treatments-xml/data/03/B0/87/03B087BA6878BF79FF01FF0D11F7FAFE.xml
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<document id="CF065686008AEDF47838EF48EB8A330B" ID-DOI="10.11646/zootaxa.2598.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="10095059" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629698685068" checkinUser="felipe" docAuthor="Ruta, Rafał &amp; Yoshitomi, Hiroyuki" docDate="2010" docId="03B087BA6878BF79FF01FF0D11F7FAFE" docLanguage="en" docName="zt02598p080.pdf" docOrigin="Zootaxa 2598 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2598.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Exochomoscirtes klausnitzeri Ruta &amp; Yoshitomi 2010, sp. nov." docType="treatment" docUuid="618F2C8C-DC30-410B-9950-280E50519073" docUuidSource="ZooBank" docVersion="4" lastPageNumber="41" masterDocId="FF89FFC2685EBF51FF96FF9A130CFF80" masterDocTitle="Revision of the genus Exochomoscirtes Pic (Coleoptera: Scirtidae: Scirtinae) 2598" masterLastPageNumber="80" masterPageNumber="1" pageNumber="39" updateTime="1699569420699" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9B409C3608D6D90CC86C893D3B4CFD6A">Revision of the genus Exochomoscirtes Pic (Coleoptera: Scirtidae: Scirtinae) 2598</mods:title>
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<mods:namePart id="2BD2A5C60F2DE6DBBAF222429090D5FD">Ruta, Rafał</mods:namePart>
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<mods:namePart id="94D18CDED3847DAEA9EE64926293B5D8">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:date id="FBA341431C52C34EB67B6284838CE681">2010</mods:date>
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<mods:classification id="B1250815070FC6B79F5671466F26FD48">journal article</mods:classification>
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<treatment id="03B087BA6878BF79FF01FF0D11F7FAFE" LSID="urn:lsid:zoobank.org:act:618F2C8C-DC30-410B-9950-280E50519073" httpUri="http://treatment.plazi.org/id/03B087BA6878BF79FF01FF0D11F7FAFE" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
<subSubSection id="C30365276878BF77FF01FF0D1153FF31" box="[151,607,151,177]" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="8BA636AC6878BF77FF01FF0D1153FF31" blockId="38.[151,607,151,212]" box="[151,607,151,177]" pageId="38" pageNumber="39">
<heading id="D0EE81C06878BF77FF01FF0D1153FF31" bold="true" box="[151,607,151,177]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
<emphasis id="B96DEABE6878BF77FF01FF0D1153FF31" bold="true" box="[151,607,151,177]" pageId="38" pageNumber="39">
<taxonomicName id="4C194D2F6878BF77FF01FF0D12F7FF31" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[151,507,151,177]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="klausnitzeri" status="sp. nov.">
<emphasis id="B96DEABE6878BF77FF01FF0D12F7FF31" bold="true" box="[151,507,151,177]" italics="true" pageId="38" pageNumber="39">Exochomoscirtes klausnitzeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56878BF77FD94FF0D1153FF31" box="[514,607,151,177]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365276878BF77FF01FF20110CFF54" box="[151,512,186,212]" pageId="38" pageNumber="39" type="description">
<paragraph id="8BA636AC6878BF77FF01FF20110CFF54" blockId="38.[151,607,151,212]" box="[151,512,186,212]" pageId="38" pageNumber="39">
(
<figureCitation id="13222A296878BF77FF09FF201204FF54" box="[159,264,186,212]" captionStart="FIGURE 5" captionStartId="9.[151,255,1360,1384]" captionTargetBox="[167,1421,160,1344]" captionTargetId="figure-294@9.[151,1436,145,1364]" captionTargetPageId="9" captionText="FIGURE 5. Abdominal details in Exochomoscirtes. A) E. dilaticornis sp. nov., ventrite 5; B) E. discoidalis (Pic), ventrite 5; C) E. jaechi sp. nov., tergite 7; D) E. klausnitzeri sp. nov., tergite 7; E) E. luteosuturalis sp. nov., tergite 7; F) E. luteosuturaloides sp. nov., tergite 7; G) E. maculatus (Waterhouse), abdomen; H) E. maculatus (Waterhouse), ventrites 35; I) E. nigromarginatus sp. nov., ventrite 5; J) E. quadripartitus sp. nov., tergite 7.; K) E. ruforotundus (Watts) (Australia), tergite 7; L) E. takizawai sp. nov., ventrite 5." figureDoi="http://doi.org/10.5281/zenodo.10095069" httpUri="https://zenodo.org/record/10095069/files/figure.png" pageId="38" pageNumber="39">Figs. 5D</figureCitation>
,
<figureCitation id="13222A296878BF77FE82FF201229FF54" box="[276,293,186,212]" captionStart="FIGURE 7" captionStartId="11.[151,255,1901,1925]" captionText="FIGURE 7. Exochomoscirtes klausnitzeri sp. nov., larva." pageId="38" pageNumber="39">7</figureCitation>
,
<figureCitation id="13222A296878BF77FEA7FF20124DFF54" box="[305,321,186,212]" captionStart="FIGURE 8" captionStartId="13.[151,255,1705,1729]" captionTargetBox="[159,1424,192,1667]" captionTargetId="figure-66@13.[151,1436,167,1679]" captionTargetPageId="13" captionText="FIGURE 8. Exochomoscirtes klausnitzeri sp. nov., details of larval morphology. A) habitus; B) maxillary palpus, dorsal view; C) maxillary palpus, lateral view; D) mandible; E) hypopharynx; F) labrum; G) epipharynx; H) foreleg; I) tergite 8; J) tergite 9; K) sternite 8; L) sternite 9. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10095073" httpUri="https://zenodo.org/record/10095073/files/figure.png" pageId="38" pageNumber="39">8</figureCitation>
,
<figureCitation id="13222A296878BF77FED9FF2012ADFF54" box="[335,417,186,212]" captionStart="FIGURE 12" captionStartId="18.[151,255,1861,1885]" captionTargetBox="[164,1428,210,1822]" captionTargetId="figure-109@18.[151,1436,197,1835]" captionTargetPageId="18" captionText="FIGURE 12. Habitus illustrations of Exochomoscirtes. A) E. maculatus (Waterhouse); B)C) E. ceylonicus sp. nov.; D) E. decemguttatus (Champion); E)G) E. klausnitzeri sp. nov., color forms F S Thailand, male, E and G Sarawak, females; H) E. cf. klausnitzeri (NW Thailand); I) E. luteosuturalis sp. nov.; J) E. cf. luteosuturalis sp. nov., female, Sulawesi; K) E. cf. luteosuturalis, female, Vietnam; L) E. luteosuturaloides sp. nov.; M) E. jaechi sp. nov.; N) E. ruforotundus (Watts); O) E. ruforotundus (Watts), Bali. Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10095081" httpUri="https://zenodo.org/record/10095081/files/figure.png" pageId="38" pageNumber="39">12EG</figureCitation>
,
<figureCitation id="13222A296878BF77FE3BFF2012C1FF54" box="[429,461,186,212]" captionStart="FIGURE 30" captionStartId="38.[151,255,1828,1852]" captionTargetBox="[170,1429,1159,1786]" captionTargetId="figure-414@38.[151,1436,1151,1801]" captionTargetPageId="38" captionText="FIGURE 30. Exochomoscirtes klausnitzeri sp. nov., male. A) sternite 9; B) tergite 8; C) tergite 8, specimen from NE India; D) tergite 9; E) tegmen; F) penis. Scale bar = 0.5 mm (AB, DF), 1.0 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10095119" httpUri="https://zenodo.org/record/10095119/files/figure.png" pageId="38" pageNumber="39">30</figureCitation>
,
<figureCitation id="13222A296878BF77FE4FFF2012FBFF54" box="[473,503,186,212]" captionStart="FIGURE 31" captionStartId="40.[151,255,945,969]" captionTargetBox="[158,1428,197,919]" captionTargetId="figure-358@40.[158,1428,197,919]" captionTargetPageId="40" captionText="FIGURE 31. Exochomoscirtes klausnitzeri sp. nov., female. A) bursal sclerite; B) prehensor and bursal sclerite." figureDoi="http://doi.org/10.5281/zenodo.10095121" httpUri="https://zenodo.org/record/10095121/files/figure.png" pageId="38" pageNumber="39">31</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365276878BF77FF01FE9011A4FC4C" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="8BA636AC6878BF77FF01FE901690FEA4" blockId="38.[151,1436,266,1092]" box="[151,1436,266,292]" pageId="38" pageNumber="39">
<emphasis id="B96DEABE6878BF77FF01FE901241FEA4" bold="true" box="[151,333,266,292]" pageId="38" pageNumber="39">Type material.</emphasis>
<materialsCitation id="3B713CF16878BF77FEC3FE901690FEA4" box="[341,1436,266,292]" collectingDate="1992-04-28" collectionCode="NMPC" collectorName="J. Horak" country="Thailand" location="Thailand" pageId="38" pageNumber="39" specimenCount="1" typeStatus="holotype">
<emphasis id="B96DEABE6878BF77FEC3FE901183FEA4" bold="true" box="[341,655,266,292]" pageId="38" pageNumber="39">
<typeStatus id="54A2880E6878BF77FEC3FE9012C5FEA4" box="[341,457,266,292]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="ED08AE696878BF77FDB4FE90118DFEA4" box="[546,641,266,292]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="38" pageNumber="39">NMPC</collectionCode>
):
</emphasis>
“S
<collectingCountry id="F30E763C6878BF77FD2AFE90102BFEA4" box="[700,807,266,292]" name="Thailand" pageId="38" pageNumber="39">Thailand</collectingCountry>
,
<date id="FFA7106C6878BF77FCA5FE9010A5FEA4" box="[819,937,266,292]" pageId="38" pageNumber="39" value="1992-04-28">
<collectingDate id="EFE3E9846878BF77FCA5FE9010A5FEA4" box="[819,937,266,292]" pageId="38" pageNumber="39" value="1992-04-28">28.4.1992</collectingDate>
</date>
/ BETONG /
<collectorName id="26EC537A6878BF77FBD8FE9017A6FEA4" box="[1102,1194,266,292]" pageId="38" pageNumber="39">J.Horák</collectorName>
lgt.” [printed label].
</materialsCitation>
</paragraph>
<paragraph id="8BA636AC6878BF77FF01FEA81292FCAC" blockId="38.[151,1436,266,1092]" pageId="38" pageNumber="39">
<materialsCitation id="3B713CF16878BF77FF01FEA81296FCAC" collectingDate="1991-12-02" collectingDateMax="2003-11-21" collectingDateMin="1991-12-02" collectionCode="MNHN, NMW, HNHM, MNHG, EUM" collectorName="Horak &amp; A. Orosz &amp; Gy. Sziraki &amp; P. Schwendinger &amp; O. Merkl &amp; G. Csorba &amp; J. Kodada &amp; Kodada &amp; C. H. S. Watts &amp; Jendek" country="Thailand" county="Male" elevation="50" location="Male" municipality="Female" pageId="38" pageNumber="39" specimenCount="283" specimenCount-male="3" stateProvince="Sarawak" typeStatus="paratype">
<emphasis id="B96DEABE6878BF77FF01FEA81217FECC" bold="true" box="[151,283,306,332]" pageId="38" pageNumber="39">
<typeStatus id="54A2880E6878BF77FF01FEA8121BFECC" box="[151,279,306,332]" pageId="38" pageNumber="39" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCounty id="62C74E206878BF77FEB3FEA8126DFECC" box="[293,353,306,332]" pageId="38" pageNumber="39">Male</collectingCounty>
(
<collectionCode id="ED08AE696878BF77FEE5FEA812D8FECC" box="[371,468,306,332]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="38" pageNumber="39">MNHN</collectionCode>
): “MUSEUM PARIS / Sumatra / Palembang / H. DONCKIER 1901” [printedlabel]; “Palembang / Sumatra” [printed label].
<emphasis id="B96DEABE6878BF77FD3FFEC0107AFEF4" bold="true" box="[681,886,346,372]" pageId="38" pageNumber="39">
<collectingMunicipality id="6BC2ACD66878BF77FD3FFEC01008FEF4" box="[681,772,346,372]" pageId="38" pageNumber="39">Female</collectingMunicipality>
(
<collectionCode id="ED08AE696878BF77FC82FEC01065FEF4" box="[788,873,346,372]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="38" pageNumber="39">NMW</collectionCode>
):
</emphasis>
“S-THAIL., Betong 1993 / Gunung Cang dun vill / Yala distr., 25.3.-22.4. / leg.
<collectorName id="26EC537A6878BF77FD80FE18116FFE1C" box="[534,611,386,412]" pageId="38" pageNumber="39">Horak</collectorName>
&amp; Strnad” [printed label].
<emphasis id="B96DEABE6878BF77FC08FE18176CFE1C" bold="true" box="[926,1120,386,412]" pageId="38" pageNumber="39">
<location id="8EC660776878BF77FC08FE1810D1FE1C" LSID="urn:lsid:plazi:treatment:03B087BA6878BF79FF01FF0D11F7FAFE:8EC660776878BF77FC08FE1810D1FE1C" box="[926,989,386,412]" country="Thailand" county="Male" municipality="Female" name="Male" pageId="38" pageNumber="39" stateProvince="Sarawak">Male</location>
(
<collectionCode id="ED08AE696878BF77FC7BFE18175FFE1C" box="[1005,1107,386,412]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="38" pageNumber="39">HNHM</collectionCode>
):
</emphasis>
<collectingCountry id="F30E763C6878BF77FBE4FE181603FE1C" box="[1138,1295,386,412]" name="Thailand" pageId="38" pageNumber="39">THAILAND</collectingCountry>
, pr. Trang. / Kao Chong / Botanical Garden.” [printed label]; “at light at staff center, /
<date id="FFA7106C6878BF77FB9FFE30179DFE44" box="[1033,1169,426,452]" pageId="38" pageNumber="39" value="2003-11-21">
<collectingDate id="EFE3E9846878BF77FB9FFE30179DFE44" box="[1033,1169,426,452]" pageId="38" pageNumber="39" value="2003-11-21">21.XI.2003</collectingDate>
</date>
, No. 13, /
<collectorName id="26EC537A6878BF77FA87FE301671FE44" box="[1297,1405,426,452]" pageId="38" pageNumber="39">A. Orosz</collectorName>
&amp;
<collectorName id="26EC537A6878BF77FF01FE48122DFE6C" box="[151,289,466,492]" pageId="38" pageNumber="39">Gy. Sziráki</collectorName>
” [printed label].
<emphasis id="B96DEABE6878BF77FE67FE4811DEFE6C" bold="true" box="[497,722,466,492]" pageId="38" pageNumber="39">
<location id="8EC660776878BF77FE67FE481140FE6C" LSID="urn:lsid:plazi:treatment:03B087BA6878BF79FF01FF0D11F7FAFE:8EC660776878BF77FE67FE481140FE6C" box="[497,588,466,492]" country="Thailand" county="Male" municipality="Female" name="Female" pageId="38" pageNumber="39" stateProvince="Sarawak">Female</location>
(
<collectionCode id="ED08AE696878BF77FDC9FE4811C9FE6C" box="[607,709,466,492]" pageId="38" pageNumber="39">MNHG</collectionCode>
):
</emphasis>
“Thailand,
<date id="FFA7106C6878BF77FCF5FE4810D5FE6C" box="[867,985,466,492]" pageId="38" pageNumber="39" value="1991-12-02">
<collectingDate id="EFE3E9846878BF77FCF5FE4810D5FE6C" box="[867,985,466,492]" pageId="38" pageNumber="39" value="1991-12-02">12.2.1991</collectingDate>
</date>
/ Huay Yang Nat. Park / Huay Yang Waterfall / Thap Sake Dist.
<quantity id="4CE19B496878BF77FE72FE601114FD94" box="[484,536,506,532]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="38" pageNumber="39" unit="m" value="50.0">
<elevation id="0034D19F6878BF77FE72FE601114FD94" box="[484,536,506,532]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="38" pageNumber="39" unit="m" value="50.0">50m</elevation>
</quantity>
/
<collectorName id="26EC537A6878BF77FDA4FE6011F1FD94" box="[562,765,506,532]" pageId="38" pageNumber="39">P. Schwendinger</collectorName>
”.
<emphasis id="B96DEABE6878BF77FC82FE6010FFFD94" bold="true" box="[788,1011,506,532]" pageId="38" pageNumber="39">
<location id="8EC660776878BF77FC82FE601063FD94" LSID="urn:lsid:plazi:treatment:03B087BA6878BF79FF01FF0D11F7FAFE:8EC660776878BF77FC82FE601063FD94" box="[788,879,506,532]" country="Thailand" county="Male" municipality="Female" name="Female" pageId="38" pageNumber="39" stateProvince="Sarawak">Female</location>
(
<collectionCode id="ED08AE696878BF77FC16FE6010EAFD94" box="[896,998,506,532]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="38" pageNumber="39">HNHM</collectionCode>
):
</emphasis>
<collectingCountry id="F30E763C6878BF77FB91FE601755FD94" box="[1031,1113,506,532]" name="Laos" pageId="38" pageNumber="39">LAOS</collectingCountry>
, Bolikhamsay Prov. / Phou Khao Kouay NBCA / Tad Leuk Waterfall,
<specimenCount id="9D1FFD256878BF77FD04FDB811D3FDBC" box="[658,735,546,572]" pageId="38" pageNumber="39" type="generic">
<quantity id="4CE19B496878BF77FD04FDB811D7FDBC" box="[658,731,546,572]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="38" pageNumber="39" unit="m" value="280.0">
<elevation id="0034D19F6878BF77FD04FDB811D7FDBC" box="[658,731,546,572]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="38" pageNumber="39" unit="m" value="280.0">280m</elevation>
</quantity>
.
</specimenCount>
” [printed label]; “at light, No.46, /
<date id="FFA7106C6878BF77FB16FDB8163CFDBC" box="[1152,1328,546,572]" pageId="38" pageNumber="39" value="1998-04-11" valueMax="1998-04-12" valueMin="1998-04-11">
<collectingDate id="EFE3E9846878BF77FB16FDB8163CFDBC" box="[1152,1328,546,572]" pageId="38" pageNumber="39" value="1998-04-11" valueMax="1998-04-12" valueMin="1998-04-11">1112.IV.1998</collectingDate>
</date>
, / leg.
<collectorName id="26EC537A6878BF77FA16FDB813D3FDE4" pageId="38" pageNumber="39">O. Merkl</collectorName>
&amp;
<collectorName id="26EC537A6878BF77FE99FDD01287FDE4" box="[271,395,586,612]" pageId="38" pageNumber="39">G. Csorba</collectorName>
” [printed label].
<emphasis id="B96DEABE6878BF77FDF4FDD01039FDE4" bold="true" box="[610,821,586,612]" pageId="38" pageNumber="39">
<location id="8EC660776878BF77FDF4FDD011B1FDE4" LSID="urn:lsid:plazi:treatment:03B087BA6878BF79FF01FF0D11F7FAFE:8EC660776878BF77FDF4FDD011B1FDE4" box="[610,701,586,612]" country="Thailand" county="Male" municipality="Female" name="Female" pageId="38" pageNumber="39" stateProvince="Sarawak">Female</location>
(
<collectionCode id="ED08AE696878BF77FD45FDD01024FDE4" box="[723,808,586,612]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="38" pageNumber="39">NMW</collectionCode>
):
</emphasis>
“SARAWAK (Borneo), / cz
<quantity id="4CE19B496878BF77FB33FDD017F8FDE4" box="[1189,1268,586,612]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="38" pageNumber="39" unit="km" value="40.0">40 km</quantity>
SE KAPIT / 03.1994,
<collectorName id="26EC537A6878BF77FE92FDE81275FD0C" box="[260,377,626,652]" pageId="38" pageNumber="39">J. Kodada</collectorName>
leg.” [printed label].
<emphasis id="B96DEABE6878BF77FDE5FDE8104CFD0C" bold="true" box="[627,832,626,652]" pageId="38" pageNumber="39">
<location id="8EC660776878BF77FDE5FDE811C2FD0C" LSID="urn:lsid:plazi:treatment:03B087BA6878BF79FF01FF0D11F7FAFE:8EC660776878BF77FDE5FDE811C2FD0C" box="[627,718,626,652]" country="Thailand" county="Male" municipality="Female" name="Female" pageId="38" pageNumber="39" stateProvince="Sarawak">Female</location>
(
<collectionCode id="ED08AE696878BF77FD4BFDE8103EFD0C" box="[733,818,626,652]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="38" pageNumber="39">NMW</collectionCode>
):
</emphasis>
<collectingCountry id="F30E763C6878BF77FCC7FDE810FCFD0C" box="[849,1008,626,652]" name="Malaysia" pageId="38" pageNumber="39">MALAYSIA</collectingCountry>
:
<collectingRegion id="49DDF84E6878BF77FC6DFDE8176CFD0C" box="[1019,1120,626,652]" country="Malaysia" name="Sarawak" pageId="38" pageNumber="39">Sarawak</collectingRegion>
/
<locationDeviation id="9A142F8E6878BF77FBE0FDE8162CFD0C" box="[1142,1312,626,652]" location="Kapit" pageId="38" pageNumber="39">
<quantity id="4CE19B496878BF77FBE0FDE817B3FD0C" box="[1142,1215,626,652]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="38" pageNumber="39" unit="km" value="25.0">25 km</quantity>
E Kapit
</locationDeviation>
/
<date id="FFA7106C6878BF77FAA0FDE81697FD0C" box="[1334,1435,626,652]" pageId="38" pageNumber="39" value="1994-03">
<collectingDate id="EFE3E9846878BF77FAA0FDE81697FD0C" box="[1334,1435,626,652]" pageId="38" pageNumber="39" value="1994-03">III. 1994</collectingDate>
</date>
/ leg.
<collectorName id="26EC537A6878BF77FF49FD001231FD34" box="[223,317,666,692]" pageId="38" pageNumber="39">Kodada</collectorName>
” [printed label].
<emphasis id="B96DEABE6878BF77FD9AFD0011D5FD34" bold="true" box="[524,729,666,692]" pageId="38" pageNumber="39">
<specimenCount id="9D1FFD256878BF77FD9AFD001160FD34" box="[524,620,666,692]" pageId="38" pageNumber="39" type="male">2 males</specimenCount>
(
<collectionCode id="ED08AE696878BF77FD16FD0011C7FD34" box="[640,715,666,692]" pageId="38" pageNumber="39">EUM</collectionCode>
):
</emphasis>
“Kebun Raya, / Bogor, Indonesia /
<collectorName id="26EC537A6878BF77FB07FD00164EFD34" box="[1169,1346,666,692]" pageId="38" pageNumber="39">C. H. S. Watts</collectorName>
/
<date id="FFA7106C6878BF77FAC8FD0013D8FD5C" pageId="38" pageNumber="39" value="2000-04">
<collectingDate id="EFE3E9846878BF77FAC8FD0013D8FD5C" pageId="38" pageNumber="39" value="2000-04">April 2000</collectingDate>
</date>
” [printed label],
<specimenCount id="9D1FFD256878BF77FE36FD581100FD5C" box="[416,524,706,732]" pageId="38" pageNumber="39" type="male">one male</specimenCount>
genitalia on slide no. HY 896.
<emphasis id="B96DEABE6878BF77FC10FD581735FD5C" bold="true" box="[902,1081,706,732]" pageId="38" pageNumber="39">
Male (
<collectionCode id="ED08AE696878BF77FC40FD581727FD5C" box="[982,1067,706,732]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="38" pageNumber="39">NMW</collectionCode>
):
</emphasis>
“NE-INDIA: Meghalaya / W Garo Hills / Balphakram NP” [printed label]; “25°11N, 90°51E /
<quantity id="4CE19B496878BF77FC35FD70172BFC84" box="[931,1063,746,772]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="38" pageNumber="39" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">
<elevation id="0034D19F6878BF77FC35FD70172BFC84" box="[931,1063,746,772]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="38" pageNumber="39" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">300500 m</elevation>
</quantity>
/ 22.
<date id="FFA7106C6878BF77FBFBFD7017EAFC84" box="[1133,1254,746,772]" pageId="38" pageNumber="39" value="1996-05-27">
<collectingDate id="EFE3E9846878BF77FBFBFD7017EAFC84" box="[1133,1254,746,772]" pageId="38" pageNumber="39" value="1996-05-27">27.5.1996</collectingDate>
</date>
/ leg.
<collectorName id="26EC537A6878BF77FABBFD701672FC84" box="[1325,1406,746,772]" pageId="38" pageNumber="39">Jendek</collectorName>
&amp; Sausa” [printed label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636AC6878BF77FF50FCA01686FCFC" blockId="38.[151,1436,266,1092]" pageId="38" pageNumber="39">
<emphasis id="B96DEABE6878BF77FF50FCA01132FCD4" bold="true" box="[198,574,826,852]" pageId="38" pageNumber="39">Additional specimens studied.</emphasis>
<materialsCitation id="3B713CF16878BF77FDD1FCA0168AFCFC" collectingDate="1996-05-27" collectionCode="NMW" collectorName="Jendek" elevation="400" location="Meghalaya" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" stateProvince="Meghalaya">
<emphasis id="B96DEABE6878BF77FDD1FCA01001FCD4" bold="true" box="[583,781,826,852]" pageId="38" pageNumber="39">
<specimenCount id="9D1FFD256878BF77FDD1FCA01196FCD4" box="[583,666,826,852]" pageId="38" pageNumber="39" type="male">1 male</specimenCount>
(
<collectionCode id="ED08AE696878BF77FD3DFCA0100CFCD4" box="[683,768,826,852]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="38" pageNumber="39">NMW</collectionCode>
):
</emphasis>
“NE-INDIA:
<collectingRegion id="49DDF84E6878BF77FC2EFCA01736FCD4" box="[952,1082,826,852]" country="India" name="Meghalaya" pageId="38" pageNumber="39">Meghalaya</collectingRegion>
/ W Garo Hills / Balphakram NP” [printed label]; “25°11N, 90°51E /
<quantity id="4CE19B496878BF77FDE9FCF81008FCFC" box="[639,772,866,892]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="38" pageNumber="39" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">
<elevation id="0034D19F6878BF77FDE9FCF81008FCFC" box="[639,772,866,892]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="38" pageNumber="39" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">300500 m</elevation>
</quantity>
/ 22.
<date id="FFA7106C6878BF77FCDDFCF810C8FCFC" box="[843,964,866,892]" pageId="38" pageNumber="39" value="1996-05-27">
<collectingDate id="EFE3E9846878BF77FCDDFCF810C8FCFC" box="[843,964,866,892]" pageId="38" pageNumber="39" value="1996-05-27">27.5.1996</collectingDate>
</date>
/ leg.
<collectorName id="26EC537A6878BF77FB9BFCF81752FCFC" box="[1037,1118,866,892]" pageId="38" pageNumber="39">Jendek</collectorName>
&amp; Sausa” [printed label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636AC6878BF77FF50FC1011A4FC4C" blockId="38.[151,1436,266,1092]" pageId="38" pageNumber="39">
<emphasis id="B96DEABE6878BF77FF50FC10111EFC24" bold="true" box="[198,530,906,932]" pageId="38" pageNumber="39">Larval specimens studied.</emphasis>
22 exx. (see remarks below), Kebun Raya, Bogor, Ind. C. H. Watts
<date id="FFA7106C6878BF77FAC8FC1013D9FC4C" pageId="38" pageNumber="39" value="2000-04">April 2000</date>
;
<specimenCount id="9D1FFD256878BF77FF77FC28123DFC4C" box="[225,305,946,972]" pageId="38" pageNumber="39" type="larva">1 larva</specimenCount>
mounted on slide no. HY 1125.
</paragraph>
</subSubSection>
<subSubSection id="C30365276878BF77FF50FC4010BEFBC4" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="8BA636AC6878BF77FF50FC4010BEFBC4" blockId="38.[151,1436,266,1092]" pageId="38" pageNumber="39">
<emphasis id="B96DEABE6878BF77FF50FC40124AFC74" bold="true" box="[198,326,986,1012]" pageId="38" pageNumber="39">Diagnosis.</emphasis>
The species is readily distinguishable thanks to the peculiar dorsal pattern (
<figureCitation id="13222A296878BF77FB45FC40166CFC74" box="[1235,1376,986,1012]" captionStart="FIGURE 12" captionStartId="18.[151,255,1861,1885]" captionTargetBox="[164,1428,210,1822]" captionTargetId="figure-109@18.[151,1436,197,1835]" captionTargetPageId="18" captionText="FIGURE 12. Habitus illustrations of Exochomoscirtes. A) E. maculatus (Waterhouse); B)C) E. ceylonicus sp. nov.; D) E. decemguttatus (Champion); E)G) E. klausnitzeri sp. nov., color forms F S Thailand, male, E and G Sarawak, females; H) E. cf. klausnitzeri (NW Thailand); I) E. luteosuturalis sp. nov.; J) E. cf. luteosuturalis sp. nov., female, Sulawesi; K) E. cf. luteosuturalis, female, Vietnam; L) E. luteosuturaloides sp. nov.; M) E. jaechi sp. nov.; N) E. ruforotundus (Watts); O) E. ruforotundus (Watts), Bali. Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10095081" httpUri="https://zenodo.org/record/10095081/files/figure.png" pageId="38" pageNumber="39">Fig. 12EG</figureCitation>
) and unmodified metatibial spurs. Genitalia are similar to that of
<taxonomicName id="4C194D2F6878BF77FC10FB981796FB9C" authority="(Champion)" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" baseAuthorityName="Champion" baseAuthorityYear="1918" box="[902,1178,1026,1052]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="retusus">
<emphasis id="B96DEABE6878BF77FC10FB9810F3FB9B" box="[902,1023,1026,1051]" italics="true" pageId="38" pageNumber="39">E. retusus</emphasis>
(Champion)
</taxonomicName>
and
<taxonomicName id="4C194D2F6878BF77FB48FB9813E3FBC4" authority="(Watts)" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" baseAuthorityName="Watts" baseAuthorityYear="2004" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="ruforotundus">
<emphasis id="B96DEABE6878BF77FB48FB981690FB9B" box="[1246,1436,1026,1051]" italics="true" pageId="38" pageNumber="39">E. ruforotundus</emphasis>
(Watts)
</taxonomicName>
, differing in the regularly rounded parameroids of the penis.
</paragraph>
</subSubSection>
<caption id="DF6666246878BF77FF01F8BE1723F8DE" ID-DOI="http://doi.org/10.5281/zenodo.10095119" ID-Zenodo-Dep="10095119" httpUri="https://zenodo.org/record/10095119/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,255,1828,1852]" targetBox="[170,1429,1159,1786]" targetPageId="38">
<paragraph id="8BA636AC6878BF77FF01F8BE1723F8DE" blockId="38.[151,1435,1828,1887]" pageId="38" pageNumber="39">
<emphasis id="B96DEABE6878BF77FF01F8BE1224F8BC" bold="true" box="[151,296,1828,1852]" pageId="38" pageNumber="39">FIGURE 30.</emphasis>
<taxonomicName id="4C194D2F6878BF77FEA7F8BF1161F8BC" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[305,621,1829,1852]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="klausnitzeri" status="sp. nov.">
<emphasis id="B96DEABE6878BF77FEA7F8BF1161F8BC" box="[305,621,1829,1852]" italics="true" pageId="38" pageNumber="39">Exochomoscirtes klausnitzeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56878BF77FDE3F8BE11C0F8BB" box="[629,716,1828,1851]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, male.
<emphasis id="B96DEABE6878BF77FC8DF8BE103BF8BC" bold="true" box="[795,823,1828,1852]" pageId="38" pageNumber="39">A)</emphasis>
sternite 9;
<emphasis id="B96DEABE6878BF77FC22F8BE10C3F8BC" bold="true" box="[948,975,1828,1852]" pageId="38" pageNumber="39">B)</emphasis>
tergite 8;
<emphasis id="B96DEABE6878BF77FBD7F8BE1751F8BC" bold="true" box="[1089,1117,1828,1852]" pageId="38" pageNumber="39">C)</emphasis>
tergite 8, specimen from NE India;
<emphasis id="B96DEABE6878BF77FF4AF8DD13F4F8DF" bold="true" box="[220,248,1863,1887]" pageId="38" pageNumber="39">D)</emphasis>
tergite 9;
<emphasis id="B96DEABE6878BF77FEF2F8DD1273F8DF" bold="true" box="[356,383,1863,1887]" pageId="38" pageNumber="39">E)</emphasis>
tegmen;
<emphasis id="B96DEABE6878BF77FE74F8DD12F7F8DF" bold="true" box="[482,507,1863,1887]" pageId="38" pageNumber="39">F)</emphasis>
penis. Scale bar = 0.5 mm (AB, DF), 1.0 mm (C).
</paragraph>
</caption>
<subSubSection id="C30365276878BF76FF50F80B11B9F972" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="description">
<paragraph id="8BA636AC6878BF76FF50F80B13E5FEAA" blockId="38.[151,1436,1937,2004]" lastBlockId="39.[151,1436,152,2018]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<emphasis id="B96DEABE6878BF77FF50F80B1252F82B" bold="true" box="[198,350,1937,1963]" pageId="38" pageNumber="39">Description.</emphasis>
<typeStatus id="54A2880E6878BF77FEFFF80812D5F82C" box="[361,473,1938,1964]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
, male. Body oval, TL/EW 1.2, somewhat flattened, covered with suberect dark brown and yellow (on orange portions of elytra) hairs. Head brown; pronotum darker than head, dark brown; scutellum brown, elytra dark brown with five orange oval areas. Orange elytral maculation consists of a circular area in basal portion, extending to the basal margin of elytra, reaching neither suture, nor lateral margin, and large oval apical spot, half of which is situated on each elytron. Antennae and mouthparts brownish, legs brown.
</paragraph>
<paragraph id="8BA636AC6879BF76FF50FEA2103CFE4A" blockId="39.[151,1436,152,2018]" pageId="39" pageNumber="40">Head with distinct punctation, punctures separated by ca. 1.0 diameter. Eyes big, protuberant; head 1.9 X wider than interocular space. Antennae filiform, reaching basal 1/3 of elytra. Antennomeres 2 and 3 of subequal lengths, antennomeres 411 a bit shorter than antennomeres 2 and 3 combined, slightly dilated, subtriangular. Palpomeres of maxillary palpi moderately wide.</paragraph>
<paragraph id="8BA636AC6879BF76FF50FE4212B0FD12" blockId="39.[151,1436,152,2018]" pageId="39" pageNumber="40">Pronotum slightly convex, covered with distinct punctation, punctures separated by 1.02.0 diameters, anterolateral corners somewhat obtuse, projecting anteriorly; lateral margins straight; posterolateral corners acute; base of pronotum subtrapezoidal, with complete margination along basal margin; PW/PL 3.0. Scutellum relatively big, punctured as on pronotum, equilaterally triangular. Angle between pronotum and elytra not marked in dorsal outline.</paragraph>
<paragraph id="8BA636AC6879BF76FF50FD3A17A7FC8A" blockId="39.[151,1436,152,2018]" pageId="39" pageNumber="40">Elytra oval, more strongly converging in posterior half, than in anterior portion, with slightly explanate lateral margins, widest slightly before the middle, punctation stronger and denser than on pronotum, punctures separated by ca. 1.0 diameter; humeri well marked; EL/EW 1.1; EL/PL 4.5; EW/PW 1.4.</paragraph>
<paragraph id="8BA636AC6879BF76FF50FC821744FCDA" blockId="39.[151,1436,152,2018]" pageId="39" pageNumber="40">Legs moderately long. Hind tibial spurs well developed, dorsal one elongated, as long as tarsomere 1 and about 2 X longer than ventral one, almost straight; ventral one is slightly curved.</paragraph>
<paragraph id="8BA636AC6879BF76FF50FCF2168AFC2A" blockId="39.[151,1436,152,2018]" pageId="39" pageNumber="40">Ventrites 1 and 2 with sparse setation in mesal portions, ventrite 2 with basal row of a few big punctures. Apex of tergite 7 with a very narrow rectangular outgrowth. Apex of ventrite 5 very shallowly emarginated.</paragraph>
<paragraph id="8BA636AC6879BF76FF50FC221726FB6A" blockId="39.[151,1436,152,2018]" pageId="39" pageNumber="40">
Sternite 9 (L 0.55,
<geoCoordinate id="EE2D506B6879BF76FE37FC2212F4FC52" box="[417,504,952,978]" degrees="0.22" direction="west" orientation="longitude" pageId="39" pageNumber="40" precision="555" value="-0.22">W 0.22</geoCoordinate>
) oblong oval, moderately sclerotized, bearing few apical setae, deeply emarginated at apex. Tergite 8 (L 0.75,
<geoCoordinate id="EE2D506B6879BF76FD99FC7A1164FC7A" box="[527,616,992,1018]" degrees="0.40" direction="west" orientation="longitude" pageId="39" pageNumber="40" precision="555" value="-0.4">W 0.40</geoCoordinate>
) large, moderately sclerotized, apical portion subtrapezoidal, bearing row of relatively long setae on apical margin, apodemes moderately long, slightly diverging basally; tergite 9 (L 0.45,
<geoCoordinate id="EE2D506B6879BF76FE96FBAA1256FBCA" box="[256,346,1072,1098]" degrees="0.24" direction="west" orientation="longitude" pageId="39" pageNumber="40" precision="555" value="-0.24">W 0.24</geoCoordinate>
) moderately sclerotized, apical portion of moderate size, with a row of short setae at apex, with a pair of short subparallel apodemes. Tegmen (L 0.45,
<geoCoordinate id="EE2D506B6879BF76FCEDFBC210DAFBF2" box="[891,982,1112,1138]" degrees="0.20" direction="west" orientation="longitude" pageId="39" pageNumber="40" precision="555" value="-0.2">W 0.20</geoCoordinate>
) well sclerotized, parameres straight, slightly widened at apices, each paramere in the middle with a small denticle pointing inwards, mesal process moderately wide, longer than half length of parameres. Penis (L 0.35,
<geoCoordinate id="EE2D506B6879BF76FC4CFB321738FB42" box="[986,1076,1192,1218]" degrees="0.13" direction="west" orientation="longitude" pageId="39" pageNumber="40" precision="555" value="-0.13">W 0.13</geoCoordinate>
) of moderate size, moderately sclerotized, pala narrow, elongate, paired parameroids much shorter than pala.
</paragraph>
<paragraph id="8BA636AC6879BF76FF50FB6D10DBFA32" blockId="39.[151,1436,152,2018]" pageId="39" pageNumber="40">
<emphasis id="B96DEABE6879BF76FF50FB6D1224FA91" bold="true" box="[198,296,1271,1297]" pageId="39" pageNumber="40">Female.</emphasis>
Punctation of pronotum and scutellum stronger, punctures separated by 1.0 diameter. Elytra with indistinct longitudinal ridges. Elytral apices subtly triangularly explanate. Palpomeres of maxillary palpi wide. Ventrite 5 shallowly emarginated at apex, with a pair of small foveae, ventrite 2 with dense setae in mesal portion, tergite 7 unmodified. Prehensor very long (L 1.1) and narrow, connected with bursal sclerite by a long accordion-shaped duct, bursal sclerite small, elongated (L 0.27).
</paragraph>
<paragraph id="8BA636AC6879BF76FF50FA25173CF9D2" blockId="39.[151,1436,152,2018]" pageId="39" pageNumber="40">
<emphasis id="B96DEABE6879BF76FF50FA25125EFA59" bold="true" box="[198,338,1471,1497]" pageId="39" pageNumber="40">Variability.</emphasis>
Males: TL/EW 1.31.4 (1.3); females: TL/EW 1.31.4 (1.3). Orange macula in basal portion of elytra variable: in some specimens (Sumatra,
<collectingCountry id="F30E763C6879BF76FD67FA721054F982" box="[753,856,1512,1538]" name="Malaysia" pageId="39" pageNumber="40">Sarawak</collectingCountry>
) interrupted in mesal portion; black portion of humeri can be narrowed, triangular (
<collectingCountry id="F30E763C6879BF76FDD8F98A11BBF9AA" box="[590,695,1552,1578]" name="Malaysia" pageId="39" pageNumber="40">Sarawak</collectingCountry>
,
<collectingCountry id="F30E763C6879BF76FD53F98A1023F9AA" box="[709,815,1552,1578]" name="Thailand" pageId="39" pageNumber="40">Thailand</collectingCountry>
) or wide and subrectangular (
<collectingCountry id="F30E763C6879BF76FB01F98A1608F9AA" box="[1175,1284,1552,1578]" name="Thailand" pageId="39" pageNumber="40">Thailand</collectingCountry>
,
<collectingCountry id="F30E763C6879BF76FA84F98A1641F9AA" box="[1298,1357,1552,1578]" name="Laos" pageId="39" pageNumber="40">Laos</collectingCountry>
,
<collectingCountry id="F30E763C6879BF76FACAF98A1694F9AA" box="[1372,1432,1552,1578]" name="Laos" pageId="39" pageNumber="40">Laos</collectingCountry>
,
<collectingCountry id="F30E763C6879BF76FF01F9A213D4F9D2" box="[151,216,1592,1618]" name="India" pageId="39" pageNumber="40">India</collectingCountry>
). No correlation was found between dorsal pattern and sex of specimens.
</paragraph>
<paragraph id="8BA636AC6879BF76FF50F9C511E5F94A" blockId="39.[151,1436,152,2018]" pageId="39" pageNumber="40">
<emphasis id="B96DEABE6879BF76FF50F9C51289F9F9" bold="true" box="[198,389,1631,1657]" pageId="39" pageNumber="40">Measurements.</emphasis>
Males (n = 4): TL 3.354.05 (3.74), PL 0.600.75 (0.69), PW 1.732.25 (2.01), EL 2.80 3.55 (3.28), EW 2.393.18 (2.80). Females (n = 5): TL 3.804.28 (3.93), PL 0.620.85 (0.70), PW 1.832.15 (1.99), EL 3.203.65 (3.45), EW 2.803.18 (2.93).
</paragraph>
<paragraph id="8BA636AC6879BF76FF50F94D11B9F972" blockId="39.[151,1436,152,2018]" box="[198,693,1751,1778]" pageId="39" pageNumber="40">
<emphasis id="B96DEABE6879BF76FF50F94D1215F971" bold="true" box="[198,281,1751,1777]" pageId="39" pageNumber="40">Larva.</emphasis>
See the redescription of the genus.
</paragraph>
</subSubSection>
<subSubSection id="C30365276879BF76FF50F965171BF8C2" pageId="39" pageNumber="40" type="distribution">
<paragraph id="8BA636AC6879BF76FF50F965171BF8C2" blockId="39.[151,1436,152,2018]" pageId="39" pageNumber="40">
<emphasis id="B96DEABE6879BF76FF50F965126BF899" bold="true" box="[198,359,1791,1817]" pageId="39" pageNumber="40">Distribution.</emphasis>
Sumatra, Borneo:
<collectingCountry id="F30E763C6879BF76FDDEF89A11BCF89A" box="[584,688,1792,1818]" name="Malaysia" pageId="39" pageNumber="40">Sarawak</collectingCountry>
,
<collectingCountry id="F30E763C6879BF76FD28F89A1026F89A" box="[702,810,1792,1818]" name="Thailand" pageId="39" pageNumber="40">Thailand</collectingCountry>
: northern part of Malayan Peninsula,
<collectingCountry id="F30E763C6879BF76FB7BF89A1626F89A" box="[1261,1322,1792,1818]" name="Laos" pageId="39" pageNumber="40">Laos</collectingCountry>
: Xaisomboun,
<collectingCountry id="F30E763C6879BF76FF76F8B2122DF8C2" box="[224,289,1832,1858]" name="India" pageId="39" pageNumber="40">India</collectingCountry>
:
<collectingRegion id="49DDF84E6879BF76FEBAF8B212A2F8C2" box="[300,430,1832,1858]" country="India" name="Meghalaya" pageId="39" pageNumber="40">Meghalaya</collectingRegion>
(see remarks below). Widely distributed in SE Asia.
</paragraph>
</subSubSection>
<subSubSection id="C30365276879BF76FF50F8D512E1F812" pageId="39" pageNumber="40" type="biology_ecology">
<paragraph id="8BA636AC6879BF76FF50F8D512E1F812" blockId="39.[151,1436,152,2018]" pageId="39" pageNumber="40">
<emphasis id="B96DEABE6879BF76FF50F8D51226F8E9" bold="true" box="[198,298,1871,1897]" pageId="39" pageNumber="40">Biology.</emphasis>
The larvae were collected from bamboo phytotelmata in Bogor (Watts pers. com.) together with larvae of
<taxonomicName id="4C194D2F6879BF76FE93F8E212E5F812" authority="(Pic)" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" baseAuthorityName="Pic" baseAuthorityYear="1918" box="[261,489,1912,1938]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="discoidalis">
<emphasis id="B96DEABE6879BF76FE93F8E212A9F811" box="[261,421,1912,1937]" italics="true" pageId="39" pageNumber="40">E. discoidalis</emphasis>
(Pic)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365276879BF76FF50F805120BF862" pageId="39" pageNumber="40" type="etymology">
<paragraph id="8BA636AC6879BF76FF50F805120BF862" blockId="39.[151,1436,152,2018]" pageId="39" pageNumber="40">
<emphasis id="B96DEABE6879BF76FF50F805125FF839" bold="true" box="[198,339,1951,1977]" pageId="39" pageNumber="40">Etymology.</emphasis>
Dedicated to Prof. Bernhard Klausnitzer (Dresden,
<collectingCountry id="F30E763C6879BF76FC5DF83A1731F83A" box="[971,1085,1952,1978]" name="Germany" pageId="39" pageNumber="40">Germany</collectingCountry>
), an outstanding specialist in
<taxonomicName id="4C194D2F6879BF76FF01F852120FF862" authorityName="Fleming" authorityYear="1821" box="[151,259,1992,2018]" class="Insecta" family="Scirtidae" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="family">Scirtidae</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF6666246876BF79FF01FC2B164EFC49" ID-DOI="http://doi.org/10.5281/zenodo.10095121" ID-Zenodo-Dep="10095121" box="[151,1346,945,969]" httpUri="https://zenodo.org/record/10095121/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,255,945,969]" targetBox="[158,1428,197,919]" targetPageId="40">
<paragraph id="8BA636AC6876BF79FF01FC2B164EFC49" blockId="40.[151,1346,945,969]" box="[151,1346,945,969]" pageId="40" pageNumber="41">
<emphasis id="B96DEABE6876BF79FF01FC2B122AFC49" bold="true" box="[151,294,945,969]" pageId="40" pageNumber="41">FIGURE 31.</emphasis>
<taxonomicName id="4C194D2F6876BF79FEBBFC28116AFC49" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[301,614,946,969]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="klausnitzeri" status="sp. nov.">
<emphasis id="B96DEABE6876BF79FEBBFC28116AFC49" box="[301,614,946,969]" italics="true" pageId="40" pageNumber="41">Exochomoscirtes klausnitzeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56876BF79FDFBFC2811CEFC49" box="[621,706,946,969]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
, female.
<emphasis id="B96DEABE6876BF79FCB4FC2B1032FC49" bold="true" box="[802,830,945,969]" pageId="40" pageNumber="41">A)</emphasis>
bursal sclerite;
<emphasis id="B96DEABE6876BF79FC7EFC2B170EFC49" bold="true" box="[1000,1026,945,969]" pageId="40" pageNumber="41">B)</emphasis>
prehensor and bursal sclerite.
</paragraph>
</caption>
<subSubSection id="C30365276876BF79FF50FC6611F7FAFE" pageId="40" pageNumber="41" type="discussion">
<paragraph id="8BA636AC6876BF79FF50FC6611F7FAFE" blockId="40.[151,1436,1020,1406]" pageId="40" pageNumber="41">
<emphasis id="B96DEABE6876BF79FF50FC661233FB96" bold="true" box="[198,319,1020,1046]" pageId="40" pageNumber="41">Remarks.</emphasis>
A male specimen from NE
<collectingCountry id="F30E763C6876BF79FD1BFC6611C5FB96" box="[653,713,1020,1046]" name="India" pageId="40" pageNumber="41">India</collectingCountry>
(not included in the
<typeStatus id="54A2880E6876BF79FC57FC6610FFFB96" box="[961,1011,1020,1046]" pageId="40" pageNumber="41">type</typeStatus>
series) with modified (elongated: L 1.50,
<geoCoordinate id="EE2D506B6876BF79FF4DFBBE123AFBBE" box="[219,310,1060,1086]" degrees="0.68" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="555" value="-0.68">W 0.68</geoCoordinate>
,
<figureCitation id="13222A296876BF79FED2FBBE12A2FBBE" box="[324,430,1060,1086]" captionStart="FIGURE 30" captionStartId="38.[151,255,1828,1852]" captionTargetBox="[170,1429,1159,1786]" captionTargetId="figure-414@38.[151,1436,1151,1801]" captionTargetPageId="38" captionText="FIGURE 30. Exochomoscirtes klausnitzeri sp. nov., male. A) sternite 9; B) tergite 8; C) tergite 8, specimen from NE India; D) tergite 9; E) tegmen; F) penis. Scale bar = 0.5 mm (AB, DF), 1.0 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.10095119" httpUri="https://zenodo.org/record/10095119/files/figure.png" pageId="40" pageNumber="41">Fig. 30C</figureCitation>
) tergite 8 is known (deposited in NMW). It was collected together with a specimen with typical tergite 8, therefore it is possible that elongation of tergite 8 may be due to intraspecific variability. A relatively small (TL 3.50) specimen (
<figureCitation id="13222A296876BF79FDEEFBEE11EAFB0E" box="[632,742,1140,1166]" captionStart="FIGURE 12" captionStartId="18.[151,255,1861,1885]" captionTargetBox="[164,1428,210,1822]" captionTargetId="figure-109@18.[151,1436,197,1835]" captionTargetPageId="18" captionText="FIGURE 12. Habitus illustrations of Exochomoscirtes. A) E. maculatus (Waterhouse); B)C) E. ceylonicus sp. nov.; D) E. decemguttatus (Champion); E)G) E. klausnitzeri sp. nov., color forms F S Thailand, male, E and G Sarawak, females; H) E. cf. klausnitzeri (NW Thailand); I) E. luteosuturalis sp. nov.; J) E. cf. luteosuturalis sp. nov., female, Sulawesi; K) E. cf. luteosuturalis, female, Vietnam; L) E. luteosuturaloides sp. nov.; M) E. jaechi sp. nov.; N) E. ruforotundus (Watts); O) E. ruforotundus (Watts), Bali. Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10095081" httpUri="https://zenodo.org/record/10095081/files/figure.png" pageId="40" pageNumber="41">Fig. 12H</figureCitation>
) from NW
<collectingCountry id="F30E763C6876BF79FCE0FBEE10D2FB0E" box="[886,990,1140,1166]" name="Thailand" pageId="40" pageNumber="41">Thailand</collectingCountry>
(
<collectingRegion id="49DDF84E6876BF79FC66FBEE178EFB0E" box="[1008,1154,1140,1166]" country="Thailand" name="Chiang Mai" pageId="40" pageNumber="41">Chiang Mai</collectingRegion>
) with uniformly brown dorsum and genitalia indistinguishable from that of
<taxonomicName id="4C194D2F6876BF79FD6AFB0710AAFB36" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[764,934,1181,1206]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="klausnitzeri">
<emphasis id="B96DEABE6876BF79FD6AFB0710AAFB36" box="[764,934,1181,1206]" italics="true" pageId="40" pageNumber="41">E. klausnitzeri</emphasis>
</taxonomicName>
was found in the collection of NMW. The identity of this specimen is uncertain. It is plausible that
<taxonomicName id="4C194D2F6876BF79FCBAFB5F10D8FB5E" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[812,980,1221,1246]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="klausnitzeri">
<emphasis id="B96DEABE6876BF79FCBAFB5F10D8FB5E" box="[812,980,1221,1246]" italics="true" pageId="40" pageNumber="41">E. klausnitzeri</emphasis>
</taxonomicName>
is in fact a complex of cryptic species, which may explain its unusually broad geographical range. More specimens will need to be studied to verify this hypothesis. Ch. Watts reared larvae of
<taxonomicName id="4C194D2F6876BF79FD04FA8F1056FAAE" box="[658,858,1301,1326]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABE6876BF79FD04FA8F1056FAAE" box="[658,858,1301,1326]" italics="true" pageId="40" pageNumber="41">Exochomoscirtes</emphasis>
</taxonomicName>
, which produced imagines of
<taxonomicName id="4C194D2F6876BF79FB56FA8F1666FAAE" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[1216,1386,1301,1326]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="klausnitzeri">
<emphasis id="B96DEABE6876BF79FB56FA8F1666FAAE" box="[1216,1386,1301,1326]" italics="true" pageId="40" pageNumber="41">E. klausnitzeri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D2F6876BF79FF01FAA7123BFAD6" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" baseAuthorityName="Pic" baseAuthorityYear="1918" box="[151,311,1341,1366]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="discoidalis">
<emphasis id="B96DEABE6876BF79FF01FAA7123BFAD6" box="[151,311,1341,1366]" italics="true" pageId="40" pageNumber="41">E. discoidalis</emphasis>
</taxonomicName>
. Specimens used in the larval description were identified (by HY) by the comparison with exuvia properly identified by association with the adult.
</paragraph>
</subSubSection>
</treatment>
</document>