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<document id="9559EA2F286425115F75B0DBDEB94E2E" ID-CLB-Dataset="8430" ID-DOI="10.11646/zootaxa.4853.1.2" ID-GBIF-Dataset="067e9fe1-910c-42f8-829f-ded1324ff8b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="4410313" ID-ZooBank="6D8FBF75-C96D-4B70-8BC6-BDB2C920C440" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609599355198" checkinUser="plazi" docAuthor="Arimoto, Kôichi" docDate="2020" docId="03B087E0FFD6FFFD7792017EFDADFD33" docLanguage="en" docName="zootaxa.4853.1.2.pdf" docOrigin="Zootaxa 4853 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calambus taiwanensis Arimoto 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="47" masterDocId="FF89FF98FFD4FFF87705056CFFD4FFB6" masterDocTitle="First record and new species of the genus Calambus Thomson (Coleoptera Elateridae, Dendrometrinae) from Taiwan" masterLastPageNumber="50" masterPageNumber="42" pageNumber="44" updateTime="1698851133678" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="80BC85D58A2A18CD6D64AF9CD84BFF3C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8DE722F72C244DC0F91A24F4E5BB9B01">
<mods:title id="B11DF4C68A2D4468032E54750A3AB5E9">First record and new species of the genus Calambus Thomson (Coleoptera Elateridae, Dendrometrinae) from Taiwan</mods:title>
</mods:titleInfo>
<mods:name id="950A6CA4D5C63AEB5CD7CE6FB68EA5E8" type="personal">
<mods:role id="1EE498AB17FA66823F1A8C2040F652DB">
<mods:roleTerm id="77DD9ADFC4E44E4EC3F7B5F692727B6E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="26AE9C4FBA171342724EE4D72C33A82A">Arimoto, Kôichi</mods:namePart>
</mods:name>
<mods:typeOfResource id="9CF2D76F1E314889AD36BC363B8EDD65">text</mods:typeOfResource>
<mods:relatedItem id="0604319A8AC2E9A8E7C6617CC97D034C" type="host">
<mods:titleInfo id="5AD374B2BD87676EA259F1E212BB8CE2">
<mods:title id="4D7179EE4DF8049C763B57A6556C2704">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B75DD6BF242CFB5DB4D71FADA15E787E">
<mods:date id="8110DCE8AE0E247C5F4E308277DD3C10">2020</mods:date>
<mods:detail id="B7B3FEAC0FEA19597B8783B6481B3D89" type="pubDate">
<mods:number id="C610E827CD4A09B10BB342B4152C02DC">2020-09-21</mods:number>
</mods:detail>
<mods:detail id="05B4DECCA72ABE172D44E7E3D2952A3A" type="volume">
<mods:number id="1D433804AC7F80422459453D4470C636">4853</mods:number>
</mods:detail>
<mods:detail id="E106F7F3BA10E213589C9143702440E2" type="issue">
<mods:number id="AAA5E95EE6A69C648896DB57EAB18EFF">1</mods:number>
</mods:detail>
<mods:extent id="71FF9F633225CD0D2EC67C464C8FFC54" unit="page">
<mods:start id="3AFF2F45C547C30587053311C8C8FAFC">42</mods:start>
<mods:end id="35A475AEC32098D8EBA8F5745D3ED392">50</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="25938C6F574B410C41ABACC5FEE23822">journal article</mods:classification>
<mods:identifier id="F65E1BFB09D9EFAA5B506C93B8AC2091" type="CLB-Dataset">8430</mods:identifier>
<mods:identifier id="9A42217475A9E2A6644A9180F8F62AC4" type="DOI">10.11646/zootaxa.4853.1.2</mods:identifier>
<mods:identifier id="509494668F9F4C86FCE2960928E27D3A" type="GBIF-Dataset">067e9fe1-910c-42f8-829f-ded1324ff8b0</mods:identifier>
<mods:identifier id="A8E91D125BA57A4EA916DE283AE5D535" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D7EA4C702C848DB178A0FAE11D57A964" type="Zenodo-Dep">4410313</mods:identifier>
<mods:identifier id="32E606CACB8F64870CFC26B43E1E8A0E" type="ZooBank">6D8FBF75-C96D-4B70-8BC6-BDB2C920C440</mods:identifier>
</mods:mods>
<treatment id="03B087E0FFD6FFFD7792017EFDADFD33" ID-DOI="http://doi.org/10.5281/zenodo.4410309" ID-GBIF-Taxon="177215456" ID-Zenodo-Dep="4410309" LSID="urn:lsid:plazi:treatment:03B087E0FFD6FFFD7792017EFDADFD33" httpUri="http://treatment.plazi.org/id/03B087E0FFD6FFFD7792017EFDADFD33" lastPageId="5" lastPageNumber="47" pageId="2" pageNumber="44">
<subSubSection id="C303657DFFD6FFFA7792017EFDB3FB9B" box="[151,615,1042,1069]" pageId="2" pageNumber="44" type="nomenclature">
<paragraph id="8BA636F6FFD6FFFA7792017EFDB3FB9B" blockId="2.[151,615,1042,1105]" box="[151,615,1042,1069]" pageId="2" pageNumber="44">
<heading id="D0EE819AFFD6FFFA7792017EFDB3FB9B" bold="true" box="[151,615,1042,1069]" fontSize="11" level="1" pageId="2" pageNumber="44" reason="1">
<emphasis id="B96DEAE4FFD6FFFA7792017EFDB3FB9B" bold="true" box="[151,615,1042,1069]" pageId="2" pageNumber="44">
<taxonomicName id="4C194D75FFD6FFFA7792017EFDCCFB9A" authority="Arimoto" authorityName="Arimoto" authorityYear="2020" box="[151,536,1042,1069]" class="Insecta" family="Elateridae" genus="Calambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="44" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="B96DEAE4FFD6FFFA7792017EFE73FB9B" bold="true" box="[151,423,1042,1069]" italics="true" pageId="2" pageNumber="44">Calambus taiwanensis</emphasis>
Arimoto
</taxonomicName>
,
<taxonomicNameLabel id="A25E579FFFD6FFFA7520017FFDB3FB9B" box="[549,615,1043,1069]" pageId="2" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C303657DFFD6FFFA7792015BFEE4FBE7" box="[151,304,1079,1105]" pageId="2" pageNumber="44" type="description">
<paragraph id="8BA636F6FFD6FFFA7792015BFEE4FBE7" blockId="2.[151,615,1042,1105]" box="[151,304,1079,1105]" pageId="2" pageNumber="44">
(
<figureCitation id="13222A73FFD6FFFA779A015BFEF3FBE7" box="[159,295,1079,1105]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1162,1186]" captionStartId-1="4.[151,250,1695,1719]" captionStartId-2="5.[151,250,1633,1657]" captionStartId-3="6.[151,250,1935,1959]" captionTargetBox-0="[164,1428,373,1143]" captionTargetBox-1="[151,1436,489,1672]" captionTargetBox-2="[159,1431,669,1601]" captionTargetBox-3="[163,1424,669,1912]" captionTargetId-0="figure-130@3.[151,1436,363,1143]" captionTargetId-1="figure-141@4.[151,1436,489,1672]" captionTargetId-2="figure-262@5.[151,1436,669,1609]" captionTargetId-3="figure-250@6.[161,1427,669,1912]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Calambus taiwanensis Arimoto sp. n. A–B: male, holotype [CTA001]; C: female, paratype [CTA002]. A, C: habitus, dorsal; B: habitus, ventral." captionText-1="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." captionText-2="FIGURE 3. Calambus taiwanensis Arimoto sp. n., male, holotype [CTA001]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of left paramere, dorsal view; H: apical part of left paramere, ventral view." captionText-3="FIGURE 4. Calambus taiwanensis Arimoto sp. n., female, paratype.A–C: [CTA004]; D–F: [CTA008]. A: tergite VIII and sternite VIII, dorsal view; B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (arrow: vagina with coiled sac extending from ventral side); E: apical part of ovipositor, dorsal side; F: ditto, ventral side." figureDoi-0="http://doi.org/10.5281/zenodo.4410315" figureDoi-1="http://doi.org/10.5281/zenodo.4410317" figureDoi-2="http://doi.org/10.5281/zenodo.4410319" figureDoi-3="http://doi.org/10.5281/zenodo.4410321" httpUri-0="https://zenodo.org/record/4410315/files/figure.png" httpUri-1="https://zenodo.org/record/4410317/files/figure.png" httpUri-2="https://zenodo.org/record/4410319/files/figure.png" httpUri-3="https://zenodo.org/record/4410321/files/figure.png" pageId="2" pageNumber="44">Figures 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C303657DFFD6FFFA77920113FC59FB2F" box="[151,909,1151,1177]" pageId="2" pageNumber="44" type="etymology">
<paragraph id="8BA636F6FFD6FFFA77920113FC59FB2F" blockId="2.[151,1437,1151,2041]" box="[151,909,1151,1177]" pageId="2" pageNumber="44">
<emphasis id="B96DEAE4FFD6FFFA77920113FECAFB2F" bold="true" box="[151,286,1151,1177]" pageId="2" pageNumber="44">Etymology.</emphasis>
Specific epithet derived from
<collectingCountry id="F30E7666FFD6FFFA75760113FD1CFB2F" box="[627,712,1151,1177]" name="Taiwan" pageId="2" pageNumber="44">Taiwan</collectingCountry>
, the
<typeStatus id="54A28854FFD6FFFA75F90113FCF8FB2F" box="[764,812,1151,1177]" pageId="2" pageNumber="44">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C303657DFFD6FFFA77C201CFFE95FA23" pageId="2" pageNumber="44" type="materials_examined">
<paragraph id="8BA636F6FFD6FFFA77C201CFFE95FA23" blockId="2.[151,1437,1151,2041]" pageId="2" pageNumber="44">
<emphasis id="B96DEAE4FFD6FFFA77C201CFFEAEFB0A" bold="true" box="[199,378,1186,1213]" pageId="2" pageNumber="44">Type material.</emphasis>
<materialsCitation id="3B713CABFFD6FFFA768301CFFED9FB57" ID-GBIF-Occurrence="3031209301" collectingDate="1971-04-04" collectorName="Kiyoshi Matsuda" country="Taiwan" location="Male" municipality="Sung Kang" pageId="2" pageNumber="44" specimenCount="1" stateProvince="Nantou" typeStatus="holotype">
<emphasis id="B96DEAE4FFD6FFFA768301CFFE27FB0B" bold="true" box="[390,499,1187,1213]" pageId="2" pageNumber="44">
<typeStatus id="54A28854FFD6FFFA768301CFFE27FB0B" box="[390,499,1187,1213]" pageId="2" pageNumber="44" type="holotype">Holotype</typeStatus>
</emphasis>
.
<location id="8EC6602DFFD6FFFA750201CFFD97FB0B" LSID="urn:lsid:plazi:treatment:03B087E0FFD6FFFD7792017EFDADFD33:8EC6602DFFD6FFFA750201CFFD97FB0B" box="[519,579,1187,1213]" country="Taiwan" municipality="Sung Kang" name="Male" pageId="2" pageNumber="44" stateProvince="Nantou">Male</location>
,
<collectingMunicipality id="6BC2AC8CFFD6FFFA755601CFFD0CFB0B" box="[595,728,1187,1213]" pageId="2" pageNumber="44">Sung Kang</collectingMunicipality>
,
<collectingRegion id="49DDF814FFD6FFFA75EC01CFFC49FB0B" box="[745,925,1187,1213]" country="Taiwan" name="Nantou" pageId="2" pageNumber="44">Nantou County</collectingRegion>
,
<collectingCountry id="F30E7666FFD6FFFA74A801CFFBD6FB0B" box="[941,1026,1187,1213]" name="Taiwan" pageId="2" pageNumber="44">Taiwan</collectingCountry>
,
<date id="FFA71036FFD6FFFA731701CFFB44FB0B" box="[1042,1168,1187,1213]" pageId="2" pageNumber="44" value="1971-04-04">
<collectingDate id="EFE3E9DEFFD6FFFA731701CFFB44FB0B" box="[1042,1168,1187,1213]" pageId="2" pageNumber="44" value="1971-04-04">4 IV 1971</collectingDate>
</date>
,
<collectorName id="26EC5320FFD6FFFA73A401CFFAB2FB0B" box="[1185,1382,1187,1213]" pageId="2" pageNumber="44">Kiyoshi Matsuda</collectorName>
leg. [CTA001]
</materialsCitation>
.
<emphasis id="B96DEAE4FFD6FFFA761C01ABFE45FB57" bold="true" box="[281,401,1223,1249]" pageId="2" pageNumber="44">
<typeStatus id="54A28854FFD6FFFA761C01ABFE45FB57" box="[281,401,1223,1249]" pageId="2" pageNumber="44" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D1FFD7FFFD6FFFA76A401ABFE28FB57" box="[417,508,1223,1249]" pageId="2" pageNumber="44" type="male">2 males</specimenCount>
,
<specimenCount id="9D1FFD7FFFD6FFFA750D01AAFDAFFB54" box="[520,635,1222,1250]" pageId="2" pageNumber="44" type="female">5 females</specimenCount>
).
<specimenCount id="9D1FFD7FFFD6FFFA758B01ABFD27FB57" box="[654,755,1223,1249]" pageId="2" pageNumber="44" type="female">1 female</specimenCount>
,
<materialsCitation id="3B713CABFFD6FFFA75FA01ABFB75FB57" ID-GBIF-Occurrence="3031209303" box="[767,1185,1222,1249]" collectingDate="1971-04-04" collectorName="Kiyoshi Matsuda" country="Taiwan" location="Male" municipality="Sung Kang" pageId="2" pageNumber="44" specimenCount="1" stateProvince="Nantou" typeStatus="paratype">same data as the holotype [CTA002]</materialsCitation>
;
<specimenCount id="9D1FFD7FFFD6FFFA73AB01ABFAC7FB57" box="[1198,1299,1223,1249]" pageId="2" pageNumber="44" type="female">1 female</specimenCount>
, same place as the
<typeStatus id="54A28854FFD6FFFA77E10187FE9DFAB3" box="[228,329,1259,1285]" pageId="2" pageNumber="44" type="holotype">holotype</typeStatus>
, S. Takeda leg. [CTA003];
<specimenCount id="9D1FFD7FFFD6FFFA75830187FD38FAB3" box="[646,748,1259,1285]" pageId="2" pageNumber="44" type="female">1 female</specimenCount>
,
<materialsCitation id="3B713CABFFD6FFFA75FF0187FE71FA9F" ID-GBIF-Occurrence="3031209305" collectingDate="1972-04-12" collectionCode="IV" collectorName="Kiyoshi Matsuda" country="Taiwan" location="Nanshanchi" pageId="2" pageNumber="44" specimenCount="1" stateProvince="Nantou" typeStatus="paratype">
<location id="8EC6602DFFD6FFFA75FF0187FC55FAB3" LSID="urn:lsid:plazi:treatment:03B087E0FFD6FFFD7792017EFDADFD33:8EC6602DFFD6FFFA75FF0187FC55FAB3" box="[762,897,1259,1285]" country="Taiwan" name="Nanshanchi" pageId="2" pageNumber="44" stateProvince="Nantou">Nanshanchi</location>
,
<collectingRegion id="49DDF814FFD6FFFA748B0187FBEBFAB3" box="[910,1087,1259,1285]" country="Taiwan" name="Nantou" pageId="2" pageNumber="44">Nantou County</collectingRegion>
,
<collectingCountry id="F30E7666FFD6FFFA73480187FB76FAB3" box="[1101,1186,1259,1285]" name="Taiwan" pageId="2" pageNumber="44">Taiwan</collectingCountry>
,
<date id="FFA71036FFD6FFFA73AA0187FAE2FAB3" box="[1199,1334,1259,1285]" pageId="2" pageNumber="44" value="1972-04-12">
<collectingDate id="EFE3E9DEFFD6FFFA73AA0187FAE2FAB3" box="[1199,1334,1259,1285]" pageId="2" pageNumber="44" value="1972-04-12">
12
<collectionCode id="ED08AE33FFD6FFFA73D00187FB26FAB3" box="[1237,1266,1259,1285]" pageId="2" pageNumber="44">IV</collectionCode>
1972
</collectingDate>
</date>
,
<collectorName id="26EC5320FFD6FFFA72460187FF23FA9F" pageId="2" pageNumber="44">Kiyoshi Matsuda</collectorName>
leg. [CTA004]
</materialsCitation>
;
<specimenCount id="9D1FFD7FFFD6FFFA76B50063FE29FA9F" box="[432,509,1295,1321]" pageId="2" pageNumber="44" type="male">1 male</specimenCount>
, same place as the former,
<date id="FFA71036FFD6FFFA742E0063FC4BFA9F" box="[811,927,1295,1321]" pageId="2" pageNumber="44" value="1973-04-07">
7
<collectionCode id="ED08AE33FFD6FFFA74450063FC89FA9F" box="[832,861,1295,1321]" pageId="2" pageNumber="44">IV</collectionCode>
1973
</date>
, Y. Kiyoyama leg. [CTA005];
<specimenCount id="9D1FFD7FFFD6FFFA73F10063FA8CFA9F" box="[1268,1368,1295,1321]" pageId="2" pageNumber="44" type="female">1 female</specimenCount>
, same place as the former,
<date id="FFA71036FFD6FFFA767E005FFE2AFAFB" box="[379,510,1331,1357]" pageId="2" pageNumber="44" value="1987-04-17">
17
<collectionCode id="ED08AE33FFD6FFFA769A005FFE68FAFB" box="[415,444,1331,1357]" pageId="2" pageNumber="44">IV</collectionCode>
1987
</date>
, C. Lo leg. [CTA006];
<specimenCount id="9D1FFD7FFFD6FFFA7406005FFC85FAFB" box="[771,849,1331,1357]" pageId="2" pageNumber="44" type="male">1 male</specimenCount>
,
<materialsCitation id="3B713CABFFD6FFFA7458005FFDABFAC7" ID-GBIF-Occurrence="3031209302" collectingDate="1985-04-25" collectionCode="IV" country="Taiwan" location="Mt. Kuan-tou Shan" pageId="2" pageNumber="44" specimenCount="1" stateProvince="Nantou" typeStatus="paratype">
<location id="8EC6602DFFD6FFFA7458005FFBECFAFB" LSID="urn:lsid:plazi:treatment:03B087E0FFD6FFFD7792017EFDADFD33:8EC6602DFFD6FFFA7458005FFBECFAFB" box="[861,1080,1331,1357]" country="Taiwan" name="Mt. Kuan-tou Shan" pageId="2" pageNumber="44" stateProvince="Nantou">Mt. Kuan-tou Shan</location>
,
<collectingRegion id="49DDF814FFD6FFFA7346005FFB25FAFB" box="[1091,1265,1331,1357]" country="Taiwan" name="Nantou" pageId="2" pageNumber="44">Nantou County</collectingRegion>
,
<collectingCountry id="F30E7666FFD6FFFA73F9005FFA85FAFB" box="[1276,1361,1331,1357]" name="Taiwan" pageId="2" pageNumber="44">Taiwan</collectingCountry>
,
<date id="FFA71036FFD6FFFA7259005EFF1BFAC7" pageId="2" pageNumber="44" value="1985-04-25">
<collectingDate id="EFE3E9DEFFD6FFFA7259005EFF1BFAC7" pageId="2" pageNumber="44" value="1985-04-25">
25
<collectionCode id="ED08AE33FFD6FFFA727A005FFA48FAFB" box="[1407,1436,1331,1357]" pageId="2" pageNumber="44">IV</collectionCode>
1985
</collectingDate>
</date>
(the collector is not stated) [CTA007]
</materialsCitation>
;
<specimenCount id="9D1FFD7FFFD6FFFA758C003BFD3AFAC7" box="[649,750,1367,1393]" pageId="2" pageNumber="44" type="female">1 female</specimenCount>
,
<materialsCitation id="3B713CABFFD6FFFA75FC003BFEEAFA23" ID-GBIF-Occurrence="3031209304" collectingDate="2005-07-11" collectorName="F. Y. Huang" country="Taiwan" location="Puli" pageId="2" pageNumber="44" specimenCount="1" stateProvince="Nantou" typeStatus="paratype">
near
<location id="8EC6602DFFD6FFFA7435003BFCB4FAC7" LSID="urn:lsid:plazi:treatment:03B087E0FFD6FFFD7792017EFDADFD33:8EC6602DFFD6FFFA7435003BFCB4FAC7" box="[816,864,1367,1393]" country="Taiwan" name="Puli" pageId="2" pageNumber="44" stateProvince="Nantou">Puli</location>
,
<collectingRegion id="49DDF814FFD6FFFA746E003BFBCDFAC7" box="[875,1049,1367,1393]" country="Taiwan" name="Nantou" pageId="2" pageNumber="44">Nantou County</collectingRegion>
,
<collectingCountry id="F30E7666FFD6FFFA7326003BFBACFAC7" box="[1059,1144,1367,1393]" name="Taiwan" pageId="2" pageNumber="44">Taiwan</collectingCountry>
,
<date id="FFA71036FFD6FFFA7386003BFAD9FAC7" box="[1155,1293,1366,1393]" pageId="2" pageNumber="44" value="2005-07-11">
<collectingDate id="EFE3E9DEFFD6FFFA7386003BFAD9FAC7" box="[1155,1293,1366,1393]" pageId="2" pageNumber="44" value="2005-07-11">11 VII 2005</collectingDate>
</date>
,
<collectorName id="26EC5320FFD6FFFA721C003BFA49FAC7" box="[1305,1437,1367,1393]" pageId="2" pageNumber="44">F. Y. Huang</collectorName>
leg. [CTA008]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C303657DFFD6FFFA77C200F2FBECF903" pageId="2" pageNumber="44" type="diagnosis">
<paragraph id="8BA636F6FFD6FFFA77C200F2FBECF903" blockId="2.[151,1437,1151,2041]" pageId="2" pageNumber="44">
<emphasis id="B96DEAE4FFD6FFFA77C200F2FE96FA0E" bold="true" box="[199,322,1438,1464]" pageId="2" pageNumber="44">Diagnosis.</emphasis>
Body surface generally shining; interspaces between fine punctures distinctly larger than puncture diameter. Body generally black, but elytra dark brown, with outer margin paler. Body covered with yellow white setae. Antennae extending beyond apex of pronotal hind angles by two apical antennomeres in male, but not reaching pronotum posterior lateral apices by apical antennomere in female; antennomere III obconical, slightly longer than II; IV shorter than II–III combined, slightly longer than V. Hind angles of prothorax short, narrow, with apex rounded. Scutellar shield widest at anterior margin; anterior margin slightly rounded; side emarginate anteriorly; posterior margin broadly rounded. Parameres of aedeagus without lateral subapical barb. Female genitalia with robust baculum; vagina with coiled sac; bursa copulatrix without sclerotized piece.
</paragraph>
</subSubSection>
<subSubSection id="C303657DFFD6FFFD77C203D3FB48FDD7" lastPageId="5" lastPageNumber="47" pageId="2" pageNumber="44" type="description">
<paragraph id="8BA636F6FFD6FFFA77C203D3FA51F8DF" blockId="2.[151,1437,1151,2041]" pageId="2" pageNumber="44">
<emphasis id="B96DEAE4FFD6FFFA77C203D3FE11F96F" bold="true" box="[199,453,1727,1753]" pageId="2" pageNumber="44">Measurements. Male</emphasis>
(n=3,
<typeStatus id="54A28854FFD6FFFA751203D3FDADF96F" box="[535,633,1727,1753]" pageId="2" pageNumber="44" type="holotype">holotype</typeStatus>
in parentheses). BL: 5.02–5.51 (5.51), BW: 1.54–1.69 (1.69), MBE: 0.58–0.63 (0.63), MAE: 0.84–0.94 (0.94), OI: 143–152 (149), PL: 1.29–1.41 (1.34), PML: 1.14–1.23 (1.16), PW: 1.29–1.40 (1.40), PI: 95.5–100 (95.5), EL: 3.40–3.86 (3.86), EW: 1.54–1.69 (1.69), EI: 221–229 (229), BI: 258–289 (289).
<emphasis id="B96DEAE4FFD6FFFA77E70247FEEDF8F3" bold="true" box="[226,313,1835,1861]" pageId="2" pageNumber="44">Female</emphasis>
(n=5). BL: 5.45–6.38, BW: 1.64–1.96, MBE: 0.61–0.71, MAE: 0.85–1.00, OI: 138–144, PL: 1.38– 1.65, PML: 1.21–1.42, PW: 1.41–1.69, PI: 95.7–108, EL: 3.72–4.41, EW: 1.64–1.96, EI: 213–226, BI: 235–279.
</paragraph>
<paragraph id="8BA636F6FFD6FFFB77C2021EFC4AFED3" blockId="2.[151,1437,1151,2041]" lastBlockId="3.[151,1437,151,357]" lastPageId="3" lastPageNumber="45" pageId="2" pageNumber="44">
<emphasis id="B96DEAE4FFD6FFFA77C2021EFE8DF83A" bold="true" box="[199,345,1906,1932]" pageId="2" pageNumber="44">Description.</emphasis>
Body surface generally smooth and shining, but elytra weakly rough; interspaces between fine punctures distinctly larger than puncture diameter, but larger punctures of anterior edge of pronotum partly overlapping (
<figureCitation id="13222A73FFD6FFFA77DD02D7FEFAF863" box="[216,302,1979,2005]" captionStart="FIGURE 1" captionStartId="3.[151,250,1162,1186]" captionTargetBox="[164,1428,373,1143]" captionTargetId="figure-130@3.[151,1436,363,1143]" captionTargetPageId="3" captionText="FIGURE 1. Calambus taiwanensis Arimoto sp. n. A–B: male, holotype [CTA001]; C: female, paratype [CTA002]. A, C: habitus, dorsal; B: habitus, ventral." figureDoi="http://doi.org/10.5281/zenodo.4410315" httpUri="https://zenodo.org/record/4410315/files/figure.png" pageId="2" pageNumber="44">Fig. 1A</figureCitation>
); large coarse punctures spread all over surface of anterior lobe of prosternum (
<figureCitation id="13222A73FFD6FFFA73AD02D7FB29F863" box="[1192,1277,1979,2005]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="2" pageNumber="44">Fig. 2F</figureCitation>
); punctures of hypomeron near pronotosternal suture large and their interspaces smaller than puncture diameter (
<figureCitation id="13222A73FFD6FFFA73E902B3FA94F84F" box="[1260,1344,2015,2041]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="2" pageNumber="44">Fig. 2F</figureCitation>
). Color. Body generally black, but elytra dark brown, with outer margin paler. (
<figureCitation id="13222A73FFD7FFFB74CF05FBFBB8FF07" box="[970,1132,151,177]" captionStart="FIGURE 1" captionStartId="3.[151,250,1162,1186]" captionTargetBox="[164,1428,373,1143]" captionTargetId="figure-130@3.[151,1436,363,1143]" captionTargetPageId="3" captionText="FIGURE 1. Calambus taiwanensis Arimoto sp. n. A–B: male, holotype [CTA001]; C: female, paratype [CTA002]. A, C: habitus, dorsal; B: habitus, ventral." figureDoi="http://doi.org/10.5281/zenodo.4410315" httpUri="https://zenodo.org/record/4410315/files/figure.png" pageId="3" pageNumber="45">Fig. 1A, B, C</figureCitation>
). Labrum black (
<figureCitation id="13222A73FFD7FFFB723605FBFA44FF07" box="[1331,1424,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2A</figureCitation>
). Antennae and mouth parts generally dark brown; mandible black, with apical and internal parts orange (
<figureCitation id="13222A73FFD7FFFB723105D7FA5AFF63" box="[1332,1422,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2A</figureCitation>
). Legs dark brown, becoming paler apically (
<figureCitation id="13222A73FFD7FFFB757B05B3FD01FF4F" box="[638,725,223,249]" captionStart="FIGURE 1" captionStartId="3.[151,250,1162,1186]" captionTargetBox="[164,1428,373,1143]" captionTargetId="figure-130@3.[151,1436,363,1143]" captionTargetPageId="3" captionText="FIGURE 1. Calambus taiwanensis Arimoto sp. n. A–B: male, holotype [CTA001]; C: female, paratype [CTA002]. A, C: habitus, dorsal; B: habitus, ventral." figureDoi="http://doi.org/10.5281/zenodo.4410315" httpUri="https://zenodo.org/record/4410315/files/figure.png" pageId="3" pageNumber="45">Fig. 1B</figureCitation>
). Tergites and sternites VIII‒X yellow brown, but median basal area of tergite VIII and posterior area of sternite VIII translucent (
<figureCitation id="13222A73FFD7FFFB7477046FFC3DFEAB" box="[882,1001,259,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[159,1431,669,1601]" captionTargetId="figure-262@5.[151,1436,669,1609]" captionTargetPageId="5" captionText="FIGURE 3. Calambus taiwanensis Arimoto sp. n., male, holotype [CTA001]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of left paramere, dorsal view; H: apical part of left paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4410319" httpUri="https://zenodo.org/record/4410319/files/figure.png" pageId="3" pageNumber="45">Fig. 3A, B</figureCitation>
). Aedeagus yellow brown (
<figureCitation id="13222A73FFD7FFFB7219046FFA5BFEAB" box="[1308,1423,259,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[159,1431,669,1601]" captionTargetId="figure-262@5.[151,1436,669,1609]" captionTargetPageId="5" captionText="FIGURE 3. Calambus taiwanensis Arimoto sp. n., male, holotype [CTA001]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of left paramere, dorsal view; H: apical part of left paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4410319" httpUri="https://zenodo.org/record/4410319/files/figure.png" pageId="3" pageNumber="45">Fig. 3E, F</figureCitation>
). Hairs. Body covered with yellow-white setae (
<figureCitation id="13222A73FFD7FFFB759F044BFCDBFEF7" box="[666,783,295,321]" captionStart="FIGURE 1" captionStartId="3.[151,250,1162,1186]" captionTargetBox="[164,1428,373,1143]" captionTargetId="figure-130@3.[151,1436,363,1143]" captionTargetPageId="3" captionText="FIGURE 1. Calambus taiwanensis Arimoto sp. n. A–B: male, holotype [CTA001]; C: female, paratype [CTA002]. A, C: habitus, dorsal; B: habitus, ventral." figureDoi="http://doi.org/10.5281/zenodo.4410315" httpUri="https://zenodo.org/record/4410315/files/figure.png" pageId="3" pageNumber="45">Fig. 1A, B</figureCitation>
). Antennae and mouth parts except for labrum with yellow to brown setae. Setae of legs yellow white becoming yellow apically.
</paragraph>
<caption id="DF66667EFFD7FFFB779201E6FE1AFB70" ID-DOI="http://doi.org/10.5281/zenodo.4410315" ID-Zenodo-Dep="4410315" httpUri="https://zenodo.org/record/4410315/files/figure.png" pageId="3" pageNumber="45" startId="3.[151,250,1162,1186]" targetBox="[164,1428,373,1143]" targetPageId="3">
<paragraph id="8BA636F6FFD7FFFB779201E6FE1AFB70" blockId="3.[151,1436,1162,1222]" pageId="3" pageNumber="45">
<emphasis id="B96DEAE4FFD7FFFB779201E6FEC7FB14" bold="true" box="[151,275,1162,1186]" pageId="3" pageNumber="45">FIGURE 1.</emphasis>
<taxonomicName id="4C194D75FFD7FFFB761C01E6FD8DFB14" authority="Arimoto" authorityName="Arimoto" authorityYear="2020" box="[281,601,1162,1186]" class="Insecta" family="Elateridae" genus="Calambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="45" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="B96DEAE4FFD7FFFB761C01E6FE2AFB14" box="[281,510,1162,1186]" italics="true" pageId="3" pageNumber="45">Calambus taiwanensis</emphasis>
Arimoto
</taxonomicName>
<emphasis id="B96DEAE4FFD7FFFB755A01E6FD4CFB14" bold="true" box="[607,664,1162,1186]" pageId="3" pageNumber="45">
<taxonomicNameLabel id="A25E579FFFD7FFFB755A01E6FD4CFB14" box="[607,664,1162,1186]" pageId="3" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A–B: male, holotype [CTA001]; C: female, paratype [CTA002]. A, C: habitus, dorsal; B: habitus, ventral.
</paragraph>
</caption>
<paragraph id="8BA636F6FFD7FFFB77C2019FFC53F9B3" blockId="3.[151,1437,1267,2039]" pageId="3" pageNumber="45">
Head. Frons without depression. Supra-antennal carina incomplete medially (
<figureCitation id="13222A73FFD7FFFB734E019FFB7EFABB" box="[1099,1194,1267,1293]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2A</figureCitation>
); frontclypeal region gradually sloping to base of labrum (
<figureCitation id="13222A73FFD7FFFB753A007AFD68FA87" box="[575,700,1302,1329]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2A, B</figureCitation>
). Antennomeres I, III, IV and IX–XI longer than wide (
<figureCitation id="13222A73FFD7FFFB723B007AFF79FAE2" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2B, C</figureCitation>
); II globular, shortest, as long as wide; III obconical, slightly longer than II; IV–X serrate; IV shorter than II–III combined, slightly longer than V.
<emphasis id="B96DEAE4FFD7FFFB75150032FD80FACE" bold="true" box="[528,596,1374,1400]" pageId="3" pageNumber="45">Male.</emphasis>
Antennae extending beyond apex of pronotal hind angles by two apical antennomeres (
<figureCitation id="13222A73FFD7FFFB762200EDFEABFA2D" box="[295,383,1409,1435]" captionStart="FIGURE 1" captionStartId="3.[151,250,1162,1186]" captionTargetBox="[164,1428,373,1143]" captionTargetId="figure-130@3.[151,1436,363,1143]" captionTargetPageId="3" captionText="FIGURE 1. Calambus taiwanensis Arimoto sp. n. A–B: male, holotype [CTA001]; C: female, paratype [CTA002]. A, C: habitus, dorsal; B: habitus, ventral." figureDoi="http://doi.org/10.5281/zenodo.4410315" httpUri="https://zenodo.org/record/4410315/files/figure.png" pageId="3" pageNumber="45">Fig. 1A</figureCitation>
); antennomere IV 1.16–1.35 x as long as wide, 1.74–2.00 x as long as III; V–VIII longer than wide; XI 2.40–2.78 x as long as wide.
<emphasis id="B96DEAE4FFD7FFFB755100C9FD66FA09" bold="true" box="[596,690,1445,1471]" pageId="3" pageNumber="45">Female.</emphasis>
Antennae not reaching pronotum posterior lateral apices by apical antennomere (
<figureCitation id="13222A73FFD7FFFB763200A4FE58FA54" box="[311,396,1480,1506]" captionStart="FIGURE 1" captionStartId="3.[151,250,1162,1186]" captionTargetBox="[164,1428,373,1143]" captionTargetId="figure-130@3.[151,1436,363,1143]" captionTargetPageId="3" captionText="FIGURE 1. Calambus taiwanensis Arimoto sp. n. A–B: male, holotype [CTA001]; C: female, paratype [CTA002]. A, C: habitus, dorsal; B: habitus, ventral." figureDoi="http://doi.org/10.5281/zenodo.4410315" httpUri="https://zenodo.org/record/4410315/files/figure.png" pageId="3" pageNumber="45">Fig. 1C</figureCitation>
); antennomere IV 1.18–1.45 x as long as wide, 1.53–1.68 x as long as III; V–VIII almost as long as wide; XI 1.82–2.14 x as long as wide. Mandible bidentate.
</paragraph>
<paragraph id="8BA636F6FFD7FFFC77C20363FE9BFE67" blockId="3.[151,1437,1267,2039]" lastBlockId="4.[151,1437,151,465]" lastPageId="4" lastPageNumber="46" pageId="3" pageNumber="45">
Prothorax sub-trapezoidal, slightly shorter than wide or as long as wide, widest at hind angles; side distinctly rounded, weakly narrowed anterior to hind angles (
<figureCitation id="13222A73FFD7FFFB75DF035EFC80F9FB" box="[730,852,1586,1613]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2D, E</figureCitation>
); anterior angles acute; lateral carina extending just behind anterior angles (
<figureCitation id="13222A73FFD7FFFB76A1033AFE2BF9C6" box="[420,511,1622,1648]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2B</figureCitation>
: arrow) to hind angles; hind angles divergent, short, narrow, straight, incurved in some specimens (
<figureCitation id="13222A73FFD7FFFB76580315FE02F922" box="[349,470,1657,1684]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2D, E</figureCitation>
), with apex rounded. Pronotum without median longitudinal depression; median basal tubercle indistinct; median basal furrow poorly developed; sublateral incision near hind angles weak; hind angles without carina. Hypomeron; mesial edge weakly rounded, with impunctate ridge next to pronotosternal suture, carinate anterolaterally (
<figureCitation id="13222A73FFD7FFFB76780388FE07F948" box="[381,467,1764,1790]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2F</figureCitation>
: black arrow); posterior margin sinuate, with large emargination near hind angles (
<figureCitation id="13222A73FFD7FFFB726A0388FF02F897" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2F, G</figureCitation>
). Prosternum in profile slightly concave anterior to mesocoxal cavity (
<figureCitation id="13222A73FFD7FFFB74F4026BFB98F897" box="[1009,1100,1799,1825]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2H</figureCitation>
: arrow); anterior lobe broadly rounded, exceeding anterior angles of prothorax (
<figureCitation id="13222A73FFD7FFFB75C80246FCF1F8F3" box="[717,805,1834,1861]" captionStart="FIGURE 1" captionStartId="3.[151,250,1162,1186]" captionTargetBox="[164,1428,373,1143]" captionTargetId="figure-130@3.[151,1436,363,1143]" captionTargetPageId="3" captionText="FIGURE 1. Calambus taiwanensis Arimoto sp. n. A–B: male, holotype [CTA001]; C: female, paratype [CTA002]. A, C: habitus, dorsal; B: habitus, ventral." figureDoi="http://doi.org/10.5281/zenodo.4410315" httpUri="https://zenodo.org/record/4410315/files/figure.png" pageId="3" pageNumber="45">Fig. 1B</figureCitation>
), in profile strongly inclined (at 39–54 degrees against prosternal horizontal line,
<figureCitation id="13222A73FFD7FFFB76C60222FDCAF8DE" box="[451,542,1870,1896]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2H</figureCitation>
: red line) and in some specimens concave just behind anterior lobe. Prosternal spine in profile strongly inclined (at 24–36 degrees against prosternal horizontal line,
<figureCitation id="13222A73FFD7FFFB7363021DFB69F83A" box="[1126,1213,1905,1932]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2H</figureCitation>
), incurved, without subapical tooth; sides in ventral view straight (
<figureCitation id="13222A73FFD7FFFB759C02F9FD24F819" box="[665,752,1941,1967]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2G</figureCitation>
); ventral margin of dorsal lobe in profile slightly sinuate (
<figureCitation id="13222A73FFD7FFFB726A02F9FF69F865" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2H</figureCitation>
: yellow line); apex in ventral view rather rounded (
<figureCitation id="13222A73FFD7FFFB75F102D5FC99F865" box="[756,845,1977,2003]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2G</figureCitation>
), in profile rounded (
<figureCitation id="13222A73FFD7FFFB733202D5FB45F865" box="[1079,1169,1977,2003]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2H</figureCitation>
). Pronotosternal sutures not grooved, moderately opened (
<figureCitation id="13222A73FFD7FFFB751502B1FDB0F841" box="[528,612,2013,2039]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2F</figureCitation>
: white arrow). Scutellar shield (
<figureCitation id="13222A73FFD7FFFB74CD02B1FBC1F841" box="[968,1045,2013,2039]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="3" pageNumber="45">Fig. 2I</figureCitation>
) 1.11–1.49 x as long as wide, widest at anterior margin, strongly inclined anterior-downwards; anterior margin slightly rounded, with transversely micro-striated ridge, laterally bent downwards; side emarginate anteriorly and then convex posteriorly; posterior margin broadly rounded. Borders of mesosternal cavity horizontal anteriorly and strongly declivous posteriorly (
<figureCitation id="13222A73FFD0FFFC77A5046FFF29FEAB" box="[160,253,259,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="4" pageNumber="46">Fig. 2H</figureCitation>
: green line); posterior margin of mesosternite between mesocoxae wider than half width of mesocoxa (
<figureCitation id="13222A73FFD0FFFC77A5044BFF23FEF7" box="[160,247,295,321]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="4" pageNumber="46">Fig. 2G</figureCitation>
). Mesepisternum reaching mesocoxal cavity (
<figureCitation id="13222A73FFD0FFFC75FE044BFC84FEF7" box="[763,848,295,321]" captionStart="FIGURE 2" captionStartId="4.[151,250,1695,1719]" captionTargetBox="[151,1436,489,1672]" captionTargetId="figure-141@4.[151,1436,489,1672]" captionTargetPageId="4" captionText="FIGURE 2. Calambus taiwanensis Arimoto sp. n. A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="4" pageNumber="46">Fig. 2F</figureCitation>
: triangle). Metasternum sulcate medially and behind mesocoxae (
<figureCitation id="13222A73FFD0FFFC76210427FEA8FED3" box="[292,380,331,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1162,1186]" captionTargetBox="[164,1428,373,1143]" captionTargetId="figure-130@3.[151,1436,363,1143]" captionTargetPageId="3" captionText="FIGURE 1. Calambus taiwanensis Arimoto sp. n. A–B: male, holotype [CTA001]; C: female, paratype [CTA002]. A, C: habitus, dorsal; B: habitus, ventral." figureDoi="http://doi.org/10.5281/zenodo.4410315" httpUri="https://zenodo.org/record/4410315/files/figure.png" pageId="4" pageNumber="46">Fig. 1B</figureCitation>
). Elytra convex; outmost interval narrowly extending sideways, forming flat plate, with a lateral carina; striae I–V grooved, but V partly reduced; remaining striae ungrooved and defined only by rows of punctures; apices rounded. Hind wings complete. Legs simple; tarsomeres I–IV becoming shorter apically; V longest.
</paragraph>
<caption id="DF66667EFFD0FFFC779203F3FF0FF861" ID-DOI="http://doi.org/10.5281/zenodo.4410317" ID-Zenodo-Dep="4410317" httpUri="https://zenodo.org/record/4410317/files/figure.png" pageId="4" pageNumber="46" startId="4.[151,250,1695,1719]" targetBox="[151,1436,489,1672]" targetPageId="4">
<paragraph id="8BA636F6FFD0FFFC779203F3FF0FF861" blockId="4.[151,1437,1695,2007]" pageId="4" pageNumber="46">
<emphasis id="B96DEAE4FFD0FFFC779203F3FEC1F901" bold="true" box="[151,277,1695,1719]" pageId="4" pageNumber="46">FIGURE 2.</emphasis>
<taxonomicName id="4C194D75FFD0FFFC761B03F3FDB7F901" authority="Arimoto" authorityName="Arimoto" authorityYear="2020" box="[286,611,1695,1719]" class="Insecta" family="Elateridae" genus="Calambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="B96DEAE4FFD0FFFC761B03F3FDD2F901" box="[286,518,1695,1719]" italics="true" pageId="4" pageNumber="46">Calambus taiwanensis</emphasis>
Arimoto
</taxonomicName>
<emphasis id="B96DEAE4FFD0FFFC756E03CCFD73F90E" bold="true" box="[619,679,1696,1720]" pageId="4" pageNumber="46">
<taxonomicNameLabel id="A25E579FFFD0FFFC756E03CCFD73F90E" box="[619,679,1696,1720]" pageId="4" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A–D, F–I: male, holotype [CTA001]; E: female, paratype [CTA004]. A: head, anterior view; B: head and pronotum in laterodorsal view and basal five antennomeres of right antenna (arrow: lateral carina of prothorax extending just behind anterior angles); C: apical four antennomeres of right antenna; D, E: left side of pronotum; F: anterior half of habitus, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterally; white arrow: pronotosternal sutures moderately opened; white triangle: mesepisternum reaching mesocoxal cavity); G: posterior half of prothorax and mesothorax, ventral view; H: anterior half of habitus, lateral view (arrow: prosternum slightly concave anterior to mesocoxal cavity; red line: anterior lobe of prosternum strongly inclined; yellow line: dorsal lobe of prosternal spine with ventral margin slightly sinuate; green line: borders of mesosternal cavity horizontal followed by strongly declivous); I: scutellar shield.
</paragraph>
</caption>
<paragraph id="8BA636F6FFD1FFFD77C205FBFED3FE3F" blockId="5.[151,1437,150,645]" pageId="5" pageNumber="47">
Abdomen. Ventrite V 0.59–0.67 x as long as wide.
<emphasis id="B96DEAE4FFD1FFFD740C05FBFC98FF07" bold="true" box="[777,844,151,177]" pageId="5" pageNumber="47">Male.</emphasis>
Tergite VIII (
<figureCitation id="13222A73FFD1FFFD74E305FBFBE2FF04" box="[998,1078,151,178]" captionStart="FIGURE 3" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[159,1431,669,1601]" captionTargetId="figure-262@5.[151,1436,669,1609]" captionTargetPageId="5" captionText="FIGURE 3. Calambus taiwanensis Arimoto sp. n., male, holotype [CTA001]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of left paramere, dorsal view; H: apical part of left paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4410319" httpUri="https://zenodo.org/record/4410319/files/figure.png" pageId="5" pageNumber="47">Fig.3A</figureCitation>
) ca. 0.70 x as long as wide; apical margin strongly prominent and broadly rounded. Sternite (
<figureCitation id="13222A73FFD1FFFD744905D7FC76FF63" box="[844,930,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[159,1431,669,1601]" captionTargetId="figure-262@5.[151,1436,669,1609]" captionTargetPageId="5" captionText="FIGURE 3. Calambus taiwanensis Arimoto sp. n., male, holotype [CTA001]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of left paramere, dorsal view; H: apical part of left paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4410319" httpUri="https://zenodo.org/record/4410319/files/figure.png" pageId="5" pageNumber="47">Fig. 3B</figureCitation>
) VIII without median notch; posterior angles slightly prominent and rounded. Tergite IX (
<figureCitation id="13222A73FFD1FFFD758B05B3FD08FF4C" box="[654,732,223,250]" captionStart="FIGURE 3" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[159,1431,669,1601]" captionTargetId="figure-262@5.[151,1436,669,1609]" captionTargetPageId="5" captionText="FIGURE 3. Calambus taiwanensis Arimoto sp. n., male, holotype [CTA001]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of left paramere, dorsal view; H: apical part of left paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4410319" httpUri="https://zenodo.org/record/4410319/files/figure.png" pageId="5" pageNumber="47">Fig.3C</figureCitation>
) longer than wide, ca. 1.4 x length of tergite X; median notch rounded. Tergite X (
<figureCitation id="13222A73FFD1FFFD7679046FFE1FFEA8" box="[380,459,259,286]" captionStart="FIGURE 3" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[159,1431,669,1601]" captionTargetId="figure-262@5.[151,1436,669,1609]" captionTargetPageId="5" captionText="FIGURE 3. Calambus taiwanensis Arimoto sp. n., male, holotype [CTA001]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of left paramere, dorsal view; H: apical part of left paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4410319" httpUri="https://zenodo.org/record/4410319/files/figure.png" pageId="5" pageNumber="47">Fig.3C</figureCitation>
) ca. 1.5 x as long as wide; posterior margin rounded. Sternite IX (
<figureCitation id="13222A73FFD1FFFD73B9046FFAD8FEA8" box="[1212,1292,259,286]" captionStart="FIGURE 3" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[159,1431,669,1601]" captionTargetId="figure-262@5.[151,1436,669,1609]" captionTargetPageId="5" captionText="FIGURE 3. Calambus taiwanensis Arimoto sp. n., male, holotype [CTA001]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of left paramere, dorsal view; H: apical part of left paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4410319" httpUri="https://zenodo.org/record/4410319/files/figure.png" pageId="5" pageNumber="47">Fig.3D</figureCitation>
) ca. 2.9 x as long as wide, rounded apically, concave laterally.
<emphasis id="B96DEAE4FFD1FFFD75C4044BFCCBFEF7" bold="true" box="[705,799,295,321]" pageId="5" pageNumber="47">Female.</emphasis>
Tergite VIII ca. 0.95 x as long as wide (
<figureCitation id="13222A73FFD1FFFD73DF044BFAE6FEF7" box="[1242,1330,295,321]" captionStart="FIGURE 4" captionStartId="6.[151,250,1935,1959]" captionTargetBox="[163,1424,669,1912]" captionTargetId="figure-250@6.[161,1427,669,1912]" captionTargetPageId="6" captionText="FIGURE 4. Calambus taiwanensis Arimoto sp. n., female, paratype.A–C: [CTA004]; D–F: [CTA008]. A: tergite VIII and sternite VIII, dorsal view; B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (arrow: vagina with coiled sac extending from ventral side); E: apical part of ovipositor, dorsal side; F: ditto, ventral side." figureDoi="http://doi.org/10.5281/zenodo.4410321" httpUri="https://zenodo.org/record/4410321/files/figure.png" pageId="5" pageNumber="47">Fig. 4B</figureCitation>
). Sternite VIII with side almost parallel on basal half (
<figureCitation id="13222A73FFD1FFFD75910427FD3AFED3" box="[660,750,331,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1935,1959]" captionTargetBox="[163,1424,669,1912]" captionTargetId="figure-250@6.[161,1427,669,1912]" captionTargetPageId="6" captionText="FIGURE 4. Calambus taiwanensis Arimoto sp. n., female, paratype.A–C: [CTA004]; D–F: [CTA008]. A: tergite VIII and sternite VIII, dorsal view; B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (arrow: vagina with coiled sac extending from ventral side); E: apical part of ovipositor, dorsal side; F: ditto, ventral side." figureDoi="http://doi.org/10.5281/zenodo.4410321" httpUri="https://zenodo.org/record/4410321/files/figure.png" pageId="5" pageNumber="47">Fig. 4C</figureCitation>
); spiculum ventrale robust, ca. 2.0 x as long as sternite VIII (
<figureCitation id="13222A73FFD1FFFD77A50403FF2EFE3F" box="[160,250,367,393]" captionStart="FIGURE 4" captionStartId="6.[151,250,1935,1959]" captionTargetBox="[163,1424,669,1912]" captionTargetId="figure-250@6.[161,1427,669,1912]" captionTargetPageId="6" captionText="FIGURE 4. Calambus taiwanensis Arimoto sp. n., female, paratype.A–C: [CTA004]; D–F: [CTA008]. A: tergite VIII and sternite VIII, dorsal view; B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (arrow: vagina with coiled sac extending from ventral side); E: apical part of ovipositor, dorsal side; F: ditto, ventral side." figureDoi="http://doi.org/10.5281/zenodo.4410321" httpUri="https://zenodo.org/record/4410321/files/figure.png" pageId="5" pageNumber="47">Fig. 4A</figureCitation>
).
</paragraph>
<paragraph id="8BA636F6FFD1FFFD77C204FFFB48FDD7" blockId="5.[151,1437,150,645]" pageId="5" pageNumber="47">
Genitalia.
<emphasis id="B96DEAE4FFD1FFFD763904FFFEADFE1B" bold="true" box="[316,377,403,429]" pageId="5" pageNumber="47">Male</emphasis>
(
<figureCitation id="13222A73FFD1FFFD768C04FFFE2AFE1B" box="[393,510,403,429]" captionStart="FIGURE 3" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[159,1431,669,1601]" captionTargetId="figure-262@5.[151,1436,669,1609]" captionTargetPageId="5" captionText="FIGURE 3. Calambus taiwanensis Arimoto sp. n., male, holotype [CTA001]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of left paramere, dorsal view; H: apical part of left paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4410319" httpUri="https://zenodo.org/record/4410319/files/figure.png" pageId="5" pageNumber="47">Fig. 3E, F</figureCitation>
). Basal piece ca. 0.25 x total length of aedeagus. Median lobe exceeding apices of parameres; basal struts ca. 0.20 x total length of median lobe. Parameres separated ventrally, without lateral subapical barb, not expanded apically; apex with several setae dorsally (
<figureCitation id="13222A73FFD1FFFD74A904B7FBD3FE43" box="[940,1031,475,501]" captionStart="FIGURE 3" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[159,1431,669,1601]" captionTargetId="figure-262@5.[151,1436,669,1609]" captionTargetPageId="5" captionText="FIGURE 3. Calambus taiwanensis Arimoto sp. n., male, holotype [CTA001]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of left paramere, dorsal view; H: apical part of left paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4410319" httpUri="https://zenodo.org/record/4410319/files/figure.png" pageId="5" pageNumber="47">Fig. 3G</figureCitation>
), with one seta ventrally (
<figureCitation id="13222A73FFD1FFFD723604B7FA5BFE43" box="[1331,1423,475,501]" captionStart="FIGURE 3" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[159,1431,669,1601]" captionTargetId="figure-262@5.[151,1436,669,1609]" captionTargetPageId="5" captionText="FIGURE 3. Calambus taiwanensis Arimoto sp. n., male, holotype [CTA001]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of left paramere, dorsal view; H: apical part of left paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4410319" httpUri="https://zenodo.org/record/4410319/files/figure.png" pageId="5" pageNumber="47">Fig. 3H</figureCitation>
), with two setae apically (
<figureCitation id="13222A73FFD1FFFD76AB0493FDF9FDAF" box="[430,557,511,537]" captionStart="FIGURE 3" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[159,1431,669,1601]" captionTargetId="figure-262@5.[151,1436,669,1609]" captionTargetPageId="5" captionText="FIGURE 3. Calambus taiwanensis Arimoto sp. n., male, holotype [CTA001]. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of left paramere, dorsal view; H: apical part of left paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4410319" httpUri="https://zenodo.org/record/4410319/files/figure.png" pageId="5" pageNumber="47">Fig. 3G, H</figureCitation>
).
<emphasis id="B96DEAE4FFD1FFFD75470493FD74FDAF" bold="true" box="[578,672,511,537]" pageId="5" pageNumber="47">Female.</emphasis>
Baculum robust, ca. 2.4 x as long as coxites (
<figureCitation id="13222A73FFD1FFFD73AA0493FADFFDAF" box="[1199,1291,511,537]" captionStart="FIGURE 4" captionStartId="6.[151,250,1935,1959]" captionTargetBox="[163,1424,669,1912]" captionTargetId="figure-250@6.[161,1427,669,1912]" captionTargetPageId="6" captionText="FIGURE 4. Calambus taiwanensis Arimoto sp. n., female, paratype.A–C: [CTA004]; D–F: [CTA008]. A: tergite VIII and sternite VIII, dorsal view; B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (arrow: vagina with coiled sac extending from ventral side); E: apical part of ovipositor, dorsal side; F: ditto, ventral side." figureDoi="http://doi.org/10.5281/zenodo.4410321" httpUri="https://zenodo.org/record/4410321/files/figure.png" pageId="5" pageNumber="47">Fig. 4D</figureCitation>
); coxites not segmented (
<figureCitation id="13222A73FFD1FFFD761B074FFE44FD8B" box="[286,400,547,573]" captionStart="FIGURE 4" captionStartId="6.[151,250,1935,1959]" captionTargetBox="[163,1424,669,1912]" captionTargetId="figure-250@6.[161,1427,669,1912]" captionTargetPageId="6" captionText="FIGURE 4. Calambus taiwanensis Arimoto sp. n., female, paratype.A–C: [CTA004]; D–F: [CTA008]. A: tergite VIII and sternite VIII, dorsal view; B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (arrow: vagina with coiled sac extending from ventral side); E: apical part of ovipositor, dorsal side; F: ditto, ventral side." figureDoi="http://doi.org/10.5281/zenodo.4410321" httpUri="https://zenodo.org/record/4410321/files/figure.png" pageId="5" pageNumber="47">Fig. 4E, F</figureCitation>
), with some setae at each side; stylus with a seta near apex. Vagina with coiled sac extending from ventral side (
<figureCitation id="13222A73FFD1FFFD7663072BFE15FDD7" box="[358,449,583,609]" captionStart="FIGURE 4" captionStartId="6.[151,250,1935,1959]" captionTargetBox="[163,1424,669,1912]" captionTargetId="figure-250@6.[161,1427,669,1912]" captionTargetPageId="6" captionText="FIGURE 4. Calambus taiwanensis Arimoto sp. n., female, paratype.A–C: [CTA004]; D–F: [CTA008]. A: tergite VIII and sternite VIII, dorsal view; B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (arrow: vagina with coiled sac extending from ventral side); E: apical part of ovipositor, dorsal side; F: ditto, ventral side." figureDoi="http://doi.org/10.5281/zenodo.4410321" httpUri="https://zenodo.org/record/4410321/files/figure.png" pageId="5" pageNumber="47">Fig. 4D</figureCitation>
: arrow); bursa copulatrix globular, without sclerotized structures.
</paragraph>
</subSubSection>
<subSubSection id="C303657DFFD1FFFD77C20706FDADFD33" box="[199,633,618,645]" pageId="5" pageNumber="47" type="distribution">
<paragraph id="8BA636F6FFD1FFFD77C20706FDADFD33" blockId="5.[151,1437,150,645]" box="[199,633,618,645]" pageId="5" pageNumber="47">
<emphasis id="B96DEAE4FFD1FFFD77C20706FEB5FD32" bold="true" box="[199,353,618,644]" pageId="5" pageNumber="47">Distribution.</emphasis>
<collectingCountry id="F30E7666FFD1FFFD76620707FE69FD33" box="[359,445,619,645]" name="Taiwan" pageId="5" pageNumber="47">Taiwan</collectingCountry>
:
<collectingRegion id="49DDF814FFD1FFFD76CD0707FDA1FD33" box="[456,629,619,645]" country="Taiwan" name="Nantou" pageId="5" pageNumber="47">Nantou County</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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