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<document id="FB4FECE9CBFC2A2867C03400BD1B0A7C" ID-DOI="10.11646/zootaxa.2598.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="10095059" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629698685068" checkinUser="felipe" docAuthor="Ruta, Rafał &amp; Yoshitomi, Hiroyuki" docDate="2010" docId="03B087BA6844BF4DFF01F99F10F7FABA" docLanguage="en" docName="zt02598p080.pdf" docOrigin="Zootaxa 2598 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2598.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Exochomoscirtes picturatus Ruta &amp; Yoshitomi 2010, sp. nov." docType="treatment" docVersion="3" lastPageNumber="29" masterDocId="FF89FFC2685EBF51FF96FF9A130CFF80" masterDocTitle="Revision of the genus Exochomoscirtes Pic (Coleoptera: Scirtidae: Scirtinae) 2598" masterLastPageNumber="80" masterPageNumber="1" pageNumber="27" updateTime="1699569420699" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F0C0554D11FFC648C1AC90CE42A14D29" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7C8E43B05965DF8ABD7ED908543B3A28">
<mods:title id="2D821F19CF4F46315946D1958746E471">Revision of the genus Exochomoscirtes Pic (Coleoptera: Scirtidae: Scirtinae) 2598</mods:title>
</mods:titleInfo>
<mods:name id="F7AF119E2C248012FEDFF33592DA8C41" type="personal">
<mods:role id="E284CE15EA74FA504DFB033DCEEC1B71">
<mods:roleTerm id="3675E19F077C7C6FF72411205499CB16">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="698FCC36D3621E6F591A6B7554D164B6">Ruta, Rafał</mods:namePart>
</mods:name>
<mods:name id="F068A48C4788CEDC9C2BFC7AEECAD090" type="personal">
<mods:role id="87357CE7117668EF1967C352CBFB89D9">
<mods:roleTerm id="7D0817B601357637F5D5CC6F0B8320DF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0978BC21D236FA72FC0E20E593332A6B">Yoshitomi, Hiroyuki</mods:namePart>
</mods:name>
<mods:typeOfResource id="BB2B23C55600137BE8CFA1A7B0B6DAE8">text</mods:typeOfResource>
<mods:relatedItem id="9C188C55E053CA7B9182A8DC11781316" type="host">
<mods:titleInfo id="BE71F0EA74782EC185BE98D17C4ED35C">
<mods:title id="837E50239BE7B5FC0F3C353D6402E1F5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="13EBAD67FE83E8A3BDD2C0CC5E4BEF8F">
<mods:date id="2F4E2CEFCD0594DCFF5C5E37C6C1AD1F">2010</mods:date>
<mods:detail id="AEDC4368CE6C1D209DB8215A59E914F6" type="pubDate">
<mods:number id="D59E73476980FB715F2F92006063A062">2010-08-31</mods:number>
</mods:detail>
<mods:detail id="CE9F7B3553B23CED6FF61F6E16C74263" type="volume">
<mods:number id="87D02A19A6FF8CF0AD9F28CD5ADCC73B">2598</mods:number>
</mods:detail>
<mods:detail id="6EB746B10095623E67A7CC0C4B11A7F1" type="issue">
<mods:number id="721019BFF25103C7AA7CA1DA717977F7">1</mods:number>
</mods:detail>
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<mods:start id="3674F3E4A50454F9C7B476895D69150F">1</mods:start>
<mods:end id="F9F53EB2B6DDFA8C21E0BC972F7F40B5">80</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="94C77534924293678D683095C9DB19B7">
<mods:url id="1B458D40AB7C10C39FA4DCD4A281F3B0">https://biotaxa.org/Zootaxa/article/view/zootaxa.2598.1.1</mods:url>
</mods:location>
<mods:classification id="CFEF9DEAC3F3780771E6B055B892A6A0">journal article</mods:classification>
<mods:identifier id="6051BBE0FEAB0CAB5988BC0BE61DD519" type="DOI">10.11646/zootaxa.2598.1.1</mods:identifier>
<mods:identifier id="81FA5E280120B41DE68AD018800403EE" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="0AFDCDAD7CE88E7B3A5186ECD35D02E9" type="Zenodo-Dep">10095059</mods:identifier>
<mods:identifier id="CFD4532AF69D725D9A80D192CBFECD62" type="ZooBank">B7D2F686-CE7A-42AE-8395-4527E587C565</mods:identifier>
</mods:mods>
<treatment id="03B087BA6844BF4DFF01F99F10F7FABA" LSID="urn:lsid:plazi:treatment:03B087BA6844BF4DFF01F99F10F7FABA" httpUri="http://treatment.plazi.org/id/03B087BA6844BF4DFF01F99F10F7FABA" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
<subSubSection id="C30365276844BF4BFF01F99F1147F99F" box="[151,587,1541,1567]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BA636AC6844BF4BFF01F99F1147F99F" blockId="26.[151,607,1541,1602]" box="[151,587,1541,1567]" pageId="26" pageNumber="27">
<heading id="D0EE81C06844BF4BFF01F99F1147F99F" bold="true" box="[151,587,1541,1567]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B96DEABE6844BF4BFF01F99F1147F99F" bold="true" box="[151,587,1541,1567]" pageId="26" pageNumber="27">
<taxonomicName id="4C194D2F6844BF4BFF01F99F12EBF99F" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[151,487,1541,1567]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="picturatus" status="sp. nov.">
<emphasis id="B96DEABE6844BF4BFF01F99F12EBF99F" bold="true" box="[151,487,1541,1567]" italics="true" pageId="26" pageNumber="27">Exochomoscirtes picturatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56844BF4BFE78F99F1147F99F" box="[494,587,1541,1567]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365276844BF4BFF01F9B21153F9C2" box="[151,607,1576,1602]" pageId="26" pageNumber="27" type="description">
<paragraph id="8BA636AC6844BF4BFF01F9B21153F9C2" blockId="26.[151,607,1541,1602]" box="[151,607,1576,1602]" pageId="26" pageNumber="27">
(
<figureCitation id="13222A296844BF4BFF09F9B21224F9C2" box="[159,296,1576,1602]" captionStart="FIGURE 1" captionStartId="6.[151,255,1902,1926]" captionTargetBox="[179,1420,474,1851]" captionTargetId="figure-179@6.[151,1436,474,1890]" captionTargetPageId="6" captionText="FIGURE 1. Morphological details of Exochomoscirtes. A) Exochomoscirtes rufomaculatus Pic, labium; B) Exochomoscirtes rufomaculatus Pic, maxilla; C) Exochomoscirtes rufomaculatus Pic, mandible; D) Exochomoscirtes picturatus sp. nov., meso- and metaventrite; E) Exochomoscirtes picturatus sp. nov., meso- and metaventrite, closeup; F) Exochomoscirtes maculatus (Waterhouse), wing; G) Exochomoscirtes dilaticornis sp. nov., wing." figureDoi="http://doi.org/10.5281/zenodo.10095061" httpUri="https://zenodo.org/record/10095061/files/figure.png" pageId="26" pageNumber="27">Figs. 1D–E</figureCitation>
,
<figureCitation id="13222A296844BF4BFEA3F9B21271F9C2" box="[309,381,1576,1602]" captionStart="FIGURE 2" captionStartId="7.[151,255,693,717]" captionTargetBox="[175,1411,170,668]" captionTargetId="figure-442@7.[167,1419,141,704]" captionTargetPageId="7" captionText="FIGURE 2. Exochomoscirtes picturatus sp. nov., maxilla. A) male; B) female." figureDoi="http://doi.org/10.5281/zenodo.10095063" httpUri="https://zenodo.org/record/10095063/files/figure.png" pageId="26" pageNumber="27">2A–B</figureCitation>
,
<figureCitation id="13222A296844BF4BFE1FF9B212A0F9C2" box="[393,428,1576,1602]" captionStart="FIGURE 3" captionStartId="8.[151,255,1079,1103]" captionTargetBox="[153,1433,194,1055]" captionTargetId="figure-129@8.[153,1433,194,1055]" captionTargetPageId="8" captionText="FIGURE 3. Metatibial spurs in Exochomoscirtes. A) E. dilaticornis sp. nov.; B) E. discoidalis (Pic); C) E. longispinus sp. nov.; D) E. nigromarginatus sp. nov.; E) E. picturatus sp. nov.; F) E. takizawai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10095065" httpUri="https://zenodo.org/record/10095065/files/figure.png" pageId="26" pageNumber="27">3E</figureCitation>
,
<figureCitation id="13222A296844BF4BFE2FF9B2110CF9C2" box="[441,512,1576,1602]" captionStart="FIGURE 11" captionStartId="17.[151,255,1886,1910]" captionTargetBox="[164,1425,210,1820]" captionTargetId="figure-69@17.[152,1434,197,1860]" captionTargetPageId="17" captionText="FIGURE 11. Habitus illustrations of Exochomoscirtes. A) E. bimaculatus Pic; B) E. basipunctatus sp. nov.; C) E. binotatus Pic; D) E. rufomaculatus Pic; E) E. ephippiatus (Champion); F)–I) E. picturatus sp. nov., color forms; J)–L) E. takizawai sp. nov., color forms; M) E. tetrastigma (Champion). Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10095079" httpUri="https://zenodo.org/record/10095079/files/figure.png" pageId="26" pageNumber="27">11F–I</figureCitation>
,
<figureCitation id="13222A296844BF4BFD9AF9B21120F9C2" box="[524,556,1576,1602]" captionStart="FIGURE 20" captionStartId="27.[151,255,1587,1611]" captionTargetBox="[183,1388,936,1544]" captionTargetId="figure-273@27.[151,1436,914,1564]" captionTargetPageId="27" captionText="FIGURE 20. Exochomoscirtes picturatus sp. nov., male. A) sternite 9; B) tergite 8; C) tergite 9; D) aedeagus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10095099" httpUri="https://zenodo.org/record/10095099/files/figure.png" pageId="26" pageNumber="27">20</figureCitation>
,
<figureCitation id="13222A296844BF4BFDAEF9B2115AF9C2" box="[568,598,1576,1602]" captionStart="FIGURE 21" captionStartId="27.[151,255,2004,2028]" captionTargetBox="[151,1436,1700,1980]" captionTargetId="figure-276@27.[151,1436,1700,1980]" captionTargetPageId="27" captionText="FIGURE 21. Exochomoscirtes picturatus sp. nov., female A)–B) prehensor and bursal sclerite." figureDoi="http://doi.org/10.5281/zenodo.10095101" httpUri="https://zenodo.org/record/10095101/files/figure.png" pageId="26" pageNumber="27">21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365276844BF4BFF01F9E81689F8D4" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8BA636AC6844BF4BFF01F9E81689F8D4" blockId="26.[151,1436,1650,2036]" pageId="26" pageNumber="27">
<emphasis id="B96DEABE6844BF4BFF01F9E81241F90C" bold="true" box="[151,333,1650,1676]" pageId="26" pageNumber="27">Type material.</emphasis>
<materialsCitation id="3B713CF16844BF4BFEC0F9E817FFF934" collectingDate="1996-04-30" collectingDateMax="1996-05-27" collectingDateMin="1996-04-30" collectionCode="NMW" collectorName="M. Strba &amp; R. Hergovits" country="Malaysia" elevation="1800" location="Banjaran Crocker Mts." pageId="26" pageNumber="27" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B96DEABE6844BF4BFEC0F9E8118AF90C" bold="true" box="[342,646,1650,1676]" pageId="26" pageNumber="27">
<typeStatus id="54A2880E6844BF4BFEC0F9E812C6F90C" box="[342,458,1650,1676]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="ED08AE696844BF4BFDB2F9E81175F90C" box="[548,633,1650,1676]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
):
</emphasis>
“
<collectingCountry id="F30E763C6844BF4BFD0FF9E81038F90C" box="[665,820,1650,1676]" name="Malaysia" pageId="26" pageNumber="27">MALAYSIA</collectingCountry>
–
<collectingRegion id="49DDF84E6844BF4BFCC5F9E810D3F90C" box="[851,991,1650,1676]" country="Malaysia" name="Sabah" pageId="26" pageNumber="27">Sabah prov.</collectingRegion>
/
<location id="8EC660776844BF4BFC61F9E81609F90C" LSID="urn:lsid:plazi:treatment:03B087BA6844BF4DFF01F99F10F7FABA:8EC660776844BF4BFC61F9E81609F90C" box="[1015,1285,1650,1676]" country="Malaysia" name="Banjaran Crocker Mts." pageId="26" pageNumber="27" stateProvince="Sabah">Banjaran Crocker Mts.</location>
/ GUNUNG ALAB peak /
<date id="FFA7106C6844BF4BFED7F900111DF934" box="[321,529,1690,1716]" pageId="26" pageNumber="27" value="1996-04-30" valueMax="1996-05-27" valueMin="1996-04-30">
<collectingDate id="EFE3E9846844BF4BFED7F900111DF934" box="[321,529,1690,1716]" pageId="26" pageNumber="27" value="1996-04-30" valueMax="1996-05-27" valueMin="1996-04-30">30.IV.–27.V.1996</collectingDate>
</date>
, 1650–
<quantity id="4CE19B496844BF4BFDFFF90011B5F934" box="[617,697,1690,1716]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="26" pageNumber="27" unit="m" value="1800.0">
<elevation id="0034D19F6844BF4BFDFFF90011B5F934" box="[617,697,1690,1716]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="26" pageNumber="27" unit="m" value="1800.0">1800m</elevation>
</quantity>
/
<collectorName id="26EC537A6844BF4BFD42F9001031F934" box="[724,829,1690,1716]" pageId="26" pageNumber="27">M. Štrba</collectorName>
&amp;
<collectorName id="26EC537A6844BF4BFCF0F90010F1F934" box="[870,1021,1690,1716]" pageId="26" pageNumber="27">R. Hergovits</collectorName>
leg.” [printed label]
</materialsCitation>
.
<materialsCitation id="3B713CF16844BF4BFB69F90011EEF95C" collectingDate="1996-04-30" collectingDateMax="1996-05-27" collectingDateMin="1996-04-30" collectionCode="NMW" collectorName="M. Strba &amp; R. Hergovits" country="Malaysia" elevation="1800" location="Banjaran Crocker Mts." pageId="26" pageNumber="27" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Female" typeStatus="paratype">
<emphasis id="B96DEABE6844BF4BFB69F90012C2F95C" bold="true" pageId="26" pageNumber="27">
<typeStatus id="54A2880E6844BF4BFB69F9001673F934" box="[1279,1407,1690,1716]" pageId="26" pageNumber="27" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="9D1FFD256844BF4BFA1BF90013ECF95C" pageId="26" pageNumber="27" type="male">2 males</specimenCount>
,
<specimenCount id="9D1FFD256844BF4BFF79F9581256F95C" box="[239,346,1730,1756]" pageId="26" pageNumber="27" type="female">1 female</specimenCount>
(
<collectionCode id="ED08AE696844BF4BFEFAF95812CDF95C" box="[364,449,1730,1756]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
):
</emphasis>
same data as holotype
</materialsCitation>
.
<materialsCitation id="3B713CF16844BF4BFD79F958168DF8D4" collectingDate="1964-03-19" collectingDateMax="1996-05-27" collectingDateMin="1964-03-19" collectionCode="DBET, NHM" collectorName="M. Strba &amp; R. Hergovits &amp; S. Kueh." country="Malaysia" elevation="1800" location="Mt. Kinabalu" municipality="Banjaran Crocker Mts." pageId="26" pageNumber="27" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="B96DEABE6844BF4BFD79F958174DF95C" bold="true" box="[751,1089,1730,1756]" pageId="26" pageNumber="27">
<specimenCount id="9D1FFD256844BF4BFD79F9581049F95C" box="[751,837,1730,1756]" pageId="26" pageNumber="27" type="male">1 male</specimenCount>
,
<specimenCount id="9D1FFD256844BF4BFCC2F95810C6F95C" box="[852,970,1730,1756]" pageId="26" pageNumber="27" type="female">3 females</specimenCount>
(
<collectionCode id="ED08AE696844BF4BFC4AF958173FF95C" box="[988,1075,1730,1756]" pageId="26" pageNumber="27">DBET</collectionCode>
):
</emphasis>
“
<collectingCountry id="F30E763C6844BF4BFBC5F95817FDF95C" box="[1107,1265,1730,1756]" name="Malaysia" pageId="26" pageNumber="27">MALAYSIA</collectingCountry>
,
<collectingRegion id="49DDF84E6844BF4BFB69F9581687F95C" box="[1279,1419,1730,1756]" country="Malaysia" name="Sabah" pageId="26" pageNumber="27">Sabah prov.</collectingRegion>
/
<collectingMunicipality id="6BC2ACD66844BF4BFF01F97012AFF884" box="[151,419,1770,1796]" pageId="26" pageNumber="27">Banjaran Crocker Mts.</collectingMunicipality>
/ GUNUNG ALAB peak
<date id="FFA7106C6844BF4BFD43F97010B1F884" box="[725,957,1770,1796]" pageId="26" pageNumber="27" value="1996-04-30" valueMax="1996-05-27" valueMin="1996-04-30">
<collectingDate id="EFE3E9846844BF4BFD43F97010B1F884" box="[725,957,1770,1796]" pageId="26" pageNumber="27" value="1996-04-30" valueMax="1996-05-27" valueMin="1996-04-30">30 IV –27 V / 1996</collectingDate>
</date>
, 1650–
<quantity id="4CE19B496844BF4BFB84F970176FF884" box="[1042,1123,1770,1796]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="26" pageNumber="27" unit="m" value="1800.0">
<elevation id="0034D19F6844BF4BFB84F970176FF884" box="[1042,1123,1770,1796]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="26" pageNumber="27" unit="m" value="1800.0">1800m</elevation>
</quantity>
/
<collectorName id="26EC537A6844BF4BFBECF97017EDF884" box="[1146,1249,1770,1796]" pageId="26" pageNumber="27">M. Štrba</collectorName>
&amp;
<collectorName id="26EC537A6844BF4BFA90F9701690F884" box="[1286,1436,1770,1796]" pageId="26" pageNumber="27">R. Hergovits</collectorName>
leg.” [printed label].
<emphasis id="B96DEABE6844BF4BFE19F888115AF8AC" bold="true" box="[399,598,1810,1836]" pageId="26" pageNumber="27">
Female (
<collectionCode id="ED08AE696844BF4BFE6DF8881145F8AC" box="[507,585,1810,1836]" pageId="26" pageNumber="27">NHM</collectionCode>
):
</emphasis>
“N BORNEO /
<location id="8EC660776844BF4BFC8DF88810B7F8AC" LSID="urn:lsid:plazi:treatment:03B087BA6844BF4DFF01F99F10F7FABA:8EC660776844BF4BFC8DF88810B7F8AC" box="[795,955,1810,1836]" country="Malaysia" municipality="Banjaran Crocker Mts." name="Mt. Kinabalu" pageId="26" pageNumber="27" stateProvince="Sabah">Mt. Kinabalu</location>
” [printed label]; “Pinosuk Plateau. /
<date id="FFA7106C6844BF4BFAE6F8881215F8D4" pageId="26" pageNumber="27" value="1964-03-19" valueMax="1964-03-28" valueMin="1964-03-19">
<collectingDate id="EFE3E9846844BF4BFAE6F8881215F8D4" pageId="26" pageNumber="27" value="1964-03-19" valueMax="1964-03-28" valueMin="1964-03-19">19– 28.iii.1964</collectingDate>
</date>
. /
<quantity id="4CE19B496844BF4BFEA2F8A01295F8D4" box="[308,409,1850,1876]" metricMagnitude="3" metricUnit="m" metricValue="1.59258" pageId="26" pageNumber="27" unit="ft" value="5225.0">
<elevation id="0034D19F6844BF4BFEA2F8A01295F8D4" box="[308,409,1850,1876]" metricMagnitude="3" metricUnit="m" metricValue="1.59258" pageId="26" pageNumber="27" unit="ft" value="5225.0">5, 225ft.</elevation>
</quantity>
” [printed label]; “Royal.Soc.Exped. / coll.
<collectorName id="26EC537A6844BF4BFC04F8A010F4F8D4" box="[914,1016,1850,1876]" pageId="26" pageNumber="27">S. Kueh.</collectorName>
/ B.M. 1964-250” [printed label]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365276844BF4BFF50F8F813FDF874" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8BA636AC6844BF4BFF50F8F813FDF874" blockId="26.[151,1436,1650,2036]" pageId="26" pageNumber="27">
<emphasis id="B96DEABE6844BF4BFF50F8F8124AF8FC" bold="true" box="[198,326,1890,1916]" pageId="26" pageNumber="27">Diagnosis.</emphasis>
Specimens with tricolor dorsal pattern cannot be mistaken with any other known
<taxonomicName id="4C194D2F6844BF4BFA80F8F913EFF824" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABE6844BF4BFA80F8F913EFF824" italics="true" pageId="26" pageNumber="27">Exochomoscirtes</emphasis>
</taxonomicName>
. Rare forms with black &amp; red pattern can be distinguished from related species by the peculiar dorsal metatibial spur in the male which is somewhat similar to the one in
<taxonomicName id="4C194D2F6844BF4BFC57F8291766F84C" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[961,1130,1971,1996]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="dilaticornis" status="sp. nov.">
<emphasis id="B96DEABE6844BF4BFC57F8291766F84C" box="[961,1130,1971,1996]" italics="true" pageId="26" pageNumber="27">E. dilaticornis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56844BF4BFBE5F82817C2F84C" box="[1139,1230,1970,1996]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C194D2F6844BF4BFA9CF8291690F84C" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[1290,1436,1971,1996]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="takizawai" status="sp. nov.">
<emphasis id="B96DEABE6844BF4BFA9CF8291690F84C" box="[1290,1436,1971,1996]" italics="true" pageId="26" pageNumber="27">E. takizawai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56844BF4BFF01F84013FDF874" box="[151,241,2010,2036]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C30365276845BF4DFF50FF0D11E5FB6A" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
<paragraph id="8BA636AC6845BF4AFF50FF0D13D2FED2" blockId="27.[151,1436,151,858]" pageId="27" pageNumber="28">
<emphasis id="B96DEABE6845BF4AFF50FF0D1252FF31" bold="true" box="[198,350,151,177]" pageId="27" pageNumber="28">Description.</emphasis>
<typeStatus id="54A2880E6845BF4AFEFCFF0212D6FF32" box="[362,474,152,178]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
, male. Body hemispherical, TL/EW 1.2, convex, covered with yellow suberect setae. Head black with brownish anterior margin of clypeus, pronotum black, elytra with color pattern. Posterior portion of elytra, humeri, and oval macula in adsutural portion of anterior half of elytra black. Remaining parts of elytra yellow and orange. Legs light brown. Mouthparts brownish, antennomeres 1–3 yellowish, 4–11 black.
</paragraph>
<paragraph id="8BA636AC6845BF4AFF50FEFA117EFD9A" blockId="27.[151,1436,151,858]" pageId="27" pageNumber="28">Head with distinct punctation, punctures separated by ca. 1.0 diameter. Eyes big, protuberant; head 1.7 X wider than interocular space. Antennae filiform, reaching basal 1/3 of elytra. Antennomere 1 cylindrical, curved, antennomere 2 short, globular, antennomere 3 distinctly longer than antennomere 2, narrow, antennomere 4–10 1.5 X longer than antennomere 3, apical antennomere ca. 1.5 X longer than preapical one. Palpomeres of maxillary palpi moderately wide.</paragraph>
<paragraph id="8BA636AC6845BF4AFF50FDB211D2FD62" blockId="27.[151,1436,151,858]" pageId="27" pageNumber="28">Pronotum small, slightly convex, covered with distinct punctation, punctures separated by 1.0–2.0 diameters, anterolateral corners almost rectangular, somewhat obtuse, projecting anteriorly; lateral margins straight; posterolateral corners acute; base of pronotum subtrapezoidal, with incomplete margination along basal margin; PW/PL 2.8. Scutellum relatively big, punctured as on pronotum, equilaterally triangular. Angle between pronotum and elytra not marked in dorsal outline.</paragraph>
<paragraph id="8BA636AC6845BF4AFF50FD6A13CFFCDA" blockId="27.[151,1436,151,858]" pageId="27" pageNumber="28">Elytra oval, with distinctly explanate lateral margins, widest in the middle, punctation stronger than on pronotum, punctures separated by 1.0–2.0 diameters; humeri well marked; EL/EW 1.1; EL/PL 5.2; EW/PW 1.6.</paragraph>
<caption id="DF6666246845BF4AFF01F9A91209F9ED" ID-DOI="http://doi.org/10.5281/zenodo.10095099" ID-Zenodo-Dep="10095099" httpUri="https://zenodo.org/record/10095099/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,255,1587,1611]" targetBox="[183,1388,936,1544]" targetPageId="27">
<paragraph id="8BA636AC6845BF4AFF01F9A91209F9ED" blockId="27.[151,1436,1587,1645]" pageId="27" pageNumber="28">
<emphasis id="B96DEABE6845BF4AFF01F9A9122BF9CB" bold="true" box="[151,295,1587,1611]" pageId="27" pageNumber="28">FIGURE 20.</emphasis>
<taxonomicName id="4C194D2F6845BF4AFEB8F9AE115BF9CB" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[302,599,1588,1611]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="picturatus" status="sp. nov.">
<emphasis id="B96DEABE6845BF4AFEB8F9AE115BF9CB" box="[302,599,1588,1611]" italics="true" pageId="27" pageNumber="28">Exochomoscirtes picturatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56845BF4AFDC8F9AE11BFF9CB" box="[606,691,1588,1611]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
, male.
<emphasis id="B96DEABE6845BF4AFC96F9A91010F9CB" bold="true" box="[768,796,1587,1611]" pageId="27" pageNumber="28">A)</emphasis>
sternite 9;
<emphasis id="B96DEABE6845BF4AFC02F9A910A3F9CB" bold="true" box="[916,943,1587,1611]" pageId="27" pageNumber="28">B)</emphasis>
tergite 8;
<emphasis id="B96DEABE6845BF4AFB8BF9A91735F9CB" bold="true" box="[1053,1081,1587,1611]" pageId="27" pageNumber="28">C)</emphasis>
tergite 9;
<emphasis id="B96DEABE6845BF4AFB30F9A917CFF9CB" bold="true" box="[1190,1219,1587,1611]" pageId="27" pageNumber="28">D)</emphasis>
aedeagus. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="DF6666246845BF4AFF01F84E1783F86B" ID-DOI="http://doi.org/10.5281/zenodo.10095101" ID-Zenodo-Dep="10095101" box="[151,1167,2004,2028]" httpUri="https://zenodo.org/record/10095101/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,255,2004,2028]" targetBox="[151,1436,1700,1980]" targetPageId="27">
<paragraph id="8BA636AC6845BF4AFF01F84E1783F86B" blockId="27.[151,1167,2004,2028]" box="[151,1167,2004,2028]" pageId="27" pageNumber="28">
<emphasis id="B96DEABE6845BF4AFF01F84E122AF86C" bold="true" box="[151,294,2004,2028]" pageId="27" pageNumber="28">FIGURE 21.</emphasis>
<taxonomicName id="4C194D2F6845BF4AFEBBF84E115AF86B" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[301,598,2004,2027]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="picturatus" status="sp. nov.">
<emphasis id="B96DEABE6845BF4AFEBBF84E115AF86B" box="[301,598,2004,2027]" italics="true" pageId="27" pageNumber="28">Exochomoscirtes picturatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C56845BF4AFDCAF84E11BDF86B" box="[604,689,2004,2027]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
, female
<emphasis id="B96DEABE6845BF4AFC9DF84E1043F86C" bold="true" box="[779,847,2004,2028]" pageId="27" pageNumber="28">A)–B)</emphasis>
prehensor and bursal sclerite.
</paragraph>
</caption>
<paragraph id="8BA636AC6842BF4DFF50FF0217A5FE82" blockId="28.[151,1436,152,1338]" pageId="28" pageNumber="29">Legs moderately long. Hind tibial spurs well developed, dorsal one elongate, apex modified – widened and tridentate, 0.9 X as long as tarsomere 1 and slightly longer than ventral spur. Ventral spur almost straight, somewhat shorter than tarsomere 1. Apex of dorsal spur widened, hooked, trifid apically.</paragraph>
<paragraph id="8BA636AC6842BF4DFF50FE8A11D4FED2" blockId="28.[151,1436,152,1338]" pageId="28" pageNumber="29">Ventrites 1 and 2 with sparse setation in mesal portion, ventrite 2 with basal row of big punctures. Apex of tergite 7 unmodified. Apex of ventrite 5 truncate.</paragraph>
<paragraph id="8BA636AC6842BF4DFF50FEFA1168FD12" blockId="28.[151,1436,152,1338]" pageId="28" pageNumber="29">
Sternite 9 (L 0.72,
<geoCoordinate id="EE2D506B6842BF4DFE3FFEFA110FFEFA" box="[425,515,352,378]" degrees="0.30" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="555" value="-0.3">W 0.30</geoCoordinate>
) subtriangular, apodemes fused basally, apex deeply emarginated, with a few apical setae. Tergite 8 (L 1.38,
<geoCoordinate id="EE2D506B6842BF4DFD83FE12117DFE22" box="[533,625,392,418]" degrees="0.50" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="555" value="-0.5">W 0.50</geoCoordinate>
) moderately sclerotized, apical portion small, deeply and narrowly emarginated, with arcuate apex, bearing row of short setae on apical margin, with a pair of very long apodemes diverging basally; tergite 9 (L 0.84,
<geoCoordinate id="EE2D506B6842BF4DFD05FE4211E0FE72" box="[659,748,472,498]" degrees="0.56" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="555" value="-0.56">W 0.56</geoCoordinate>
) moderately sclerotized, apical portion small, with minute setae at apex, with a pair of long apodemes diverging basally. Tegmen (L 0.78,
<geoCoordinate id="EE2D506B6842BF4DFBFCFD9A17C9FD9A" box="[1130,1221,512,538]" degrees="0.61" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="555" value="-0.61">W 0.61</geoCoordinate>
) well sclerotized, parameres straight, widening apically, inwardly pointed at apices, mesal process very narrow and short, much shorter than parameres. Penis (L 0.39,
<geoCoordinate id="EE2D506B6842BF4DFDC9FDCA11B4FDEA" box="[607,696,592,618]" degrees="0.09" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="555" value="-0.09">W 0.09</geoCoordinate>
) small, moderately sclerotized, pala very narrow and elongate, apical process evenly shorter than pala.
</paragraph>
<paragraph id="8BA636AC6842BF4DFF50FD051252FC8A" blockId="28.[151,1436,152,1338]" pageId="28" pageNumber="29">
<emphasis id="B96DEABE6842BF4DFF50FD051224FD39" bold="true" box="[198,296,671,697]" pageId="28" pageNumber="29">Female.</emphasis>
Metatibial spurs unmodified, dorsal spur almost straight, slightly shorter than tarsomere 1, ventral spur half as long as dorsal one. Dorsal pattern lighter (see: variability). Prehensor tubular, narrow and moderately long.
</paragraph>
<paragraph id="8BA636AC6842BF4DFF50FC8D1048FBF2" blockId="28.[151,1436,152,1338]" pageId="28" pageNumber="29">
<emphasis id="B96DEABE6842BF4DFF50FC8D125EFCB1" bold="true" box="[198,338,791,817]" pageId="28" pageNumber="29">Variability.</emphasis>
TL/EW in males 1.2, TL/EW in females 1.2–1.3 (1.3). Dorsal pattern extremely variable. Males with black pronotum and usually tricolor elytral pattern: humeral portion, postscutellar macula and posterior portion of elytra black, stripes connecting black maculae orange, remaining area yellow. One specimen is dark with bicolor elytra, each elytron black with two orange maculae: an oval basal one and a transversely oval one in the middle of the elytra. Females with pronotum yellow to brown, elytra orange with black pattern consisting of humeral maculae, postscutellar macula and transverse stripe in posterior portion of elytra. One female specimen (coll. NHM) is darker, anterior 2/3 of elytra are brownish-black with oval orange macula in basal portion of elytra and transversely oval macula in the middle length of elytra, posterior 1/3 of elytra orange. Several color variations are shown on
<figureCitation id="13222A296842BF4DFD20FBC2104CFBF2" box="[694,832,1112,1138]" captionStart="FIGURE 11" captionStartId="17.[151,255,1886,1910]" captionTargetBox="[164,1425,210,1820]" captionTargetId="figure-69@17.[152,1434,197,1860]" captionTargetPageId="17" captionText="FIGURE 11. Habitus illustrations of Exochomoscirtes. A) E. bimaculatus Pic; B) E. basipunctatus sp. nov.; C) E. binotatus Pic; D) E. rufomaculatus Pic; E) E. ephippiatus (Champion); F)–I) E. picturatus sp. nov., color forms; J)–L) E. takizawai sp. nov., color forms; M) E. tetrastigma (Champion). Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10095079" httpUri="https://zenodo.org/record/10095079/files/figure.png" pageId="28" pageNumber="29">Figs. 11F–I</figureCitation>
.
</paragraph>
<paragraph id="8BA636AC6842BF4DFF50FBE511E5FB6A" blockId="28.[151,1436,152,1338]" pageId="28" pageNumber="29">
<emphasis id="B96DEABE6842BF4DFF50FBE51289FB19" bold="true" box="[198,389,1151,1177]" pageId="28" pageNumber="29">Measurements.</emphasis>
Males (n = 4): TL 3.78–4.50 (4.25), PL 0.65–0.80 (0.73), PW 1.85–2.35 (2.13), EL 3.40– 4.15 (3.83), EW 3.13–3.70 (3.50). Females (n = 4): TL 4.00–4.27 (4.17), PL 0.70–0.75 (0.74), PW 1.95–2.13 (2.08), EL 3.68–3.92 (3.81), EW 3.20–3.40 (3.28).
</paragraph>
</subSubSection>
<subSubSection id="C30365276842BF4DFF50FB6D12C0FA92" box="[198,460,1271,1298]" pageId="28" pageNumber="29" type="distribution">
<paragraph id="8BA636AC6842BF4DFF50FB6D12C0FA92" blockId="28.[151,1436,152,1338]" box="[198,460,1271,1298]" pageId="28" pageNumber="29">
<emphasis id="B96DEABE6842BF4DFF50FB6D126BFA91" bold="true" box="[198,359,1271,1297]" pageId="28" pageNumber="29">Distribution.</emphasis>
Borneo.
</paragraph>
</subSubSection>
<subSubSection id="C30365276842BF4DFF50FA8510F7FABA" box="[198,1019,1311,1338]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8BA636AC6842BF4DFF50FA8510F7FABA" blockId="28.[151,1436,152,1338]" box="[198,1019,1311,1338]" pageId="28" pageNumber="29">
<emphasis id="B96DEABE6842BF4DFF50FA85125BFAB9" bold="true" box="[198,343,1311,1337]" pageId="28" pageNumber="29">Etymology:</emphasis>
Lat.
<taxonomicName id="4C194D2F6842BF4DFE02FABA1107FAB9" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[404,523,1312,1337]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="picturatus">
<emphasis id="B96DEABE6842BF4DFE02FABA1107FAB9" box="[404,523,1312,1337]" italics="true" pageId="28" pageNumber="29">picturatus</emphasis>
</taxonomicName>
= painted, from colorful pattern on elytra.
</paragraph>
</subSubSection>
</treatment>
</document>
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