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<mods:title id="004A05418DE360DA2C9B48FC7F1874F0">On a collection of thoracotreme crabs (Crustacea: Brachyura: Ocypodidae, Macrophthalmidae, Dotillidae) from two offshore islands of Papua, eastern Indonesia, with descriptions of two new species</mods:title>
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<mods:namePart id="B6399A6D77FB571EB860470077096168">Murniati, Dewi Citra</mods:namePart>
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<emphasis id="B96DEAC24E49FF8360C821404E48FDD5" bold="true" box="[229,518,605,629]" italics="true" pageId="4" pageNumber="465">Ocypode ceratophthalmus</emphasis>
(
<bibRefCitation id="EF884B214E49FF83623521404EEAFDD5" author="Pallas SP" box="[536,676,605,629]" pageId="4" pageNumber="465" pagination="83 - 84" refId="ref19118" refString="Pallas SP (1772) Spicelegia Zoologica. Quibus novae imprimis et obscurae Animalium Species. Iconibus, descriptionibus atque commentariis illustrantor. Berolini [= Berlin]. Fasculus nonus, 9: 83 - 84, pl. 5" type="journal article" year="1772">Pallas, 1772</bibRefCitation>
)
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<paragraph id="8BA636D04E49FF8361B321604DBEFD35" blockId="4.[229,683,605,661]" box="[414,496,637,661]" pageId="4" pageNumber="465">
(
<figureCitation id="13222A554E49FF83618B21604DA6FD35" box="[422,488,637,661]" captionStart="Fig" captionStartId="3.[113,148,2006,2028]" captionTargetBox="[126,1438,189,1991]" captionTargetId="figure-11@3.[126,1438,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Ocypode ceratophthalmus (Pallas, 1772). MZB.Cru.5017, male, 38.7 × 34.1 mm; Papua: Bepondi Island. A, dorsal habitus, preserved colouration. B–K, left appendages. Chela: B, inner surface; C, outer surface. P2–P5: D–G, anterior surface; H–K, posterior surface; L, stridulating ridge on inner surface of palm of chela. G1: M, whole morphology; N, apical portion. Scale bars: A, 10 mm; B–K, 5 mm; L–N, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.7399908" httpUri="https://zenodo.org/record/7399908/files/figure.png" pageId="4" pageNumber="465">Fig. 2</figureCitation>
)
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<taxonomicName id="4C194D534E49FF8360A521A14E5CFD71" authority="Pallas, 1772: 83" authorityName="Pallas" authorityPageNumber="83" authorityYear="1772" box="[136,530,700,722]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B96DEAC24E49FF8360A521A14D3AFD71" box="[136,372,700,721]" italics="true" pageId="4" pageNumber="465">Cancer ceratophthalmus</emphasis>
<bibRefCitation id="EF884B214E49FF83615521A14E5CFD71" author="Pallas SP" box="[376,530,700,722]" pageId="4" pageNumber="465" pagination="83 - 84" refId="ref19118" refString="Pallas SP (1772) Spicelegia Zoologica. Quibus novae imprimis et obscurae Animalium Species. Iconibus, descriptionibus atque commentariis illustrantor. Berolini [= Berlin]. Fasculus nonus, 9: 83 - 84, pl. 5" type="journal article" year="1772">Pallas, 1772: 83</bibRefCitation>
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, pl. 5, figs. 7, 8 (unknown locality); 
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<bibRefCitation id="EF884B214E49FF83612121C44D97FD4E" author="Fabricius JC" box="[268,473,729,751]" pageId="4" pageNumber="465" refId="ref17825" refString="Fabricius JC (1781) Species Insectorum exhibentes eorum differentias specificas, Synonyma auctorum, Loca natalia, Metamorphosis adiectis Observationibus, Descriptionibus. Volume 1. Hamburg &amp; Kilonii. viii + 552 pp." type="book" year="1781">Fabricius, 1781: 499</bibRefCitation>
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.
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<taxonomicName id="4C194D534E49FF8360A521E84D33FCAA" ID-CoL="99397" baseAuthorityName="Pallas" baseAuthorityYear="1872" box="[136,381,757,778]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="ceratophthalma">
<emphasis id="B96DEAC24E49FF8360A521E84D33FCAA" box="[136,381,757,778]" italics="true" pageId="4" pageNumber="465">Ocypode ceratophthalma</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EF884B214E49FF8361BF21E84EEEFCAB" author="Sakai K &amp; Turkay M" box="[402,672,757,779]" pageId="4" pageNumber="465" pagination="665 - 794" refId="ref19495" refString="Sakai K &amp; Turkay M (2013) Revision of the genus Ocypode with the description of a new genus, Hoplocypode (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum, 56 (2): 665 - 794." type="journal article" year="2013">Sakai &amp; Türkay, 2013: 685</bibRefCitation>
(Sumatra; Nias Island; We Island; Padang; Bengkulu; Lampung; Java Sea; Jakarta Bay, Java; Lesser Sunda Islands; Lombok Island; 
<collectingCountry id="F30E76404E49FF8362E620324CA5FCC2" name="Flores Island" pageId="4" pageNumber="465">Flores Island</collectingCountry>
; Sumba Island; Dobo Island; 
<collectingCountry id="F30E76404E49FF8361D220514EC7FCC2" box="[511,649,844,866]" name="Ambon Island" pageId="4" pageNumber="465">Ambon Island</collectingCountry>
; Kalimantan; Cendrawasih Bay, Papua), figs. 1D–I, 10A–E, 32A–C.
</paragraph>
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<taxonomicName id="4C194D534E49FF8360A520984D33FC3A" ID-CoL="99397" baseAuthorityName="Pallas" baseAuthorityYear="1772" box="[136,381,901,922]" class="Malacostraca" family="Ocypodidae" genus="Ocypoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="ceratophthalma">
<emphasis id="B96DEAC24E49FF8360A520984D33FC3A" box="[136,381,901,922]" italics="true" pageId="4" pageNumber="465">Ocypoda ceratophthalma</emphasis>
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– 
<bibRefCitation id="EF884B214E49FF8361B820984E61FC3B" author="Tesch JJ" box="[405,559,901,923]" pageId="4" pageNumber="465" pagination="1 - 6" refId="ref19962" refString="Tesch JJ (1918) The Decapoda Brachyura of the Siboga-Expedition. I. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga-Expedition Monograph, 39 c (82): 1 - 148, pls 1 - 6." type="journal article" year="1918">Tesch, 1918: 36</bibRefCitation>
(Labuan Bajo, 
<collectingCountry id="F30E76404E49FF8362E920984F4DFC3B" box="[708,771,901,923]" name="Indonesia" pageId="4" pageNumber="465">Flores</collectingCountry>
; Solor Island, 
<collectingRegion id="49DDF8324E49FF83616F20BE4E50FC18" box="[322,542,931,952]" country="Indonesia" name="Nusa Tenggara Timur" pageId="4" pageNumber="465">East Nusa Tenggara</collectingRegion>
; Makassar, 
<collectingRegion id="49DDF8324E49FF83628F20BF4F4CFC18" box="[674,770,930,952]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="465">Sulawesi</collectingRegion>
; Kaniungan Island, east of 
<collectingRegion id="49DDF8324E49FF83618F20A24E5AFC75" box="[418,532,959,981]" country="Indonesia" name="Kalimantan Utara" pageId="4" pageNumber="465">Kalimantan</collectingRegion>
; Sulu Archipelago; Talaut Islands, Karakelang; west coast of Seram Island; Kei Island); 
<bibRefCitation id="EF884B214E49FF83608320E44D66FBAE" author="Wada K" box="[174,296,1017,1039]" pageId="4" pageNumber="465" refId="ref20063" refString="Wada K (2019) Brachyuran species recorded from mangrove swamp in Indonesia in 1984 - 2001. Cancer, 28: e 144 - e 147. [in Japanese]" type="book" year="2019">Wada, 2019</bibRefCitation>
: e147 (Lukupang, 
<collectingRegion id="49DDF8324E49FF8361CD20E44E75FBAF" box="[480,571,1017,1039]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="465">Sulawesi</collectingRegion>
).
</paragraph>
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<treatmentCitationGroup id="AB0911FE4E49FF8360A527084F49FB8B" box="[136,775,1045,1067]" pageId="4" pageNumber="465">
<taxonomicName id="4C194D534E49FF8360A527084DCDFB8A" ID-CoL="48LW3" baseAuthorityName="Pallas" baseAuthorityYear="1772" box="[136,387,1045,1066]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="ceratophthalmus">
<emphasis id="B96DEAC24E49FF8360A527084DCDFB8A" box="[136,387,1045,1066]" italics="true" pageId="4" pageNumber="465">Ocypode ceratophthalmus</emphasis>
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– 
<treatmentCitation id="0AB810C14E49FF8361BB27084E10FB8B" author="Shih HT &amp; Chan BKK &amp; Teng SJ &amp; Wong KJH" box="[406,606,1045,1067]" page="159" pageId="4" pageNumber="465" year="2015">
<bibRefCitation id="EF884B214E49FF8361BB27084E10FB8B" author="Shih HT &amp; Chan BKK &amp; Teng SJ &amp; Wong KJH" box="[406,606,1045,1067]" pageId="4" pageNumber="465" refId="ref19758" refString="Shih HT, Chan BKK, Teng SJ &amp; Wong KJH (2015) Crustacean Fauna of Taiwan. Brachyuran Crabs. Volume 2: Ocypodoidea. National Chung Hsing University, Taichung, 320 pp." type="book" year="2015">Shih et al., 2015: 159</bibRefCitation>
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(Southeast Asia).
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<emphasis id="B96DEAC24E49FF8360A5274E4D39FBCB" bold="true" box="[136,375,1107,1131]" pageId="4" pageNumber="465">Material examined.</emphasis>
<specimenCount id="9D1FFD594E49FF8361AE27494D93FBCC" box="[387,477,1108,1132]" pageId="4" pageNumber="465" type="male">9 males</specimenCount>
(13.7 × 
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– 26.1 × 
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), 
<materialsCitation id="3B713C8D4E49FF83610C27694D78FAAC" ID-GBIF-Occurrence="4010507322" collectingDate="2018-11-22" collectionCode="MZB" collectorName="DC Murniati; Raharjo &amp; Melati" county="Cru." latitude="-1.6242944" location="Sarmi District" longLatPrecision="1" longitude="138.74165" municipality="Liki Village" pageId="4" pageNumber="465" specimenCount="3" specimenCount-female="3" stateProvince="Papua Barat">
<specimenCount id="9D1FFD594E49FF83610C27694DC0FB2C" box="[289,398,1140,1164]" pageId="4" pageNumber="465" type="female">3 females</specimenCount>
(16.8 × 13.7 – 21.9 × 
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) (
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. 
<collectingCounty id="62C74E5C4E49FF8360FA27894D44FB0C" box="[215,266,1172,1196]" pageId="4" pageNumber="465">Cru.</collectingCounty>
4913), 
<collectingMunicipality id="6BC2ACAA4E49FF83614C27894DB8FB0C" box="[353,502,1172,1196]" pageId="4" pageNumber="465">Liki Village</collectingMunicipality>
, 
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, 
<location id="8EC6600B4E49FF8362EE27894D59FB6C" LSID="urn:lsid:plazi:treatment:03B087C64E49FF8360C82140498DFE55:8EC6600B4E49FF8362EE27894D59FB6C" county="Cru." latitude="-1.6242944" longLatPrecision="1" longitude="138.74165" municipality="Liki Village" name="Sarmi Municipality" pageId="4" pageNumber="465" stateProvince="Papua Barat">Sarmi Municipality</location>
, 
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, 
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, 
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<collectingDate id="EFE3E9F84E49FF83616427C94E66FB4C" box="[329,552,1236,1260]" pageId="4" pageNumber="465" value="2018-11-22">22 November 2018</collectingDate>
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, 
<collectorName id="26EC53064E49FF83621A27C94E82FB4C" box="[567,716,1236,1260]" pageId="4" pageNumber="465">DC Murniati</collectorName>
, SNI 
<collectorName id="26EC53064E49FF8360A427E94CAEFAAC" box="[137,224,1268,1292]" pageId="4" pageNumber="465">Raharjo</collectorName>
, 
<collectorName id="26EC53064E49FF8360C027E94D78FAAC" box="[237,310,1268,1292]" pageId="4" pageNumber="465">Melati</collectorName>
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; 
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<specimenCount id="9D1FFD594E49FF83616E27E94DEBFAAC" box="[323,421,1268,1292]" pageId="4" pageNumber="465" type="male">10 males</specimenCount>
(14.1 × 12.0 mm – 38.7 × 
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. 
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5017), 
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, 
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, 
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, 
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, 
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, 
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&amp; SNI 
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(32.7 × 
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. 
<collectingMunicipality id="6BC2ACAA4E49FF8360A526894CF6FA0C" box="[136,184,1428,1452]" pageId="4" pageNumber="465">Cru.</collectingMunicipality>
532), 
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<location id="8EC6600B4E49FF8361E626894E41FA0C" LSID="urn:lsid:plazi:treatment:03B087C64E49FF8360C82140498DFE55:8EC6600B4E49FF8361E626894E41FA0C" box="[459,527,1428,1452]" municipality="Cru." name="Island" pageId="4" pageNumber="465" stateProvince="Lampung, Sumatra">Island</location>
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<collectingDate id="EFE3E9F84E49FF83623726894EA2FA0C" box="[538,748,1428,1452]" pageId="4" pageNumber="465" value="1975-11-08">08 November 1975</collectingDate>
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, 
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(22.6 × 
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) (
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. 
<collectingMunicipality id="6BC2ACAA4E49FF83626926A94E3AFA6C" box="[580,628,1460,1484]" pageId="4" pageNumber="465">Cru.</collectingMunicipality>
1918), 
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, near 
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<date id="FFA710104E49FF83614226C94E4FFA4C" box="[367,513,1492,1516]" pageId="4" pageNumber="465" value="1972-05-21">
<collectingDate id="EFE3E9F84E49FF83614226C94E4FFA4C" box="[367,513,1492,1516]" pageId="4" pageNumber="465" value="1972-05-21">21 May 1972</collectingDate>
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, 
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<specimenCount id="9D1FFD594E49FF8362AD26C94E86FA4C" box="[640,712,1492,1516]" pageId="4" pageNumber="465" type="male">1 male</specimenCount>
(41.7 × 
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) (
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. 
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1634), 
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, 
<collectingRegion id="49DDF8324E49FF8362B526E94CA1F98C" country="Indonesia" name="Nusa Tenggara Timur" pageId="4" pageNumber="465">East Nusa Tenggara</collectingRegion>
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<date id="FFA710104E49FF8360D725094DC2F98C" box="[250,396,1556,1580]" pageId="4" pageNumber="465" value="1982-06-02">
<collectingDate id="EFE3E9F84E49FF8360D725094DC2F98C" box="[250,396,1556,1580]" pageId="4" pageNumber="465" value="1982-06-02">02 June 1982</collectingDate>
</date>
, 
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(31.7 × 
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) (
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. 
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2683), 
<collectingMunicipality id="6BC2ACAA4E49FF83616025294DACF9EC" box="[333,482,1588,1612]" pageId="4" pageNumber="465">Teluk Terima</collectingMunicipality>
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, 
<location id="8EC6600B4E49FF83624825294E86F9EC" LSID="urn:lsid:plazi:treatment:03B087C64E49FF8360C82140498DFE55:8EC6600B4E49FF83624825294E86F9EC" box="[613,712,1588,1612]" county="Cru." municipality="Teluk Terima" name="Buleleng" pageId="4" pageNumber="465" stateProvince="Bali">Buleleng</location>
, 
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, 
<date id="FFA710104E49FF8360A525494D52F9CC" box="[136,284,1620,1644]" pageId="4" pageNumber="465" value="2009-07-18">
<collectingDate id="EFE3E9F84E49FF8360A525494D52F9CC" box="[136,284,1620,1644]" pageId="4" pageNumber="465" value="2009-07-18">18 July 2009</collectingDate>
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, 
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&amp; 
<collectorName id="26EC53064E49FF8361A525494E57F9CC" box="[392,537,1620,1644]" pageId="4" pageNumber="465">DC Murniati</collectorName>
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; 
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(40.2 × 
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) (
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. 
<collectingCounty id="62C74E5C4E49FF83612625694D75F92C" box="[267,315,1652,1676]" pageId="4" pageNumber="465">Cru.</collectingCounty>
1996), 
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, 
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, 
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, 
<collectingRegion id="49DDF8324E49FF8360CD25894D6AF90B" box="[224,292,1684,1707]" country="Indonesia" name="Papua Barat" pageId="4" pageNumber="465">Papua</collectingRegion>
, 
<date id="FFA710104E49FF83611D25894D8BF90C" box="[304,453,1684,1708]" pageId="4" pageNumber="465" value="2007-06-12">
<collectingDate id="EFE3E9F84E49FF83611D25894D8BF90C" box="[304,453,1684,1708]" pageId="4" pageNumber="465" value="2007-06-12">12 June 2007</collectingDate>
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, 
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.
</paragraph>
</subSubSection>
<subSubSection id="C303655B4E49FF8360A525C9498DFE55" pageId="4" pageNumber="465" type="discussion">
<paragraph id="8BA636D04E49FF8360A525C9497DFF55" blockId="4.[136,776,1748,2092]" lastBlockId="4.[835,1474,189,245]" pageId="4" pageNumber="465">
<emphasis id="B96DEAC24E49FF8360A525C94CBEF94C" bold="true" box="[136,240,1748,1772]" pageId="4" pageNumber="465">Remarks</emphasis>
. The morphology of the present specimens agrees well with the description and figures by 
<bibRefCitation id="EF884B214E49FF83627E25E94C9CF88C" author="Sakai K &amp; Turkay M" pageId="4" pageNumber="465" pagination="665 - 794" refId="ref19495" refString="Sakai K &amp; Turkay M (2013) Revision of the genus Ocypode with the description of a new genus, Hoplocypode (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum, 56 (2): 665 - 794." type="journal article" year="2013">Sakai &amp; Türkay (2013)</bibRefCitation>
. The inner surface of the major palm has a stridulating ridge which consists of 30–49 elements (tubercles and striae) (
<figureCitation id="13222A554E49FF8360BD24494CA9F8CC" box="[144,231,1876,1900]" captionStart="Fig" captionStartId="3.[113,148,2006,2028]" captionTargetBox="[126,1438,189,1991]" captionTargetId="figure-11@3.[126,1438,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Ocypode ceratophthalmus (Pallas, 1772). MZB.Cru.5017, male, 38.7 × 34.1 mm; Papua: Bepondi Island. A, dorsal habitus, preserved colouration. B–K, left appendages. Chela: B, inner surface; C, outer surface. P2–P5: D–G, anterior surface; H–K, posterior surface; L, stridulating ridge on inner surface of palm of chela. G1: M, whole morphology; N, apical portion. Scale bars: A, 10 mm; B–K, 5 mm; L–N, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.7399908" httpUri="https://zenodo.org/record/7399908/files/figure.png" pageId="4" pageNumber="465">Fig. 2B</figureCitation>
), and the G1 is three sided proximally and has a small thumb-shaped projection subdistally (
<figureCitation id="13222A554E49FF83625524694EB5F82C" box="[632,763,1908,1932]" captionStart="Fig" captionStartId="3.[113,148,2006,2028]" captionTargetBox="[126,1438,189,1991]" captionTargetId="figure-11@3.[126,1438,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Ocypode ceratophthalmus (Pallas, 1772). MZB.Cru.5017, male, 38.7 × 34.1 mm; Papua: Bepondi Island. A, dorsal habitus, preserved colouration. B–K, left appendages. Chela: B, inner surface; C, outer surface. P2–P5: D–G, anterior surface; H–K, posterior surface; L, stridulating ridge on inner surface of palm of chela. G1: M, whole morphology; N, apical portion. Scale bars: A, 10 mm; B–K, 5 mm; L–N, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.7399908" httpUri="https://zenodo.org/record/7399908/files/figure.png" pageId="4" pageNumber="465">Fig. 2M–N</figureCitation>
). There are variations in the number of stridulatory elements on the inner surface of the palm of the major cheliped (35–51) and minor chelipeds (0–10). These variations are related to size of the carapaces. Larger crabs have a larger number of stridulatory elements. On the inner surface of the palm of the minor cheliped, only specimens with more than 40.0 mm cw have this element (
<bibRefCitation id="EF884B214E49FF83641423C04966FF55" author="Fadhillah NF &amp; Prihatini W &amp; Murniati DC" box="[1081,1320,221,245]" pageId="4" pageNumber="465" refId="ref17861" refString="Fadhillah NF, Prihatini W &amp; Murniati DC (2018) Laporan praktik kerja magang di Bidang Zoologi (Museum Zoologicum Bogoriense) Pusat Penelitian Biologi, LIPI. Unpublished internship report. Program Studi Biologi, Fakultas Matematika dan Ilmu Pengetahuan Alam, Universitas Pakuan, Bogor, 27 pp. [In Indonesian]" type="book" year="2018">Fadhillah et al., 2018</bibRefCitation>
).
</paragraph>
<paragraph id="8BA636D04E49FF83636E2200498DFE55" blockId="4.[834,1475,285,501]" pageId="4" pageNumber="465">
This species inhabits high intertidal areas with sandy substrate and is typically nocturnal. They are widely distributed in the Indonesian Archipelago; i.e., 
<collectingRegion id="49DDF8324E49FF8364A7224048ADFED5" box="[1162,1251,349,373]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="465">Sumatra</collectingRegion>
Island, 
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Island, 
<collectingRegion id="49DDF8324E49FF83636E22604FEAFE35" box="[835,932,381,405]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="465">Sulawesi</collectingRegion>
Island, the 
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, the 
<collectingCountry id="F30E76404E49FF83657E226049F0FE35" box="[1363,1470,381,405]" name="Indonesia" pageId="4" pageNumber="465">Moluccas</collectingCountry>
, the 
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, and 
<collectingRegion id="49DDF8324E49FF8364022283483EFE15" box="[1071,1136,414,437]" country="Indonesia" name="Papua Barat" pageId="4" pageNumber="465">Papua</collectingRegion>
(
<bibRefCitation id="EF884B214E49FF8364AC22804969FE15" author="Rathbun MJ" box="[1153,1319,413,437]" pageId="4" pageNumber="465" pagination="305 - 317" refId="ref19413" refString="Rathbun MJ (1910) Decapod crustaceans collected in Dutch East India and elsewhere by Mr. Thomas Barbour in 1906 - 1907. Bulletin of the Museum of Comparative Zoology at Harvard College, 52 (16): 305 - 317." type="journal article" year="1910">Rathbun, 1910</bibRefCitation>
; 
<bibRefCitation id="EF884B214E49FF836518228049F1FE15" author="Tesch JJ" box="[1333,1471,413,437]" pageId="4" pageNumber="465" pagination="1 - 6" refId="ref19962" refString="Tesch JJ (1918) The Decapoda Brachyura of the Siboga-Expedition. I. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga-Expedition Monograph, 39 c (82): 1 - 148, pls 1 - 6." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
; 
<bibRefCitation id="EF884B214E49FF83636E22A0480BFE75" author="Sakai K &amp; Turkay M" box="[835,1093,445,469]" pageId="4" pageNumber="465" pagination="665 - 794" refId="ref19495" refString="Sakai K &amp; Turkay M (2013) Revision of the genus Ocypode with the description of a new genus, Hoplocypode (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum, 56 (2): 665 - 794." type="journal article" year="2013">Sakai &amp; Türkay, 2013</bibRefCitation>
; 
<bibRefCitation id="EF884B214E49FF83647822A048ACFE75" author="Wada K" box="[1109,1250,445,469]" pageId="4" pageNumber="465" refId="ref20063" refString="Wada K (2019) Brachyuran species recorded from mangrove swamp in Indonesia in 1984 - 2001. Cancer, 28: e 144 - e 147. [in Japanese]" type="book" year="2019">Wada, 2019</bibRefCitation>
). The present study gives a new distribution record of the species in 
<collectingCountry id="F30E76404E49FF83657922C049F1FE55" box="[1364,1471,477,501]" name="Indonesia" pageId="4" pageNumber="465">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>