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<document id="FE8C1027D277CF25FD7AFE407D6264C0" ID-DOI="10.11646/zootaxa.5380.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="10216220" ID-ZooBank="6B51DC3F-67C4-4833-92C7-3387297985A6" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1701246344783" checkinUser="plazi" docAuthor="Gouvea, Bruno, Valente, Roberta M. &amp; Asenjo, Angélico" docDate="2023" docId="03B087E3214AF253FF29FF34AFB5170C" docLanguage="en" docName="zootaxa.5380.2.2.pdf" docOrigin="Zootaxa 5380 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5380.2.2/52383" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oxarthrius Reitter 1883" docType="treatment" docVersion="1" lastPageNumber="148" masterDocId="FF89FF9B2147F25DFFBEFFA3AA3D156D" masterDocTitle="A new species from the eastern Amazon rainforest with lectotype designations and key for Brazilian Oxarthrius Reitter, 1883 (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="150" masterPageNumber="134" pageNumber="147" updateTime="1701732240379" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="19B097303913B3BE6051AE0DC3FF605B">A new species from the eastern Amazon rainforest with lectotype designations and key for Brazilian Oxarthrius Reitter, 1883 (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:affiliation id="47DB0AD16A199890B38A51BCC2F87E5C">Universidade Federal do Pará, Instituto de Ciências Biológicas, Laboratório de Invertebrados, Belém, Pará, Brazil. brunoleandro 555 @ gmail. com; https: // orcid. org / 0009 - 0002 - 1540 - 9303</mods:affiliation>
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<mods:namePart id="711A7C3195183D6A96A359212C6DF02D">Valente, Roberta M.</mods:namePart>
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<mods:affiliation id="10D570C04D930A9C34A8B61B2A20EF2F">Universidade Federal do Pará, Instituto de Ciências Biológicas, Laboratório de Invertebrados, Belém, Pará, Brazil. roberta. mvalente @ gmail. com; https: // orcid. org / 0000 - 0002 - 8249 - 5937</mods:affiliation>
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<mods:affiliation id="4CB82CCAE351FC4AEFE4222565932B8D">Instituto Tecnológico VALE, R. Boaventura da Silva 955, Nazaré, 66055 - 090 Belém, PA. Brazil. &amp; Departamento de Entomología, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Av. Arenales 1256, Jesús María, Lima 14, Perú.</mods:affiliation>
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<paragraph id="8BA636F5214AF250FF29FF34A92815DC" blockId="13.[151,789,151,178]" box="[151,789,151,178]" pageId="13" pageNumber="147">
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Key to Brazilian species of
<taxonomicName id="4C194D76214AF250FE57FF34A92815DC" ID-CoL="8X7MV" authority="Reitter, 1883" authorityName="Reitter" authorityYear="1883" box="[489,789,151,178]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE7214AF250FE57FF34A85215DF" bold="true" box="[489,623,151,178]" italics="true" pageId="13" pageNumber="147">Oxarthrius</emphasis>
<bibRefCitation id="EF884B04214AF250FDC8FF34A92815DC" author="Reitter, E." box="[630,789,151,177]" pageId="13" pageNumber="147" pagination="371 - 386" refId="ref10163" refString="Reitter, E. (1883) Neue Pselaphiden und Scydmaeniden aus Central- und Sudamerika. Verhandlungen der KaiserlichKoniglichen Zoologisch-Botanischen Gesellschaft in Wien, 32, 371 - 386. [Note: nominal year of publication 1882]" type="journal article" year="1883">Reitter, 1883</bibRefCitation>
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<paragraph id="8BA636F5214AF250FF29FF7FAFA1159C" blockId="13.[151,1437,220,594]" box="[151,1436,220,242]" pageId="13" pageNumber="147">
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1. Antennal segments 39 slightly longer than wide, or subquadrate (
<figureCitation id="13222A70214AF250FCDEFF7FA993159C" box="[864,942,220,242]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[151,1435,191,1852]" captionTargetId="figure-20@5.[151,1435,179,1852]" captionTargetPageId="5" captionText="FIGURE 2. Oxarthrius spiculatus sp. nov. (A-E. Male holotype - STP0599; F. Female paratype STP0267) A. Habitus, ventral view. B. Head, dorsal view. C. Habitus, lateral view. D. Antennae. E. Abdomen ventral view. F. Abdomen ventral view. Scale bars = 1 mm (A, C); 0.5 mm (B, E, F); 200 µm (D)." figureDoi="http://doi.org/10.5281/zenodo.10216226" httpUri="https://zenodo.org/record/10216226/files/figure.png" pageId="13" pageNumber="147">Fig. 2D</figureCitation>
). Vertexal sulcus [vs] present (
<figureCitation id="13222A70214AF250FB6DFF7FAF23159C" box="[1235,1310,220,242]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="13" pageNumber="147">Fig. 3A</figureCitation>
)........ 2
</keyLead>
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<paragraph id="8BA636F5214AF250FF29FF59AFA11441" blockId="13.[151,1437,220,594]" pageId="13" pageNumber="147">
<keyLead id="30E895C0214AF250FF29FF59AFA11441" pageId="13" pageNumber="147">
- Antennal segments 3-9 extremely elongate, at least twice as long as wide (
<bibRefCitation id="EF884B04214AF250FC18FF5AAE641462" author="Asenjo, A. &amp; Zampaulo, R. de &amp; Ferreira, R. L." box="[934,1113,249,271]" pageId="13" pageNumber="147" pagination="404 - 414" refId="ref8207" refString="Asenjo, A., Zampaulo, R. de A. &amp; Ferreira, R. L. (2018) Two new troglobitic species of Oxarthrius Reitter (Coleoptera, Staphylinidae, Pselaphinae) from Brazil. Zootaxa, 4462 (3), 404 - 414. https: // doi. org / 10.11646 / zootaxa. 4462.3.6" type="journal article" year="2018">
Asenjo
<emphasis id="B96DEAE7214AF250FC4FFF59AE261463" box="[1009,1051,249,271]" italics="true" pageId="13" pageNumber="147">et al</emphasis>
. 2018
</bibRefCitation>
; figs 12, 1213). Vertexal sulcus [vs] absent (
<bibRefCitation id="EF884B04214AF250FEE9FEB4A8361441" author="Asenjo, A. &amp; Zampaulo, R. de &amp; Ferreira, R. L." box="[343,523,279,301]" pageId="13" pageNumber="147" pagination="404 - 414" refId="ref8207" refString="Asenjo, A., Zampaulo, R. de A. &amp; Ferreira, R. L. (2018) Two new troglobitic species of Oxarthrius Reitter (Coleoptera, Staphylinidae, Pselaphinae) from Brazil. Zootaxa, 4462 (3), 404 - 414. https: // doi. org / 10.11646 / zootaxa. 4462.3.6" type="journal article" year="2018">
Asenjo
<emphasis id="B96DEAE7214AF250FE1CFEB4ABF01441" box="[418,461,279,300]" italics="true" pageId="13" pageNumber="147">et al</emphasis>
. 2018
</bibRefCitation>
; figs 3, 14)................................................................. 4
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<paragraph id="8BA636F5214AF250FF29FE97AFA11427" blockId="13.[151,1437,220,594]" box="[151,1436,308,330]" pageId="13" pageNumber="147">
<keyLead id="30E895C0214AF250FF29FE97AFA11427" box="[151,1436,308,330]" pageId="13" pageNumber="147">
2. Carina connecting pronotal setae lacking. Pronotum lacking median basal carina...........
<taxonomicName id="4C194D76214AF250FBCEFE97AFA11427" ID-CoL="4BBS4" authority="(Reitter)" baseAuthorityName="Reitter" baseAuthorityYear="1883" box="[1136,1436,308,330]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B96DEAE7214AF250FBCEFE97AF781424" box="[1136,1349,308,329]" italics="true" pageId="13" pageNumber="147">Oxarthrius bispinosus</emphasis>
(Reitter)
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</paragraph>
<paragraph id="8BA636F5214AF250FF29FEF1AFA1140A" blockId="13.[151,1437,220,594]" box="[151,1436,337,359]" pageId="13" pageNumber="147">
<keyLead id="30E895C0214AF250FF29FEF1AFA1140A" box="[151,1436,337,359]" pageId="13" pageNumber="147">
- Pronotum with sinuous carina connecting base of setae (
<figureCitation id="13222A70214AF250FD40FEF2A974140A" box="[766,841,337,359]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="13" pageNumber="147">Fig. 3B</figureCitation>
). Pronotum with median basal carina (
<figureCitation id="13222A70214AF250FB02FEF2AF3B140A" box="[1212,1286,337,359]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="13" pageNumber="147">Fig. 3B</figureCitation>
).......... 3
</keyLead>
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<keyStep id="30ED2E50214AF250FF29FECCAFA114B1" pageId="13" pageNumber="147">
<paragraph id="8BA636F5214AF250FF29FECCAFA014CF" blockId="13.[151,1437,220,594]" pageId="13" pageNumber="147">
<keyLead id="30E895C0214AF250FF29FECCAFA014CF" pageId="13" pageNumber="147">
3. Lateral antebasal foveae [laf] wide (
<figureCitation id="13222A70214AF250FD82FECCA8B714E9" box="[572,650,367,389]" captionStart="FIGURE 7" captionStartId="11.[151,250,1831,1855]" captionTargetBox="[151,1435,179,1804]" captionTargetId="figure-20@11.[151,1436,179,1804]" captionTargetPageId="11" captionText="FIGURE 7. Oxarthrius anthicoides (L.W.Schaufuss, 1887), Lectotype (MNHN, EC21543). A. habitus, dorsal view. B. habitus, ventral view. C. original labels. D. habitus, lateral view. Credits of the photographs: Christophe Rivier (Muséum National dHistoire Naturelle, Paris, France). Scale bars = 1 mm (A, B, D)" figureDoi="http://doi.org/10.5281/zenodo.10216242" httpUri="https://zenodo.org/record/10216242/files/figure.png" pageId="13" pageNumber="147">Fig. 7D</figureCitation>
). Vertexal sulcus [vs], deep and wide, laterally carinate (
<figureCitation id="13222A70214AF250FB19FECCAF2514E9" box="[1191,1304,367,389]" captionStart="FIGURE 7" captionStartId="11.[151,250,1831,1855]" captionTargetBox="[151,1435,179,1804]" captionTargetId="figure-20@11.[151,1436,179,1804]" captionTargetPageId="11" captionText="FIGURE 7. Oxarthrius anthicoides (L.W.Schaufuss, 1887), Lectotype (MNHN, EC21543). A. habitus, dorsal view. B. habitus, ventral view. C. original labels. D. habitus, lateral view. Credits of the photographs: Christophe Rivier (Muséum National dHistoire Naturelle, Paris, France). Scale bars = 1 mm (A, B, D)" figureDoi="http://doi.org/10.5281/zenodo.10216242" httpUri="https://zenodo.org/record/10216242/files/figure.png" pageId="13" pageNumber="147">Figs. 7AB</figureCitation>
)............................................................................
<taxonomicName id="4C194D76214AF250FBA6FE2FAFA014CF" ID-CoL="4BBRY" authority="(L. W. Schaufuss)" baseAuthorityName="L. W. Schaufuss" box="[1048,1437,396,418]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="anthicoides">
<emphasis id="B96DEAE7214AF250FBA6FE2FAECB14CC" box="[1048,1270,396,417]" italics="true" pageId="13" pageNumber="147">Oxarthrius anthicoides</emphasis>
(L.W.Schaufuss)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA636F5214AF250FF29FE09AFA114B1" blockId="13.[151,1437,220,594]" pageId="13" pageNumber="147">
<keyLead id="30E895C0214AF250FF29FE09AFA114B1" pageId="13" pageNumber="147">
- Lateral antebasal foveae [laf] narrow (
<figureCitation id="13222A70214AF250FDE9FE0AA89E14D2" box="[599,675,425,447]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="13" pageNumber="147">Fig. 3B</figureCitation>
). Vertexal sulcus [vs] narrow, not laterally carinate (
<figureCitation id="13222A70214AF250FB26FE0AAEDE14D2" box="[1176,1251,425,447]" captionStart="FIGURE 3" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[152,1436,179,1463]" captionTargetId="figure-19@6.[152,1436,179,1463]" captionTargetPageId="6" captionText="FIGURE 3. Oxarthrius spiculatus sp. nov., paratype male- STP0269. A. Head, dorsal view. B. Pronotum, dorsal view. C. Elytra, dorsal view. D. Abdomen, dorsal view. Scale bars = 100 μm (A, B, D); 200 μm (C). Abbreviations: vf, vertexal foveae; laf, lateral antebasal foveae; oblf, outer basolateral foveae; iblf, inner basolateral foveae; bef, basal elytral foveae; ss, sutural stria; mbf, mediobasal foveae; blf, basolateral foveae." figureDoi="http://doi.org/10.5281/zenodo.10216228" httpUri="https://zenodo.org/record/10216228/files/figure.png" pageId="13" pageNumber="147">Fig. 3A</figureCitation>
).........................................................................................
<emphasis id="B96DEAE7214AF250FBCCFE65AFA114B1" bold="true" box="[1138,1436,454,476]" pageId="13" pageNumber="147">
<taxonomicName id="4C194D76214AF250FBCCFE65AF7414B1" authority="Gouvea &amp; Valente &amp; Asenjo, 2023" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[1138,1353,454,476]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="spiculatus" status="sp. nov.">
<emphasis id="B96DEAE7214AF250FBCCFE65AF7414B1" bold="true" box="[1138,1353,454,476]" italics="true" pageId="13" pageNumber="147">Oxarthrius spiculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E579C214AF250FAEEFE64AFA114B1" box="[1360,1436,455,476]" pageId="13" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30ED2E50214AF250FF29FE47AFA1173F" pageId="13" pageNumber="147">
<paragraph id="8BA636F5214AF250FF29FE47AFA1177A" blockId="13.[151,1437,220,594]" pageId="13" pageNumber="147">
<keyLead id="30E895C0214AF250FF29FE47AFA1177A" pageId="13" pageNumber="147">
4. Antennal segment 10 twice as long as wide (
<bibRefCitation id="EF884B04214AF250FD2AFE47A9761494" author="Asenjo, A. &amp; Zampaulo, R. de &amp; Ferreira, R. L." box="[660,843,484,506]" pageId="13" pageNumber="147" pagination="404 - 414" refId="ref8207" refString="Asenjo, A., Zampaulo, R. de A. &amp; Ferreira, R. L. (2018) Two new troglobitic species of Oxarthrius Reitter (Coleoptera, Staphylinidae, Pselaphinae) from Brazil. Zootaxa, 4462 (3), 404 - 414. https: // doi. org / 10.11646 / zootaxa. 4462.3.6" type="journal article" year="2018">
Asenjo
<emphasis id="B96DEAE7214AF250FD5FFE46A9311494" box="[737,780,484,506]" italics="true" pageId="13" pageNumber="147">et al</emphasis>
. 2018
</bibRefCitation>
; figs 12). Basal half of male mesofemora with an elongated and sinuated spine (
<bibRefCitation id="EF884B04214AF250FE21FDA2A869177A" author="Asenjo, A. &amp; Zampaulo, R. de &amp; Ferreira, R. L." box="[415,596,513,535]" pageId="13" pageNumber="147" pagination="404 - 414" refId="ref8207" refString="Asenjo, A., Zampaulo, R. de A. &amp; Ferreira, R. L. (2018) Two new troglobitic species of Oxarthrius Reitter (Coleoptera, Staphylinidae, Pselaphinae) from Brazil. Zootaxa, 4462 (3), 404 - 414. https: // doi. org / 10.11646 / zootaxa. 4462.3.6" type="journal article" year="2018">
Asenjo
<emphasis id="B96DEAE7214AF250FE55FDA1A828177B" box="[491,533,513,535]" italics="true" pageId="13" pageNumber="147">et al</emphasis>
. 2018
</bibRefCitation>
; fig. 5).....................................
<taxonomicName id="4C194D76214AF250FBD2FDA2AFA1177A" ID-CoL="4BBS7" authority="Asenjo" authorityName="Asenjo" authorityYear="2018" box="[1132,1436,513,535]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="inexpectatus">
<emphasis id="B96DEAE7214AF250FBD2FDA2AF69177A" box="[1132,1364,513,535]" italics="true" pageId="13" pageNumber="147">Oxarthrius inexpectatus</emphasis>
Asenjo
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA636F5214AF250FF29FDBCAFA1173F" blockId="13.[151,1437,220,594]" pageId="13" pageNumber="147">
<keyLead id="30E895C0214AF250FF29FDBCAFA1173F" pageId="13" pageNumber="147">
- Antennal segment 10 three times as long as wide (
<bibRefCitation id="EF884B04214AF250FD67FDBCA9A81759" author="Asenjo, A. &amp; Zampaulo, R. de &amp; Ferreira, R. L." box="[729,917,543,565]" pageId="13" pageNumber="147" pagination="404 - 414" refId="ref8207" refString="Asenjo, A., Zampaulo, R. de A. &amp; Ferreira, R. L. (2018) Two new troglobitic species of Oxarthrius Reitter (Coleoptera, Staphylinidae, Pselaphinae) from Brazil. Zootaxa, 4462 (3), 404 - 414. https: // doi. org / 10.11646 / zootaxa. 4462.3.6" type="journal article" year="2018">
Asenjo
<emphasis id="B96DEAE7214AF250FC99FDBCA9691759" box="[807,852,543,564]" italics="true" pageId="13" pageNumber="147">et al</emphasis>
. 2018
</bibRefCitation>
; figs 1213). Basal half of male mesofemora with a truncate spine (
<bibRefCitation id="EF884B04214AF250FECAFD9FA814173C" author="Asenjo, A. &amp; Zampaulo, R. de &amp; Ferreira, R. L." box="[372,553,572,594]" pageId="13" pageNumber="147" pagination="404 - 414" refId="ref8207" refString="Asenjo, A., Zampaulo, R. de A. &amp; Ferreira, R. L. (2018) Two new troglobitic species of Oxarthrius Reitter (Coleoptera, Staphylinidae, Pselaphinae) from Brazil. Zootaxa, 4462 (3), 404 - 414. https: // doi. org / 10.11646 / zootaxa. 4462.3.6" type="journal article" year="2018">
Asenjo
<emphasis id="B96DEAE7214AF250FE7EFD9EABD7173C" box="[448,490,572,594]" italics="true" pageId="13" pageNumber="147">et al</emphasis>
. 2018
</bibRefCitation>
; fig. 16).............................................
<taxonomicName id="4C194D76214AF250FB1CFD9FAFA1173F" ID-CoL="4BBS3" authority="Asenjo" authorityName="Asenjo" authorityYear="2018" box="[1186,1436,572,594]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="aurora">
<emphasis id="B96DEAE7214AF250FB1CFD9FAF69173F" box="[1186,1364,572,594]" italics="true" pageId="13" pageNumber="147">Oxarthrius aurora</emphasis>
Asenjo
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
<subSubSection id="C303657E214AF250FF29FD07A84411FE" pageId="13" pageNumber="147" type="discussion">
<paragraph id="8BA636F5214AF250FF29FD07AB2617D3" blockId="13.[151,283,676,702]" box="[151,283,676,702]" pageId="13" pageNumber="147">
<heading id="D0EE8199214AF250FF29FD07AB2617D3" bold="true" box="[151,283,676,702]" fontSize="11" level="1" pageId="13" pageNumber="147" reason="1">
<emphasis id="B96DEAE7214AF250FF29FD07AB2617D3" bold="true" box="[151,283,676,702]" pageId="13" pageNumber="147">Discussion</emphasis>
</heading>
</paragraph>
<paragraph id="8BA636F5214AF250FF29FD4EAB27166A" blockId="13.[151,282,749,775]" box="[151,282,749,775]" pageId="13" pageNumber="147">
<heading id="D0EE8199214AF250FF29FD4EAB27166A" bold="true" box="[151,282,749,775]" fontSize="11" level="1" pageId="13" pageNumber="147" reason="1">
<emphasis id="B96DEAE7214AF250FF29FD4EAB27166A" bold="true" box="[151,282,749,775]" pageId="13" pageNumber="147">Taxonomy</emphasis>
</heading>
</paragraph>
<paragraph id="8BA636F5214AF250FF29FC97A84411FE" blockId="13.[151,1437,820,1171]" pageId="13" pageNumber="147">
The new species described here belongs to the genus
<taxonomicName id="4C194D76214AF250FD41FC97A9471623" authorityName="Reitter" authorityYear="1883" box="[767,890,820,846]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE7214AF250FD41FC97A9471623" box="[767,890,820,846]" italics="true" pageId="13" pageNumber="147">Oxarthrius</emphasis>
</taxonomicName>
due to the pronotum having a pair of antebasal spines, and lacking an antebasal transverse sulcus. It is placed in the subgenus
<taxonomicName id="4C194D76214AF250FB85FCFBAE8B161F" box="[1083,1206,856,882]" pageId="13" pageNumber="147" rank="subGenus" subGenus="Oxarthrius">
<emphasis id="B96DEAE7214AF250FB85FCFBAE8B161F" box="[1083,1206,856,882]" italics="true" pageId="13" pageNumber="147">Oxarthrius</emphasis>
</taxonomicName>
as defined by
<bibRefCitation id="EF884B04214AF250FAD7FCFBAADF16FA" author="Park, O." pageId="13" pageNumber="147" refId="ref9661" refString="Park, O. (1942) A study in neotropical Pselaphidae. Northwestern University Studies in the Biological Sciences and Medicine, Number 1. Northwestern University, Evanston and Chicago, x + 403 pp. https: // doi. org / 10.5962 / bhl. title. 6838" type="book" year="1942">Park (1942)</bibRefCitation>
based on the characters “maxillary palpi with simple unscared external face” and “each elytron with a single basal fovea”. Currently, only
<taxonomicName id="4C194D76214AF250FE5BFC03A96C16D6" authority="Asenjo, 2018" authorityName="Asenjo" authorityYear="2018" box="[485,849,928,955]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="aurora">
<emphasis id="B96DEAE7214AF250FE5BFC03A88816D7" box="[485,693,928,954]" italics="true" pageId="13" pageNumber="147">Oxarthrius aurora</emphasis>
<bibRefCitation id="EF884B04214AF250FD05FC03A96C16D6" author="Asenjo, A. &amp; Zampaulo, R. de &amp; Ferreira, R. L." box="[699,849,928,955]" pageId="13" pageNumber="147" pagination="404 - 414" refId="ref8207" refString="Asenjo, A., Zampaulo, R. de A. &amp; Ferreira, R. L. (2018) Two new troglobitic species of Oxarthrius Reitter (Coleoptera, Staphylinidae, Pselaphinae) from Brazil. Zootaxa, 4462 (3), 404 - 414. https: // doi. org / 10.11646 / zootaxa. 4462.3.6" type="journal article" year="2018">Asenjo, 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C194D76214AF250FC34FC03AF0816D6" authority="Asenjo, 2018" authorityName="Asenjo" authorityYear="2018" box="[906,1333,928,955]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="inexpectatus">
<emphasis id="B96DEAE7214AF250FC34FC03AEA516D7" box="[906,1176,928,954]" italics="true" pageId="13" pageNumber="147">Oxarthrius inexpectatus</emphasis>
<bibRefCitation id="EF884B04214AF250FB21FC03AF0816D6" author="Asenjo, A. &amp; Zampaulo, R. de &amp; Ferreira, R. L." box="[1183,1333,928,955]" pageId="13" pageNumber="147" pagination="404 - 414" refId="ref8207" refString="Asenjo, A., Zampaulo, R. de A. &amp; Ferreira, R. L. (2018) Two new troglobitic species of Oxarthrius Reitter (Coleoptera, Staphylinidae, Pselaphinae) from Brazil. Zootaxa, 4462 (3), 404 - 414. https: // doi. org / 10.11646 / zootaxa. 4462.3.6" type="journal article" year="2018">Asenjo, 2018</bibRefCitation>
</taxonomicName>
have the male genitalia described. Photographs of
<typeStatus id="54A28857214AF250FDCDFC66A89E16B2" box="[627,675,965,991]" pageId="13" pageNumber="147">type</typeStatus>
specimens of
<taxonomicName id="4C194D76214AF250FCF5FC66A9DB16B3" baseAuthorityName="Reitter" baseAuthorityYear="1883" box="[843,998,964,991]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B96DEAE7214AF250FCF5FC66A9DB16B3" box="[843,998,964,991]" italics="true" pageId="13" pageNumber="147">O. bispinosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D76214AF250FB9FFC66AEFA16B3" box="[1057,1223,964,991]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="anthicoides">
<emphasis id="B96DEAE7214AF250FB9FFC66AEFA16B3" box="[1057,1223,964,991]" italics="true" pageId="13" pageNumber="147">O. anthicoides</emphasis>
</taxonomicName>
were fundamental for definition of
<taxonomicName id="4C194D76214AF250FEEAFC4AABD4116F" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[340,489,1000,1027]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="spiculatus">
<emphasis id="B96DEAE7214AF250FEEAFC4AABD4116F" box="[340,489,1000,1027]" italics="true" pageId="13" pageNumber="147">O. spiculatus</emphasis>
</taxonomicName>
as a new species considering that they are very similar, and the original description was insufficiently detailed to distinguish the two species. For first time the female genital complex is described for a species of
<taxonomicName id="4C194D76214AF250FE98FB93AB9C1127" authorityName="Reitter" authorityYear="1883" box="[294,417,1072,1098]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE7214AF250FE98FB93AB9C1127" box="[294,417,1072,1098]" italics="true" pageId="13" pageNumber="147">Oxarthrius</emphasis>
</taxonomicName>
so it is not possible to compare it with any described species. As far as is known, this is the first species of
<taxonomicName id="4C194D76214AF250FECEFBF7ABD61103" authorityName="Reitter" authorityYear="1883" box="[368,491,1108,1134]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE7214AF250FECEFBF7ABD61103" box="[368,491,1108,1134]" italics="true" pageId="13" pageNumber="147">Oxarthrius</emphasis>
</taxonomicName>
that the male have a spine on the metatrochanter since the male is unknown for
<taxonomicName id="4C194D76214AF250FA3FFBF6AB2511FF" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="anthicoides">
<emphasis id="B96DEAE7214AF250FA3FFBF6AB2511FF" italics="true" pageId="13" pageNumber="147">O. anthicoides</emphasis>
</taxonomicName>
,
<taxonomicName id="4C194D76214AF250FE98FBDAAB8311FE" authorityName="Raffray" authorityYear="1898" box="[294,446,1145,1171]" class="Insecta" family="Staphylinidae" genus="Batrisus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="forticornis">
<emphasis id="B96DEAE7214AF250FE98FBDAAB8311FE" box="[294,446,1145,1171]" italics="true" pageId="13" pageNumber="147">O. forticornis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C194D76214AF250FE73FBDAA84F11FF" authorityName="Newton" authorityYear="2017" box="[461,626,1144,1171]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="simplexides">
<emphasis id="B96DEAE7214AF250FE73FBDAA84F11FF" box="[461,626,1144,1171]" italics="true" pageId="13" pageNumber="147">O. simplexides</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C303657E214AF253FF29FB47AFB5170C" lastPageId="14" lastPageNumber="148" pageId="13" pageNumber="147" type="distribution">
<paragraph id="8BA636F5214AF250FF29FB47ABE01192" blockId="13.[151,477,1252,1279]" box="[151,477,1252,1279]" pageId="13" pageNumber="147">
<heading id="D0EE8199214AF250FF29FB47ABE01192" bold="true" box="[151,477,1252,1279]" fontSize="11" level="1" pageId="13" pageNumber="147" reason="1">
<emphasis id="B96DEAE7214AF250FF29FB47ABE01192" bold="true" box="[151,477,1252,1279]" pageId="13" pageNumber="147">
Distribution of
<taxonomicName id="4C194D76214AF250FEE9FB47ABE01192" ID-CoL="8X7MV" authorityName="Reitter" authorityYear="1883" box="[343,477,1252,1279]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE7214AF250FEE9FB47ABE01192" bold="true" box="[343,477,1252,1279]" italics="true" pageId="13" pageNumber="147">Oxarthrius</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA636F5214AF250FF29FA8FAB78130A" blockId="13.[151,1437,1324,2035]" pageId="13" pageNumber="147">
The genus is widely distributed throughout South America (
<collectingCountry id="F30E7665214AF250FC8EFA8FA999102B" box="[816,932,1324,1350]" name="Argentina" pageId="13" pageNumber="147">Argentina</collectingCountry>
,
<collectingCountry id="F30E7665214AF250FC11FA8FAE39102B" box="[943,1028,1324,1350]" name="Bolivia" pageId="13" pageNumber="147">Bolivia</collectingCountry>
,
<collectingCountry id="F30E7665214AF250FBB0FA8FAE68102B" box="[1038,1109,1324,1350]" name="Brazil" pageId="13" pageNumber="147">Brazil</collectingCountry>
,
<collectingCountry id="F30E7665214AF250FBDEFA8EAEF5102A" box="[1120,1224,1325,1351]" name="Paraguay" pageId="13" pageNumber="147">Paraguay</collectingCountry>
and
<collectingCountry id="F30E7665214AF250FB43FA8EAF0F102A" box="[1277,1330,1325,1351]" name="Peru" pageId="13" pageNumber="147">Peru</collectingCountry>
) and also in Central America it is known from
<collectingCountry id="F30E7665214AF250FDFDFAF3A8811007" box="[579,700,1360,1386]" name="Guatemala" pageId="13" pageNumber="147">Guatemala</collectingCountry>
and
<collectingCountry id="F30E7665214AF250FD49FAF2A96D1006" box="[759,848,1361,1387]" name="Panama" pageId="13" pageNumber="147">Panama</collectingCountry>
(
<bibRefCitation id="EF884B04214AF250FCDEFAF3A9E41006" author="Park, O." box="[864,985,1360,1387]" pageId="13" pageNumber="147" refId="ref9661" refString="Park, O. (1942) A study in neotropical Pselaphidae. Northwestern University Studies in the Biological Sciences and Medicine, Number 1. Northwestern University, Evanston and Chicago, x + 403 pp. https: // doi. org / 10.5962 / bhl. title. 6838" type="book" year="1942">Park 1942</bibRefCitation>
;
<bibRefCitation id="EF884B04214AF250FC58FAF3AE831006" author="Asenjo, A. &amp; Zampaulo, R. de &amp; Ferreira, R. L." box="[998,1214,1360,1387]" pageId="13" pageNumber="147" pagination="404 - 414" refId="ref8207" refString="Asenjo, A., Zampaulo, R. de A. &amp; Ferreira, R. L. (2018) Two new troglobitic species of Oxarthrius Reitter (Coleoptera, Staphylinidae, Pselaphinae) from Brazil. Zootaxa, 4462 (3), 404 - 414. https: // doi. org / 10.11646 / zootaxa. 4462.3.6" type="journal article" year="2018">
Asenjo
<emphasis id="B96DEAE7214AF250FB80FAF2AE4C1007" box="[1086,1137,1360,1386]" italics="true" pageId="13" pageNumber="147">et al</emphasis>
. 2018
</bibRefCitation>
). The northernmost record is for
<taxonomicName id="4C194D76214AF250FE99FAD6ABEE10E3" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[295,467,1396,1423]" class="Insecta" family="Staphylinidae" genus="Batrisus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="hamaticollis">
<emphasis id="B96DEAE7214AF250FE99FAD6ABEE10E3" box="[295,467,1396,1423]" italics="true" pageId="13" pageNumber="147">O. hamaticollis</emphasis>
</taxonomicName>
, which was collected at an elevation of
<quantity id="4CE19B10214AF250FC2BFAD6AE3810E3" box="[917,1029,1396,1423]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="13" pageNumber="147" unit="ft" value="3000.0">3,000 feet</quantity>
at Las Mercedes,
<collectingCountry id="F30E7665214AF250FB6FFAD7AF7710E3" box="[1233,1354,1396,1422]" name="Guatemala" pageId="13" pageNumber="147">Guatemala</collectingCountry>
(
<bibRefCitation id="EF884B04214AF250FAE7FAD7AAEF10DE" author="Sharp, D. S." pageId="13" pageNumber="147" refId="ref10412" refString="Sharp, D. S. (1887) Biologia Centrali-Americana, Insecta, Coleoptera, Pselaphidae, Scydmaenidae. Vol. 2 (1). pp. 1 - 71, pls. 1 - 2." type="book" year="1887">Sharp 1887</bibRefCitation>
;
<bibRefCitation id="EF884B04214AF250FF60FA3BAB6810DE" author="Park, O." box="[222,341,1432,1459]" pageId="13" pageNumber="147" refId="ref9661" refString="Park, O. (1942) A study in neotropical Pselaphidae. Northwestern University Studies in the Biological Sciences and Medicine, Number 1. Northwestern University, Evanston and Chicago, x + 403 pp. https: // doi. org / 10.5962 / bhl. title. 6838" type="book" year="1942">Park 1942</bibRefCitation>
). From the
<collectingCountry id="F30E7665214AF250FE68FA3AA81210DE" box="[470,559,1433,1459]" name="Panama" pageId="13" pageNumber="147">Panama</collectingCountry>
Canal Zone (Barro Colorado Island, Gatun Lake) specimens of
<taxonomicName id="4C194D76214AF250FABEFA3AAFA110DF" authorityName="Park" authorityYear="1942" box="[1280,1436,1432,1459]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="sternadens">
<emphasis id="B96DEAE7214AF250FABEFA3AAFA110DF" box="[1280,1436,1432,1459]" italics="true" pageId="13" pageNumber="147">O. sternadens</emphasis>
</taxonomicName>
were collected from beneath bark of a log, and specimens of
<taxonomicName id="4C194D76214AF250FCDBFA1EA9D310BB" authorityName="Park" authorityYear="1942" box="[869,1006,1468,1495]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="escharus">
<emphasis id="B96DEAE7214AF250FCDBFA1EA9D310BB" box="[869,1006,1468,1495]" italics="true" pageId="13" pageNumber="147">O. escharus</emphasis>
</taxonomicName>
were taken from a decaying log and galleries of termites (probably
<taxonomicName id="4C194D76214AF250FE42FA42A94F1096" authority="Snyder, 1922" authorityName="Snyder" authorityYear="1922" box="[508,882,1504,1531]" class="Insecta" family="Rhinotermitidae" genus="Coptotermes" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B96DEAE7214AF250FE42FA42A8ED1097" box="[508,720,1505,1531]" italics="true" pageId="13" pageNumber="147">Coptotermes niger</emphasis>
<bibRefCitation id="EF884B04214AF250FD64FA43A94F1096" author="Snyder, T. E." box="[730,882,1504,1531]" pageId="13" pageNumber="147" pagination="1 - 32" refId="ref10451" refString="Snyder, T. E. (1922) New termites from Hawaii, Central and South America, and the Antilles. Proceedings of the United States National Museum, 60, 1 - 32 + 5 pls. https: // doi. org / 10.5479 / si. 00963801.61 - 2441.1" type="journal article" year="1922">Snyder, 1922</bibRefCitation>
</taxonomicName>
) in a fallen tree (
<bibRefCitation id="EF884B04214AF250FB81FA43AE871096" author="Park, O." box="[1087,1210,1504,1531]" pageId="13" pageNumber="147" refId="ref9661" refString="Park, O. (1942) A study in neotropical Pselaphidae. Northwestern University Studies in the Biological Sciences and Medicine, Number 1. Northwestern University, Evanston and Chicago, x + 403 pp. https: // doi. org / 10.5962 / bhl. title. 6838" type="book" year="1942">Park 1942</bibRefCitation>
). The southernmost record for the genus is of
<emphasis id="B96DEAE7214AF250FE7AF9A6A8621373" box="[452,607,1540,1567]" italics="true" pageId="13" pageNumber="147">
<taxonomicName id="4C194D76214AF250FE7AF9A6A8611373" authorityName="Bruch" authorityYear="1933" box="[452,604,1540,1567]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="attaphilus">O. attaphilus</taxonomicName>
,
</emphasis>
which was collected at Loreto, Misiones,
<collectingCountry id="F30E7665214AF250FBF9F9A7AE8B1373" box="[1095,1206,1540,1566]" name="Argentina" pageId="13" pageNumber="147">Argentina</collectingCountry>
in an ant nest (
<taxonomicName id="4C194D76214AF250FAD0F9A6AAD3132F" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Formicidae" genus="Atta" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="sexdens">
<emphasis id="B96DEAE7214AF250FAD0F9A6AAD3132F" italics="true" pageId="13" pageNumber="147">Atta sexdens</emphasis>
</taxonomicName>
L.) (
<bibRefCitation id="EF884B04214AF250FE92F98BAB8A132E" author="Bruch, C." box="[300,439,1576,1603]" pageId="13" pageNumber="147" pagination="12 - 37" refId="ref8490" refString="Bruch, C. (1933) Coleopteros mirmecofilos de Misiones (Staph. Pselaph. Hister.). Revista de Entomologia, 3 (1), 12 - 37, pl. 1." type="journal article" year="1933">Bruch 1933</bibRefCitation>
;
<bibRefCitation id="EF884B04214AF250FE78F98BA802132E" author="Park, O." box="[454,575,1576,1603]" pageId="13" pageNumber="147" refId="ref9661" refString="Park, O. (1942) A study in neotropical Pselaphidae. Northwestern University Studies in the Biological Sciences and Medicine, Number 1. Northwestern University, Evanston and Chicago, x + 403 pp. https: // doi. org / 10.5962 / bhl. title. 6838" type="book" year="1942">Park 1942</bibRefCitation>
). The southwest record is for
<taxonomicName id="4C194D76214AF250FC21F98AAE75132F" authorityName="Raffray" authorityYear="1918" box="[927,1096,1576,1603]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="rugosicollis">
<emphasis id="B96DEAE7214AF250FC21F98AAE75132F" box="[927,1096,1576,1603]" italics="true" pageId="13" pageNumber="147">O. rugosicollis</emphasis>
</taxonomicName>
from “
<collectingRegion id="49DDF817214AF250FB23F98BAF30132F" box="[1181,1293,1576,1602]" country="Paraguay" name="Asuncion" pageId="13" pageNumber="147">Asunción</collectingRegion>
” (
<collectingCountry id="F30E7665214AF250FA97F98AAFA8132E" box="[1321,1429,1577,1603]" name="Paraguay" pageId="13" pageNumber="147">Paraguay</collectingCountry>
) (
<bibRefCitation id="EF884B04214AF250FF20F9EFAB04130A" author="Raffray, A." box="[158,313,1612,1639]" pageId="13" pageNumber="147" pagination="473 - 502" refId="ref10062" refString="Raffray, A. (1918) Nouvelles especes de Pselaphides (Paraguay-Laos-Philippines). Annales de la Societe Entomologique de France, 86, 473 - 502. [Note: nominal year of publication 1917]" type="journal article" year="1918">Raffray 1918</bibRefCitation>
).
</paragraph>
<paragraph id="8BA636F5214AF250FF79F9D2A8B812C6" blockId="13.[151,1437,1324,2035]" pageId="13" pageNumber="147">
Four species were collected from
<collectingCountry id="F30E7665214AF250FDF1F9D3A89B13E7" box="[591,678,1648,1674]" name="Bolivia" pageId="13" pageNumber="147">Bolivia</collectingCountry>
: O.
<taxonomicName id="4C194D76214AF250FD69F9D3A97A13E7" authorityName="Newton" authorityYear="2017" box="[727,839,1648,1674]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="armatides">
<emphasis id="B96DEAE7214AF250FD69F9D3A97A13E7" box="[727,839,1648,1674]" italics="true" pageId="13" pageNumber="147">armatides</emphasis>
</taxonomicName>
,
<taxonomicName id="4C194D76214AF250FCE6F9D2AE3D13E7" authorityName="Newton" authorityYear="2017" box="[856,1024,1648,1675]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="simplexides">
<emphasis id="B96DEAE7214AF250FCE6F9D2AE3D13E7" box="[856,1024,1648,1675]" italics="true" pageId="13" pageNumber="147">O. simplexides</emphasis>
</taxonomicName>
,
<taxonomicName id="4C194D76214AF250FBAEF9D2AF7813E6" authority="(Raffray, 1898)" baseAuthorityName="Raffray" baseAuthorityYear="1898" box="[1040,1349,1648,1675]" class="Insecta" family="Staphylinidae" genus="Batrisus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="B96DEAE7214AF250FBAEF9D2AEB313E7" box="[1040,1166,1649,1675]" italics="true" pageId="13" pageNumber="147">O. rugosus</emphasis>
(
<bibRefCitation id="EF884B04214AF250FB20F9D3AF0013E6" author="Raffray, A." box="[1182,1341,1648,1675]" pageId="13" pageNumber="147" pagination="198 - 273" refId="ref9877" refString="Raffray, A. (1898) Notes sur les Pselaphides. Revision generique de la tribi des Euplectini. Descriptions'd'especes nouvelles. Revue'd'Entomologie, 17, 198 - 273." type="journal article" year="1898">Raffray, 1898</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C194D76214AF250FA3FF9D2ABF913C2" authority="(Raffray, 1898)" baseAuthorityName="Raffray" baseAuthorityYear="1898" class="Insecta" family="Staphylinidae" genus="Batrisus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="forticornis">
<emphasis id="B96DEAE7214AF250FA3FF9D2AB3013C2" italics="true" pageId="13" pageNumber="147">O. forticornis</emphasis>
(
<bibRefCitation id="EF884B04214AF250FEA3F937AB8113C2" author="Raffray, A." box="[285,444,1684,1711]" pageId="13" pageNumber="147" pagination="198 - 273" refId="ref9877" refString="Raffray, A. (1898) Notes sur les Pselaphides. Revision generique de la tribi des Euplectini. Descriptions'd'especes nouvelles. Revue'd'Entomologie, 17, 198 - 273." type="journal article" year="1898">Raffray, 1898</bibRefCitation>
)
</taxonomicName>
, but there is no precise information about their
<typeStatus id="54A28857214AF250FC60F936AE3313C2" box="[990,1038,1685,1711]" pageId="13" pageNumber="147">type</typeStatus>
localities since they were recorded as “
<emphasis id="B96DEAE7214AF250FF7BF91BA85613BE" box="[197,619,1720,1747]" italics="true" pageId="13" pageNumber="147">Haute-bolivie: triubu des Yuracaris”</emphasis>
(
<bibRefCitation id="EF884B04214AF250FDC3F91BA92413BE" author="Raffray, A." box="[637,793,1720,1747]" pageId="13" pageNumber="147" pagination="198 - 273" refId="ref9877" refString="Raffray, A. (1898) Notes sur les Pselaphides. Revision generique de la tribi des Euplectini. Descriptions'd'especes nouvelles. Revue'd'Entomologie, 17, 198 - 273." type="journal article" year="1898">Raffray 1898</bibRefCitation>
;
<bibRefCitation id="EF884B04214AF250FC99F91BA99C13BE" author="Park, O." box="[807,929,1720,1747]" pageId="13" pageNumber="147" refId="ref9661" refString="Park, O. (1942) A study in neotropical Pselaphidae. Northwestern University Studies in the Biological Sciences and Medicine, Number 1. Northwestern University, Evanston and Chicago, x + 403 pp. https: // doi. org / 10.5962 / bhl. title. 6838" type="book" year="1942">Park 1942</bibRefCitation>
). We believe that these species occur in the Bolivian Amazon, because the Yuracaré indigenous people lived until the 1940s in a large area of tropical rainforest at the foot of the Andes Mountains and along rivers from
<collectingRegion id="49DDF817214AF250FC99F8A2A99E1276" box="[807,931,1793,1819]" country="Bolivia" name="Santa Cruz" pageId="13" pageNumber="147">Santa Cruz</collectingRegion>
to
<collectingRegion id="49DDF817214AF250FC74F8A3AE611277" box="[970,1116,1792,1818]" country="Bolivia" name="Cochabamba" pageId="13" pageNumber="147">Cochabamba</collectingRegion>
(
<bibRefCitation id="EF884B04214AF250FBD2F8A3AFA51276" author="D'Orbigny, A. D." box="[1132,1432,1792,1819]" pageId="13" pageNumber="147" refId="ref8720" refString="D'Orbigny, A. D. (1839) L'homme americain (de l'Amerique Meridionale): considere sous ses rapports physiologiques et moraux, volume 2. Pitois-Levrault." type="book" year="1839">DOrbigny 1839: 354-355</bibRefCitation>
,
<bibRefCitation id="EF884B04214AF250FF29F886AB4D1252" author="Metraux, A." box="[151,368,1828,1855]" pageId="13" pageNumber="147" pagination="465 - 504" refId="ref9302" refString="Metraux, A. (1948) Tribes of the Eastern Slopes of the Bolivian Andes. In: Steward J. H. (eds.), Handbook of South American Indians, volume 3: The Tropical Forest Tribes: 465 - 504. Smithsonian Institute, Washington." type="book chapter" year="1948">Metraux 1948: 485</bibRefCitation>
), which belongs to the Bolivian Amazon region (
<bibRefCitation id="EF884B04214AF250FC33F887AE441252" author="Plaza Martinez, P." box="[909,1145,1828,1855]" pageId="13" pageNumber="147" refId="ref9713" refString="Plaza Martinez, P. (2011) Historia, lengua, cultura y educacion em la nacion yurakare. Cochabamba, Bolivia." type="book" year="2011">Plaza Martinez 2011</bibRefCitation>
). Additionally,
<specimenCount id="9D1FFD7C214AF250FA9CF886AFA11253" box="[1314,1436,1828,1855]" count="1" pageId="13" pageNumber="147" type="female">one female</specimenCount>
of
<taxonomicName id="4C194D76214AF250FF09F8EAAB63120F" authorityName="Newton" authorityYear="2017" box="[183,350,1864,1891]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="simplexides">
<emphasis id="B96DEAE7214AF250FF09F8EAAB63120F" box="[183,350,1864,1891]" italics="true" pageId="13" pageNumber="147">O. simplexides</emphasis>
</taxonomicName>
was subsequently collected from
<collectingCountry id="F30E7665214AF250FD5FF8EAA929120E" box="[737,788,1865,1891]" name="Peru" pageId="13" pageNumber="147">Peru</collectingCountry>
at an elevation of
<quantity id="4CE19B10214AF250FC55F8EAAE60120F" box="[1003,1117,1864,1891]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="13" pageNumber="147" unit="ft" value="2000.0">2,000 feet</quantity>
from
<taxonomicName id="4C194D76214AF250FB1AF8EBAF53120E" authority="el Perene" authorityName="el Perene" box="[1188,1390,1864,1891]" class="Insecta" family="Staphylinidae" genus="Colonia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="genus">Colonia el Perené</taxonomicName>
“El campamiento” (Chanchamayo,
<collectingRegion id="49DDF817214AF250FE44F8CEA80A12EA" box="[506,567,1901,1927]" country="Peru" name="Junin" pageId="13" pageNumber="147">Junín</collectingRegion>
Department) (
<bibRefCitation id="EF884B04214AF250FD65F8CFA94612EA" author="Fletcher, F. C." box="[731,891,1900,1927]" pageId="13" pageNumber="147" pagination="203 - 231" refId="ref8822" refString="Fletcher, F. C. (1928) Pselaphidae collected by Dr. Alfons Dampf in Central America. Annals of the Entomological Society of America, 21 (2), 203 - 231. https: // doi. org / 10.1093 / aesa / 21.2.203" type="journal article" year="1928">Fletcher 1928</bibRefCitation>
;
<bibRefCitation id="EF884B04214AF250FC39F8CFA9C012EA" author="Park, O." box="[903,1021,1900,1927]" pageId="13" pageNumber="147" refId="ref9661" refString="Park, O. (1942) A study in neotropical Pselaphidae. Northwestern University Studies in the Biological Sciences and Medicine, Number 1. Northwestern University, Evanston and Chicago, x + 403 pp. https: // doi. org / 10.5962 / bhl. title. 6838" type="book" year="1942">Park 1942</bibRefCitation>
;
<bibRefCitation id="EF884B04214AF250FBB7F8CEAEF612EB" author="Rasmussen, C." box="[1033,1227,1900,1927]" pageId="13" pageNumber="147" pagination="137 - 147" refId="ref10100" refString="Rasmussen, C. (2016) J. C. Bradley's narrative of the Cornell Entomological Expedition to South America (1919 - 1920): collecting localities and entomological travel details. Journal of the History of Collections, 28 (1), 137 - 147. https: // doi. org / 10.1093 / jhc / fhu 074" type="journal article" year="2016">Rasmussen 2016</bibRefCitation>
), which is situated in the Peruvian Amazon region (
<bibRefCitation id="EF884B04214AF250FDBCF832A84712C6" author="IGN &amp; Instituto Geografico Nacional" box="[514,634,1937,1963]" pageId="13" pageNumber="147" refId="ref9037" refString="IGN - Instituto Geografico Nacional. (2018) Atlas del Peru, Lima-Peru. Available from: https: // app 4. ign. gob. pe / capitulos / ' / regiones. php (accessed 29 July 2023)" type="url" year="2018">IGN 2018</bibRefCitation>
).
</paragraph>
<paragraph id="8BA636F5214AF253FF79F817A9511774" blockId="13.[151,1437,1324,2035]" lastBlockId="14.[151,1437,151,609]" lastPageId="14" lastPageNumber="148" pageId="13" pageNumber="147">
Previously, four species had been recorded from
<collectingCountry id="F30E7665214AF250FCB2F817A96912A3" box="[780,852,1972,1998]" name="Brazil" pageId="13" pageNumber="147">Brazil</collectingCountry>
. The species
<taxonomicName id="4C194D76214AF250FC49F816AE5612A3" authorityName="Asenjo" authorityYear="2018" box="[1015,1131,1973,1999]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="aurora">
<emphasis id="B96DEAE7214AF250FC49F816AE5612A3" box="[1015,1131,1973,1999]" italics="true" pageId="13" pageNumber="147">O. aurora</emphasis>
</taxonomicName>
was taken in the state of
<collectingRegion id="49DDF817214AF250FF29F87BAB3E129F" box="[151,259,2008,2034]" country="Brazil" name="Tocantins" pageId="13" pageNumber="147">Tocantins</collectingRegion>
and
<taxonomicName id="4C194D76214AF250FE80F87AABD2129F" authorityName="Asenjo" authorityYear="2018" box="[318,495,2009,2035]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="147" phylum="Arthropoda" rank="species" species="inexpectatus">
<emphasis id="B96DEAE7214AF250FE80F87AABD2129F" box="[318,495,2009,2035]" italics="true" pageId="13" pageNumber="147">O. inexpectatus</emphasis>
</taxonomicName>
was collected in the state of
<collectingRegion id="49DDF817214AF250FCFCF87BA9E0129F" box="[834,989,2008,2034]" country="Brazil" name="Minas Gerais" pageId="13" pageNumber="147">Minas Gerais</collectingRegion>
, both being the first troglobitic species of
<taxonomicName id="4C194D762149F253FF08FF34AB0C15DC" authorityName="Reitter" authorityYear="1883" box="[182,305,151,177]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE72149F253FF08FF34AB0C15DC" box="[182,305,151,177]" italics="true" pageId="14" pageNumber="148">Oxarthrius</emphasis>
</taxonomicName>
, and which were taken from caves (
<bibRefCitation id="EF884B042149F253FD76FF34A9A715DC" author="Asenjo, A. &amp; Zampaulo, R. de &amp; Ferreira, R. L." box="[712,922,151,177]" pageId="14" pageNumber="148" pagination="404 - 414" refId="ref8207" refString="Asenjo, A., Zampaulo, R. de A. &amp; Ferreira, R. L. (2018) Two new troglobitic species of Oxarthrius Reitter (Coleoptera, Staphylinidae, Pselaphinae) from Brazil. Zootaxa, 4462 (3), 404 - 414. https: // doi. org / 10.11646 / zootaxa. 4462.3.6" type="journal article" year="2018">
Asenjo
<emphasis id="B96DEAE72149F253FC9EFF34A96F15DC" box="[800,850,151,177]" italics="true" pageId="14" pageNumber="148">et al</emphasis>
. 2018
</bibRefCitation>
) situated in the Cerrado Biome (
<bibRefCitation id="EF884B042149F253FAB7FF34AFAC15DC" author="IBGE &amp; Instituto Brasileiro de Geografia" box="[1289,1425,151,177]" pageId="14" pageNumber="148" pagination="45" refId="ref8974" refString="IBGE - Instituto Brasileiro de Geografia e Estatistica. (2019) Biomas e Sistema Costeiro-Marinho do Brasil: compativel com a escala: 1: 250.000. Serie Relatorio Metodologicos, 45. Brasilia. Available from: https: // www. ibge. gov. br / apps / biomas / # / home (accessed 25 July 2023)" type="journal article" year="2019">IBGE 2019</bibRefCitation>
). The distribution of
<taxonomicName id="4C194D762149F253FED1FF18A82F15B8" box="[367,530,187,213]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="148" phylum="Arthropoda" rank="species" species="anthicoides">
<emphasis id="B96DEAE72149F253FED1FF18A82F15B8" box="[367,530,187,213]" italics="true" pageId="14" pageNumber="148">O. anthicoides</emphasis>
</taxonomicName>
is problematic, since it is recorded from “Brasilia” (L.W.Shauffus 1887), and the distribution of
<taxonomicName id="4C194D762149F253FE80FF7CABEB1594" baseAuthorityName="Reitter" baseAuthorityYear="1883" box="[318,470,223,249]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="148" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B96DEAE72149F253FE80FF7CABEB1594" box="[318,470,223,249]" italics="true" pageId="14" pageNumber="148">O. bispinosus</emphasis>
</taxonomicName>
is uncertain also since it was based on material from “Brasilia; Santa-Rita; September (Salhb.[Reinhold Ferdinand Sahlberg])” (
<bibRefCitation id="EF884B042149F253FDDAFEA0A9111470" author="Reitter, E." box="[612,812,259,285]" pageId="14" pageNumber="148" pagination="371 - 386" refId="ref10163" refString="Reitter, E. (1883) Neue Pselaphiden und Scydmaeniden aus Central- und Sudamerika. Verhandlungen der KaiserlichKoniglichen Zoologisch-Botanischen Gesellschaft in Wien, 32, 371 - 386. [Note: nominal year of publication 1882]" type="journal article" year="1883">Reitter 1883: 376</bibRefCitation>
). However, in same work
<bibRefCitation id="EF884B042149F253FBF2FEA0AF1D1470" author="Reitter, E." box="[1100,1312,259,285]" pageId="14" pageNumber="148" pagination="371 - 386" refId="ref10163" refString="Reitter, E. (1883) Neue Pselaphiden und Scydmaeniden aus Central- und Sudamerika. Verhandlungen der KaiserlichKoniglichen Zoologisch-Botanischen Gesellschaft in Wien, 32, 371 - 386. [Note: nominal year of publication 1882]" type="journal article" year="1883">Reitter (1883: 379)</bibRefCitation>
mentioned in a key
<taxonomicName id="4C194D762149F253FF4EFE84A819142C" authority="Rttr. Petropolis" authorityName="Rttr. Petropolis" box="[240,548,295,321]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="148" phylum="Arthropoda" rank="species" species="bispinosus">“bispinosus Rttr. Petropolis</taxonomicName>
”, Petropólis is a municipality in the state of
<collectingRegion id="49DDF8172149F253FC40FE84AEA2142C" box="[1022,1183,295,321]" country="Brazil" name="Rio de Janeiro" pageId="14" pageNumber="148">Rio de Janeiro</collectingRegion>
. Subsequently,
<bibRefCitation id="EF884B042149F253FAF9FE84AB311408" author="Raffray, A." pageId="14" pageNumber="148" pagination="198 - 273" refId="ref9877" refString="Raffray, A. (1898) Notes sur les Pselaphides. Revision generique de la tribi des Euplectini. Descriptions'd'especes nouvelles. Revue'd'Entomologie, 17, 198 - 273." type="journal article" year="1898">Raffray (1898:438</bibRefCitation>
,
<bibRefCitation id="EF884B042149F253FEA5FEE8ABBB1408" author="Raffray, A." box="[283,390,331,357]" pageId="14" pageNumber="148" pagination="1 - 487" refId="ref9981" refString="Raffray, A. (1908) Coleoptera. Fam. Pselaphidae. In: Wytsman P. (ed.), Genera Insectorum. Verteneuil &amp; L. Desmet, Bruxelles, pp. 1 - 487." type="book chapter" year="1908">1908:134</bibRefCitation>
) and
<bibRefCitation id="EF884B042149F253FE75FEE8A8B81408" author="Park, O." box="[459,645,331,357]" pageId="14" pageNumber="148" refId="ref9661" refString="Park, O. (1942) A study in neotropical Pselaphidae. Northwestern University Studies in the Biological Sciences and Medicine, Number 1. Northwestern University, Evanston and Chicago, x + 403 pp. https: // doi. org / 10.5962 / bhl. title. 6838" type="book" year="1942">Park (1942:246)</bibRefCitation>
mentioned the locality of
<taxonomicName id="4C194D762149F253FC04FEE8AE6B1408" box="[954,1110,331,357]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="148" phylum="Arthropoda" rank="species" species="bispinosus">
<emphasis id="B96DEAE72149F253FC04FEE8AE6B1408" box="[954,1110,331,357]" italics="true" pageId="14" pageNumber="148">O. bispinosus</emphasis>
</taxonomicName>
as “Santa Rita, Blumenau”. According to
<bibRefCitation id="EF884B042149F253FE8AFECCABCC14E4" author="Papavero, N." box="[308,497,367,393]" pageId="14" pageNumber="148" refId="ref9608" refString="Papavero, N. (1973) Essays on the history of Neotropical dipterology: with special reference to collectors (1750 - 1905), Vol. II. Museu de Zoologia, Universidade de Sao Paulo. https: // doi. org / 10.5962 / bhl. title. 101597" type="book" year="1973">Papavero (1973)</bibRefCitation>
, Sahlberg was in
<collectingRegion id="49DDF8172149F253FD02FECCA95D14E4" box="[700,864,367,393]" country="Brazil" name="Rio de Janeiro" pageId="14" pageNumber="148">Rio de Janeiro</collectingRegion>
in
<date id="FFA710352149F253FC3BFECCAE2B14E4" box="[901,1046,366,393]" pageId="14" pageNumber="148" value="1850-08">August 1850</date>
and collected for about 3 months, however, in the same paper it is mentioned that Sahlberg also collected in the municipality of
<collectingRegion id="49DDF8172149F253FB78FE30AF6014C0" box="[1222,1373,403,429]" country="Brazil" name="Minas Gerais" pageId="14" pageNumber="148">Minas Gerais</collectingRegion>
from Chapéu dUvas to Diamantino, presenting the possibility of Santa Rita being on his itinerary. Considering that there is no mention of Santa Rita during Sahlbergs visit to
<collectingRegion id="49DDF8172149F253FD57FE78A9B11498" box="[745,908,475,501]" country="Brazil" name="Rio de Janeiro" pageId="14" pageNumber="148">Rio de Janeiro</collectingRegion>
, it is possible that Santa Rita corresponds to the current district of Santa Rita Durão, in the state of
<collectingRegion id="49DDF8172149F253FD70FE5CA9551774" box="[718,872,511,537]" country="Brazil" name="Minas Gerais" pageId="14" pageNumber="148">Minas Gerais</collectingRegion>
.
</paragraph>
<paragraph id="8BA636F52149F253FF79FD80AFB5170C" blockId="14.[151,1437,151,609]" pageId="14" pageNumber="148">
Therefore,
<taxonomicName id="4C194D762149F253FEFAFD80A8041750" authority="Gouvea &amp; Valente &amp; Asenjo, 2023" authorityName="Gouvea &amp; Valente &amp; Asenjo" authorityYear="2023" box="[324,569,547,573]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="148" phylum="Arthropoda" rank="species" species="spiculatus" status="sp. nov.">
<emphasis id="B96DEAE72149F253FEFAFD80A8041750" box="[324,569,547,573]" italics="true" pageId="14" pageNumber="148">Oxarthrius spiculatus</emphasis>
</taxonomicName>
<emphasis id="B96DEAE72149F253FDFEFD80A8A61750" bold="true" box="[576,667,547,573]" pageId="14" pageNumber="148">
<taxonomicNameLabel id="A25E579C2149F253FDFEFD80A8A61750" box="[576,667,547,573]" pageId="14" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
recorded from Paragominas, State of
<collectingRegion id="49DDF8172149F253FBF6FD80AE401750" box="[1096,1149,547,573]" country="Brazil" name="Para" pageId="14" pageNumber="148">Pará</collectingRegion>
, is the first species of the genus recorded from the Brazilian Amazon. Herein we expand the distribution of the genus to
<location id="8EC6602E2149F253FB07FDE4AFB9170C" LSID="urn:lsid:plazi:treatment:03B087E3214AF253FF29FF34AFB5170C:8EC6602E2149F253FB07FDE4AFB9170C" box="[1209,1412,583,609]" name="eastern Amazonia" pageId="14" pageNumber="148" stateProvince="Para">eastern Amazonia</location>
.
</paragraph>
</subSubSection>
</treatment>
</document>