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<mods:title id="2BFD3BF26576A1AC30581BD90D1552DB">A review of the wasp mimicking spider genus Coenoptychus Simon, 1885 (Araneae: Corinnidae: Castianeirinae)</mods:title>
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<mods:namePart id="535D9F1A2633408F41A51C56D4CEA10D">Sebastian, Pothalil A.</mods:namePart>
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<treatment id="03B087C4FF8D551C96A44535FE15FDB3" ID-DOI="http://doi.org/10.5281/zenodo.5979934" ID-GBIF-Taxon="143619735" ID-Zenodo-Dep="5979934" LSID="urn:lsid:plazi:treatment:03B087C4FF8D551C96A44535FE15FDB3" httpUri="http://treatment.plazi.org/id/03B087C4FF8D551C96A44535FE15FDB3" lastPageId="8" lastPageNumber="171" pageId="7" pageNumber="170">
<subSubSection id="C3036559FF8D551396A44535FC9EF818" box="[151,784,1915,1942]" pageId="7" pageNumber="170" type="nomenclature">
<paragraph id="8BA636D2FF8D551396A44535FC9EF818" blockId="7.[151,784,1915,1942]" box="[151,784,1915,1942]" pageId="7" pageNumber="170">
<heading id="D0EE81BEFF8D551396A44535FC9EF818" bold="true" box="[151,784,1915,1942]" fontSize="11" level="1" pageId="7" pageNumber="170" reason="1">
<emphasis id="B96DEAC0FF8D551396A44535FC9EF818" bold="true" box="[151,784,1915,1942]" pageId="7" pageNumber="170">
<taxonomicName id="4C194D51FF8D551396A44535FD0DF818" ID-CoL="5ZJH4" authority="(Haddad, 2004)" authorityName="Paul &amp; Sankaran &amp; Sebastian &amp; Joseph" authorityYear="2018" baseAuthorityName="Haddad" baseAuthorityYear="2004" box="[151,643,1915,1942]" class="Arachnida" family="Corinnidae" genus="Coenoptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="170" phylum="Arthropoda" rank="species" species="tropicalis" status="comb. nov.">
<emphasis id="B96DEAC0FF8D551396A44535FE34F81B" bold="true" box="[151,442,1915,1941]" italics="true" pageId="7" pageNumber="170">Coenoptychus tropicalis</emphasis>
(
<bibRefCitation id="EF884B23FF8D551397FA4532FDF4F818" box="[457,634,1916,1942]" pageId="7" pageNumber="170" refId="ref4745" refString="Haddad, C. R. (2004) A revision of the spider genus Graptartia Simon, 1896 (Araneae: Corinnidae) in the Afrotropical region. African Entomology, 12, 71 - 81." type="journal article">Haddad, 2004</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25E57BBFF8D551394B94532FC9EF818" box="[650,784,1916,1942]" pageId="7" pageNumber="170">comb. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3036559FF8D551C96A445F2FE71FE09" lastPageId="8" lastPageNumber="171" pageId="7" pageNumber="170" type="reference_group">
<paragraph id="8BA636D2FF8D551396A445F2FD52F87E" blockId="7.[151,1436,1980,2032]" pageId="7" pageNumber="170">
<materialsCitation id="3B713C8FFF8D551396A445F2FD52F87E" ID-GBIF-Occurrence="1838188835" collectingDate="1991-11-09" collectionCode="MRAC" collectorName="R. Jocque" country="Tanzania" location="Kyela" pageId="7" pageNumber="170" specimenCode="MRAC 173236" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<treatmentCitationGroup id="AB0911FCFF8D551396A445F2FD52F87E" infered="true" pageId="7" pageNumber="170">
<taxonomicName id="4C194D51FF8D551396A445F2FDBAF85C" ID-CoL="934H2" authority="Haddad, 2004: 78" authorityName="Haddad" authorityPageNumber="78" authorityYear="2004" box="[151,564,1980,2002]" class="Arachnida" family="Corinnidae" genus="Graptartia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="170" phylum="Arthropoda" rank="species" species="tropicalis">
<emphasis id="B96DEAC0FF8D551396A445F2FEE0F85C" box="[151,366,1980,2002]" italics="true" pageId="7" pageNumber="170">Graptartia tropicalis</emphasis>
<treatmentCitation id="0AB810C3FF8D5513974445F2FDBAF85C" author="" box="[375,564,1980,2002]" page="78" pageId="7" pageNumber="170">
<bibRefCitation id="EF884B23FF8D5513974445F2FD82F85C" box="[375,524,1980,2002]" pageId="7" pageNumber="170" refId="ref4745" refString="Haddad, C. R. (2004) A revision of the spider genus Graptartia Simon, 1896 (Araneae: Corinnidae) in the Afrotropical region. African Entomology, 12, 71 - 81." type="journal article">Haddad, 2004</bibRefCitation>
: 78
</treatmentCitation>
</taxonomicName>
, figs 3, 6, 9, 12, 1416, 2933 (
<typeStatus id="54A28870FF8D551395BE45F2FC63F85C" box="[909,1005,1980,2002]" pageId="7" pageNumber="170" type="holotype">Holotype</typeStatus>
<specimenCount id="9D1FFD5BFF8D551395C545F3FB85F85C" box="[1014,1035,1981,2002]" pageId="7" pageNumber="170" type="female"></specimenCount>
:
<collectingCountry id="F30E7642FF8D5513922945F2FB11F85F" box="[1050,1183,1980,2001]" name="Tanzania" pageId="7" pageNumber="170">TANZANIA</collectingCountry>
:
<location id="8EC66009FF8D5513929945F2FB68F85C" LSID="urn:lsid:plazi:treatment:03B087C4FF8D551C96A44535FE15FDB3:8EC66009FF8D5513929945F2FB68F85C" box="[1194,1254,1980,2002]" country="Tanzania" name="Kyela" pageId="7" pageNumber="170">Kyela</location>
(garden), leg.
<collectorName id="26EC5304FF8D551393B745F3FE9EF87E" pageId="7" pageNumber="170">R. Jocqué</collectorName>
,
<collectingDate id="EFE3E9FAFF8D551397294594FE0CF87E" box="[282,386,2010,2032]" pageId="7" pageNumber="170" value="1991-11-09">
<date id="FFA71012FF8D551397294594FE0CF87E" box="[282,386,2010,2032]" pageId="7" pageNumber="170" value="1991-11-09">9.XI.1991</date>
</collectingDate>
,
<specimenCode id="DBBF9EA9FF8D551397BE4594FDA7F87E" box="[397,553,2010,2032]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Mus<75>e Royal de l<>Afrique Centrale" pageId="7" pageNumber="170">MRAC 173236</specimenCode>
—not examined).
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
<paragraph id="8BA636D2FF82551C96A442F2FE71FE09" blockId="8.[151,1436,188,573]" pageId="8" pageNumber="171">
<emphasis id="B96DEAC0FF82551C96A442F2FE30FF5B" bold="true" box="[151,446,188,213]" pageId="8" pageNumber="171">Justification for transfer</emphasis>
. The original description and illustrations of
<taxonomicName id="4C194D51FF82551C95F742F3FA87FF5A" ID-CoL="934H2" authority="(Haddad, 2004)" authorityName="Haddad" authorityYear="2004" baseAuthorityName="Haddad" baseAuthorityYear="2004" box="[964,1289,188,213]" class="Arachnida" family="Corinnidae" genus="Graptartia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="171" phylum="Arthropoda" rank="species" species="tropicalis">
<emphasis id="B96DEAC0FF82551C95F742F3FBC0FF5B" box="[964,1102,188,213]" italics="true" pageId="8" pageNumber="171">G. tropicalis</emphasis>
(
<bibRefCitation id="EF884B23FF82551C926C42F2FA8FFF5A" box="[1119,1281,188,213]" pageId="8" pageNumber="171" refId="ref4745" refString="Haddad, C. R. (2004) A revision of the spider genus Graptartia Simon, 1896 (Araneae: Corinnidae) in the Afrotropical region. African Entomology, 12, 71 - 81." type="journal article">Haddad, 2004</bibRefCitation>
)
</taxonomicName>
indicate that the diagnostic features of the species match the somatic and genitalic features of
<taxonomicName id="4C194D51FF82551C921F4291FB44FF76" ID-CoL="3S2H" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[1068,1226,223,248]" class="Arachnida" family="Corinnidae" genus="Coenoptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAC0FF82551C921F4291FB44FF76" box="[1068,1226,223,248]" italics="true" pageId="8" pageNumber="171">Coenoptychus</emphasis>
</taxonomicName>
: male smaller and less robust than female, recurved PER, abdomen and legs with feathery setae, promargin of chelicerae with three teeth, subtriangular spermathecae I and reniform spermathecae I, lack of RTA in the male palp, and spiralled embolus with two turns (
<bibRefCitation id="EF884B23FF82551C979F4302FCD6FEEA" box="[428,856,332,357]" pageId="8" pageNumber="171" refId="ref4745" refString="Haddad, C. R. (2004) A revision of the spider genus Graptartia Simon, 1896 (Araneae: Corinnidae) in the Afrotropical region. African Entomology, 12, 71 - 81." type="journal article">Haddad 2004: figs 3, 9, 12, 14, 2933</bibRefCitation>
). Based on these characters, we propose the transfer of this species to
<taxonomicName id="4C194D51FF82551C976A4321FE79FE06" ID-CoL="3S2H" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[345,503,367,392]" class="Arachnida" family="Corinnidae" genus="Coenoptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAC0FF82551C976A4321FE79FE06" box="[345,503,367,392]" italics="true" pageId="8" pageNumber="171">Coenoptychus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036559FF82551C96F443DAFE15FDB3" pageId="8" pageNumber="171" type="distribution">
<paragraph id="8BA636D2FF82551C96F443DAFB7DFE23" blockId="8.[151,1436,188,573]" box="[199,1267,404,429]" pageId="8" pageNumber="171">
<emphasis id="B96DEAC0FF82551C96F443DAFEEEFE23" bold="true" box="[199,352,404,429]" pageId="8" pageNumber="171">Distribution.</emphasis>
<collectingCountry id="F30E7642FF82551C975443DAFD7FFE22" box="[359,753,404,429]" name="Democratic Republic of the Congo" pageId="8" pageNumber="171">
Democratic Republic of the
<collectingRegion id="49DDF830FF82551C949743DAFD7FFE22" box="[676,753,404,428]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="8" pageNumber="171">Congo</collectingRegion>
</collectingCountry>
,
<collectingCountry id="F30E7642FF82551C94CE43DBFC0AFE22" box="[765,900,404,429]" name="Ivory Coast" pageId="8" pageNumber="171">
Ivory
<collectingRegion id="49DDF830FF82551C957243DAFC0AFE22" box="[833,900,404,428]" country="Tanzania" name="Pwani" pageId="8" pageNumber="171">Coast</collectingRegion>
</collectingCountry>
,
<collectingCountry id="F30E7642FF82551C95BC43DAFBAFFE23" box="[911,1057,404,429]" name="South Africa" pageId="8" pageNumber="171">
<collectingRegion id="49DDF830FF82551C95BC43DAFC5EFE23" box="[911,976,404,429]" country="Tanzania" name="Kusini Unguja" pageId="8" pageNumber="171">South</collectingRegion>
Africa
</collectingCountry>
,
<collectingCountry id="F30E7642FF82551C921F43DAFB1FFE23" box="[1068,1169,404,429]" name="Tanzania" pageId="8" pageNumber="171">Tanzania</collectingCountry>
(
<figureCitation id="13222A57FF82551C929343DAFB6BFE23" box="[1184,1253,404,429]" captionStart="FIGURE 5" captionStartId="7.[151,250,1290,1312]" captionTargetBox="[151,1436,636,1279]" captionTargetId="figure@7.[151,1436,636,1279]" captionTargetPageId="7" captionText="FIGURE 5. Distribution records of Coenoptychus pulcher (circles), C. mutillicus (squares) and C. tropicalis (triangles): type localities (black), new records (red), other published records (white)." httpUri="https://zenodo.org/record/1221994/files/figure.png" pageId="8" pageNumber="171">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BA636D2FF82551C96F443F9FE15FDB3" blockId="8.[151,1436,188,573]" pageId="8" pageNumber="171">
<emphasis id="B96DEAC0FF82551C96F443F9FE88FE5E" bold="true" box="[199,262,439,464]" pageId="8" pageNumber="171">Note.</emphasis>
The transfer of both
<taxonomicName id="4C194D51FF82551C97C743F6FDF9FE5E" baseAuthorityName="Haddad" baseAuthorityYear="2004" box="[500,631,439,464]" class="Arachnida" family="Corinnidae" genus="Graptartia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="171" phylum="Arthropoda" rank="species" species="mutillica">
<emphasis id="B96DEAC0FF82551C97C743F6FDF9FE5E" box="[500,631,439,464]" italics="true" pageId="8" pageNumber="171">G. mutillica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D51FF82551C949E43F6FCB9FE5E" baseAuthorityName="Haddad" baseAuthorityYear="2004" box="[685,823,439,464]" class="Arachnida" family="Corinnidae" genus="Graptartia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="171" phylum="Arthropoda" rank="species" species="tropicalis">
<emphasis id="B96DEAC0FF82551C949E43F6FCB9FE5E" box="[685,823,439,464]" italics="true" pageId="8" pageNumber="171">G. tropicalis</emphasis>
</taxonomicName>
to
<taxonomicName id="4C194D51FF82551C956843F9FC74FE5E" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[859,1018,439,464]" class="Arachnida" family="Corinnidae" genus="Coenoptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAC0FF82551C956843F9FC74FE5E" box="[859,1018,439,464]" italics="true" pageId="8" pageNumber="171">Coenoptychus</emphasis>
</taxonomicName>
is supported by the cladistic analysis of Afrotropical
<taxonomicName id="4C194D51FF82551C97614392FE7CFE7B" authorityName="Reiskind" authorityYear="1969" box="[338,498,476,501]" class="Arachnida" family="Corinnidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="171" phylum="Arthropoda" rank="subFamily" subFamily="Castianeirinae">Castianeirinae</taxonomicName>
by
<bibRefCitation id="EF884B23FF82551C94144392FD56FE7A" box="[551,728,476,501]" pageId="8" pageNumber="171" refId="ref4783" refString="Haddad, C. R. (2013) Taxonomic notes on the spider genus Messapus Simon, 1898 (Araneae, Corinnidae), with the description of the new genera Copuetta and Wasaka and the first cladistic analysis of Afrotropical Castianeirinae. Zootaxa, 3688 (1), 1 - 79. https: // doi. org / 10.11646 / zootaxa. 3688.1.1" type="journal article">Haddad (2013)</bibRefCitation>
, wherein these two species were in a clade paraphyletic to
<taxonomicName id="4C194D51FF82551C96A4404EFDAAFD96" authority="Simon, 1896" authorityName="Simon" authorityYear="1896" box="[151,548,511,536]" class="Arachnida" family="Corinnidae" genus="Graptartia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="171" phylum="Arthropoda" rank="species" species="granulosa">
<emphasis id="B96DEAC0FF82551C96A4404EFE04FD96" box="[151,394,511,536]" italics="true" pageId="8" pageNumber="171">Graptartia granulosa</emphasis>
Simon, 1896
</taxonomicName>
. He suggested that these two species may belong to
<taxonomicName id="4C194D51FF82551C924743B1FA9DFD96" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[1140,1299,511,536]" class="Arachnida" family="Corinnidae" genus="Coenoptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAC0FF82551C924743B1FA9DFD96" box="[1140,1299,511,536]" italics="true" pageId="8" pageNumber="171">Coenoptychus</emphasis>
</taxonomicName>
, but did not formally transfer them.
</paragraph>
</subSubSection>
</treatment>
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