treatments-xml/data/03/B0/87/03B087CBFFC6FFADFE18FA28FE0EE7F7.xml

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<mods:title id="117654D5117999DD511E2690920E39B8">Taxonomic revision of the Hydroporus bodemeyeri species complex (Coleoptera: Dytiscidae) with a geometric morphometric analysis of body shape within the group</mods:title>
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<mods:namePart id="0FA383524B758280FB5935374E554B12">Hájek, Jirí</mods:namePart>
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<mods:namePart id="7082628111519724C9BB033CB74474FE">Fikácek, Martin</mods:namePart>
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<mods:title id="199D172D065166B6ECC8C943D3312027">Journal of Natural History</mods:title>
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<mods:date id="F706B9F488370FA5ABC02B7E4BD8350F">2010</mods:date>
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<taxonomicName id="4C194D5EFFC6FFA2FE18FA28FCC2E4C7" ID-CoL="5HZWT" authority="Hájek &amp; Fikácek, 2010" authorityName="Hájek &amp; Fikácek" authorityYear="2010" box="[472,811,1447,1471]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1635" phylum="Arthropoda" rank="subSpecies" species="anatolicus" status="subsp. nov." subSpecies="koksali">
<emphasis id="B96DEACFFFC6FFA2FE18FA28FCC2E4C7" bold="true" box="[472,811,1447,1471]" italics="true" pageId="6" pageNumber="1635">Hydroporus anatolicus koksali</emphasis>
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<taxonomicNameLabel id="A25E57B4FFC6FFA2FCF2FA28FC66E4C7" box="[818,911,1447,1471]" pageId="6" pageNumber="1635" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
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<treatmentCitation id="0AB810CCFFC6FFA2FF14FA67FD02E487" author="Erman OK &amp; Erman O" box="[212,747,1511,1536]" page="31" pageId="6" pageNumber="1635" year="2007">
<taxonomicName id="4C194D5EFFC6FFA2FF14FA67FD02E487" ID-CoL="3N8PR" authority=": Erman et al. 2007: 31" authorityName="Erman" authorityPageNumber="31" authorityYear="2007" box="[212,747,1511,1536]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1635" phylum="Arthropoda" rank="species" species="anatolicus">
<emphasis id="B96DEACFFFC6FFA2FF14FA67FE3DE778" box="[212,468,1512,1536]" italics="true" pageId="6" pageNumber="1635">Hydroporus anatolicus</emphasis>
: 
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(distribution); 
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<bibRefCitation id="EF884B2CFFC6FFA2FC60FA68FEA5E767" author="Darilmaz MC &amp; Kiyak S" pageId="6" pageNumber="1635" pagination="1585 - 1636" refId="ref13176" refString="Darilmaz MC, Kiyak S. 2009. Checklist of Gyrinidae, Haliplidae, Noteridae and Dytiscidae of Turkey (Coleoptera: Adephaga). J Nat Hist. 43 (25 - 26): 1585 - 1636." type="journal article" year="2009">Darilmaz and Kiyak 2009: 1614</bibRefCitation>
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(catalogue; partim.).
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locality.
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, 
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, 
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West Borçka
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, 
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[pass], ca. 
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, 
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.
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<caption id="DF666655FFC7FFA3FF50FA7CFD0FE706" ID-DOI="http://doi.org/10.5281/zenodo.5210005" ID-Zenodo-Dep="5210005" httpUri="https://zenodo.org/record/5210005/files/figure.png" pageId="7" pageNumber="1636" startId="7.[144,212,1523,1545]" targetBox="[145,1102,176,1490]" targetPageId="7">
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Figure 2(A–L). Median lobe in lateral and ventral view of (A) 
<taxonomicName id="4C194D5EFFC7FFA3FCC7FA7CFBA0E771" authority="anatolicus" authorityName="anatolicus J. Balfour-Browne" authorityYear="1963" box="[775,1097,1523,1545]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1636" phylum="Arthropoda" rank="subSpecies" species="anatolicus" subSpecies="anatolicus">
<emphasis id="B96DEACFFFC7FFA3FCC7FA7CFBA0E771" box="[775,1097,1523,1545]" italics="true" pageId="7" pageNumber="1636">Hydroporus anatolicus anatolicus</emphasis>
</taxonomicName>
; (C) 
<taxonomicName id="4C194D5EFFC7FFA3FF78F99EFE61E75F" authorityName="Hájek &amp; Fikácek" authorityYear="2010" box="[184,392,1553,1575]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1636" phylum="Arthropoda" rank="subSpecies" species="anatolicus" subSpecies="koksali">
<emphasis id="B96DEACFFFC7FFA3FF78F99EFE61E75F" box="[184,392,1553,1575]" italics="true" pageId="7" pageNumber="1636">H. anatolicus koksali</emphasis>
</taxonomicName>
ssp. nov.; (E) 
<taxonomicName id="4C194D5EFFC7FFA3FDDBF99EFCC8E75F" authority="bodemeyeri" authorityName="bodemeyeri Ganglbauer" authorityYear="1900" box="[539,801,1553,1575]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1636" phylum="Arthropoda" rank="subSpecies" species="bodemeyeri" subSpecies="bodemeyeri">
<emphasis id="B96DEACFFFC7FFA3FDDBF99EFCC8E75F" box="[539,801,1553,1575]" italics="true" pageId="7" pageNumber="1636">H. bodemeyeri bodemeyeri</emphasis>
</taxonomicName>
; (G) 
<taxonomicName id="4C194D5EFFC7FFA3FC99F99EFBA6E75F" authorityName="Hájek &amp; Fikácek" authorityYear="2010" box="[857,1103,1553,1575]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1636" phylum="Arthropoda" rank="subSpecies" species="bodemeyeri" subSpecies="cariaensis">
<emphasis id="B96DEACFFFC7FFA3FC99F99EFBA6E75F" box="[857,1103,1553,1575]" italics="true" pageId="7" pageNumber="1636">H. bodemeyeri cariaensis</emphasis>
</taxonomicName>
ssp. nov.; (H) 
<taxonomicName id="4C194D5EFFC7FFA3FEDEF9A1FDEFE73C" authorityName="Hájek &amp; Fikácek" authorityYear="2010" baseAuthorityName="Gschwendtner" baseAuthorityYear="1935" box="[286,518,1582,1604]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1636" phylum="Arthropoda" rank="subSpecies" species="bodemeyeri" subSpecies="guignoti">
<emphasis id="B96DEACFFFC7FFA3FEDEF9A1FDEFE73C" box="[286,518,1582,1604]" italics="true" pageId="7" pageNumber="1636">H. bodemeyeri guignoti</emphasis>
</taxonomicName>
; (I) 
<taxonomicName id="4C194D5EFFC7FFA3FDF1F9A1FD53E73C" authorityName="Miller and Fery" authorityYear="1995" box="[561,698,1582,1604]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1636" phylum="Arthropoda" rank="species" species="cuprescens">
<emphasis id="B96DEACFFFC7FFA3FDF1F9A1FD53E73C" box="[561,698,1582,1604]" italics="true" pageId="7" pageNumber="1636">H. cuprescens</emphasis>
</taxonomicName>
; (K) 
<taxonomicName id="4C194D5EFFC7FFA3FD2FF9A1FC7AE73C" authority="Hájek &amp; Fikácek, 2010" authorityName="Hájek &amp; Fikácek" authorityYear="2010" box="[751,915,1582,1604]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1636" phylum="Arthropoda" rank="species" species="kurdistanicus" status="sp. nov.">
<emphasis id="B96DEACFFFC7FFA3FD2FF9A1FC7AE73C" box="[751,915,1582,1604]" italics="true" pageId="7" pageNumber="1636">H. kurdistanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57B4FFC7FFA3FC58F9A0FC00E73C" box="[920,1001,1583,1604]" pageId="7" pageNumber="1636" rank="species">sp. nov.</taxonomicNameLabel>
Paramere of (B) 
<taxonomicName id="4C194D5EFFC7FFA3FF12F9C4FDC9E719" authority="anatolicus" authorityName="anatolicus J. Balfour-Browne" authorityYear="1963" box="[210,544,1611,1633]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1636" phylum="Arthropoda" rank="subSpecies" species="anatolicus" subSpecies="anatolicus">
<emphasis id="B96DEACFFFC7FFA3FF12F9C4FDC9E719" box="[210,544,1611,1633]" italics="true" pageId="7" pageNumber="1636">Hydroporus anatolicus anatolicus</emphasis>
</taxonomicName>
; (D) 
<taxonomicName id="4C194D5EFFC7FFA3FD96F9C4FCC3E719" authorityName="Hájek &amp; Fikácek" authorityYear="2010" box="[598,810,1611,1633]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1636" phylum="Arthropoda" rank="subSpecies" species="anatolicus" subSpecies="koksali">
<emphasis id="B96DEACFFFC7FFA3FD96F9C4FCC3E719" box="[598,810,1611,1633]" italics="true" pageId="7" pageNumber="1636">H. anatolicus koksali</emphasis>
</taxonomicName>
ssp. nov.; (F) 
<taxonomicName id="4C194D5EFFC7FFA3FC7DF9C4FEEBE707" authority="bodemeyeri" authorityName="bodemeyeri Ganglbauer" authorityYear="1900" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1636" phylum="Arthropoda" rank="subSpecies" species="bodemeyeri" subSpecies="bodemeyeri">
<emphasis id="B96DEACFFFC7FFA3FC7DF9C4FEEBE707" italics="true" pageId="7" pageNumber="1636">H. bodemeyeri bodemeyeri</emphasis>
</taxonomicName>
; (J) 
<taxonomicName id="4C194D5EFFC7FFA3FEEFF9E6FE50E707" authorityName="Miller and Fery" authorityYear="1995" box="[303,441,1641,1663]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1636" phylum="Arthropoda" rank="species" species="cuprescens">
<emphasis id="B96DEACFFFC7FFA3FEEFF9E6FE50E707" box="[303,441,1641,1663]" italics="true" pageId="7" pageNumber="1636">H. cuprescens</emphasis>
</taxonomicName>
; (L) 
<taxonomicName id="4C194D5EFFC7FFA3FE2BF9E6FD66E707" authority="Hájek &amp; Fikácek, 2010" authorityName="Hájek &amp; Fikácek" authorityYear="2010" box="[491,655,1641,1663]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1636" phylum="Arthropoda" rank="species" species="kurdistanicus" status="sp. nov.">
<emphasis id="B96DEACFFFC7FFA3FE2BF9E6FD66E707" box="[491,655,1641,1663]" italics="true" pageId="7" pageNumber="1636">H. kurdistanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57B4FFC7FFA3FD55F9E6FD0FE706" box="[661,742,1641,1662]" pageId="7" pageNumber="1636" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
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<emphasis id="B96DEACFFFC8FFACFF14FF28FE9EE1C7" box="[212,375,167,191]" italics="true" pageId="8" pageNumber="1637">Type material.</emphasis>
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<typeStatus id="54A2887FFFC8FFACFEBDFF28FE0CE1C7" box="[381,485,167,191]" pageId="8" pageNumber="1637" type="holotype">Holotype</typeStatus>
male (
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): “ 
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, 
<collectingRegion id="49DDF83FFFC8FFACFCE4FF28FC59E1C7" box="[804,944,167,191]" country="Turkey" name="Artvin" pageId="8" pageNumber="1637">Artvin prov.</collectingRegion>
, / 
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W Borçka
</locationDeviation>
, / CANKURTARAN GEÇIDI [pass], / 
<location id="8EC66006FFC8FFACFD84FF48FDBEE1A7" LSID="urn:lsid:plazi:treatment:03B087CBFFC6FFADFE18FA28FE0EE7F7:8EC66006FFC8FFACFD84FF48FDBEE1A7" box="[580,599,199,223]" country="Turkey" latitude="41.395" longLatPrecision="115" longitude="41.533333" name="Ji" pageId="8" pageNumber="1637" stateProvince="Artvin">Ji</location>
<emphasis id="B96DEACFFFC8FFACFD97FF48FD88E1A7" bold="true" box="[599,609,199,223]" pageId="8" pageNumber="1637">r</emphasis>
í Hájek &amp; Jan R 
<emphasis id="B96DEACFFFC8FFACFCE1FF48FCD9E1A7" bold="true" box="[801,816,199,223]" pageId="8" pageNumber="1637">8</emphasis>
ži 
<emphasis id="B96DEACFFFC8FFACFC82FF48FCA7E1A7" bold="true" box="[834,846,199,223]" pageId="8" pageNumber="1637">C</emphasis>
ka leg. [printed] // 
<date id="FFA7101DFFC8FFACFBE3FF48FEE3E187" pageId="8" pageNumber="1637" value="2004-06-29" valueMax="2004-06-30" valueMin="2004-06-29">
<collectingDate id="EFE3E9F5FFC8FFACFBE3FF48FEE3E187" pageId="8" pageNumber="1637" value="2004-06-29" valueMax="2004-06-30" valueMin="2004-06-29">29.–30.VI. 2004</collectingDate>
</date>
, 730– 
<quantity id="4CE19B38FFC8FFACFE87FF68FE62E187" box="[327,395,231,255]" metricMagnitude="2" metricUnit="m" metricValue="7.75" pageId="8" pageNumber="1637" unit="m" value="775.0">
<elevation id="0034D1EEFFC8FFACFE87FF68FE62E187" box="[327,395,231,255]" metricMagnitude="2" metricUnit="m" metricValue="7.75" pageId="8" pageNumber="1637" unit="m" value="775.0">775 m</elevation>
</quantity>
, / 
<geoCoordinate id="EE2D501AFFC8FFACFE62FF68FDE6E187" box="[418,527,230,255]" degrees="41" direction="north" minutes="23.7" orientation="latitude" pageId="8" pageNumber="1637" precision="92" value="41.395">41°23.7′N</geoCoordinate>
<geoCoordinate id="EE2D501AFFC8FFACFDD4FF68FD63E187" box="[532,650,230,255]" degrees="041" direction="east" minutes="32.0" orientation="longitude" pageId="8" pageNumber="1637" precision="92" value="41.533333">041°32.0′E</geoCoordinate>
– / 
<geoCoordinate id="EE2D501AFFC8FFACFD6FFF68FCF2E187" box="[687,795,230,255]" degrees="41" direction="north" minutes="24.3" orientation="latitude" pageId="8" pageNumber="1637" precision="92" value="41.405">41°24.3′N</geoCoordinate>
<geoCoordinate id="EE2D501AFFC8FFACFCE1FF68FC7EE187" box="[801,919,230,255]" degrees="041" direction="east" minutes="31.9" orientation="longitude" pageId="8" pageNumber="1637" precision="92" value="41.531666">041°31.9′E</geoCoordinate>
, / margin of deciduous forest / (with dominant 
<taxonomicName id="4C194D5EFFC8FFACFE1AFE88FD64E067" authority=", Castanea" authorityName="Castanea" box="[474,653,263,287]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="1637" phylum="Tracheophyta" rank="genus">
<emphasis id="B96DEACFFFC8FFACFE1AFE88FD64E067" box="[474,653,263,287]" italics="true" pageId="8" pageNumber="1637">Fagus, Castanea</emphasis>
</taxonomicName>
) [printed] // 
<typeStatus id="54A2887FFFC8FFACFCD6FE88FC44E067" box="[790,941,263,287]" pageId="8" pageNumber="1637" type="holotype">HOLOTYPE</typeStatus>
/ 
<taxonomicName id="4C194D5EFFC8FFACFC01FE88FB6FE067" authorityName="Clairville" authorityYear="1806" box="[961,1158,263,287]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1637" phylum="Arthropoda" rank="genus">HYDROPORUS</taxonomicName>
/ anatolicus / koksali ssp. nov. / 
<collectorName id="26EC530BFFC8FFACFDF1FEA7FD78E047" box="[561,657,295,319]" pageId="8" pageNumber="1637">J. Hájek</collectorName>
&amp; 
<collectorName id="26EC530BFFC8FFACFD74FEA7FCFBE047" box="[692,786,295,319]" pageId="8" pageNumber="1637">M. Fiká</collectorName>
<emphasis id="B96DEACFFFC8FFACFCD2FEA8FCF7E047" bold="true" box="[786,798,295,319]" pageId="8" pageNumber="1637">C</emphasis>
ek det. 2007 [printed, red label]”
</materialsCitation>
. 
<materialsCitation id="3B713C80FFC8FFACFF14FEC7FEDAE007" collectingDate="2004-06-29" collectingDateMax="2004-06-30" collectingDateMin="2004-06-29" collectionCode="GWCW, NMPC" collectorName="J. Hajek &amp; M. Fika" country="Turkey" elevation="775" latitude="41.395" location="Ji" longLatPrecision="115" longitude="41.533333" pageId="8" pageNumber="1637" specimenCount="12" specimenCount-female="11" specimenCount-male="1" stateProvince="Artvin" typeStatus="paratype">
<typeStatus id="54A2887FFFC8FFACFF14FEC7FEA1E018" box="[212,328,328,352]" pageId="8" pageNumber="1637" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D1FFD54FFC8FFACFE92FEC8FE24E027" box="[338,461,327,351]" pageId="8" pageNumber="1637" type="male">four males</specimenCount>
, 
<specimenCount id="9D1FFD54FFC8FFACFE19FEC8FDB9E027" box="[473,592,327,351]" pageId="8" pageNumber="1637" type="female">11 females</specimenCount>
(nos. 1–15), same label data as holotype (
<collectionCode id="ED08AE18FFC8FFACFBEDFEC8FB66E027" box="[1069,1167,327,351]" pageId="8" pageNumber="1637">GWCW</collectionCode>
, 
<collectionCode id="ED08AE18FFC8FFACFF14FEE8FEC2E007" box="[212,299,359,383]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="1637">NMPC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B713C80FFC8FFACFE8BFEE7FB66E0C7" collectionCode="NHMW" collectorName="F. Schubert" country="Turkey" determinerName="Wewalka" elevation="1500" latitude="41.35" location="Br." longLatPrecision="1156" longitude="41.683334" municipality="Borcka" pageId="8" pageNumber="1637" specimenCount="2" specimenCount-female="2" stateProvince="Artvin" typeStatus="paratype">
<specimenCount id="9D1FFD54FFC8FFACFE8BFEE7FE3AE007" box="[331,467,359,383]" pageId="8" pageNumber="1637" type="female">one female</specimenCount>
(no. 16), “ 
<collectingMunicipality id="6BC2ACA7FFC8FFACFDB1FEE8FD2EE007" box="[625,711,359,383]" pageId="8" pageNumber="1637">Borçka</collectingMunicipality>
[
<collectingCountry id="F30E764DFFC8FFACFD20FEE8FCDFE007" box="[736,822,359,383]" name="Turkey" pageId="8" pageNumber="1637">Turkey</collectingCountry>
, 
<collectingRegion id="49DDF83FFFC8FFACFC8FFEE8FC04E007" box="[847,1005,359,383]" country="Turkey" name="Artvin" pageId="8" pageNumber="1637">Artvin prov.</collectingRegion>
, ca. 
<geoCoordinate id="EE2D501AFFC8FFACFBF9FEE8FB7DE007" box="[1081,1172,358,383]" degrees="41" direction="north" minutes="21" orientation="latitude" pageId="8" pageNumber="1637" precision="925" value="41.35">41°21′N</geoCoordinate>
<geoCoordinate id="EE2D501AFFC8FFACFF14FE08FEC3E0E7" box="[212,298,390,415]" degrees="41" direction="east" minutes="41" orientation="longitude" pageId="8" pageNumber="1637" precision="925" value="41.683334">41°41′E</geoCoordinate>
],Asm. [= Asia minor]NO. [NE] / 
<quantity id="4CE19B38FFC8FFACFD10FE08FCC2E0E7" box="[720,811,391,415]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="1637" unit="m" value="1500.0">
<elevation id="0034D1EEFFC8FFACFD10FE08FCC2E0E7" box="[720,811,391,415]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="1637" unit="m" value="1500.0">1500 m</elevation>
</quantity>
, 4/7.6.[19]69 / leg. 
<collectorName id="26EC530BFFC8FFACFBCCFE08FB7DE0E7" box="[1036,1172,391,415]" pageId="8" pageNumber="1637">F. Schubert</collectorName>
[printed] // 
<taxonomicName id="4C194D5EFFC8FFACFE90FE28FE35E0C7" authorityName="Clairville" authorityYear="1806" box="[336,476,423,447]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1637" phylum="Arthropoda" rank="genus">Hydroporus</taxonomicName>
<specimenCount id="9D1FFD54FFC8FFACFE22FE29FE11E0C7" box="[482,504,422,447]" pageId="8" pageNumber="1637" type="female">♀</specimenCount>
? / anatolicus B.- 
<location id="8EC66006FFC8FFACFD71FE27FD3FE0C7" LSID="urn:lsid:plazi:treatment:03B087CBFFC6FFADFE18FA28FE0EE7F7:8EC66006FFC8FFACFD71FE27FD3FE0C7" box="[689,726,424,447]" country="Turkey" latitude="41.35" longLatPrecision="1156" longitude="41.683334" municipality="Borcka" name="Br." pageId="8" pageNumber="1637" stateProvince="Artvin">Br.</location>
/ det. 
<determinerName id="7E7ADA19FFC8FFACFCD1FE28FC97E0C7" box="[785,894,423,447]" pageId="8" pageNumber="1637">Wewalka</determinerName>
98 [printed]” (
<collectionCode id="ED08AE18FFC8FFACFBE2FE28FB6EE0C7" box="[1058,1159,423,447]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="1637" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B713C80FFC8FFACFF14FE47FD8AE347" collectingDate="1986-06-11" collectionCode="HFCB" collectorName="J. Balfour-Browne" country="Turkey" determinerName="Fery" elevation="1600" latitude="41.25" location="Pirnalli" longLatPrecision="1157" longitude="42.05" pageId="8" pageNumber="1637" specimenCount="1" specimenCount-male="1" stateProvince="Artvin" typeStatus="paratype">
<specimenCount id="9D1FFD54FFC8FFACFF14FE47FED7E0A7" box="[212,318,455,479]" pageId="8" pageNumber="1637" type="male">one male</specimenCount>
(no. 17), “TURQUIE [
<collectingCountry id="F30E764DFFC8FFACFD9CFE48FD5DE0A7" box="[604,692,455,479]" name="Turkey" pageId="8" pageNumber="1637">Turkey</collectingCountry>
]: 
<collectingRegion id="49DDF83FFFC8FFACFD09FE48FCFDE0A7" box="[713,788,455,479]" country="Turkey" name="Artvin" pageId="8" pageNumber="1637">Artvin</collectingRegion>
[prov.] / 
<location id="8EC66006FFC8FFACFC49FE48FC35E0A7" LSID="urn:lsid:plazi:treatment:03B087CBFFC6FFADFE18FA28FE0EE7F7:8EC66006FFC8FFACFC49FE48FC35E0A7" box="[905,988,455,479]" country="Turkey" latitude="41.25" longLatPrecision="1157" longitude="42.05" name="Pirnalli" pageId="8" pageNumber="1637" stateProvince="Artvin">Pirnalli</location>
[= P 
<emphasis id="B96DEACFFFC8FFACFBDDFE48FBCDE0A7" bold="true" box="[1053,1060,455,479]" pageId="8" pageNumber="1637">i</emphasis>
rnall 
<emphasis id="B96DEACFFFC8FFACFB99FE48FB8EE0A7" bold="true" box="[1113,1127,455,479]" pageId="8" pageNumber="1637">i,</emphasis>
ca. 
<geoCoordinate id="EE2D501AFFC8FFACFF14FE68FED9E087" box="[212,304,486,511]" degrees="41" direction="north" minutes="15" orientation="latitude" pageId="8" pageNumber="1637" precision="925" value="41.25">41°15′N</geoCoordinate>
<geoCoordinate id="EE2D501AFFC8FFACFEFBFE68FE7CE087" box="[315,405,486,511]" degrees="42" direction="east" minutes="03" orientation="longitude" pageId="8" pageNumber="1637" precision="925" value="42.05">42°03′E</geoCoordinate>
], massif / du Karkal Da 
<emphasis id="B96DEACFFFC8FFACFD78FE68FD25E087" bold="true" box="[696,716,487,511]" pageId="8" pageNumber="1637">gi</emphasis>
[mts] / 
<quantity id="4CE19B38FFC8FFACFCEBFE68FC6DE087" box="[811,900,487,511]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="1637" unit="m" value="1600.0">
<elevation id="0034D1EEFFC8FFACFCEBFE68FC6DE087" box="[811,900,487,511]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="1637" unit="m" value="1600.0">1600 m</elevation>
</quantity>
; 
<date id="FFA7101DFFC8FFACFC53FE68FBE6E087" box="[915,1039,487,511]" pageId="8" pageNumber="1637" value="1986-06-11">
<collectingDate id="EFE3E9F5FFC8FFACFC53FE68FBE6E087" box="[915,1039,487,511]" pageId="8" pageNumber="1637" value="1986-06-11">11.VI.1986</collectingDate>
</date>
[printed] // Besuchet-Löbl / Burckhardt [printed] // 
<taxonomicName id="4C194D5EFFC8FFACFD56FD88FC74E367" box="[662,925,519,543]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1637" phylum="Arthropoda" rank="species" species="anatolicus">Hydroporus anatolicus</taxonomicName>
/ 
<collectorName id="26EC530BFFC8FFACFC73FD87FB6CE367" box="[947,1157,519,543]" pageId="8" pageNumber="1637">J. Balfour-Browne</collectorName>
/ 
<determinerName id="7E7ADA19FFC8FFACFF14FDA7FEE0E338" box="[212,265,552,576]" pageId="8" pageNumber="1637">Fery</determinerName>
det. 1998 [printed]” (
<collectionCode id="ED08AE18FFC8FFACFDCBFDA8FDB5E347" box="[523,604,551,575]" pageId="8" pageNumber="1637">HFCB</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B713C80FFC8FFACFDB3FDA8FDC6E3E7" collectingDate="1999-09-10" collectionCode="HFCB, KECE, NMPC" collectorName="O. K. Erman" country="Turkey" county="Ozdere Koyu" location="Yedigoller" municipality="Yedi" pageId="8" pageNumber="1637" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Erzurum" typeStatus="paratype">
<specimenCount id="9D1FFD54FFC8FFACFDB3FDA8FD09E347" box="[627,736,551,575]" pageId="8" pageNumber="1637" type="male">six males</specimenCount>
, 
<specimenCount id="9D1FFD54FFC8FFACFD30FDA8FC90E347" box="[752,889,551,575]" pageId="8" pageNumber="1637" type="female">five females</specimenCount>
(nos. 18–28), “[
<collectingCountry id="F30E764DFFC8FFACFBF6FDA8FB66E347" box="[1078,1167,551,575]" name="Turkey" pageId="8" pageNumber="1637">Turkey</collectingCountry>
, 
<collectingRegion id="49DDF83FFFC8FFACFF14FDC7FE69E327" box="[212,384,584,607]" country="Turkey" name="Erzurum" pageId="8" pageNumber="1637">Erzurum prov.</collectingRegion>
] 
<collectingCounty id="62C74E51FFC8FFACFE50FDC8FDC2E318" box="[400,555,583,608]" pageId="8" pageNumber="1637">Özdere Köyü</collectingCounty>
[vill.], 
<collectingMunicipality id="6BC2ACA7FFC8FFACFDBEFDC8FD5DE327" box="[638,692,583,607]" pageId="8" pageNumber="1637">Yedi</collectingMunicipality>
/ göller [
<location id="8EC66006FFC8FFACFCE0FDC8FC70E327" LSID="urn:lsid:plazi:treatment:03B087CBFFC6FFADFE18FA28FE0EE7F7:8EC66006FFC8FFACFCE0FDC8FC70E327" box="[800,921,583,607]" country="Turkey" county="Ozdere Koyu" municipality="Yedi" name="Yedigoller" pageId="8" pageNumber="1637" stateProvince="Erzurum">Yedigöller</location>
] deresi [stream], 
<location id="8EC66006FFC8FFACFBA0FDC8FB7AE327" LSID="urn:lsid:plazi:treatment:03B087CBFFC6FFADFE18FA28FE0EE7F7:8EC66006FFC8FFACFBA0FDC8FB7AE327" box="[1120,1171,583,607]" country="Turkey" county="Ozdere Koyu" municipality="Yedi" name="Oltu" pageId="8" pageNumber="1637" stateProvince="Erzurum">Oltu</location>
[distr.] / 
<date id="FFA7101DFFC8FFACFEF6FDE8FE48E307" box="[310,417,615,639]" pageId="8" pageNumber="1637" value="1999-09-10">
<collectingDate id="EFE3E9F5FFC8FFACFEF6FDE8FE48E307" box="[310,417,615,639]" pageId="8" pageNumber="1637" value="1999-09-10">9.10.1999</collectingDate>
</date>
/ 
<collectorName id="26EC530BFFC8FFACFE7BFDE8FDA8E307" box="[443,577,615,639]" pageId="8" pageNumber="1637">Ö.K.Erman</collectorName>
[handwritten] // 
<taxonomicName id="4C194D5EFFC8FFACFCC4FDE8FC66E307" authorityName="Clairville" authorityYear="1806" box="[772,911,615,639]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1637" phylum="Arthropoda" rank="genus">Hydroporus</taxonomicName>
/ obsoletus [handwritten]” (
<collectionCode id="ED08AE18FFC8FFACFEDAFD08FE83E3E7" box="[282,362,647,671]" pageId="8" pageNumber="1637">HFCB</collectionCode>
, 
<collectionCode id="ED08AE18FFC8FFACFEB6FD08FE2CE3E7" box="[374,453,647,671]" pageId="8" pageNumber="1637">KECE</collectionCode>
, 
<collectionCode id="ED08AE18FFC8FFACFE10FD08FDCEE3E7" box="[464,551,647,671]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="1637">NMPC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B713C80FFC8FFACFDFAFD07FBCDE3C7" collectionCode="HFCB" country="Turkey" county="Yolu" location="Yan" municipality="Km Ce" pageId="8" pageNumber="1637" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D1FFD54FFC8FFACFDFAFD07FD49E3E7" box="[570,672,647,671]" pageId="8" pageNumber="1637" type="male">one male</specimenCount>
(no. 29), “TR [
<collectingCountry id="F30E764DFFC8FFACFC8DFD08FC4BE3E7" box="[845,930,647,671]" name="Turkey" pageId="8" pageNumber="1637">Turkey</collectingCountry>
]-Erzurum- 
<emphasis id="B96DEACFFFC8FFACFBDFFD08FBC0E3E7" bold="true" box="[1055,1065,647,671]" pageId="8" pageNumber="1637">I</emphasis>
spir 
<collectingCounty id="62C74E51FFC8FFACFB9AFD08FB7AE3E7" box="[1114,1171,647,671]" pageId="8" pageNumber="1637">Yolu</collectingCounty>
/ 55 
<collectingMunicipality id="6BC2ACA7FFC8FFACFEC2FD27FEBAE3C7" box="[258,339,679,703]" pageId="8" pageNumber="1637">Km Çe</collectingMunicipality>
<emphasis id="B96DEACFFFC8FFACFE93FD28FEB4E3C7" bold="true" box="[339,349,679,703]" pageId="8" pageNumber="1637">s</emphasis>
menin 
<location id="8EC66006FFC8FFACFE69FD27FE33E3C7" LSID="urn:lsid:plazi:treatment:03B087CBFFC6FFADFE18FA28FE0EE7F7:8EC66006FFC8FFACFE69FD27FE33E3C7" box="[425,474,680,703]" country="Turkey" county="Yolu" municipality="Km Ce" name="Yan" pageId="8" pageNumber="1637">Yan</location>
<emphasis id="B96DEACFFFC8FFACFE1AFD28FE08E3C7" bold="true" box="[474,481,679,703]" pageId="8" pageNumber="1637">i</emphasis>
/ Eylül-2000 / Leg. Erman,Ö.K. [printed]” (
<collectionCode id="ED08AE18FFC8FFACFC0BFD28FBF5E3C7" box="[971,1052,679,703]" pageId="8" pageNumber="1637">HFCB</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B713C80FFC8FFADFBEEFD27FD63E1C7" collectionCode="NHMW" country="Turkey" determinerName="G. Wewalka" lastPageId="9" lastPageNumber="1638" location="Jach" municipality="Tortum-Narman" pageId="8" pageNumber="1637" specimenCount="1" specimenCount-male="1" stateProvince="Erzurum" typeStatus="paratype">
<specimenCount id="9D1FFD54FFC8FFACFBEEFD27FB7AE3C7" box="[1070,1171,679,703]" pageId="8" pageNumber="1637" type="male">one male</specimenCount>
(no. 30), “TR [
<collectingCountry id="F30E764DFFC8FFACFE43FD48FE30E3A7" box="[387,473,711,735]" name="Turkey" pageId="8" pageNumber="1637">Turkey</collectingCountry>
]- 
<collectingRegion id="49DDF83FFFC8FFACFE27FD48FD93E3A7" box="[487,634,711,735]" country="Turkey" name="Erzurum" pageId="8" pageNumber="1637">ERZURUM</collectingRegion>
[prov.] 9.6. / 
<collectingMunicipality id="6BC2ACA7FFC8FFACFCDCFD47FC35E3A7" box="[796,988,712,735]" pageId="8" pageNumber="1637">Tortum-Narman</collectingMunicipality>
/ lg. 
<location id="8EC66006FFC8FFACFBD0FD48FBAFE3A7" LSID="urn:lsid:plazi:treatment:03B087CBFFC6FFADFE18FA28FE0EE7F7:8EC66006FFC8FFACFBD0FD48FBAFE3A7" box="[1040,1094,711,735]" country="Turkey" municipality="Tortum-Narman" name="Jach" pageId="8" pageNumber="1637" stateProvince="Erzurum">Jäch</location>
[19]89 (72) [printed] // 
<taxonomicName id="4C194D5EFFC8FFACFE54FD68FDC9E387" authorityName="Clairville" authorityYear="1806" box="[404,544,743,767]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1637" phylum="Arthropoda" rank="genus">Hydroporus</taxonomicName>
/ anatolicus B-Br / det. 
<determinerName id="7E7ADA19FFC8FFACFCFEFD68FC21E387" box="[830,968,743,767]" pageId="8" pageNumber="1637">G.Wewalka</determinerName>
89 [handwritten, name of determinator printed]” (
<collectionCode id="ED08AE18FFC9FFADFDDDFF28FD6BE1C7" box="[541,642,167,191]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="9" pageNumber="1638" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B713C80FFC9FFADFD5CFF27FE8DE047" collectingDate="2004-07-11" collectingDateMax="2004-07-12" collectingDateMin="2004-07-11" collectionCode="NMPC" country="Turkey" elevation="2388" latitude="40.345" location="Ji" longLatPrecision="116" longitude="41.695" pageId="9" pageNumber="1638" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Erzurum" typeStatus="paratype">
<specimenCount id="9D1FFD54FFC9FFADFD5CFF27FCE2E1C7" box="[668,779,167,191]" pageId="9" pageNumber="1638" type="male">one male</specimenCount>
, 
<specimenCount id="9D1FFD54FFC9FFADFCDDFF27FC43E1C7" box="[797,938,167,191]" pageId="9" pageNumber="1638" type="female">two females</specimenCount>
(nos. 31–33), “ 
<collectingCountry id="F30E764DFFC9FFADFF5AFF48FEFDE1A7" box="[154,276,199,223]" name="Turkey" pageId="9" pageNumber="1638">TURKEY</collectingCountry>
, 
<collectingRegion id="49DDF83FFFC9FFADFEDEFF47FE28E1A7" box="[286,449,200,223]" country="Turkey" name="Erzurum" pageId="9" pageNumber="1638">Erzurum prov.</collectingRegion>
, / 
<locationDeviation id="9A142FFFFFC9FFADFE17FF48FD77E1A7" box="[471,670,199,223]" location="Narman" pageId="9" pageNumber="1638">
<quantity id="4CE19B38FFC9FFADFE17FF48FDF3E1A7" box="[471,538,199,223]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="1638" unit="km" value="20.0">20 km</quantity>
W Narman
</locationDeviation>
, / KIREÇLI GEÇIDI [pass], / 
<quantity id="4CE19B38FFC9FFADFC7BFF48FBA5E1A7" box="[955,1100,199,223]" metricMagnitude="3" metricUnit="m" metricValue="2.3875" metricValueMax="2.4" metricValueMin="2.375" pageId="9" pageNumber="1638" unit="m" value="2387.5" valueMax="2400.0" valueMin="2375.0">
<elevation id="0034D1EEFFC9FFADFC7BFF48FBA5E1A7" box="[955,1100,199,223]" metricMagnitude="3" metricUnit="m" metricValue="2.3875" metricValueMax="2.4" metricValueMin="2.375" pageId="9" pageNumber="1638" unit="m" value="2387.5" valueMax="2400.0" valueMin="2375.0">2375–2400 m</elevation>
</quantity>
, 
<date id="FFA7101DFFC9FFADFF50FF68FEACE187" box="[144,325,231,255]" pageId="9" pageNumber="1638" value="2004-07-11" valueMax="2004-07-12" valueMin="2004-07-11">
<collectingDate id="EFE3E9F5FFC9FFADFF50FF68FEACE187" box="[144,325,231,255]" pageId="9" pageNumber="1638" value="2004-07-11" valueMax="2004-07-12" valueMin="2004-07-11">11.–12.VII.2004</collectingDate>
</date>
/ 
<location id="8EC66006FFC9FFADFE9CFF68FE86E187" LSID="urn:lsid:plazi:treatment:03B087CBFFC6FFADFE18FA28FE0EE7F7:8EC66006FFC9FFADFE9CFF68FE86E187" box="[348,367,231,255]" country="Turkey" latitude="40.345" longLatPrecision="116" longitude="41.695" name="Ji" pageId="9" pageNumber="1638" stateProvince="Erzurum">Ji</location>
<emphasis id="B96DEACFFFC9FFADFEAFFF68FE90E187" bold="true" box="[367,377,231,255]" pageId="9" pageNumber="1638">r</emphasis>
í Hájek &amp; Jan R 
<emphasis id="B96DEACFFFC9FFADFDF7FF68FDAFE187" bold="true" box="[567,582,231,255]" pageId="9" pageNumber="1638">8</emphasis>
ži 
<emphasis id="B96DEACFFFC9FFADFD98FF68FD8DE187" bold="true" box="[600,612,231,255]" pageId="9" pageNumber="1638">C</emphasis>
ka leg. [printed] // 
<geoCoordinate id="EE2D501AFFC9FFADFCF7FF68FC40E187" box="[823,937,230,255]" degrees="40" direction="north" minutes="20.7" orientation="latitude" pageId="9" pageNumber="1638" precision="92" value="40.345">40°20.7′N</geoCoordinate>
<geoCoordinate id="EE2D501AFFC9FFADFC71FF68FBC2E187" box="[945,1067,230,255]" degrees="041" direction="east" minutes="41.7" orientation="longitude" pageId="9" pageNumber="1638" precision="92" value="41.695">041°41.7′E</geoCoordinate>
– / 
<geoCoordinate id="EE2D501AFFC9FFADFF50FE88FEE8E067" box="[144,257,262,287]" degrees="40" direction="north" minutes="20.2" orientation="latitude" pageId="9" pageNumber="1638" precision="92" value="40.336666">40°20.2′N</geoCoordinate>
<geoCoordinate id="EE2D501AFFC9FFADFEC9FE88FE6CE067" box="[265,389,262,287]" degrees="041" direction="east" minutes="41.7" orientation="longitude" pageId="9" pageNumber="1638" precision="92" value="41.695">041°41.7′E</geoCoordinate>
, steppe and / pastures, wet vegetation / near spring on E slope [printed]” (
<collectionCode id="ED08AE18FFC9FFADFEC6FEA8FEB5E047" box="[262,348,295,319]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="1638">NMPC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B713C80FFC9FFADFEADFEA8FE03E007" country="Turkey" location="Each" pageId="9" pageNumber="1638" specimenCount="4" stateProvince="Erzurum" typeStatus="paratype">
<location id="8EC66006FFC9FFADFEADFEA8FE4CE047" LSID="urn:lsid:plazi:treatment:03B087CBFFC6FFADFE18FA28FE0EE7F7:8EC66006FFC9FFADFEADFEA8FE4CE047" box="[365,421,295,319]" country="Turkey" name="Each" pageId="9" pageNumber="1638" stateProvince="Erzurum">Each</location>
<typeStatus id="54A2887FFFC9FFADFE6AFEA7FDE3E038" box="[426,522,296,320]" pageId="9" pageNumber="1638" type="paratype">paratype</typeStatus>
is provided with a red label similar to that of the 
<typeStatus id="54A2887FFFC9FFADFBE2FEA8FF3BE018" pageId="9" pageNumber="1638" type="holotype">holotype</typeStatus>
, except “ 
<typeStatus id="54A2887FFFC9FFADFEF4FEC8FE39E027" box="[308,464,327,351]" pageId="9" pageNumber="1638" type="paratype">PARATYPE</typeStatus>
” is wrtitten instead of “ 
<typeStatus id="54A2887FFFC9FFADFD17FEC8FC90E027" box="[727,889,327,351]" pageId="9" pageNumber="1638" type="holotype">HOLOTYPE</typeStatus>
”, the respective sex symbol and collection number
</materialsCitation>
.
</paragraph>
<caption id="DF666655FFC8FFACFF14F9C5FB6DE7E3" ID-DOI="http://doi.org/10.5281/zenodo.5210007" ID-Zenodo-Dep="5210007" httpUri="https://zenodo.org/record/5210007/files/figure.png" pageId="8" pageNumber="1637" startId="8.[212,279,1610,1632]" targetBox="[213,1170,887,1581]" targetPageId="8">
<paragraph id="8BA636DDFFC8FFACFF14F9C5FB6DE7E3" blockId="8.[212,1171,1610,1691]" pageId="8" pageNumber="1637">
Figure 3. Average landmark configuration and body shape of (A) 
<taxonomicName id="4C194D5EFFC8FFACFC6CF9C4FED1E706" authority="anatolicus" authorityName="anatolicus J. Balfour-Browne" authorityYear="1963" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1637" phylum="Arthropoda" rank="subSpecies" species="anatolicus" subSpecies="anatolicus">
<emphasis id="B96DEACFFFC8FFACFC6CF9C4FED1E706" italics="true" pageId="8" pageNumber="1637">Hydroporus anatolicus anatolicus</emphasis>
</taxonomicName>
; (B) 
<taxonomicName id="4C194D5EFFC8FFACFEACF9E7FDA9E706" authorityName="Hájek &amp; Fikácek" authorityYear="2010" box="[364,576,1640,1662]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1637" phylum="Arthropoda" rank="subSpecies" species="anatolicus" subSpecies="koksali">
<emphasis id="B96DEACFFFC8FFACFEACF9E7FDA9E706" box="[364,576,1640,1662]" italics="true" pageId="8" pageNumber="1637">H. anatolicus koksali</emphasis>
</taxonomicName>
ssp. nov.; (C) 
<taxonomicName id="4C194D5EFFC8FFACFD14F9E7FC90E706" authority="Hájek &amp; Fikácek, 2010" authorityName="Hájek &amp; Fikácek" authorityYear="2010" box="[724,889,1640,1662]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1637" phylum="Arthropoda" rank="species" species="kurdistanicus" status="sp. nov.">
<emphasis id="B96DEACFFFC8FFACFD14F9E7FC90E706" box="[724,889,1640,1662]" italics="true" pageId="8" pageNumber="1637">H. kurdistanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57B4FFC8FFACFC40F9E6FC3AE706" box="[896,979,1641,1662]" pageId="8" pageNumber="1637" rank="species">sp. nov.</taxonomicNameLabel>
; (D) 
<taxonomicName id="4C194D5EFFC8FFACFBC7F9E7FEC8E7E3" authorityName="Hájek &amp; Fikácek" authorityYear="2010" baseAuthorityName="Gschwendtner" baseAuthorityYear="1935" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1637" phylum="Arthropoda" rank="subSpecies" species="bodemeyeri" subSpecies="guignoti">
<emphasis id="B96DEACFFFC8FFACFBC7F9E7FEC8E7E3" italics="true" pageId="8" pageNumber="1637">H. bodemeyeri guignoti</emphasis>
</taxonomicName>
; (E) 
<taxonomicName id="4C194D5EFFC8FFACFE91F90AFDA6E7E3" authority="bodemeyeri" authorityName="bodemeyeri Ganglbauer" authorityYear="1900" box="[337,591,1669,1691]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1637" phylum="Arthropoda" rank="subSpecies" species="bodemeyeri" subSpecies="bodemeyeri">
<emphasis id="B96DEACFFFC8FFACFE91F90AFDA6E7E3" box="[337,591,1669,1691]" italics="true" pageId="8" pageNumber="1637">H. bodemeyeri bodemeyeri</emphasis>
</taxonomicName>
; (F) 
<taxonomicName id="4C194D5EFFC8FFACFDBEF90AFC85E7E3" authorityName="Hájek &amp; Fikácek" authorityYear="2010" box="[638,876,1669,1691]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1637" phylum="Arthropoda" rank="subSpecies" species="bodemeyeri" subSpecies="cariaensis">
<emphasis id="B96DEACFFFC8FFACFDBEF90AFC85E7E3" box="[638,876,1669,1691]" italics="true" pageId="8" pageNumber="1637">H. bodemeyeri cariaensis</emphasis>
</taxonomicName>
ssp. nov.; (G) 
<taxonomicName id="4C194D5EFFC8FFACFC3AF90AFB97E7E3" authorityName="Miller and Fery" authorityYear="1995" box="[1018,1150,1669,1691]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1637" phylum="Arthropoda" rank="species" species="cuprescens">
<emphasis id="B96DEACFFFC8FFACFC3AF90AFB97E7E3" box="[1018,1150,1669,1691]" italics="true" pageId="8" pageNumber="1637">H. cuprescens</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA636DDFFC9FFADFF50FE28FDB5E087" blockId="9.[144,1105,423,511]" pageId="9" pageNumber="1638">
<emphasis id="B96DEACFFFC9FFADFF50FE28FE36E0C7" box="[144,479,423,447]" italics="true" pageId="9" pageNumber="1638">Additional material examined.</emphasis>
<materialsCitation id="3B713C80FFC9FFADFE25FE27FDB1E087" collectingDate="1992-08-12" collectionCode="HFCB" collectorName="P. Mazzoldi" country="Turkey" elevation="1625" latitude="40.933334" location="Ayder - Libler Golu" longLatPrecision="1159" longitude="41.233334" municipality="Caml" pageId="9" pageNumber="1638" specimenCount="1" specimenCount-female="1" stateProvince="Rize">
<collectingCountry id="F30E764DFFC9FFADFE25FE27FD8AE0C7" box="[485,611,424,447]" name="Turkey" pageId="9" pageNumber="1638">TURKEY</collectingCountry>
: 
<specimenCount id="9D1FFD54FFC9FFADFDACFE27FD33E0C7" box="[620,730,423,447]" pageId="9" pageNumber="1638" type="female">one female</specimenCount>
, 
<collectingRegion id="49DDF83FFFC9FFADFD21FE28FCA2E0C7" box="[737,843,423,447]" country="Turkey" name="Rize" pageId="9" pageNumber="1638">Rize prov.</collectingRegion>
, south-east 
<collectingMunicipality id="6BC2ACA7FFC9FFADFC7CFE28FC1AE0C7" box="[956,1011,423,447]" pageId="9" pageNumber="1638">Çaml</collectingMunicipality>
<emphasis id="B96DEACFFFC9FFADFC33FE28FC13E0C7" bold="true" box="[1011,1018,423,447]" pageId="9" pageNumber="1638">i</emphasis>
shem 
<emphasis id="B96DEACFFFC9FFADFBEFFE28FBD1E0C7" bold="true" box="[1071,1080,423,447]" pageId="9" pageNumber="1638">s</emphasis>
in, small streams on road 
<location id="8EC66006FFC9FFADFE5BFE48FD66E0A7" LSID="urn:lsid:plazi:treatment:03B087CBFFC6FFADFE18FA28FE0EE7F7:8EC66006FFC9FFADFE5BFE48FD66E0A7" box="[411,655,455,479]" country="Turkey" latitude="40.933334" longLatPrecision="1159" longitude="41.233334" municipality="Caml" name="Ayder - Libler Golu" pageId="9" pageNumber="1638" stateProvince="Rize">Ayder – Libler Gölü</location>
, ca. 
<geoCoordinate id="EE2D501AFFC9FFADFD05FE48FCC6E0A7" box="[709,815,454,479]" degrees="40" direction="north" minutes="56" orientation="latitude" pageId="9" pageNumber="1638" precision="925" value="40.933334">40°56′ N</geoCoordinate>
, 
<geoCoordinate id="EE2D501AFFC9FFADFCFBFE48FC48E0A7" box="[827,929,454,479]" degrees="41" direction="east" minutes="14" orientation="longitude" pageId="9" pageNumber="1638" precision="925" value="41.233334">41°14′ E</geoCoordinate>
, 
<quantity id="4CE19B38FFC9FFADFC6EFE48FBA2E0A7" box="[942,1099,455,479]" metricMagnitude="3" metricUnit="m" metricValue="1.625" metricValueMax="1.65" metricValueMin="1.6" pageId="9" pageNumber="1638" unit="m" value="1625.0" valueMax="1650.0" valueMin="1600.0">
<elevation id="0034D1EEFFC9FFADFC6EFE48FBA2E0A7" box="[942,1099,455,479]" metricMagnitude="3" metricUnit="m" metricValue="1.625" metricValueMax="1.65" metricValueMin="1.6" pageId="9" pageNumber="1638" unit="m" value="1625.0" valueMax="1650.0" valueMin="1600.0">1600–1650 m</elevation>
</quantity>
, 
<date id="FFA7101DFFC9FFADFF50FE68FED2E087" box="[144,315,487,511]" pageId="9" pageNumber="1638" value="1992-08-12">
<collectingDate id="EFE3E9F5FFC9FFADFF50FE68FED2E087" box="[144,315,487,511]" pageId="9" pageNumber="1638" value="1992-08-12">12 August 1992</collectingDate>
</date>
, 
<collectorName id="26EC530BFFC9FFADFE85FE67FE21E087" box="[325,456,487,511]" pageId="9" pageNumber="1638">P. Mazzoldi</collectorName>
leg. (
<collectionCode id="ED08AE18FFC9FFADFDC1FE68FDB9E087" box="[513,592,487,511]" pageId="9" pageNumber="1638">HFCB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636DDFFC9FFADFF50FDA8FC89E327" blockId="9.[144,1103,551,607]" pageId="9" pageNumber="1638">
<emphasis id="B96DEACFFFC9FFADFF50FDA8FED0E347" box="[144,313,551,575]" italics="true" pageId="9" pageNumber="1638">Measurements.</emphasis>
Body length: 
<quantity id="4CE19B38FFC9FFADFE1BFDA8FD9AE347" box="[475,627,551,575]" metricMagnitude="-3" metricUnit="m" metricValue="3.375" metricValueMax="3.5" metricValueMin="3.25" pageId="9" pageNumber="1638" unit="mm" value="3.375" valueMax="3.5" valueMin="3.25">3.25–3.50 mm</quantity>
(mean value 
<quantity id="4CE19B38FFC9FFADFCC8FDA8FC80E347" box="[776,873,551,575]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="9" pageNumber="1638" unit="mm" value="3.4">3.40 mm</quantity>
, 
<typeStatus id="54A2887FFFC9FFADFCB5FDA8FC3FE347" box="[885,982,551,575]" pageId="9" pageNumber="1638" type="holotype">holotype</typeStatus>
<quantity id="4CE19B38FFC9FFADFC1EFDA8FBADE347" box="[990,1092,551,575]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="9" pageNumber="1638" unit="mm" value="3.3">3.30 mm</quantity>
), width: 
<quantity id="4CE19B38FFC9FFADFF1EFDC8FE94E327" box="[222,381,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.65" metricValueMin="1.45" pageId="9" pageNumber="1638" unit="mm" value="1.5499999999999998" valueMax="1.65" valueMin="1.45">1.45–1.65 mm</quantity>
(mean value 
<quantity id="4CE19B38FFC9FFADFDD3FDC8FD91E327" box="[531,632,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="9" pageNumber="1638" unit="mm" value="1.58">1.58 mm</quantity>
, 
<typeStatus id="54A2887FFFC9FFADFD43FDC8FD0EE327" box="[643,743,583,607]" pageId="9" pageNumber="1638" type="holotype">holotype</typeStatus>
<quantity id="4CE19B38FFC9FFADFD2DFDC8FCBAE327" box="[749,851,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="1638" unit="mm" value="1.5">1.50 mm</quantity>
).
</paragraph>
<paragraph id="8BA636DDFFC9FFADFF50FD08FEFDE267" blockId="9.[144,1104,647,1151]" pageId="9" pageNumber="1638">
<emphasis id="B96DEACFFFC9FFADFF50FD08FEFEE3E7" box="[144,279,647,671]" italics="true" pageId="9" pageNumber="1638">Description.</emphasis>
Body flat, elongate and subparallel (
<figureCitation id="13222A58FFC9FFADFD22FD08FC89E3E7" box="[738,864,647,671]" captionStart="Figure 3" captionStartId="8.[212,279,1610,1632]" captionTargetBox="[213,1170,887,1581]" captionTargetId="figure-339@8.[213,1170,884,1581]" captionTargetPageId="8" captionText="Figure 3. Average landmark configuration and body shape of (A) Hydroporus anatolicus anatolicus; (B) H. anatolicus koksali ssp. nov.; (C) H. kurdistanicus sp. nov.; (D) H. bodemeyeri guignoti; (E) H. bodemeyeri bodemeyeri; (F) H. bodemeyeri cariaensis ssp. nov.; (G) H. cuprescens." figureDoi="http://doi.org/10.5281/zenodo.5210007" httpUri="https://zenodo.org/record/5210007/files/figure.png" pageId="9" pageNumber="1638">Figure 3B</figureCitation>
), similar to that of nominotypical subspecies, but elytra even more tapering to apex, with maximum width close to basal third of elytra. Elytra narrowed also to base of pronotum so pronoto-elytral angle is more distinct than in nominotypical subspecies.
</paragraph>
<paragraph id="8BA636DDFFC9FFADFF7AFCA8FC24E207" blockId="9.[144,1104,647,1151]" pageId="9" pageNumber="1638">Elytra with two slightly distinct punctured lines on disc. Punctation between puncture lines comprising smaller and sparser punctures than in nominotypical subspecies. Distance between punctures more than double their diameter.</paragraph>
<paragraph id="8BA636DDFFC9FFADFF7AFC07FE27E2C7" blockId="9.[144,1104,647,1151]" pageId="9" pageNumber="1638">Legs: protibiae and mesotibiae narrow basally, then slightly curved on ventral side and broadened to apex.</paragraph>
<paragraph id="8BA636DDFFC9FFADFF7AFC48FDF7E547" blockId="9.[144,1104,647,1151]" pageId="9" pageNumber="1638">
Male: dorsal surface shiny. Claws of fore tarsi only slightly hooked. Median lobe of aedeagus in ventral view (
<figureCitation id="13222A58FFC9FFADFE26FC68FDB6E287" box="[486,607,999,1023]" captionStart="Figure 2" captionStartId="7.[144,212,1523,1545]" captionTargetBox="[145,1102,176,1490]" captionTargetId="figure-91@7.[145,1102,176,1490]" captionTargetPageId="7" captionText="Figure 2(A–L). Median lobe in lateral and ventral view of (A) Hydroporus anatolicus anatolicus; (C) H. anatolicus koksali ssp. nov.; (E) H. bodemeyeri bodemeyeri; (G) H. bodemeyeri cariaensis ssp. nov.; (H) H. bodemeyeri guignoti; (I) H. cuprescens; (K) H. kurdistanicus sp. nov. Paramere of (B) Hydroporus anatolicus anatolicus; (D) H. anatolicus koksali ssp. nov.; (F) H. bodemeyeri bodemeyeri; (J) H. cuprescens; (L) H. kurdistanicus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5210005" httpUri="https://zenodo.org/record/5210005/files/figure.png" pageId="9" pageNumber="1638">Figure 2C</figureCitation>
) with apex shorter than in nominotypical subspecies. In lateral view, median lobe only weakly inflexed, apex short, curved upward. Paramere as in 
<figureCitation id="13222A58FFC9FFADFE62FBA8FDF3E547" box="[418,538,1063,1087]" captionStart="Figure 2" captionStartId="7.[144,212,1523,1545]" captionTargetBox="[145,1102,176,1490]" captionTargetId="figure-91@7.[145,1102,176,1490]" captionTargetPageId="7" captionText="Figure 2(A–L). Median lobe in lateral and ventral view of (A) Hydroporus anatolicus anatolicus; (C) H. anatolicus koksali ssp. nov.; (E) H. bodemeyeri bodemeyeri; (G) H. bodemeyeri cariaensis ssp. nov.; (H) H. bodemeyeri guignoti; (I) H. cuprescens; (K) H. kurdistanicus sp. nov. Paramere of (B) Hydroporus anatolicus anatolicus; (D) H. anatolicus koksali ssp. nov.; (F) H. bodemeyeri bodemeyeri; (J) H. cuprescens; (L) H. kurdistanicus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5210005" httpUri="https://zenodo.org/record/5210005/files/figure.png" pageId="9" pageNumber="1638">Figure 2D</figureCitation>
.
</paragraph>
<paragraph id="8BA636DDFFC9FFADFF7AFBC8FD2AE507" blockId="9.[144,1104,647,1151]" pageId="9" pageNumber="1638">Female: pronotum slightly broader than in male. Microreticulation of elytra more distinct than in males so surface subopaque.</paragraph>
<paragraph id="8BA636DDFFC9FFADFF50FB38FC8EE437" blockId="9.[144,1104,1207,1360]" pageId="9" pageNumber="1638">
<emphasis id="B96DEACFFFC9FFADFF50FB38FE6CE5B7" box="[144,389,1207,1231]" italics="true" pageId="9" pageNumber="1638">Differential diagnosis.</emphasis>
The new subspecies differs from nominotypical subspecies by the oval elytra tapering strongly to apex and slightly to base, sparser and finer punctation of elytra, protibiae and mesotibiae with sinuation on ventral side, and shape of median lobe. Apex of median lobe short and curved upwards in 
<emphasis id="B96DEACFFFC9FFADFC47FA98FBF5E457" box="[903,1052,1303,1327]" italics="true" pageId="9" pageNumber="1638">H. a. koksali</emphasis>
ssp. nov., compared with long and nearly straight in 
<emphasis id="B96DEACFFFC9FFADFD71FAB7FC89E428" box="[689,864,1336,1360]" italics="true" pageId="9" pageNumber="1638">
H. a. 
<taxonomicName id="4C194D5EFFC9FFADFD30FAB7FC89E428" box="[752,864,1336,1360]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1638" phylum="Arthropoda" rank="species" species="anatolicus">anatolicus</taxonomicName>
</emphasis>
.
</paragraph>
<paragraph id="8BA636DDFFC9FFADFF50FA07FD12E4C7" blockId="9.[144,1104,1415,1471]" pageId="9" pageNumber="1638">
<emphasis id="B96DEACFFFC9FFADFF50FA07FEF9E4D8" box="[144,272,1416,1440]" italics="true" pageId="9" pageNumber="1638">Etymology.</emphasis>
The new subspecies is dedicated to our friend Ömer Köksal Erman (
<collectingRegion id="49DDF83FFFC9FFADFF57FA28FEE9E4C7" box="[151,256,1447,1471]" country="Turkey" name="Erzurum" pageId="9" pageNumber="1638">Erzurum</collectingRegion>
, 
<collectingCountry id="F30E764DFFC9FFADFECCFA28FE8BE4C7" box="[268,354,1447,1471]" name="Turkey" pageId="9" pageNumber="1638">Turkey</collectingCountry>
), a specialist on Turkish 
<taxonomicName id="4C194D5EFFC9FFADFDBAFA28FD1FE4C7" authorityName="Leach" authorityYear="1815" box="[634,758,1447,1471]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1638" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
.
</paragraph>
<paragraph id="8BA636DDFFC9FFADFF50FA77FD1DE757" blockId="9.[144,1104,1527,1583]" pageId="9" pageNumber="1638">
<emphasis id="B96DEACFFFC9FFADFF50FA77FF07E768" box="[144,238,1528,1552]" italics="true" pageId="9" pageNumber="1638">Habitat.</emphasis>
Collected among gravel in small temporary streams in a mountain area (
<quantity id="4CE19B38FFC9FFADFF57F998FEC9E757" box="[151,288,1559,1583]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="2.4" metricValueMin="0.7" pageId="9" pageNumber="1638" unit="m" value="1550.0" valueMax="2400.0" valueMin="700.0">700–2400 m</quantity>
above sea level) in north-eastern 
<collectingCountry id="F30E764DFFC9FFADFD5AF998FD19E757" box="[666,752,1559,1583]" name="Turkey" pageId="9" pageNumber="1638">Turkey</collectingCountry>
.
</paragraph>
<paragraph id="8BA636DDFFC9FFADFF50F9D7FE0EE7F7" blockId="9.[144,1104,1623,1679]" pageId="9" pageNumber="1638">
<emphasis id="B96DEACFFFC9FFADFF50F9D7FEF3E708" box="[144,282,1624,1648]" italics="true" pageId="9" pageNumber="1638">Distribution.</emphasis>
Known from several localities in north-eastern 
<collectingCountry id="F30E764DFFC9FFADFCF2F9D8FC6AE717" box="[818,899,1623,1647]" name="Turkey" pageId="9" pageNumber="1638">Turkey</collectingCountry>
(
<collectingRegion id="49DDF83FFFC9FFADFC53F9D8FC09E717" box="[915,992,1623,1647]" country="Turkey" name="Artvin" pageId="9" pageNumber="1638">Artvin</collectingRegion>
, 
<collectingRegion id="49DDF83FFFC9FFADFC2CF9D7FBB9E717" box="[1004,1104,1624,1647]" country="Turkey" name="Erzurum" pageId="9" pageNumber="1638">Erzurum</collectingRegion>
and 
<collectingRegion id="49DDF83FFFC9FFADFF00F9F8FF1BE7F7" box="[192,242,1655,1679]" country="Turkey" name="Rize" pageId="9" pageNumber="1638">Rize</collectingRegion>
provinces) (
<figureCitation id="13222A58FFC9FFADFEB9F9F8FE33E7F7" box="[377,474,1655,1679]" captionStart="Figure 6" captionStartId="23.[144,211,645,667]" captionTargetBox="[145,1102,176,615]" captionTargetId="figure-397@23.[145,1103,176,615]" captionTargetPageId="23" captionText="Figure 6. Distribution of the species of the H. bodemeyeri species complex." figureDoi="http://doi.org/10.5281/zenodo.5210015" httpUri="https://zenodo.org/record/5210015/files/figure.png" pageId="9" pageNumber="1638">Figure 6</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>