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<document id="EE79ABE833985A05344E4C8612536A2D" ID-CLB-Dataset="46311" ID-DOI="10.5281/zenodo.278746" ID-GBIF-Dataset="290b4c66-1488-4f52-ba67-7f7942a01328" ID-ISSN="1175-5326" ID-Zenodo-Dep="278746" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460391410718" checkinUser="plazi" docAuthor="Nguyen, Anh D." docDate="2011" docId="03B087BF60781F4AFF3AFDE5AAABFA1B" docLanguage="en" docName="zt03036p068.pdf" docOrigin="Zootaxa 3036" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tonkinosoma Jeekel 1953, sp. nov." docType="treatment" docVersion="10" lastPageNumber="67" masterDocId="FF89FFC7607E1F43FFADFD72AB3BFF90" masterDocTitle="A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam" masterLastPageNumber="68" masterPageNumber="58" pageNumber="64" updateTime="1698298273915" updateUser="ExternalLinkService">
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<mods:title id="BA76D1A0ECDC4ED6C0661C2E1F58F326">A review of the millipede tribe Tonkinosomatini (Diplopoda: Polydesmida: Paradoxosomatidae) from Vietnam</mods:title>
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<mods:namePart id="28AFF66F774DE20AE0C10718C32DDFD3">Nguyen, Anh D.</mods:namePart>
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<mods:date id="D9FD7C48EF5CF231943F878F3A4BB51F">2011</mods:date>
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<treatment id="03B087BF60781F4AFF3AFDE5AAABFA1B" ID-DOI="http://doi.org/10.5281/zenodo.6189320" ID-GBIF-Taxon="119416885" ID-Zenodo-Dep="6189320" LSID="urn:lsid:plazi:treatment:03B087BF60781F4AFF3AFDE5AAABFA1B" httpUri="http://treatment.plazi.org/id/03B087BF60781F4AFF3AFDE5AAABFA1B" lastPageId="9" lastPageNumber="67" pageId="6" pageNumber="64">
<subSubSection id="C303652260781F45FF3AFDE5AA53FE87" pageId="6" pageNumber="64" type="nomenclature">
<paragraph id="8BA636A960781F45FF3AFDE5A914FF21" blockId="6.[151,559,151,177]" box="[151,559,151,177]" pageId="6" pageNumber="64">
<heading id="D0EE81C560781F45FF3AFDE5A914FF21" bold="true" box="[151,559,151,177]" fontSize="11" level="1" pageId="6" pageNumber="64" reason="1">
<emphasis id="B96DEABB60781F45FF3AFDE5A914FF21" bold="true" box="[151,559,151,177]" pageId="6" pageNumber="64">
Genus
<taxonomicName id="4C194D2A60781F45FF43FDE5A914FF21" ID-CoL="7Q4P5" authority="Jeekel, 1953" authorityName="Jeekel" authorityYear="1953" box="[238,559,151,177]" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABB60781F45FF43FDE5AAA9FF21" bold="true" box="[238,402,151,177]" italics="true" pageId="6" pageNumber="64">Tonkinosoma</emphasis>
Jeekel, 1953
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA636A960781F45FF3AFDAFAA53FE87" blockId="6.[151,496,221,279]" pageId="6" pageNumber="64">
<taxonomicName id="4C194D2A60781F45FF3AFDAFAAABFF67" box="[151,400,221,247]" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="64" phylum="Arthropoda" rank="species" species="jeekeli" status="sp. nov.">
<emphasis id="B96DEABB60781F45FF3AFDAFAAABFF67" bold="true" box="[151,400,221,247]" italics="true" pageId="6" pageNumber="64">Tonkinosoma jeekeli</emphasis>
</taxonomicName>
<emphasis id="B96DEABB60781F45FE3AFDACAACBFF66" bold="true" box="[407,496,222,246]" pageId="6" pageNumber="64">
<taxonomicNameLabel id="A25E57C060781F45FE3AFDACAACBFF66" box="[407,496,222,246]" pageId="6" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13222A2C60781F45FF32FD8DAA30FE87" box="[159,267,255,279]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="6.[151,250,1571,1594]" captionStartId-1="7.[151,250,1957,1980]" captionStartId-2="8.[151,250,1120,1143]" captionTargetBox-0="[264,1322,909,1549]" captionTargetBox-1="[300,1290,487,1914]" captionTargetBox-2="[264,1323,193,1098]" captionTargetId-0="figure@6.[264,1323,909,1550]" captionTargetId-1="figure@7.[264,1323,481,1936]" captionTargetId-2="figure@8.[264,1323,193,1099]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURE 6. Tonkinosoma jeekeli sp. nov., holotype body: anterior part, lateral view (A), posterior part, lateral view (C), whole body, dorsal view (B). Scale bar = 10 mm." captionText-1="FIGURE 7. Tonkinosoma jeekeli sp. nov., holotype, body ring 10 11, dorsal view (A); sternite V, ventral view (B); hypop-roct, ventral view (C); epiproct, ventral view (D); left gonopod, lateral view (E), mesal view (F). Scale bar = 1 mm." captionText-2="FIGURE 8. Tonkinosoma jeekeli sp. nov., paratype, left gonopod, mesal view (A), lateral view (B); Tip of gonopod, lateral view (C)." httpUri-0="https://zenodo.org/record/278752/files/figure.png" httpUri-1="https://zenodo.org/record/278753/files/figure.png" httpUri-2="https://zenodo.org/record/278754/files/figure.png" pageId="6" pageNumber="64">Figs. 68</figureCitation>
. Map 1)
</paragraph>
</subSubSection>
<subSubSection id="C303652260781F45FF3AFC35AF5EFE5C" pageId="6" pageNumber="64" type="materials_examined">
<paragraph id="8BA636A960781F45FF3AFC35A99AFE14" blockId="6.[151,1437,327,857]" pageId="6" pageNumber="64">
<emphasis id="B96DEABB60781F45FF3AFC35AABAFEF0" bold="true" box="[151,385,327,352]" pageId="6" pageNumber="64">Material examined.</emphasis>
<typeStatus id="54A2880B60781F45FE24FC35A926FECF" box="[393,541,327,351]" pageId="6" pageNumber="64" type="holotype">HOLOTYPE</typeStatus>
: 13 (
<emphasis id="B96DEABB60781F45FDCFFC35A9CBFEF0" bold="true" box="[610,752,327,352]" pageId="6" pageNumber="64">IEBR-128H</emphasis>
),
<collectingCountry id="F30E763960781F45FCA5FC35A851FECF" box="[776,874,327,351]" name="Vietnam" pageId="6" pageNumber="64">Vietnam</collectingCountry>
, Ninh Binh Province, Cuc Phuong National Park, forest,
<date id="FFA7106960781F45FF48FC1EAAFBFE14" box="[229,448,364,388]" pageId="6" pageNumber="64" value="2006-09-15">15 September 2006</date>
, leg. Luu Van Hien.
</paragraph>
<paragraph id="8BA636A960781F45FF68FCFDAF5EFE5C" blockId="6.[151,1437,327,857]" pageId="6" pageNumber="64">
<typeStatus id="54A2880B60781F45FF68FCFDAA66FE37" box="[197,349,399,423]" pageId="6" pageNumber="64" type="paratype">PARATYPES</typeStatus>
: 23, 3ƤƤ (
<emphasis id="B96DEABB60781F45FDA4FCFDA9A9FE38" bold="true" box="[521,658,399,424]" pageId="6" pageNumber="64">IEBR-128P</emphasis>
), same data as
<typeStatus id="54A2880B60781F45FCE1FCFDA889FE37" box="[844,946,399,423]" pageId="6" pageNumber="64" type="holotype">holotype</typeStatus>
; 3 3, (
<emphasis id="B96DEABB60781F45FBB3FCFDAFADFE38" bold="true" box="[1054,1174,399,424]" pageId="6" pageNumber="64">IEBR-129</emphasis>
),
<collectingCountry id="F30E763960781F45FB1CFCFDAE28FE37" box="[1201,1299,399,423]" name="Vietnam" pageId="6" pageNumber="64">Vietnam</collectingCountry>
, Hai Phong Province, Cat Ba National Park, primary forest,
<date id="FFA7106960781F45FD1FFCC6A853FE5C" box="[690,872,436,460]" pageId="6" pageNumber="64" value="2006-08-10">10 August 2006</date>
, leg. Nguyen Tri Tien.
</paragraph>
</subSubSection>
<subSubSection id="C303652260781F45FF6AFCAAA91CFD84" pageId="6" pageNumber="64" type="etymology">
<paragraph id="8BA636A960781F45FF6AFCAAA91CFD84" blockId="6.[151,1437,327,857]" pageId="6" pageNumber="64">
<emphasis id="B96DEABB60781F45FF6AFCAAAA75FE60" bold="true" box="[199,334,472,496]" pageId="6" pageNumber="64">Etymology.</emphasis>
This species is dedicated to the late Dr. C.A.W. Jeekel, a prominent specialist in Diplopoda, who established the genus
<taxonomicName id="4C194D2A60781F45FE21FC8EA91BFD84" box="[396,544,508,532]" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABB60781F45FE21FC8EA91BFD84" box="[396,544,508,532]" italics="true" pageId="6" pageNumber="64">Tonkinosoma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C303652260781F45FF6AFF6DAA44FD34" pageId="6" pageNumber="64" type="diagnosis">
<paragraph id="8BA636A960781F45FF6AFF6DAA44FD34" blockId="6.[151,1437,327,857]" pageId="6" pageNumber="64">
<emphasis id="B96DEABB60781F45FF6AFF6DAA7AFDA8" bold="true" box="[199,321,543,568]" pageId="6" pageNumber="64">Diagnosis.</emphasis>
This species differs from its congener
<taxonomicName id="4C194D2A60781F45FD56FF52AFB8FDA7" authority="Jeekel, 1953" authorityName="Jeekel" authorityYear="1953" box="[763,1155,543,568]" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="64" phylum="Arthropoda" rank="species" species="flexipes">
<emphasis id="B96DEABB60781F45FD56FF52A8D7FDA8" box="[763,1004,544,568]" italics="true" pageId="6" pageNumber="64">Tonkinosoma flexipes</emphasis>
Jeekel, 1953
</taxonomicName>
, in following characters: sternite V with two independent small lamina between coxae 4, and without any modification between coxae 5; gonopod femorite much broader and constricted distally; solenophore with both well-developed
<emphasis id="B96DEABB60781F45FB4BFF1AAEA7FD10" box="[1254,1436,616,640]" italics="true" pageId="6" pageNumber="64">lamina medialis</emphasis>
and
<emphasis id="B96DEABB60781F45FF6BFFFEAA43FD34" box="[198,376,652,676]" italics="true" pageId="6" pageNumber="64">lamina lateralis</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C303652260781F4BFF6AFFDDAA88FA45" lastPageId="8" lastPageNumber="66" pageId="6" pageNumber="64" type="description">
<paragraph id="8BA636A960781F45FF6AFFDDA896FC9F" blockId="6.[151,1437,327,857]" pageId="6" pageNumber="64">
<emphasis id="B96DEABB60781F45FF6AFFDDAA62FD58" bold="true" box="[199,345,687,712]" pageId="6" pageNumber="64">Description.</emphasis>
Size: body
<quantity id="4CE19B4C60781F45FE70FFDDA943FD57" box="[477,632,687,711]" metricMagnitude="-2" metricUnit="m" metricValue="4.505" metricValueMax="4.75" metricValueMin="4.26" pageId="6" pageNumber="64" unit="mm" value="45.05" valueMax="47.5" valueMin="42.6">42.647.5mm</quantity>
(male),
<quantity id="4CE19B4C60781F45FD7AFFDDA824FD57" box="[727,799,687,711]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="6" pageNumber="64" unit="mm" value="57.0">57mm</quantity>
(female) in length, width of midbody pro- and metazona
<quantity id="4CE19B4C60781F45FF3AFFA6AA2DFD7C" box="[151,278,724,748]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.3" metricValueMin="4.2" pageId="6" pageNumber="64" unit="mm" value="4.25" valueMax="4.3" valueMin="4.2">4.24.3mm</quantity>
(male), 4.8 (female) and
<quantity id="4CE19B4C60781F45FD99FFA6A988FD7C" box="[564,691,724,748]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.8" metricValueMin="5.3" pageId="6" pageNumber="64" unit="mm" value="5.55" valueMax="5.8" valueMin="5.3">5.35.8mm</quantity>
(male),
<quantity id="4CE19B4C60781F45FCBFFFA6A85BFD7C" box="[786,864,724,748]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="6" pageNumber="64" unit="mm" value="7.2">7.2mm</quantity>
(female), respectively.
<typeStatus id="54A2880B60781F45FBC5FFA6AFEBFD7C" box="[1128,1232,724,748]" pageId="6" pageNumber="64" type="holotype">Holotype</typeStatus>
<quantity id="4CE19B4C60781F45FB7AFFA6AE08FD7C" box="[1239,1331,724,748]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" pageId="6" pageNumber="64" unit="mm" value="47.5">47.5mm</quantity>
long and width of midbody pro- and metazoan
<quantity id="4CE19B4C60781F45FD93FF85A9B6FC9F" box="[574,653,759,783]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="6" pageNumber="64" unit="mm" value="4.3">4.3mm</quantity>
and
<quantity id="4CE19B4C60781F45FD6EFF85A82FFC9F" box="[707,788,759,783]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="6" pageNumber="64" unit="mm" value="5.8">5.8mm</quantity>
, respectively.
</paragraph>
<paragraph id="8BA636A960781F45FF68FE6EA9DDFCC7" blockId="6.[151,1437,327,857]" pageId="6" pageNumber="64">
Coloration: body in general reddish brown; legs, antenna and sterna yellowish brown; paraterga, marginally rear part of metaterga and pleura paler (
<figureCitation id="13222A2C60781F45FDFFFE4DA99BFCC7" box="[594,672,831,855]" captionStart="FIGURE 6" captionStartId="6.[151,250,1571,1594]" captionTargetBox="[264,1322,909,1549]" captionTargetId="figure@6.[264,1323,909,1550]" captionTargetPageId="6" captionText="FIGURE 6. Tonkinosoma jeekeli sp. nov., holotype body: anterior part, lateral view (A), posterior part, lateral view (C), whole body, dorsal view (B). Scale bar = 10 mm." httpUri="https://zenodo.org/record/278752/files/figure.png" pageId="6" pageNumber="64">Figs. 6</figureCitation>
AC).
</paragraph>
<caption id="DF66662160781F45FF3AFB51A97EF9C8" httpUri="https://zenodo.org/record/278752/files/figure.png" pageId="6" pageNumber="64" targetBox="[264,1322,909,1549]" targetPageId="6">
<paragraph id="8BA636A960781F45FF3AFB51A97EF9C8" blockId="6.[151,1436,1571,1625]" pageId="6" pageNumber="64">
<emphasis id="B96DEABB60781F45FF3AFB51AA29F9A9" bold="true" box="[151,274,1571,1594]" pageId="6" pageNumber="64">FIGURE 6.</emphasis>
<taxonomicName id="4C194D2A60781F45FEB4FB56AADCF9AA" box="[281,487,1572,1594]" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="64" phylum="Arthropoda" rank="species" species="jeekeli" status="sp. nov.">
<emphasis id="B96DEABB60781F45FEB4FB56AADCF9AA" box="[281,487,1572,1594]" italics="true" pageId="6" pageNumber="64">Tonkinosoma jeekeli</emphasis>
</taxonomicName>
<emphasis id="B96DEABB60781F45FE40FB56A905F9A9" bold="true" box="[493,574,1572,1593]" pageId="6" pageNumber="64">
<taxonomicNameLabel id="A25E57C060781F45FE40FB56A905F9A9" box="[493,574,1572,1593]" pageId="6" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype body: anterior part, lateral view (A), posterior part, lateral view (C), whole body, dorsal view (B). Scale bar = 10 mm.
</paragraph>
</caption>
<paragraph id="8BA636A960781F45FF68FB0DA99AF94F" blockId="6.[151,1436,1663,2012]" pageId="6" pageNumber="64">
Head: a bit smaller than collum, modestly setose on labrum, but sparsely on area between antennal sockets. Frons slightly convex. Epicranical suture obviously distinct. Antenna short, claviform, ca. reaching to body ring 3 laterally. Antennomere 1 &lt;7&lt;6&lt;2=5 (
<figureCitation id="13222A2C60781F45FD97FBB5A946F94F" box="[570,637,1735,1759]" captionStart="FIGURE 6" captionStartId="6.[151,250,1571,1594]" captionTargetBox="[264,1322,909,1549]" captionTargetId="figure@6.[264,1323,909,1550]" captionTargetPageId="6" captionText="FIGURE 6. Tonkinosoma jeekeli sp. nov., holotype body: anterior part, lateral view (A), posterior part, lateral view (C), whole body, dorsal view (B). Scale bar = 10 mm." httpUri="https://zenodo.org/record/278752/files/figure.png" pageId="6" pageNumber="64">Fig. 6</figureCitation>
A).
</paragraph>
<paragraph id="8BA636A960781F45FF68FB9EA9D8F8DC" blockId="6.[151,1436,1663,2012]" pageId="6" pageNumber="64">Collum: slightly broader than body ring 2, subtrapeziform, surface shining, smooth, without any rugosity or wrinkles, and with traces of 2 rows of 4+4 and 3+3 setae in front margin and in middle, respectively. Paratergum subtriangular, poorly-developed. Axial line wanting.</paragraph>
<paragraph id="8BA636A960781F44FF68FA25A846FF44" blockId="6.[151,1436,1663,2012]" lastBlockId="7.[151,1436,151,429]" lastPageId="7" lastPageNumber="65" pageId="6" pageNumber="64">
Body rings: Width of body ring 4&lt;3&lt;2=5, parallel-sided on body rings 516, thereafter tapering gradually posterior. Surface smooth, shining, with traces of a row of 3+3 setae before transverse sulcus, which starts on metatergum 5, however, very superficial and vague (
<figureCitation id="13222A2C60781F45FD0FFAEDA9D3F827" box="[674,744,1951,1975]" captionStart="FIGURE 7" captionStartId="7.[151,250,1957,1980]" captionTargetBox="[300,1290,487,1914]" captionTargetId="figure@7.[264,1323,481,1936]" captionTargetPageId="7" captionText="FIGURE 7. Tonkinosoma jeekeli sp. nov., holotype, body ring 10 11, dorsal view (A); sternite V, ventral view (B); hypop-roct, ventral view (C); epiproct, ventral view (D); left gonopod, lateral view (E), mesal view (F). Scale bar = 1 mm." httpUri="https://zenodo.org/record/278753/files/figure.png" pageId="6" pageNumber="64">Fig. 7</figureCitation>
A). Paraterga modestly developed, set lower than metatergal surface (
<figureCitation id="13222A2C60781F45FF54FAB6AA05F84C" box="[249,318,1988,2012]" captionStart="FIGURE 6" captionStartId="6.[151,250,1571,1594]" captionTargetBox="[264,1322,909,1549]" captionTargetId="figure@6.[264,1323,909,1550]" captionTargetPageId="6" captionText="FIGURE 6. Tonkinosoma jeekeli sp. nov., holotype body: anterior part, lateral view (A), posterior part, lateral view (C), whole body, dorsal view (B). Scale bar = 10 mm." httpUri="https://zenodo.org/record/278752/files/figure.png" pageId="6" pageNumber="64">Fig. 6</figureCitation>
A&amp;C). Caudal corner acute, surpassing rear contour (
<figureCitation id="13222A2C60781F45FC0EFAB6A8C8F84C" box="[931,1011,1988,2012]" captionStart="FIGURE 6" captionStartId="6.[151,250,1571,1594]" captionTargetBox="[264,1322,909,1549]" captionTargetId="figure@6.[264,1323,909,1550]" captionTargetPageId="6" captionText="FIGURE 6. Tonkinosoma jeekeli sp. nov., holotype body: anterior part, lateral view (A), posterior part, lateral view (C), whole body, dorsal view (B). Scale bar = 10 mm." httpUri="https://zenodo.org/record/278752/files/figure.png" pageId="6" pageNumber="64">Figs. 6</figureCitation>
A&amp;C, 7A). Pleurosternal carinae like small, oblique keels, present until body ring 10, and missing on subsequent ones (
<figureCitation id="13222A2C60791F44FB86FDE5AF55FF3F" box="[1067,1134,151,175]" captionStart="FIGURE 6" captionStartId="6.[151,250,1571,1594]" captionTargetBox="[264,1322,909,1549]" captionTargetId="figure@6.[264,1323,909,1550]" captionTargetPageId="6" captionText="FIGURE 6. Tonkinosoma jeekeli sp. nov., holotype body: anterior part, lateral view (A), posterior part, lateral view (C), whole body, dorsal view (B). Scale bar = 10 mm." httpUri="https://zenodo.org/record/278752/files/figure.png" pageId="7" pageNumber="65">Fig. 6</figureCitation>
A&amp;C). Axial line thin, visible. Stricture dividing pro- and metazona thin, narrow and striate.
</paragraph>
<paragraph id="8BA636A960791F44FF68FDADAF20FE8C" blockId="7.[151,1436,151,429]" pageId="7" pageNumber="65">
Sterna: modestly setose, cross-impression moderate in terms of both longitudinal and transverse suture, without any modification except two independent lamina between coxae 4 (
<figureCitation id="13222A2C60791F44FC18FC76A8C3FE8C" box="[949,1016,260,284]" captionStart="FIGURE 7" captionStartId="7.[151,250,1957,1980]" captionTargetBox="[300,1290,487,1914]" captionTargetId="figure@7.[264,1323,481,1936]" captionTargetPageId="7" captionText="FIGURE 7. Tonkinosoma jeekeli sp. nov., holotype, body ring 10 11, dorsal view (A); sternite V, ventral view (B); hypop-roct, ventral view (C); epiproct, ventral view (D); left gonopod, lateral view (E), mesal view (F). Scale bar = 1 mm." httpUri="https://zenodo.org/record/278753/files/figure.png" pageId="7" pageNumber="65">Fig. 7</figureCitation>
B).
</paragraph>
<paragraph id="8BA636A960791F44FF68FC5AA97DFEF4" blockId="7.[151,1436,151,429]" pageId="7" pageNumber="65">Legs: rather short, but slender, about 1.1 times (male) as long as body height. No adenostyle or any other modification. Tarsi without setose brushes.</paragraph>
<paragraph id="8BA636A960791F44FF68FC1DAE2CFE3C" blockId="7.[151,1436,151,429]" pageId="7" pageNumber="65">
Telson: Epiproct short, broadly truncated with four spinnerets at tip (
<figureCitation id="13222A2C60791F44FC6AFC1DAF30FE17" box="[967,1035,367,391]" captionStart="FIGURE 7" captionStartId="7.[151,250,1957,1980]" captionTargetBox="[300,1290,487,1914]" captionTargetId="figure@7.[264,1323,481,1936]" captionTargetPageId="7" captionText="FIGURE 7. Tonkinosoma jeekeli sp. nov., holotype, body ring 10 11, dorsal view (A); sternite V, ventral view (B); hypop-roct, ventral view (C); epiproct, ventral view (D); left gonopod, lateral view (E), mesal view (F). Scale bar = 1 mm." httpUri="https://zenodo.org/record/278753/files/figure.png" pageId="7" pageNumber="65">Fig. 7</figureCitation>
D). Paraproct sub-semicircular with two setiferous knobs on disk. Hypoproct subtriangular with two distolateral setiferous knobs (
<figureCitation id="13222A2C60791F44FB1FFCE6AFCEFE3C" box="[1202,1269,404,428]" captionStart="FIGURE 7" captionStartId="7.[151,250,1957,1980]" captionTargetBox="[300,1290,487,1914]" captionTargetId="figure@7.[264,1323,481,1936]" captionTargetPageId="7" captionText="FIGURE 7. Tonkinosoma jeekeli sp. nov., holotype, body ring 10 11, dorsal view (A); sternite V, ventral view (B); hypop-roct, ventral view (C); epiproct, ventral view (D); left gonopod, lateral view (E), mesal view (F). Scale bar = 1 mm." httpUri="https://zenodo.org/record/278753/files/figure.png" pageId="7" pageNumber="65">Fig. 7</figureCitation>
C).
</paragraph>
<caption id="DF66662160791F44FF3AFAD7AFCEF84A" httpUri="https://zenodo.org/record/278753/files/figure.png" pageId="7" pageNumber="65" targetBox="[300,1290,487,1914]" targetPageId="7">
<paragraph id="8BA636A960791F44FF3AFAD7AFCEF84A" blockId="7.[151,1436,1957,2011]" pageId="7" pageNumber="65">
<emphasis id="B96DEABB60791F44FF3AFAD7AA29F82B" bold="true" box="[151,274,1957,1980]" pageId="7" pageNumber="65">FIGURE 7.</emphasis>
<taxonomicName id="4C194D2A60791F44FEB5FAD4AADEF82C" box="[280,485,1958,1980]" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="65" phylum="Arthropoda" rank="species" species="jeekeli" status="sp. nov.">
<emphasis id="B96DEABB60791F44FEB5FAD4AADEF82C" box="[280,485,1958,1980]" italics="true" pageId="7" pageNumber="65">Tonkinosoma jeekeli</emphasis>
</taxonomicName>
<emphasis id="B96DEABB60791F44FE41FAD4A900F82B" bold="true" box="[492,571,1958,1979]" pageId="7" pageNumber="65">
<taxonomicNameLabel id="A25E57C060791F44FE41FAD4A900F82B" box="[492,571,1958,1979]" pageId="7" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, body ring 1011, dorsal view (A); sternite V, ventral view (B); hypop-roct, ventral view (C); epiproct, ventral view (D); left gonopod, lateral view (E), mesal view (F). Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="DF66662160761F4BFF3AF912ABCCFB05" httpUri="https://zenodo.org/record/278754/files/figure.png" pageId="8" pageNumber="66" targetBox="[264,1323,193,1098]" targetPageId="8">
<paragraph id="8BA636A960761F4BFF3AF912ABCCFB05" blockId="8.[151,1436,1120,1173]" pageId="8" pageNumber="66">
<emphasis id="B96DEABB60761F4BFF3AF912AA2FFBE5" bold="true" box="[151,276,1120,1143]" pageId="8" pageNumber="66">FIGURE 8.</emphasis>
<taxonomicName id="4C194D2A60761F4BFEB1F912AAD6FBE6" box="[284,493,1120,1142]" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="species" species="jeekeli" status="sp. nov.">
<emphasis id="B96DEABB60761F4BFEB1F912AAD6FBE6" box="[284,493,1120,1142]" italics="true" pageId="8" pageNumber="66">Tonkinosoma jeekeli</emphasis>
</taxonomicName>
<emphasis id="B96DEABB60761F4BFE58F913A973FBE6" bold="true" box="[501,584,1121,1142]" pageId="8" pageNumber="66">
<taxonomicNameLabel id="A25E57C060761F4BFE58F913A973FBE6" box="[501,584,1121,1142]" pageId="8" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, left gonopod, mesal view (A), lateral view (B); Tip of gonopod, lateral view (C).
</paragraph>
</caption>
<paragraph id="8BA636A960761F4BFF68F9B3A943FA1D" blockId="8.[151,1436,1217,1925]" pageId="8" pageNumber="66">
Gonopods: (
<figureCitation id="13222A2C60761F4BFEE2F9B3AAA0FB49" box="[335,411,1217,1241]" captionStart="FIGURE 7" captionStartId="7.[151,250,1957,1980]" captionTargetBox="[300,1290,487,1914]" captionTargetId="figure@7.[264,1323,481,1936]" captionTargetPageId="7" captionText="FIGURE 7. Tonkinosoma jeekeli sp. nov., holotype, body ring 10 11, dorsal view (A); sternite V, ventral view (B); hypop-roct, ventral view (C); epiproct, ventral view (D); left gonopod, lateral view (E), mesal view (F). Scale bar = 1 mm." httpUri="https://zenodo.org/record/278753/files/figure.png" pageId="8" pageNumber="66">Figs. 7</figureCitation>
EF, 8AC) relatively simple. Coxite subcylindrical, slender and as long as femorite. Prefemur subrectangular, densely setose as usual, set off from femorite by a transversal suture (
<emphasis id="B96DEABB60761F4BFB0CF994AF80FB6E" bold="true" box="[1185,1211,1254,1278]" pageId="8" pageNumber="66">su</emphasis>
) laterally. Femorite (
<emphasis id="B96DEABB60761F4BFF0DF87BAB8DFAB2" bold="true" box="[160,182,1289,1314]" pageId="8" pageNumber="66">fe</emphasis>
) stout, expending in middle with a smaller distal part, without any femoral processes. Postfemoral region (= solenophore,
<emphasis id="B96DEABB60761F4BFE9FF85CAA67FAD6" bold="true" box="[306,348,1326,1350]" pageId="8" pageNumber="66">sph</emphasis>
) separated from femorite by an oblique sulcus laterally, curved down ventrally, and strongly twisted ventromesad. Both
<emphasis id="B96DEABB60761F4BFE65F823A940FAF9" box="[456,635,1361,1385]" italics="true" pageId="8" pageNumber="66">lamina medialis</emphasis>
(
<emphasis id="B96DEABB60761F4BFD26F820A991FAFA" bold="true" box="[651,682,1362,1386]" pageId="8" pageNumber="66">lm</emphasis>
) and
<emphasis id="B96DEABB60761F4BFD44F823A8A1FAF9" box="[745,922,1361,1385]" italics="true" pageId="8" pageNumber="66">lamina lateralis</emphasis>
(
<emphasis id="B96DEABB60761F4BFC07F820A881FAFA" bold="true" box="[938,954,1362,1386]" pageId="8" pageNumber="66">ll</emphasis>
) present. The former with a triangular lobe (
<emphasis id="B96DEABB60761F4BFF0DF804AB8AFA1E" bold="true" box="[160,177,1398,1422]" pageId="8" pageNumber="66">tl</emphasis>
) at distal part. Tip of solenophore acute.
</paragraph>
<paragraph id="8BA636A960761F4BFF68F8EBAA88FA45" blockId="8.[151,1436,1217,1925]" pageId="8" pageNumber="66">Seminal groove running entirely on mesal side before entering flagiliform solenomere, which is almost sheathed by solenophore.</paragraph>
</subSubSection>
<subSubSection id="C303652260761F4AFF68F890AAABFA1B" lastPageId="9" lastPageNumber="67" pageId="8" pageNumber="66" type="discussion">
<paragraph id="8BA636A960761F4BFF68F890AE6FF98D" blockId="8.[151,1436,1217,1925]" pageId="8" pageNumber="66">
<emphasis id="B96DEABB60761F4BFF68F890AA02FA6A" bold="true" box="[197,313,1506,1530]" pageId="8" pageNumber="66">Remarks.</emphasis>
Jeekel (1953) proposed a monotypic genus
<taxonomicName id="4C194D2A60761F4BFC95F893A8F7FA69" box="[824,972,1505,1529]" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABB60761F4BFC95F893A8F7FA69" box="[824,972,1505,1529]" italics="true" pageId="8" pageNumber="66">Tonkinosoma</emphasis>
</taxonomicName>
for a new species
<taxonomicName id="4C194D2A60761F4BFB05F893AA1EF98D" authority="Jeekel, 1953" authorityName="Jeekel" authorityYear="1953" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="species" species="flexipes">
<emphasis id="B96DEABB60761F4BFB05F893AEA7FA69" box="[1192,1436,1505,1529]" italics="true" pageId="8" pageNumber="66">Tonkinosoma flexipes</emphasis>
Jeekel, 1953
</taxonomicName>
, from Mauson Mt. (Lang Son province). The generic diagnosis was given to its species as well.
</paragraph>
<paragraph id="8BA636A960761F4BFF68FB5BA9B5F8AD" blockId="8.[151,1436,1217,1925]" pageId="8" pageNumber="66">
<bibRefCitation id="EF884B5860761F4BFF68FB5BAABAF9D1" author="Hoffman" box="[197,385,1577,1601]" pageId="8" pageNumber="66" refString="Hoffman, R. L. (1961) Two new diplopod genera from western China (Polydesmida: Strongylosomatidae). Annals and Magazine of Natural History, {13} 3 (33), 533 - 543." type="journal article" year="1961">Hoffman (1961)</bibRefCitation>
, when he created a new genus
<taxonomicName id="4C194D2A60761F4BFD43FB5BA8B9F9D1" box="[750,898,1577,1601]" class="Diplopoda" family="Paradoxosomatidae" genus="Szechuanella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABB60761F4BFD43FB5BA8B9F9D1" box="[750,898,1577,1601]" italics="true" pageId="8" pageNumber="66">Szechuanella</emphasis>
</taxonomicName>
for a new species
<taxonomicName id="4C194D2A60761F4BFBCDFB58AFEFF9D1" box="[1120,1236,1577,1601]" class="Diplopoda" family="Paradoxosomatidae" genus="Sellanucheza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="species" species="tenebra">
<emphasis id="B96DEABB60761F4BFBCDFB58AFEFF9D1" box="[1120,1236,1577,1601]" italics="true" pageId="8" pageNumber="66">S. tenebra</emphasis>
</taxonomicName>
from
<collectingCountry id="F30E763960761F4BFAB3FB5BAE5FF9D1" box="[1310,1380,1577,1601]" name="China" pageId="8" pageNumber="66">China</collectingCountry>
, discussed and compared both genera,
<taxonomicName id="4C194D2A60761F4BFDB2FB3CA98FF9F6" box="[543,692,1614,1638]" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABB60761F4BFDB2FB3CA98FF9F6" box="[543,692,1614,1638]" italics="true" pageId="8" pageNumber="66">Tonkinosoma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D2A60761F4BFD47FB3CA845F9F6" box="[746,894,1614,1638]" class="Diplopoda" family="Paradoxosomatidae" genus="Szechuanella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABB60761F4BFD47FB3CA845F9F6" box="[746,894,1614,1638]" italics="true" pageId="8" pageNumber="66">Szechuanella</emphasis>
</taxonomicName>
. He suggested that
<taxonomicName id="4C194D2A60761F4BFBF7FB3CAFF0F9F6" box="[1114,1227,1614,1638]" class="Diplopoda" family="Paradoxosomatidae" genus="Sellanucheza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="species" species="tenebra">
<emphasis id="B96DEABB60761F4BFBF7FB3CAFF0F9F6" box="[1114,1227,1614,1638]" italics="true" pageId="8" pageNumber="66">S. tenebra</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D2A60761F4BFAAFFB3DAE49F9F6" box="[1282,1394,1614,1638]" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="species" species="flexipes">
<emphasis id="B96DEABB60761F4BFAAFFB3DAE49F9F6" box="[1282,1394,1614,1638]" italics="true" pageId="8" pageNumber="66">T. flexipes</emphasis>
</taxonomicName>
are very similar in gonopod structure except for the degree of curvature of the solenophore, a presence of pleurosternal carinae in
<taxonomicName id="4C194D2A60761F4BFEA0FBE4AA9AF93E" box="[269,417,1686,1710]" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABB60761F4BFEA0FBE4AA9AF93E" box="[269,417,1686,1710]" italics="true" pageId="8" pageNumber="66">Tonkinosoma</emphasis>
</taxonomicName>
, and also in small detail of the solenophore structure. Later
<bibRefCitation id="EF884B5860761F4BFBE4FBE7AE3BF93D" author="Hoffman" box="[1097,1280,1685,1709]" pageId="8" pageNumber="66" refString="Hoffman, R. L. (1963) A contribution to the knowledge of Asiatic Strongylosomoid Diplopoda (Polydesmida: Strongylosomatidae). Annals and Magazine of Natural History, {13} 5 (58), 577 - 593." type="journal article" year="1963">Hoffman (1963)</bibRefCitation>
placed a species
<taxonomicName id="4C194D2A60761F4BFF64FBCBA902F941" authority="Attems, 1953" authorityName="Attems" authorityYear="1953" box="[201,569,1721,1746]" class="Diplopoda" family="Paradoxosomatidae" genus="Nedyopus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="species" species="variatus">
<emphasis id="B96DEABB60761F4BFF64FBCBAAA0F941" box="[201,411,1721,1745]" italics="true" pageId="8" pageNumber="66">Nedyopus variatus</emphasis>
<bibRefCitation id="EF884B5860761F4BFE0FFBC8A902F941" author="Attems" box="[418,569,1721,1746]" pageId="8" pageNumber="66" refString="Attems, C. (1953) Myriopoden von Indochina. Expedition von Dr. C Dawydoff (1938 - 1939). Memoires du Museum National d'Histoire Naturelle, {N. S., Ser. A, Zool.}, 5 (3), 133 - 230." type="journal article" year="1953">Attems, 1953</bibRefCitation>
</taxonomicName>
into his genus
<taxonomicName id="4C194D2A60761F4BFD4FFBCBA84DF941" box="[738,886,1721,1745]" class="Diplopoda" family="Paradoxosomatidae" genus="Szechuanella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABB60761F4BFD4FFBCBA84DF941" box="[738,886,1721,1745]" italics="true" pageId="8" pageNumber="66">Szechuanella</emphasis>
</taxonomicName>
, and maintained his separation of the two genera
<taxonomicName id="4C194D2A60761F4BFF3AFBACAA10F966" box="[151,299,1758,1782]" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABB60761F4BFF3AFBACAA10F966" box="[151,299,1758,1782]" italics="true" pageId="8" pageNumber="66">Tonkinosoma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D2A60761F4BFECCFBACAACEF966" box="[353,501,1758,1782]" class="Diplopoda" family="Paradoxosomatidae" genus="Szechuanella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABB60761F4BFECCFBACAACEF966" box="[353,501,1758,1782]" italics="true" pageId="8" pageNumber="66">Szechuanella</emphasis>
</taxonomicName>
. However, based on fresh material, the two genera are obviously distinguished by a presence/absence of additional processes in the postfemoral region of the gonopod, femorite enlarged or not, and the presence of pleurosternal carinae as well.
</paragraph>
<paragraph id="8BA636A960761F4BFF68FA38AA65F815" blockId="8.[151,1436,1217,1925]" pageId="8" pageNumber="66">
In brief, the genus
<taxonomicName id="4C194D2A60761F4BFE36FA3BA9FFF8F1" authority="Jeekel, 1953" authorityName="Jeekel" authorityYear="1953" box="[411,708,1865,1889]" class="Diplopoda" family="Paradoxosomatidae" genus="Tonkinosoma" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABB60761F4BFE36FA3BA914F8F1" box="[411,559,1865,1889]" italics="true" pageId="8" pageNumber="66">Tonkinosoma</emphasis>
Jeekel, 1953
</taxonomicName>
currently comprises two species and is distribute in the north of
<collectingCountry id="F30E763960761F4BFF3AFA1FABCDF815" box="[151,246,1901,1925]" name="Vietnam" pageId="8" pageNumber="66">Vietnam</collectingCountry>
(Map 1).
</paragraph>
<paragraph id="8BA636A960771F4AFF3AF988AAABFA1B" blockId="9.[151,1436,1274,1419]" pageId="9" pageNumber="67">
<emphasis id="B96DEABB60771F4AFF3AF988ABD0FA80" bold="true" box="[151,235,1274,1296]" pageId="9" pageNumber="67">MAP 1.</emphasis>
Distribution of all Tonkinosomatini species. 1: Wushan, Szechuan Province,
<collectingCountry id="F30E763960771F4AFBA8F988AF7EFA80" box="[1029,1093,1274,1296]" name="China" pageId="9" pageNumber="67">China</collectingCountry>
; 2: Sapa, Lao Cai province,
<collectingCountry id="F30E763960771F4AFAC4F988ABFDFABF" name="Vietnam" pageId="9" pageNumber="67">Vietnam</collectingCountry>
; 3: Mau Son Mt., Lang Son Province,
<collectingCountry id="F30E763960771F4AFDFEF86BA997FABF" box="[595,684,1305,1327]" name="Vietnam" pageId="9" pageNumber="67">Vietnam</collectingCountry>
; 4: Xuan Son National Park (NP), Phu Tho Province,
<collectingCountry id="F30E763960771F4AFB7AF86BAE0BFABF" box="[1239,1328,1305,1327]" name="Vietnam" pageId="9" pageNumber="67">Vietnam</collectingCountry>
; 5: Cat Ba Island, Hai Phong Province,
<collectingCountry id="F30E763960771F4AFE6FF845A920FADD" box="[450,539,1335,1357]" name="Vietnam" pageId="9" pageNumber="67">Vietnam</collectingCountry>
; 6: Cuc Phuong NP, Ninh Binh Province,
<collectingCountry id="F30E763960771F4AFC7EF845AF17FADD" box="[979,1068,1335,1357]" name="Vietnam" pageId="9" pageNumber="67">Vietnam</collectingCountry>
; 7: Pu Mat NP, Nghe An Province,
<collectingCountry id="F30E763960771F4AFF3AF824ABCBFAFC" box="[151,240,1366,1388]" name="Vietnam" pageId="9" pageNumber="67">Vietnam</collectingCountry>
; 8: Huong Son, Ha Tinh Province,
<collectingCountry id="F30E763960771F4AFDFBF824A994FAFC" box="[598,687,1366,1388]" name="Vietnam" pageId="9" pageNumber="67">Vietnam</collectingCountry>
; 9: Phong Nha Ke Bang NP, Quang Binh Province; 10: Loxo pass, Kon Tum Province,
<collectingCountry id="F30E763960771F4AFE9EF807AAB7FA1B" box="[307,396,1397,1419]" name="Vietnam" pageId="9" pageNumber="67">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>