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<document id="BBFAC05FE1219AF3C28117199374DDEC" ID-DOI="10.11646/zootaxa.2598.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="10095059" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629698685068" checkinUser="felipe" docAuthor="Ruta, Rafał &amp; Yoshitomi, Hiroyuki" docDate="2010" docId="03B087BA6867BF6BFF01FA0810D3F86A" docLanguage="en" docName="zt02598p080.pdf" docOrigin="Zootaxa 2598 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2598.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Exochomoscirtes nigromarginatus Ruta &amp; Yoshitomi 2010, sp. nov." docType="treatment" docUuid="B9B22817-3743-4CF0-B24F-04B78492C0A6" docUuidSource="ZooBank" docVersion="4" lastPageNumber="59" masterDocId="FF89FFC2685EBF51FF96FF9A130CFF80" masterDocTitle="Revision of the genus Exochomoscirtes Pic (Coleoptera: Scirtidae: Scirtinae) 2598" masterLastPageNumber="80" masterPageNumber="1" pageNumber="58" updateTime="1699569420699" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D84736210B1B2805C15D3D9B98406794" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="227356FD914725586DC3B63B8F528C27">Revision of the genus Exochomoscirtes Pic (Coleoptera: Scirtidae: Scirtinae) 2598</mods:title>
</mods:titleInfo>
<mods:name id="3834DFBF9536B56E6E0217C0CA1A1018" type="personal">
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</mods:role>
<mods:namePart id="D03CB70BB49403F532D3C7831F0ADCA4">Ruta, Rafał</mods:namePart>
</mods:name>
<mods:name id="C19C64FF52799FC32FB8FD945B674D02" type="personal">
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<mods:namePart id="DF6D7A8A524A87B1A4B6EB8D04B18EBE">Yoshitomi, Hiroyuki</mods:namePart>
</mods:name>
<mods:typeOfResource id="39AB707C1EEE277C4863A10D61EE657F">text</mods:typeOfResource>
<mods:relatedItem id="6A31BF865838B4C6478F9FD7FBED017C" type="host">
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<mods:title id="833BEFF663EF3D74F8BDD3A3667409C8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="75196535BA50242AA68DBFA39B2B7128">
<mods:date id="9F573DCC7DD63232DE93C03BBFD4F786">2010</mods:date>
<mods:detail id="6674F9F6983BE4A232A8043EF7832166" type="pubDate">
<mods:number id="BA3294473E310C36E234341A60B814DB">2010-08-31</mods:number>
</mods:detail>
<mods:detail id="D91E29E0B21F2F0ED5ECF4816AB216A8" type="volume">
<mods:number id="2B94B2545CEF70EDA81A7CC7B4F365F1">2598</mods:number>
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<mods:url id="30E16A1F014435F17440F9B3B39E41E5">https://biotaxa.org/Zootaxa/article/view/zootaxa.2598.1.1</mods:url>
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<mods:classification id="7721598679BF2D58452CD186026F97BF">journal article</mods:classification>
<mods:identifier id="925CB85D349FCE5BE22D268DA2CAF3F4" type="DOI">10.11646/zootaxa.2598.1.1</mods:identifier>
<mods:identifier id="94DD5E5B9B15F3AD719AF622A9622BB7" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="46E6B5A85B1737E454E899785D7160E3" type="Zenodo-Dep">10095059</mods:identifier>
<mods:identifier id="4675B22D4F941662070E8E5DC4E997B3" type="ZooBank">B7D2F686-CE7A-42AE-8395-4527E587C565</mods:identifier>
</mods:mods>
<treatment id="03B087BA6867BF6BFF01FA0810D3F86A" LSID="urn:lsid:zoobank.org:act:B9B22817-3743-4CF0-B24F-04B78492C0A6" httpUri="http://treatment.plazi.org/id/03B087BA6867BF6BFF01FA0810D3F86A" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
<subSubSection id="C30365276867BF68FF01FA081191FA2C" box="[151,669,1426,1452]" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="8BA636AC6867BF68FF01FA081191FA2C" blockId="57.[151,669,1426,1486]" box="[151,669,1426,1452]" pageId="57" pageNumber="58">
<heading id="D0EE81C06867BF68FF01FA081191FA2C" bold="true" box="[151,669,1426,1452]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
<emphasis id="B96DEABE6867BF68FF01FA081191FA2C" bold="true" box="[151,669,1426,1452]" pageId="57" pageNumber="58">
<taxonomicName id="4C194D2F6867BF68FF01FA081134FA2C" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[151,568,1426,1452]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="nigromarginatus" status="sp. nov.">
<emphasis id="B96DEABE6867BF68FF01FA081134FA2C" bold="true" box="[151,568,1426,1452]" italics="true" pageId="57" pageNumber="58">Exochomoscirtes nigromarginatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56867BF68FDD6FA081191FA2C" box="[576,669,1426,1452]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365276867BF68FF01FA2E1290FA4E" box="[151,412,1460,1486]" pageId="57" pageNumber="58" type="description">
<paragraph id="8BA636AC6867BF68FF01FA2E1290FA4E" blockId="57.[151,669,1426,1486]" box="[151,412,1460,1486]" pageId="57" pageNumber="58">
<heading id="D0EE81C06867BF68FF01FA2E1290FA4E" bold="true" box="[151,412,1460,1486]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
<emphasis id="B96DEABE6867BF68FF01FA2E1290FA4E" bold="true" box="[151,412,1460,1486]" pageId="57" pageNumber="58">
(
<figureCitation id="13222A296867BF68FF09FA2E1204FA4E" box="[159,264,1460,1486]" captionStart="FIGURE 3" captionStartId="8.[151,255,1079,1103]" captionTargetBox="[153,1433,194,1055]" captionTargetId="figure-129@8.[153,1433,194,1055]" captionTargetPageId="8" captionText="FIGURE 3. Metatibial spurs in Exochomoscirtes. A) E. dilaticornis sp. nov.; B) E. discoidalis (Pic); C) E. longispinus sp. nov.; D) E. nigromarginatus sp. nov.; E) E. picturatus sp. nov.; F) E. takizawai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10095065" httpUri="https://zenodo.org/record/10095065/files/figure.png" pageId="57" pageNumber="58">Figs. 3D</figureCitation>
,
<figureCitation id="13222A296867BF68FE80FA2E123DFA4E" box="[278,305,1460,1486]" captionStart="FIGURE 5" captionStartId="9.[151,255,1360,1384]" captionTargetBox="[167,1421,160,1344]" captionTargetId="figure-294@9.[151,1436,145,1364]" captionTargetPageId="9" captionText="FIGURE 5. Abdominal details in Exochomoscirtes. A) E. dilaticornis sp. nov., ventrite 5; B) E. discoidalis (Pic), ventrite 5; C) E. jaechi sp. nov., tergite 7; D) E. klausnitzeri sp. nov., tergite 7; E) E. luteosuturalis sp. nov., tergite 7; F) E. luteosuturaloides sp. nov., tergite 7; G) E. maculatus (Waterhouse), abdomen; H) E. maculatus (Waterhouse), ventrites 3–5; I) E. nigromarginatus sp. nov., ventrite 5; J) E. quadripartitus sp. nov., tergite 7.; K) E. ruforotundus (Watts) (Australia), tergite 7; L) E. takizawai sp. nov., ventrite 5." figureDoi="http://doi.org/10.5281/zenodo.10095069" httpUri="https://zenodo.org/record/10095069/files/figure.png" pageId="57" pageNumber="58">5I</figureCitation>
,
<figureCitation id="13222A296867BF68FEA8FA2E1264FA4E" box="[318,360,1460,1486]" captionStart="FIGURE 13" captionStartId="19.[151,244,1921,1942]" captionTargetBox="[156,1436,160,1902]" captionTargetId="figure-95@19.[151,1436,155,1932]" captionTargetPageId="19" captionText="FIGURE 13. Habitus illustrations of Exochomoscirtes. A) E. quadripartitus sp. nov.; B) E. discoidalis (Pic), male; C) E. discoidalis (Pic), female; D) E. thailandicus sp. nov.; E) E. serricornis sp. nov.; F) E. cambodiensis sp. nov.; G) E. bezdeki sp. nov., male; H) E. bezdeki sp. nov., female; I) E. nigromarginatus sp. nov.; J) E. sphaericus (Champion); K) E. globosus sp. nov.; L) E. buechei sp. nov.; M) E. dilaticornis sp. nov., male. Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10095085" httpUri="https://zenodo.org/record/10095085/files/figure.png" pageId="57" pageNumber="58">13I</figureCitation>
,
<figureCitation id="13222A296867BF68FEE0FA2E1299FA4E" box="[374,405,1460,1486]" captionStart="FIGURE 47" captionStartId="58.[151,255,1131,1155]" captionTargetBox="[164,1426,495,1096]" captionTargetId="figure-385@58.[151,1436,477,1104]" captionTargetPageId="58" captionText="FIGURE 47. Exochomoscirtes nigromarginatus sp. nov., male. A) sternite 8; B) sternite 9; C) tergite 8; D) tergite 9; E) tegmen; F) penis. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10095156" httpUri="https://zenodo.org/record/10095156/files/figure.png" pageId="57" pageNumber="58">47</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365276867BF68FF01F99E117AF9C7" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="8BA636AC6867BF68FF01F99E117AF9C7" blockId="57.[151,1436,1540,2007]" pageId="57" pageNumber="58">
<emphasis id="B96DEABE6867BF68FF01F99E125DF99E" bold="true" box="[151,337,1540,1566]" pageId="57" pageNumber="58">Type material.</emphasis>
<materialsCitation id="3B713CF16867BF68FECBF99E117AF9C7" collectingDate="2007-03-14" collectionCode="DBET" country="Malaysia" location="Pahang prov." pageId="57" pageNumber="58" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
<emphasis id="B96DEABE6867BF68FECBF99E1194F99E" bold="true" box="[349,664,1540,1566]" pageId="57" pageNumber="58">
<typeStatus id="54A2880E6867BF68FECBF99E12DCF99E" box="[349,464,1540,1566]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="ED08AE696867BF68FDA2F99E1187F99E" box="[564,651,1540,1566]" pageId="57" pageNumber="58">DBET</collectionCode>
):
</emphasis>
“
<collectingCountry id="F30E763C6867BF68FD38F99F1013F99F" box="[686,799,1541,1567]" name="Malaysia" pageId="57" pageNumber="58">Malaysia</collectingCountry>
,
<collectingRegion id="49DDF84E6867BF68FCA6F99F10C2F99F" box="[816,974,1541,1567]" country="Malaysia" name="Pahang" pageId="57" pageNumber="58">Pahang prov.</collectingRegion>
/ Taman Negara N.P. / Kuala Tahan,
<date id="FFA7106C6867BF68FF01F9B71203F9C7" box="[151,271,1581,1607]" pageId="57" pageNumber="58" value="2007-03-14">
<collectingDate id="EFE3E9846867BF68FF01F9B71203F9C7" box="[151,271,1581,1607]" pageId="57" pageNumber="58" value="2007-03-14">14.3.2007</collectingDate>
</date>
, / lgt. Jiři Foit” [printed label].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30365276867BF68FF50F9CE1000F967" pageId="57" pageNumber="58" type="diagnosis">
<paragraph id="8BA636AC6867BF68FF50F9CE1000F967" blockId="57.[151,1436,1540,2007]" pageId="57" pageNumber="58">
<emphasis id="B96DEABE6867BF68FF50F9CE124AF9EE" bold="true" box="[198,326,1620,1646]" pageId="57" pageNumber="58">Diagnosis.</emphasis>
Body shape is similar to
<taxonomicName id="4C194D2F6867BF68FDE1F9CF11FDF9EE" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[631,753,1621,1646]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="bezdeki" status="sp. nov.">
<emphasis id="B96DEABE6867BF68FDE1F9CF11FDF9EE" box="[631,753,1621,1646]" italics="true" pageId="57" pageNumber="58">E. bezdeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56867BF68FD6CF9CF1054F9EF" box="[762,856,1621,1647]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
, but the dorsum is redish-brown with black margination on elytra (entirely black in
<taxonomicName id="4C194D2F6867BF68FDC5F9E711C3F916" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[595,719,1661,1686]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="bezdeki">
<emphasis id="B96DEABE6867BF68FDC5F9E711C3F916" box="[595,719,1661,1686]" italics="true" pageId="57" pageNumber="58">E. bezdeki</emphasis>
</taxonomicName>
).
<taxonomicName id="4C194D2F6867BF68FD7AF9E7178DF916" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[748,1153,1661,1686]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="nigromarginatus" status="sp. nov.">
<emphasis id="B96DEABE6867BF68FD7AF9E7178DF916" box="[748,1153,1661,1686]" italics="true" pageId="57" pageNumber="58">Exochomoscirtes nigromarginatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56867BF68FB1BF9E717E0F917" box="[1165,1260,1661,1687]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="4C194D2F6867BF68FEAAF93F12B9F93E" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[316,437,1701,1726]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="bezdeki" status="sp. nov.">
<emphasis id="B96DEABE6867BF68FEAAF93F12B9F93E" box="[316,437,1701,1726]" italics="true" pageId="57" pageNumber="58">E. bezdeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56867BF68FE28F93F1116F93F" box="[446,538,1701,1727]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
on the basis of the morphology of the tegmen, which has strongly widened parameroids (slightly widened in
<taxonomicName id="4C194D2F6867BF68FDB7F9571195F966" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[545,665,1741,1766]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="bezdeki" status="sp. nov.">
<emphasis id="B96DEABE6867BF68FDB7F9571195F966" box="[545,665,1741,1766]" italics="true" pageId="57" pageNumber="58">E. bezdeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56867BF68FD36F95711F3F967" box="[672,767,1741,1767]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365276867BF6BFF50F96E1756F81A" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="description">
<paragraph id="8BA636AC6867BF68FF50F96E100DF807" blockId="57.[151,1436,1540,2007]" pageId="57" pageNumber="58">
<emphasis id="B96DEABE6867BF68FF50F96E1252F88E" bold="true" box="[198,350,1780,1806]" pageId="57" pageNumber="58">Description.</emphasis>
<typeStatus id="54A2880E6867BF68FEFCF96F12D6F88F" box="[362,474,1781,1807]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
, male. Body oval, TL/EW 1.3, slightly flattened, covered with brownish setae. Head brown, anterior portion dark brown, pronotum brown with brownish-black margins, elytra brown with blackish margins. Mouthparts and antennae yellow. Ventrum brown, legs brown with the exception of apices of metafemora, which are darkened, brownish black.
</paragraph>
<paragraph id="8BA636AC6867BF6BFF50F80F1041FF5A" blockId="57.[151,1436,1540,2007]" lastBlockId="58.[151,1436,152,418]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
Head covered with small but dense punctures, punctures separated by ca. 1.0 diameter. Eyes big, protuberant; head 2.0 X wider than interocular space. Antennae filiform, antennomeres 9–11 missing in the
<typeStatus id="54A2880E6867BF68FAB8F8271694F857" box="[1326,1432,1981,2007]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
. Antennomeres 2 and 3 of subequal lengths, antennomeres 4–11 slightly longer than antennomeres 2 and 3 combined. Palpomeres of maxillary palpi slightly widened.
</paragraph>
<paragraph id="8BA636AC6864BF6BFF50FF7211D2FE22" blockId="58.[151,1436,152,418]" pageId="58" pageNumber="59">Pronotum slightly convex, covered with distinct punctation, stronger than on head, separated by ca. 1.0 (rarely 2.0) diameters, anterolateral corners almost right-angled, slightly obtuse, projecting anteriorly; lateral margins straight; posterolateral corners acute; base of pronotum subtrapezoidal, with incomplete margination along basal margin; PW/PL 3.0. Scutellum punctured as on pronotum, equilaterally triangular. Angle between pronotum and elytra not marked in dorsal outline.</paragraph>
<caption id="DF6666246864BF6BFF01FBF1113CFB24" ID-DOI="http://doi.org/10.5281/zenodo.10095156" ID-Zenodo-Dep="10095156" httpUri="https://zenodo.org/record/10095156/files/figure.png" pageId="58" pageNumber="59" startId="58.[151,255,1131,1155]" targetBox="[164,1426,495,1096]" targetPageId="58">
<paragraph id="8BA636AC6864BF6BFF01FBF1113CFB24" blockId="58.[151,1436,1131,1189]" pageId="58" pageNumber="59">
<emphasis id="B96DEABE6864BF6BFF01FBF1122BFB03" bold="true" box="[151,295,1131,1155]" pageId="58" pageNumber="59">FIGURE 47.</emphasis>
<taxonomicName id="4C194D2F6864BF6BFEB8FBF11190FB02" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[302,668,1131,1154]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="nigromarginatus" status="sp. nov.">
<emphasis id="B96DEABE6864BF6BFEB8FBF11190FB02" box="[302,668,1131,1154]" italics="true" pageId="58" pageNumber="59">Exochomoscirtes nigromarginatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56864BF6BFD32FBF111F5FB02" box="[676,761,1131,1154]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
, male.
<emphasis id="B96DEABE6864BF6BFCD3FBF1106EFB03" bold="true" box="[837,866,1131,1155]" pageId="58" pageNumber="59">A)</emphasis>
sternite 8;
<emphasis id="B96DEABE6864BF6BFC4CFBF110F9FB03" bold="true" box="[986,1013,1131,1155]" pageId="58" pageNumber="59">B)</emphasis>
sternite 9;
<emphasis id="B96DEABE6864BF6BFBFBFBF11785FB03" bold="true" box="[1133,1161,1131,1155]" pageId="58" pageNumber="59">C)</emphasis>
tergite 8;
<emphasis id="B96DEABE6864BF6BFB61FBF11618FB03" bold="true" box="[1271,1300,1131,1155]" pageId="58" pageNumber="59">D)</emphasis>
tergite 9;
<emphasis id="B96DEABE6864BF6BFA17FBF11690FB03" bold="true" box="[1409,1436,1131,1155]" pageId="58" pageNumber="59">E)</emphasis>
tegmen;
<emphasis id="B96DEABE6864BF6BFF65FB171200FB25" bold="true" box="[243,268,1165,1189]" pageId="58" pageNumber="59">F)</emphasis>
penis. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BA636AC6864BF6BFF50FA9A1671FAC2" blockId="58.[151,1436,1280,2026]" pageId="58" pageNumber="59">Elytra oval, with slightly explanate lateral margins, widest in the middle, punctation stronger than on pronotum, punctures separated by ca. 1.0 diameter; humeri well marked; EL/EW 1.1; EL/PL 4.9; EW/PW 1.5.</paragraph>
<paragraph id="8BA636AC6864BF6BFF01FACA13FFFA62" blockId="58.[151,1436,1280,2026]" pageId="58" pageNumber="59">Legs moderately long. Hind tibial spurs well developed, dorsal one curved, as long as 2/3 length of tarsomere 1, and nearly twice as long as ventral one, slightly widened subapically, forming small, flattened lobe; ventral spur slightly curved. Tarsomere 1 of posterior legs widened apically, with bulbous, microreticulated dorsal portion.</paragraph>
<paragraph id="8BA636AC6864BF6BFF50FA6A10A5F9B2" blockId="58.[151,1436,1280,2026]" pageId="58" pageNumber="59">Ventrite 1 with sparse setation in mesal portion, ventrite 2 with basal row of big punctures. Apex of tergite 7 unmodified. Apex of ventrite 5 shallowly emarginated, bisinuate.</paragraph>
<paragraph id="8BA636AC6864BF6BFF50F9DA1658F8CA" blockId="58.[151,1436,1280,2026]" pageId="58" pageNumber="59">
Sternite 8 (L 0.12,
<geoCoordinate id="EE2D506B6864BF6BFE3AF9DA110BF9DA" box="[428,519,1600,1626]" degrees="0.29" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="555" value="-0.29">W 0.29</geoCoordinate>
) widely U-shaped; sternite 9 (L 0.82,
<geoCoordinate id="EE2D506B6864BF6BFC40F9DA173DF9DA" box="[982,1073,1600,1626]" degrees="0.38" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="555" value="-0.38">W 0.38</geoCoordinate>
) subtriangular, emarginated at apex, apodemes fused basally, with several very short setae at apex. Tergite 8 (L 1.60,
<geoCoordinate id="EE2D506B6864BF6BFB03F9F217E2F902" box="[1173,1262,1640,1666]" degrees="1.20" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="555" value="-1.2">W 1.20</geoCoordinate>
) large, moderately sclerotized, with arcuate apical margin, bearing row of moderately long setae on apical margin, apodemes long, diverging basally; tergite 9 (L 0.98,
<geoCoordinate id="EE2D506B6864BF6BFD74F9221030F952" box="[738,828,1720,1746]" degrees="0.65" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="555" value="-0.65">W 0.65</geoCoordinate>
) moderately sclerotized, apical portion small, with short setae at apex, with a pair of apodemes diverging basally. Tegmen (L 0.87,
<geoCoordinate id="EE2D506B6864BF6BFBFDF97A17CAF97A" box="[1131,1222,1760,1786]" degrees="0.73" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="555" value="-0.73">W 0.73</geoCoordinate>
) well sclerotized, parameres strongly widened apically, mesal process narrow, as long as half length of parameres. Penis (L 0.45,
<geoCoordinate id="EE2D506B6864BF6BFF4FF8AA123DF8CA" box="[217,305,1840,1866]" degrees="0.16" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="555" value="-0.16">W 0.16</geoCoordinate>
) small, moderately sclerotized, pala subtriangular, apical processes almost as long as pala.
</paragraph>
<paragraph id="8BA636AC6864BF6BFF50F8C21295F8F2" blockId="58.[151,1436,1280,2026]" box="[198,409,1880,1906]" pageId="58" pageNumber="59">Female unknown.</paragraph>
<paragraph id="8BA636AC6864BF6BFF50F81A1756F81A" blockId="58.[151,1436,1280,2026]" box="[198,1114,1920,1946]" pageId="58" pageNumber="59">
<emphasis id="B96DEABE6864BF6BFF50F81A1289F81A" bold="true" box="[198,389,1920,1946]" pageId="58" pageNumber="59">Measurements.</emphasis>
Male (n = 1): TL 4.64, PL 0.85, PW 2.55, EL 4.15, EW 3.70.
</paragraph>
</subSubSection>
<subSubSection id="C30365276864BF6BFF50F832114AF842" box="[198,582,1960,1986]" pageId="58" pageNumber="59" type="distribution">
<paragraph id="8BA636AC6864BF6BFF50F832114AF842" blockId="58.[151,1436,1280,2026]" box="[198,582,1960,1986]" pageId="58" pageNumber="59">
<emphasis id="B96DEABE6864BF6BFF50F832126BF842" bold="true" box="[198,359,1960,1986]" pageId="58" pageNumber="59">Distribution.</emphasis>
<collectingRegion id="49DDF84E6864BF6BFEF8F83212CBF842" box="[366,455,1960,1986]" country="Malaysia" name="Pahang" pageId="58" pageNumber="59">Pahang</collectingRegion>
,
<collectingCountry id="F30E763C6864BF6BFE45F832114EF842" box="[467,578,1960,1986]" name="Malaysia" pageId="58" pageNumber="59">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365276864BF6BFF50F84A10D3F86A" box="[198,991,2000,2026]" pageId="58" pageNumber="59" type="etymology">
<paragraph id="8BA636AC6864BF6BFF50F84A10D3F86A" blockId="58.[151,1436,1280,2026]" box="[198,991,2000,2026]" pageId="58" pageNumber="59">
<emphasis id="B96DEABE6864BF6BFF50F84A125FF86A" bold="true" box="[198,339,2000,2026]" pageId="58" pageNumber="59">Etymology.</emphasis>
Specific epithet referring to dark margination of elytra.
</paragraph>
</subSubSection>
</treatment>
</document>
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