEAE15B07FFF1FB69F557FB2DF807" box="[1159,1179,1914,1937]" italics="true" pageId="98" pageNumber="99">H</emphasis>
.
<emphasis id="B96DEAE15B07FFF1FB5FF554FA8DF807" box="[1201,1339,1913,1937]" italics="true" pageId="98" pageNumber="99">inplacopaca</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D705B07FFF1FA6FF557FEBFF823" authorityName="Perkins" authorityYear="2011" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="labropaca">
<emphasis id="B96DEAE15B07FFF1FA6FF557FA23F807" box="[1409,1429,1914,1937]" italics="true" pageId="98" pageNumber="99">H</emphasis>
.
<emphasis id="B96DEAE15B07FFF1FF79F5B0FEBFF823" box="[151,265,1949,1973]" italics="true" pageId="98" pageNumber="99">labropaca</emphasis>
</taxonomicName>
in the absence of metaventral plaques; differing from
<taxonomicName id="4C194D705B07FFF1FC93F5B3FB9DF823" authorityName="Perkins" authorityYear="2011" box="[893,1067,1949,1973]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="inplacopaca">
<emphasis id="B96DEAE15B07FFF1FC93F5B3FC27F823" box="[893,913,1950,1973]" italics="true" pageId="98" pageNumber="99">H</emphasis>
.
<emphasis id="B96DEAE15B07FFF1FC4FF5B0FB9DF823" box="[929,1067,1949,1973]" italics="true" pageId="98" pageNumber="99">inplacopaca</emphasis>
</taxonomicName>
in the smaller size (ca. 1.68 vs.
<quantity id="4CE19B165B07FFF1FF79F5ECFEB7F84F" box="[151,257,1985,2009]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" pageId="98" pageNumber="99" unit="mm" value="2.07">2.07 mm</quantity>
), the less dense and slightly coarser pronotal punctation, and the less apparent dorsal setae (
<figureCitation id="13222A765B07FFF1FAC7F5ECFA21F84F" box="[1321,1431,1985,2009]" captionStart="FIGURE 167" captionStartId="270.[151,250,1753,1776]" captionTargetBox="[203,1388,209,1717]" captionTargetId="figure-29@270.[187,1400,193,1732]" captionTargetPageId="270" captionText="FIGURE 167. Hydraena labropaca, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283732" httpUri="https://zenodo.org/record/5283732/files/figure.png" pageId="98" pageNumber="99">Figs. 167</figureCitation>
,
</paragraph>
<paragraph id="8BA636F35B06FFF0FF79F2BAFD1AFF61" blockId="99.[151,1436,151,1292]" pageId="99" pageNumber="100">
<figureCitation id="13222A765B06FFF0FF79F2BAFF75FF39" box="[151,195,151,175]" captionStart="FIGURE 170" captionStartId="272.[151,250,1753,1776]" captionTargetBox="[246,1325,198,1718]" captionTargetId="figure-29@272.[238,1348,193,1732]" captionTargetPageId="272" captionText="FIGURE 170. Hydraena implacopaca, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283736" httpUri="https://zenodo.org/record/5283736/files/figure.png" pageId="99" pageNumber="100">170</figureCitation>
,
<figureCitation id="13222A765B06FFF0FF3FF2BAFF48FF39" box="[209,254,151,175]" captionStart="FIGURE 171" captionStartId="273.[151,250,1753,1776]" captionTargetBox="[219,1367,214,1720]" captionTargetId="figure-25@273.[200,1387,193,1732]" captionTargetPageId="273" captionText="FIGURE 171. Hydraena transvallis, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283738" httpUri="https://zenodo.org/record/5283738/files/figure.png" pageId="99" pageNumber="100">171</figureCitation>
). Males of
<taxonomicName id="4C194D705B06FFF0FE6AF2B4FDA9FF26" authorityName="Perkins" authorityYear="2011" box="[388,543,152,176]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="transvallis">
<emphasis id="B96DEAE15B06FFF0FE6AF2B4FE2EFF26" box="[388,408,153,176]" italics="true" pageId="99" pageNumber="100">H</emphasis>
.
<emphasis id="B96DEAE15B06FFF0FE47F2B5FDA9FF26" box="[425,543,152,176]" italics="true" pageId="99" pageNumber="100">transvallis</emphasis>
</taxonomicName>
are differentiated from both species by the widened mesotibiae; and from
<taxonomicName id="4C194D705B06FFF0FA6FF2B4FEBFFF42" authorityName="Perkins" authorityYear="2011" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="labropaca">
<emphasis id="B96DEAE15B06FFF0FA6FF2B4FA23FF26" box="[1409,1429,153,176]" italics="true" pageId="99" pageNumber="100">H</emphasis>
.
<emphasis id="B96DEAE15B06FFF0FF79F291FEBFFF42" box="[151,265,188,212]" italics="true" pageId="99" pageNumber="100">labropaca</emphasis>
</taxonomicName>
by the differently shaped labrum. The male genitalia of the three species share a basic form, but differ in several significant details (
<figureCitation id="13222A765B06FFF0FE2DF2F2FD98FF61" box="[451,558,223,247]" captionStart="FIGURES 168169" captionStartId="271.[151,264,1650,1673]" captionText="FIGURES 168169. Aedeagi of holotypes. 168. H. balkei. 169. H. labropaca." pageId="99" pageNumber="100">Figs. 169</figureCitation>
,
<figureCitation id="13222A765B06FFF0FDD4F2F2FD29FF61" box="[570,671,223,247]" captionStart="FIGURES 172173" captionStartId="274.[151,264,1650,1673]" captionText="FIGURES 172173. Aedeagi of holotypes. 172. H. implacopaca. 173. H. transvallis." pageId="99" pageNumber="100">172, 173</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B06FFF0FF29F32EFAF7FBCE" pageId="99" pageNumber="100" type="description">
<paragraph id="8BA636F35B06FFF0FF29F32EFC5CFEF2" blockId="99.[151,1436,151,1292]" pageId="99" pageNumber="100">
<emphasis id="B96DEAE15B06FFF0FF29F32EFEEEFE8A" bold="true" box="[199,344,259,284]" inLineHeading="true" pageId="99" pageNumber="100" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B06FFF0FE42F329FDB8FE8A" box="[428,526,260,284]" pageId="99" pageNumber="100" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.68/0.78; head 0.25/0.44; pronotum 0.41/0.57, PA 0.49, PB 0.48; elytra 1.04/0.78. Dorsum brown to dark brown, head and pronotal disc slightly darker, palpi light brown, except distal 1/2 of last palpomere darker.
</paragraph>
<paragraph id="8BA636F35B06FFF0FF29F35DFCE2FDC9" blockId="99.[151,1436,151,1292]" pageId="99" pageNumber="100">Dorsum of head and pronotum densely punctulate, dull; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 1xpd or slightly larger, each puncture with short fine seta; punctures of clypeus obsolete. Labrum apicomedially with deep V-shaped notch, anterior margin at sides of notch with tiny point, free margins very weakly upturned. Mentum finely, moderately densely punctate, shining; postmentum anterior 1/2 with punctation similar to mentum, posterior 1/2 densely micropunctulate. Genae raised, posterior ridge strong. Pronotum with anterior margin weakly emarginate over middle 1/2; emargination with very narrow hyaline margin; pronotal foveae PF1 and PF2 moderately deep, PF3 and PF4 very deep.</paragraph>
<paragraph id="8BA636F35B06FFF0FF29F041FAEFFD79" blockId="99.[151,1436,151,1292]" pageId="99" pageNumber="100">Elytra dull, punctures ca. 1xpd or slightly larger than largest pronotal punctures, each puncture with short fine seta and three granules, one at anterior margin and one on each side. Intervals not raised, width ca. 1xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another. Explanate margin well developed, about as wide as widest part of metatibia.</paragraph>
<paragraph id="8BA636F35B06FFF0FF29F0D1FD16FC51" blockId="99.[151,1436,151,1292]" pageId="99" pageNumber="100">Metaventrite markedly tumid on each side of median impression, plaques absent, hydrofuge pubescence dense. P1 laminate; median carina sinuate in profile. P2 moderately narrow, l/w ca. 2/1, sides parallel, apex blunt. No midlongitudinal carina between mesoventral intercoxal process and median impression. AIS flat, width at straight posterior margin ca. 2x P2. Protibia (male) widened, weakly arcuate, medial surface flat; mesotibia widened, widest part, slightly past midlength, wider than metatibia; metatibia very gradually widening from base to apex, very weakly arcuate. Abdominal apex symmetrical.</paragraph>
<paragraph id="8BA636F35B06FFF0FF29F1F9FAF7FBCE" blockId="99.[151,1436,151,1292]" pageId="99" pageNumber="100">
Aedeagus (
<figureCitation id="13222A765B06FFF0FEAAF1F9FE15FC7A" box="[324,419,980,1004]" captionStart="FIGURES 172173" captionStartId="274.[151,264,1650,1673]" captionText="FIGURES 172173. Aedeagi of holotypes. 172. H. implacopaca. 173. H. transvallis." pageId="99" pageNumber="100">Fig. 173</figureCitation>
) main piece with large distal lobe, in ventral aspect, weakly arching from right side toward left side, apically rounded, small lobe on left side; left paramere smaller than distal lobe, widest at midlength, with two small lobes, each with a cluster of setae; right paramere longer than and apically slightly narrower than distal lobe of main piece, apically blunt; distal piece, in right lateral view, with large lobe of characteristic shape.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B06FFF0FF29F649FF6FFB36" pageId="99" pageNumber="100" type="etymology">
<paragraph id="8BA636F35B06FFF0FF29F649FF6FFB36" blockId="99.[151,1436,151,1292]" pageId="99" pageNumber="100">
<emphasis id="B96DEAE15B06FFF0FF29F649FEFBFBEA" bold="true" box="[199,333,1124,1148]" inLineHeading="true" pageId="99" pageNumber="100" reason="1">Etymology.</emphasis>
Named in reference to the geographical distribution, two localities separated by extensive lowlands.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B06FFF0FF29F686FBE3FA9A" pageId="99" pageNumber="100" type="distribution">
<paragraph id="8BA636F35B06FFF0FF29F686FBE3FA9A" blockId="99.[151,1436,151,1292]" pageId="99" pageNumber="100">
<emphasis id="B96DEAE15B06FFF0FF29F686FED6FB52" bold="true" box="[199,352,1195,1220]" inLineHeading="true" pageId="99" pageNumber="100" reason="1">Distribution.</emphasis>
Currently known from one locality in the southeastern part of Area 1, at the border with Area 13 (
<quantity id="4CE19B165B06FFF0FF71F6FDFF4EFB7E" box="[159,248,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="99" pageNumber="100" unit="m" value="1000.0">1000 m</quantity>
), and one locality in the Finisterre Mts., Area 5 at the border with Area 3 (
<quantity id="4CE19B165B06FFF0FBD5F6FDFB25FB7E" box="[1083,1171,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.082" pageId="99" pageNumber="100" unit="m" value="1082.0">1082 m</quantity>
) (
<figureCitation id="13222A765B06FFF0FB45F6FDFABBFB7E" box="[1195,1293,1232,1256]" captionStart="FIGURES 503506" captionStartId="401.[151,264,1650,1673]" captionTargetBox="[165,1421,222,1609]" captionTargetId="figure-20@401.[165,1421,193,1629]" captionTargetPageId="401" captionText="FIGURES 503506. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5299476" httpUri="https://zenodo.org/record/5299476/files/figure.png" pageId="99" pageNumber="100">Fig. 506</figureCitation>
). This is one of ten species with distributions exclusively shared between Areas 1 and 5 (
<figureCitation id="13222A765B06FFF0FC09F6D9FBF0FA9A" box="[999,1094,1268,1292]" captionStart="FIGURES 421422" captionStartId="378.[151,263,1752,1775]" captionTargetBox="[228,1326,203,1731]" captionTargetId="figure-46@378.[221,1367,193,1731]" captionTargetPageId="378" captionText="FIGURES 421422. 421. All known collecting localities for Papua New Guinea Hydraena. 422. Ten species, representing eight species groups, with distributions exclusively shared between Areas of Endemism 1 and 5." figureDoi="http://doi.org/10.5281/zenodo.5284047" httpUri="https://zenodo.org/record/5284047/files/figure.png" pageId="99" pageNumber="100">Fig. 422</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>